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<document id="EF55A349FB5556B69B2339366B757630" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong &amp; Prokop, Jakub" docDate="2012" docId="B970A055FFC3FFE1FEA52362FCF4FAC8" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Hydrophilopsia longitarsalis Ponomarenko 1987" docType="treatment" docVersion="4" lastPageNumber="92" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="92" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="77D861A615F205D05ED3A1B94880AA56" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9D3DF27185326162A46C6C558C2627CA">
<mods:title id="BC1501EF2C5CCF97809EF5BA9BE8D81A">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:name id="110CE455DC3BE6D94E4069DDB16A1383" type="personal">
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<mods:namePart id="2F23FE584D562B9524D29D81FD535854">Fikáček, Martin</mods:namePart>
<mods:affiliation id="27D299E3CA7BCF5CE1083E85514E0830">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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</mods:role>
<mods:namePart id="B5BA47300C3FFD23CCFC5017BE4A68B9">Prokin, Alexander</mods:namePart>
<mods:affiliation id="E6AD2FD890E10042E59C4AA04B9FC2E1">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru &amp; Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
</mods:name>
<mods:name id="62E227BFE0227A45BC6B5F2EABFEA151" type="personal">
<mods:role id="59E93AE14BDA123FB195A54FA87F1B9E">
<mods:roleTerm id="D2274486B951901C26831E4385814705">Author</mods:roleTerm>
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<mods:namePart id="670A4A88359580956D7C87563CF18C13">Angus, Robert B.</mods:namePart>
<mods:affiliation id="E833AD4D6DEF9C84D15952E50655C59A">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
</mods:name>
<mods:name id="84175E81BC8BE3E74D5DE72993449B3C" type="personal">
<mods:role id="3DC5DCA7B09F6303B68A5214D238B444">
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<mods:namePart id="DBA1F1AE1BE8F98085AB71FD72571816">Pono, Alexander</mods:namePart>
</mods:name>
<mods:name id="9D886144222C26A4AC2404B6410B1648" type="personal">
<mods:role id="FB820B411ACA9C26E916B1834EFF5BD1">
<mods:roleTerm id="7F72F097B9B78022BECD58E782CC49B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B2A2DC263897967C9FF8E8AAAB91E19">Marenko</mods:namePart>
<mods:affiliation id="8DB23B0F530FAF08046D12A2E2849F9A">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
</mods:name>
<mods:name id="EE885C60F9ADC72310E54405E3DE3711" type="personal">
<mods:role id="85D994CBFB1AFBA08D1B746CE0AEF880">
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</mods:role>
<mods:namePart id="870C8E636BEEE13116ECF7ABC67BBC66">Yue, Yanli</mods:namePart>
<mods:affiliation id="FD3A6CE244E8F7AB80032821D17BDA58">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
</mods:name>
<mods:name id="7AB4D65804B5E0647C166B851968F72A" type="personal">
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</mods:role>
<mods:namePart id="39298BE6136B7548F5699A01A949DACB">Ren, Dong</mods:namePart>
</mods:name>
<mods:name id="63A70FB5953EF100C477B33A143A3418" type="personal">
<mods:role id="2CAD628E8299DC829AA1237D3F8B0850">
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</mods:role>
<mods:namePart id="8C15FD4051FD7019DF91AB02A2AECD40">Prokop, Jakub</mods:namePart>
<mods:affiliation id="6186F0D2FFF036362B67EE504D2C6BFE">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:typeOfResource id="3EB223A4B0CFDC2D8972AED457E96FB7">text</mods:typeOfResource>
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<mods:title id="E7941ECB175372FB335D962562759C04">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="83D1CDAF88034B676A35BEC175CA7465">
<mods:date id="5F599F43EABEAE45999DB090F49584D2">2012</mods:date>
<mods:detail id="0788A37421EBA45D9C433E549F910252" type="pubDate">
<mods:number id="17B494974504C178C1F774769EE92CEA">2012-06-30</mods:number>
