184 lines
18 KiB
XML
184 lines
18 KiB
XML
<document id="0E6394D1C1BAA19EB6238686AEE52C09" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong & Prokop, Jakub" docDate="2012" docId="B970A055FFC4FFE0FD91220DFE7AFCE8" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Laetopsia Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop 2012, gen. nov." docType="treatment" docVersion="4" lastPageNumber="94" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="93" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="A2C8A8247421164A7ED95CE0E5DFE405">
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<mods:title id="AF9F2ABE18FBDE1EE65A943E4A91ADDB">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:name id="A932AF1EFA1B46CF051B2DB57B5DDF97" type="personal">
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<mods:roleTerm id="71C4A08FAF2BEEF9981F0B61C7A3A241">Author</mods:roleTerm>
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<mods:namePart id="5206B77C67B472CD0C4A702B3783B5E0">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="22A1F11C1C2FC1D9F8F1BCAACDC632BB">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com & Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:name id="29E43F24EF4A26E6BA41876779F42A31" type="personal">
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<mods:roleTerm id="C5648603F8A57D96BA09450CA9A0A87E">Author</mods:roleTerm>
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<mods:namePart id="3459D99365B734DF9B27D0C2889285B3">Prokin, Alexander</mods:namePart>
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<mods:affiliation id="55C730CA656C9B29B0A6EC285DBE0710">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru & Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="1FED9CB52ED389A45649B9E6CA4CAD90" type="personal">
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<mods:roleTerm id="CBD943646F2570E0FF6A3A664EC058EA">Author</mods:roleTerm>
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<mods:namePart id="0068D2B715929E13F70DB67DE9A57C36">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="3E84603E26A589BBCF60A91076246926">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
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</mods:name>
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<mods:name id="6FB17DD4D0FE42EFB408089B7455B800" type="personal">
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<mods:namePart id="8CFACA458F8F94BB68F76B5F76B0DA95">Pono, Alexander</mods:namePart>
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</mods:name>
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<mods:name id="8174C0E277EE737E6E65866C48C47643" type="personal">
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<mods:roleTerm id="17A8274094A7B5E1A5A203C8B7D0947D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DB40C133124B3DB57D75F5CD67A3DF27">Marenko</mods:namePart>
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<mods:affiliation id="31CCE4277B36C5ADB000D0991260159D">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
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</mods:name>
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<mods:name id="36E665CD68958353577EE7044D77F588" type="personal">
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<mods:roleTerm id="2C5D1EB0CA40E4F4A21D1D5B121C1202">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8F469DD5C5E6D269AE73C3ADA0EB0317">Yue, Yanli</mods:namePart>
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<mods:affiliation id="69796991EDD275430019D2F0422A0467">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
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<mods:name id="CB05D287D6E982801D4F42A988FF4908" type="personal">
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<mods:namePart id="B46CC3D93489D6EAF2355F416E0ADBC1">Ren, Dong</mods:namePart>
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<mods:name id="D1B8DB60F690EBE0850B43702ED03A6E" type="personal">
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<mods:roleTerm id="EEE40B536CAC75B39E600F5F2A298508">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9B97FBC9EEF1CF5982636157CA69A1E1">Prokop, Jakub</mods:namePart>
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<mods:affiliation id="AEA901399767E641A5C5FF69661A599E">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:title id="B3FD7C348F61CE3C304A80E1D468A25A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="470FC44D37DAC18A6723CE26E317D378">
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<mods:date id="E097D78CD626B32E91C75C71FE2668B6">2012</mods:date>
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<mods:number id="3ED09526432179D3DA57417AF1B19C08">2012-06-30</mods:number>
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<mods:detail id="7A35200E858C9BB793009D19E731FF17" type="volume">
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<mods:number id="11B530637A997FF05EF743B935962F84">52</mods:number>
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<mods:number id="962A320DDE894736F32ECBC0FF855293">1</mods:number>
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<mods:start id="2225C686C916FF39AC55B4AE3A1DEBF5">89</mods:start>
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<mods:end id="A2AAD0E2427A888D0D22435C97BB35A8">127</mods:end>
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<mods:classification id="CEFF83819A42B4824D378DB6DD346850">journal article</mods:classification>
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<mods:identifier id="260094D278AE894F8FD69EC16CC97ABC" type="DOI">10.5281/zenodo.5330604</mods:identifier>
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<mods:identifier id="9BDC4637CDCBB5D4170278591D8D7D16" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="2C2F98DA0B572A5CD92429E9758E6A8F" type="Zenodo-Dep">5330604</mods:identifier>
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<treatment id="B970A055FFC4FFE0FD91220DFE7AFCE8" LSID="urn:lsid:plazi:treatment:B970A055FFC4FFE0FD91220DFE7AFCE8" httpUri="http://treatment.plazi.org/id/B970A055FFC4FFE0FD91220DFE7AFCE8" lastPageId="5" lastPageNumber="94" pageId="4" pageNumber="93">