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<mods:detail id="36C77687F68974AE72609154055DB499" type="volume">
<mods:number id="43803EBAC35E78941724298691194A41">52</mods:number>
</mods:detail>
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<mods:number id="C3AE44BE4E84D90DBAD26E3C015E5729">1</mods:number>
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<mods:start id="7D564E73054E3636DADEE5A8828C858B">89</mods:start>
<mods:end id="08F2181B9B604D212E22E3CA01A296B9">127</mods:end>
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<mods:classification id="935E7C0647007D520F681F75411B4013">journal article</mods:classification>
<mods:identifier id="5D7AF0A523642D30B5E1429E48090B4A" type="DOI">10.5281/zenodo.5330604</mods:identifier>
<mods:identifier id="90AD44A6324DEA4F9EB3D338CC227A2D" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="D11AFC58766033AAE4768B0CA1CAF0F2" type="Zenodo-Dep">5330604</mods:identifier>
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<treatment id="B970A055FFC3FFE1FEA52362FCF4FAC8" LSID="urn:lsid:plazi:treatment:B970A055FFC3FFE1FEA52362FCF4FAC8" httpUri="http://treatment.plazi.org/id/B970A055FFC3FFE1FEA52362FCF4FAC8" lastPageId="4" lastPageNumber="92" pageId="3" pageNumber="92">
<subSubSection id="79C342C8FFC3FFE6FEA52362FCCDFBB7" box="[318,884,1075,1100]" pageId="3" pageNumber="92" type="nomenclature">
<paragraph id="31661143FFC3FFE6FEA52362FCCDFBB7" blockId="3.[318,884,1075,1100]" box="[318,884,1075,1100]" pageId="3" pageNumber="92">
<heading id="6A2EA62FFFC3FFE6FEA52362FCCDFBB7" bold="true" box="[318,884,1075,1100]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="92" reason="3">
<emphasis id="03ADCD51FFC3FFE6FEA52362FCCDFBB7" bold="true" box="[318,884,1075,1100]" pageId="3" pageNumber="92">
†
<taxonomicName id="F6D96AC0FFC3FFE6FEC92365FCCDFBB7" ID-CoL="3N7WD" authority="Ponomarenko, 1987" authorityName="Ponomarenko" authorityYear="1987" box="[338,884,1075,1100]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="longitarsalis">
<emphasis id="03ADCD51FFC3FFE6FEC92365FD32FBB7" bold="true" box="[338,651,1075,1100]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia longitarsalis</emphasis>
<bibRefCitation id="55486CB2FFC3FFE6FD092362FCCDFBB7" author="PONOMARENKO A. G." box="[658,884,1076,1100]" pageId="3" pageNumber="92" pagination="83 - 97" refId="ref15732" refString="PONOMARENKO A. G. 1987: Novye mezozoiskye vodnye zhestkokrylye (Insecta, Coleoptera) iz Azii. [New Mesozoic aquatic beetles (Insecta, Coleoptera) from Asia]. Paleontologischeskiy Zhurnal 1987: 83 - 97 (in Russian)." type="journal article" year="1987">Ponomarenko, 1987</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE6FD912304FD11FB9E" box="[522,680,1106,1125]" pageId="3" pageNumber="92" type="description">
<paragraph id="31661143FFC3FFE6FD912304FD11FB9E" blockId="3.[113,680,1106,1160]" box="[522,680,1106,1125]" pageId="3" pageNumber="92">
(
<figureCitation id="A9E20DC6FFC3FFE6FD8B2304FDDBFB9E" box="[528,610,1106,1125]" captionStart="Figs" captionStartId="4.[151,190,907,926]" captionTargetBox="[217,1122,220,883]" captionTargetId="figure-188@4.[201,1129,216,894]" captionTargetPageId="4" captionText="Figs. 1–4. Hydrophilopsia longitarsalis Ponomarenko, 1987 (holotype). 1 – piece, general view; 2 – counterpiece, general view; 3 – detail of antenna; 4 – detail of the aedeagus. Scale bar: 2 mm (corresponds to Figs. 1–2). Abbreviations: acl – antennal club; aed – aedeagus; ant – antenna; cup – cupule; ins – intermediate antennomeres; ped – pedicel; sc – scapus." figureDoi="http://doi.org/10.5281/zenodo.5330606" httpUri="https://zenodo.org/record/5330606/files/figure.png" pageId="3" pageNumber="92">Figs. 1–4</figureCitation>
,
<figureCitation id="A9E20DC6FFC3FFE6FDF02304FD1BFB9E" box="[619,674,1106,1125]" captionStart="Figs" captionStartId="22.[151,190,1538,1557]" captionTargetBox="[167,1113,217,1527]" captionTargetId="figure-10@22.[166,1113,217,1527]" captionTargetPageId="22" captionText="Figs. 42–52. Representatives of Hydrophilopsia Ponomarenko, 1986 and Laetopsia gen. nov. 42–48 – Hydrophilopsia longitarsalis Ponomarenko, 1986, holotype (42–43 – piece and couterpiece, general views; 44 – detail of the head and thorax, photo under alcohol; 45 – same, photo in dry state; 46 – detail of aedeagus; 47 – legs of piece, 48 – legs of counterpiece). 49–52 – Laetopsia shatrovskiyi (Prokin, Ren &amp; Fikáček, 2010) (49, 51 – CNU 2009199; 50, 52 – CNU 2010242)." figureDoi="http://doi.org/10.5281/zenodo.5330643" httpUri="https://zenodo.org/record/5330643/files/figure.png" pageId="3" pageNumber="92">42–48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE6FFEA2320FDF9FB73" box="[113,576,1140,1160]" pageId="3" pageNumber="92" type="reference_group">
<paragraph id="31661143FFC3FFE6FFEA2320FDF9FB73" blockId="3.[113,680,1106,1160]" box="[113,576,1140,1160]" pageId="3" pageNumber="92">
<treatmentCitationGroup id="11C9366DFFC3FFE6FFEA2320FDF9FB73" box="[113,576,1140,1160]" pageId="3" pageNumber="92">
<treatmentCitation id="B0783752FFC3FFE6FFEA2320FD85FB73" author="PONOMARENKO A. G." box="[113,572,1140,1160]" page="94" pageId="3" pageNumber="92" year="1987">
<taxonomicName id="F6D96AC0FFC3FFE6FFEA2320FD85FB73" ID-CoL="3N7WD" authority="Ponomarenko, 1987: 94" authorityName="Ponomarenko" authorityPageNumber="94" authorityYear="1987" box="[113,572,1140,1160]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="longitarsalis">
<emphasis id="03ADCD51FFC3FFE6FFEA2320FED1FB73" box="[113,360,1142,1160]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia longitarsalis</emphasis>
<bibRefCitation id="55486CB2FFC3FFE6FEF62322FD85FB73" author="PONOMARENKO A. G." box="[365,572,1140,1160]" pageId="3" pageNumber="92" pagination="83 - 97" refId="ref15732" refString="PONOMARENKO A. G. 1987: Novye mezozoiskye vodnye zhestkokrylye (Insecta, Coleoptera) iz Azii. [New Mesozoic aquatic beetles (Insecta, Coleoptera) from Asia]. Paleontologischeskiy Zhurnal 1987: 83 - 97 (in Russian)." type="journal article" year="1987">Ponomarenko, 1987: 94</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE6FFEA23C8FC95FAFA" pageId="3" pageNumber="92" type="materials_examined">
<paragraph id="31661143FFC3FFE6FFEA23C8FEE2FB1C" blockId="3.[113,1090,1181,1282]" pageId="3" pageNumber="92">
<materialsCitation id="81B11B1EFFC3FFE6FFEA23C8FEEEFB1C" collectorName="Vasilenko &amp; A. Prokin" country="Russia" location="Chita Region" pageId="3" pageNumber="92" specimenCount="1" typeStatus="holotype">
<emphasis id="03ADCD51FFC3FFE6FFEA23C8FE81FB4A" bold="true" box="[113,312,1181,1201]" pageId="3" pageNumber="92">
<typeStatus id="EE62AFE1FFC3FFE6FFEA23C8FF27FB4A" box="[113,158,1182,1201]" pageId="3" pageNumber="92">Type</typeStatus>
locality and age.
</emphasis>
<collectingCountry id="49CE51D3FFC3FFE6FEA523C8FEC0FB4A" box="[318,377,1182,1201]" name="Russia" pageId="3" pageNumber="92">Russia</collectingCountry>
,
<location id="34064798FFC3FFE6FE1923C8FE4CFB4A" LSID="urn:lsid:plazi:treatment:B970A055FFC3FFE1FEA52362FCF4FAC8:34064798FFC3FFE6FE1923C8FE4CFB4A" box="[386,501,1182,1201]" country="Russia" name="Chita Region" pageId="3" pageNumber="92">Chita Region</location>
, Daya outcrops [left bank of the Daya river above the Shiviya falls]. Latest Jurassic, Tithonian, ca. 151–146 mya (
<bibRefCitation id="55486CB2FFC3FFE6FE6223EFFD3CFB37" author="RASNITSYN A. P." box="[505,645,1209,1228]" pageId="3" pageNumber="92" refId="ref15925" refString="RASNITSYN A. P. 1990: Pozdne-mezozoiskie nasekomye Vostochnogo Zabaikalya. Nauka, Moskva, 200 pp (in Russian)." type="book" year="1990">RASNITSYN 1990</bibRefCitation>
, SINITSHENKOVA &amp; ZHERIKHIN 1996;
<collectorName id="9C2C7495FFC3FFE6FC2323EFFBB7FB37" box="[952,1038,1209,1228]" pageId="3" pageNumber="92">Vasilenko</collectorName>
, pers. comm. to
<collectorName id="9C2C7495FFC3FFE6FF5D2382FEA2FB1C" box="[198,283,1236,1255]" pageId="3" pageNumber="92">A. Prokin</collectorName>
, 2011)
</materialsCitation>
.