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<subSubSection id="79C342C8FFC4FFE1FD91220DFD4DFA88" box="[522,756,1371,1395]" pageId="4" pageNumber="93" type="nomenclature">
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<paragraph id="31661143FFC4FFE1FD91220DFD4DFA88" blockId="4.[522,756,1371,1395]" box="[522,756,1371,1395]" pageId="4" pageNumber="93">
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<heading id="6A2EA62FFFC4FFE1FD91220DFD4DFA88" bold="true" box="[522,756,1371,1395]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="93" reason="3">
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<emphasis id="03ADCD51FFC4FFE1FD91220DFD4DFA88" bold="true" box="[522,756,1371,1395]" pageId="4" pageNumber="93">
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†
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<taxonomicName id="F6D96AC0FFC4FFE1FD85220DFD30FA88" ID-CoL="8XM77" authority="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop, 2012" authorityName="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop" authorityYear="2012" box="[542,649,1371,1395]" class="Insecta" family="Helophoridae" genus="Laetopsia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="03ADCD51FFC4FFE1FD85220DFD30FA88" bold="true" box="[542,649,1371,1395]" italics="true" pageId="4" pageNumber="93">Laetopsia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="189E702AFFC4FFE1FD14220DFD4DFA88" box="[655,756,1371,1395]" pageId="4" pageNumber="93" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="79C342C8FFC4FFE1FF0C22DCFD10FA18" pageId="4" pageNumber="93" type="type_taxon">
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<paragraph id="31661143FFC4FFE1FF0C22DCFF53FA39" blockId="4.[151,1126,1418,1507]" pageId="4" pageNumber="93">
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||
<emphasis id="03ADCD51FFC4FFE1FF0C22DCFE92FA59" bold="true" box="[151,299,1418,1442]" pageId="4" pageNumber="93">
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<typeStatus id="EE62AFE1FFC4FFE1FF0C22DCFF76FA59" box="[151,207,1418,1442]" pageId="4" pageNumber="93">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="F6D96AC0FFC4FFE1FEA922DDFC26FA59" ID-CoL="3N7WF" authority="Prokin, Ren & Fikacek, 2011" authorityName="Prokin, Ren & Fikacek" authorityYear="2011" box="[306,927,1418,1442]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="shatrovskiyi">
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<emphasis id="03ADCD51FFC4FFE1FEA922DDFDD9FA59" box="[306,608,1419,1442]" italics="true" pageId="4" pageNumber="93">Hydrophilopsia shatrovskiyi</emphasis>
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Prokin, Ren & Fikáček, 2011
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</taxonomicName>
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(by present designation).
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</paragraph>
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<paragraph id="31661143FFC4FFE1FF0C229CFD10FA18" blockId="4.[151,1126,1418,1507]" box="[151,681,1482,1507]" pageId="4" pageNumber="93">
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<emphasis id="03ADCD51FFC4FFE1FF0C229CFE98FA19" bold="true" box="[151,289,1482,1506]" pageId="4" pageNumber="93">Time range.</emphasis>
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Early Cretaceous, ca. 145–112 mya.
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</paragraph>
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</subSubSection>
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<subSubSection id="79C342C8FFC4FFE0FF0C22AFFD34FE6F" lastPageId="5" lastPageNumber="94" pageId="4" pageNumber="93" type="diagnosis">
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<paragraph id="31661143FFC4FFE0FF0C22AFFD34FE6F" blockId="4.[151,1127,1528,1618]" lastBlockId="5.[113,1089,219,787]" lastPageId="5" lastPageNumber="94" pageId="4" pageNumber="93">
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<emphasis id="03ADCD51FFC4FFE1FF0C22AFFEE2F9EA" bold="true" box="[151,347,1528,1553]" pageId="4" pageNumber="93">
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Diagnosis.
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<emphasis id="03ADCD51FFC4FFE1FE8E22AEFEEBF9EA" bold="true" box="[277,338,1528,1553]" italics="true" pageId="4" pageNumber="93">Adult</emphasis>
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:
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</emphasis>
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Head and pronotum without setiferous granules, with grooved median portion of frontal suture, gula wide, pronotal flanks narrow, elytra without elevated costae, pale with dark stripes along striae; pronotum with a median longitudinal groove. Within the helophorid lineage, the genus is easily recognized according to the pale elytra with dark longitudinal stripes along elytral striae (unique character), narrow pronotal flanks (in contrast to wide flanks in
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<taxonomicName id="F6D96AC0FFC5FFE0FE92264BFE16FECF" authorityName="Ponomarenko" authorityYear="1987" box="[265,431,285,308]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FE92264BFE16FECF" box="[265,431,285,308]" italics="true" pageId="5" pageNumber="94">Hydrophilopsia</emphasis>
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</taxonomicName>
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) and a pronotum with a single median longitudinal groove and without setiferous granules (in
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<taxonomicName id="F6D96AC0FFC5FFE0FE36266BFD91FEAF" authorityName="Fabricius" authorityYear="1775" box="[429,552,317,340]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FE36266BFD91FEAF" box="[429,552,317,340]" italics="true" pageId="5" pageNumber="94">Helophorus</emphasis>
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</taxonomicName>
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, the subgenera