</paragraph>
<paragraph id="31661143FFC3FFE6FFEA23BBFC95FAFA" blockId="3.[113,1090,1181,1282]" box="[113,812,1261,1282]" pageId="3" pageNumber="92">
<emphasis id="03ADCD51FFC3FFE6FFEA23BBFEA7FAFA" bold="true" box="[113,286,1261,1281]" pageId="3" pageNumber="92">Material examined</emphasis>
(
<specimenCount id="27DFDACAFFC3FFE6FEB023B8FED1FAFA" box="[299,360,1262,1281]" pageId="3" pageNumber="92" type="generic">1 spec.</specimenCount>
).
<typeStatus id="EE62AFE1FFC3FFE6FEEC23B9FE72FAFB" box="[375,459,1263,1282]" pageId="3" pageNumber="92" type="holotype">HOLOTYPE</typeStatus>
: PIN 3063/614 (piece and counterpiece).
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE1FFEA224CFCF4FAC8" lastPageId="4" lastPageNumber="93" pageId="3" pageNumber="92" type="description">
<paragraph id="31661143FFC3FFE1FFEA224CFCF4FAC8" blockId="3.[113,1089,1306,1618]" lastBlockId="4.[151,1127,1051,1331]" lastPageId="4" lastPageNumber="93" pageId="3" pageNumber="92">
<emphasis id="03ADCD51FFC3FFE6FFEA224CFEA9FAC9" bold="true" box="[113,272,1306,1330]" pageId="3" pageNumber="92">Redescription</emphasis>
(male). Body elongate oval,
<quantity id="F621BCA6FFC3FFE6FDDF224CFD2DFACA" box="[580,660,1306,1330]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="92" unit="mm" value="9.7">9.7 mm</quantity>
long. Head with distinct lateral portions of frontoclypeal suture, median portion weakly preserved and probably not grooved, dorsal surface of head finely punctate, without setiferous granules. Narrow postocular bridge present. Mentum transverse, 1.5× wider than long, finely punctate. Gula wide, with fine median longitudinal ridge. Antenna with 9 antennomeres (i.e. with three intermediate antennomeres,
<figureCitation id="A9E20DC6FFC3FFE6FFEA22ECFF17FA29" box="[113,174,1466,1490]" captionStart="Figs" captionStartId="4.[151,190,907,926]" captionTargetBox="[217,1122,220,883]" captionTargetId="figure-188@4.[201,1129,216,894]" captionTargetPageId="4" captionText="Figs. 1–4. Hydrophilopsia longitarsalis Ponomarenko, 1987 (holotype). 1 – piece, general view; 2 – counterpiece, general view; 3 – detail of antenna; 4 – detail of the aedeagus. Scale bar: 2 mm (corresponds to Figs. 1–2). Abbreviations: acl – antennal club; aed – aedeagus; ant – antenna; cup – cupule; ins – intermediate antennomeres; ped – pedicel; sc – scapus." figureDoi="http://doi.org/10.5281/zenodo.5330606" httpUri="https://zenodo.org/record/5330606/files/figure.png" pageId="3" pageNumber="92">Fig. 3</figureCitation>
), scape long, terminal three antennomeres forming a club. Pronotum slightly narrowing anteriad, arcuate laterally, with protruding anterolateral corners, longitudinal grooves not preserved (probably absent), lateral margins smooth; pronotal flanks very wide, the ridge delimiting them from inner portion of hypomeron arcuate in shape. Mesanepisterna not meeting medially, mesoventrite strongly narrowing anteriad, very narrow at anterior margin, bearing a transverse ridge posteriorly. Mesocoxal cavities only very narrowly divided from each other. Elytron with 10 elytral series, scutellary stria not preserved, all elytral intervals of the same height, epipleuron narrow, subdivided into wider inner and narrower outer parts. Metaventrite much longer than mesoventrite, metanepisternum with oblique transverse ridge anteriorly. Mesotibia with sparsely arranged swimming hairs, its apex with several spines; mesotarsus with five tarsomeres, each with series of swimming hairs dorsally, tarsomere 1 shortest, tarsomere 2 longer than tarsomere 3, claws long, arcuate.Abdomen with five ventrites, posterior margin of ventrite 5 possibly with rectangular teeth.Aedeagus similar to
<taxonomicName id="F6D96AC0FFC4FFE1FC7223AAFBDFFAE8" authorityName="Fabricius" authorityYear="1775" box="[1001,1126,1276,1299]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC4FFE1FC7223AAFBDFFAE8" box="[1001,1126,1276,1299]" italics="true" pageId="4" pageNumber="93">Helophorus</emphasis>
</taxonomicName>
in general shape, phallobase with acute symmetrical manubrium.
</paragraph>
</subSubSection>
</treatment>
</document>
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