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<taxonomicName id="F6D96AC0FFC5FFE0FD5E266BFCDDFEAF" authorityName="Orchymont" authorityYear="1927" box="[709,868,317,340]" class="Insecta" family="Helophoridae" genus="Orphelophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FD5E266BFCDDFEAF" box="[709,868,317,340]" italics="true" pageId="5" pageNumber="94">Orphelophorus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F6D96AC0FFC5FFE0FC08266BFBF8FEAF" authorityName="Ponomarenko" authorityYear="1977" box="[915,1089,317,340]" class="Insecta" family="Helophoridae" genus="Mesohelophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FC08266BFBF8FEAF" box="[915,1089,317,340]" italics="true" pageId="5" pageNumber="94">Mesohelophorus</emphasis>
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</taxonomicName>
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bear a reduced number (i.e. less than five) of longitudinal pronotal grooves but always bear very distinct setiferous granules on the pronotum).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="79C342C8FFC5FFE0FFEA26CAFE7AFCE8" pageId="5" pageNumber="94" type="etymology">
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<paragraph id="31661143FFC5FFE0FFEA26CAFD15FE08" blockId="5.[113,1089,219,787]" pageId="5" pageNumber="94">
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<emphasis id="03ADCD51FFC5FFE0FFEA26CAFF4BFE4F" bold="true" box="[113,242,412,436]" pageId="5" pageNumber="94">Etymology.</emphasis>
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From
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<emphasis id="03ADCD51FFC5FFE0FEAD26CBFECFFE4F" box="[310,374,413,436]" italics="true" pageId="5" pageNumber="94">laetus</emphasis>
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(Latin), meaning joyful, pleasant, referring to the colourful elytra of this genus, plus the ending -
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<emphasis id="03ADCD51FFC5FFE0FE0226EBFE7CFE2F" box="[409,453,445,468]" italics="true" pageId="5" pageNumber="94">psia</emphasis>
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derived from the genus
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<taxonomicName id="F6D96AC0FFC5FFE0FD5E26EBFCD2FE2F" authorityName="Ponomarenko" authorityYear="1987" box="[709,875,445,468]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FD5E26EBFCD2FE2F" box="[709,875,445,468]" italics="true" pageId="5" pageNumber="94">Hydrophilopsia</emphasis>
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</taxonomicName>
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to which all species of the genus were assigned before. Gender: feminine.
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</paragraph>
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<paragraph id="31661143FFC5FFE0FFEA26ADFE7AFCE8" blockId="5.[113,1089,219,787]" pageId="5" pageNumber="94">
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<emphasis id="03ADCD51FFC5FFE0FFEA26ADFE92FDEF" bold="true" box="[113,299,507,532]" pageId="5" pageNumber="94">Taxonomic note.</emphasis>
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All species included herein share plesiomorphic characters with
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<taxonomicName id="F6D96AC0FFC5FFE0FC4926ABFF0BFDCF" authorityName="Ponomarenko" authorityYear="1987" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FC4926ABFF0BFDCF" italics="true" pageId="5" pageNumber="94">Hydrophilopsia</emphasis>
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</taxonomicName>
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but lack its only autapomorphy, i.e. the extremely wide pronotal flanks. In addition, all of them are readily distinguishable by pale elytra bearing dark stripes along elytral striae, a character which may represent an unique autapomorphy of the genus (but is unfortunately not preserved in
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<taxonomicName id="F6D96AC0FFC5FFE0FEBF252BFE73FD6F" authorityName="Ponomarenko" authorityYear="1987" box="[292,458,637,660]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FEBF252BFE73FD6F" box="[292,458,637,660]" italics="true" pageId="5" pageNumber="94">Hydrophilopsia</emphasis>
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</taxonomicName>
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). In absence of any additional data, both above differences indicate that the species with striped elytra form an easily diagnosable lineage separate from
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<taxonomicName id="F6D96AC0FFC5FFE0FFEA25EBFEAEFD2F" authorityName="Ponomarenko" authorityYear="1987" box="[113,279,701,724]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="03ADCD51FFC5FFE0FFEA25EBFEAEFD2F" box="[113,279,701,724]" italics="true" pageId="5" pageNumber="94">Hydrophilopsia</emphasis>
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</taxonomicName>
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, which is here described as
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<taxonomicName id="F6D96AC0FFC5FFE0FDCB25EBFD00FD2F" authority="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop, 2012" authorityName="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop" authorityYear="2012" box="[592,697,701,724]" class="Insecta" family="Helophoridae" genus="Laetopsia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="03ADCD51FFC5FFE0FDCB25EBFD00FD2F" box="[592,697,701,724]" italics="true" pageId="5" pageNumber="94">Laetopsia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="189E702AFFC5FFE0FD5925EAFC9FFD28" box="[706,806,700,724]" pageId="5" pageNumber="94" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
The genus seems to have coexisted with
|
||
<taxonomicName id="F6D96AC0FFC5FFE0FE97258BFE31FD0F" authorityName="Fabricius" authorityYear="1775" box="[268,392,733,756]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFC5FFE0FE97258BFE31FD0F" box="[268,392,733,756]" italics="true" pageId="5" pageNumber="94">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
until the early Cretaceous and likely represents an extinct Mesozoic clade of the helophorid lineage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |