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<mods:title id="7D5CE63F3768CEF4919244979924E469">Reclassification of the doryctine tribe Rhaconotini (Hymenoptera, Braconidae)</mods:title>
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Genus
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<emphasis id="15C02323AD1C5F76FDA8E3E638DB3CBE" bold="true" box="[542,767,479,506]" italics="true" pageId="101" pageNumber="102">Pseudorhaconotus</emphasis>
<bibRefCitation id="432582C0AD1C5F76FCB0E3D93E4C3CBE" author="van Achterberg C. &amp; Shaw M. R." box="[774,1128,480,506]" pageId="101" pageNumber="102" pagination="1 - 5" refId="ref80765" refString="van Achterberg C. &amp; Shaw M. R. 2010. Pseudorhaconotus enervatus, a new genus and species from Spain (Hymenoptera: Braconidae: Doryctinae). Zoologische Mededelingen 84 (1 - 9): 1 - 5." type="journal article" year="2010">van Achterberg &amp; Shaw, 2010</bibRefCitation>
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<taxonomicName id="E0B484B2AD1C5F76FF0BE01B39053F78" authority="van Achterberg &amp; Shaw, 2010: 2" authorityName="van Achterberg &amp; Shaw" authorityPageNumber="2" authorityYear="2010" box="[189,801,546,572]" class="Insecta" family="Braconidae" genus="Pseudorhaconotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="15C02323AD1C5F76FF0BE01B3BB33F78" box="[189,407,546,572]" italics="true" pageId="101" pageNumber="102">Pseudorhaconotus</emphasis>
<bibRefCitation id="432582C0AD1C5F76FE28E01B39053F78" author="van Achterberg C. &amp; Shaw M. R." box="[414,801,546,572]" pageId="101" pageNumber="102" pagination="1 - 5" refId="ref80765" refString="van Achterberg C. &amp; Shaw M. R. 2010. Pseudorhaconotus enervatus, a new genus and species from Spain (Hymenoptera: Braconidae: Doryctinae). Zoologische Mededelingen 84 (1 - 9): 1 - 5." type="journal article" year="2010">van Achterberg &amp; Shaw, 2010: 2</bibRefCitation>
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<taxonomicName id="E0B484B2AD1C5F76FF0BE05D3BB33F3A" authorityName="van Achterberg &amp; Shaw" authorityYear="2010" box="[189,407,612,638]" class="Insecta" family="Braconidae" genus="Pseudorhaconotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="15C02323AD1C5F76FF0BE05D3BB33F3A" box="[189,407,612,638]" italics="true" pageId="101" pageNumber="102">Pseudorhaconotus</emphasis>
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<bibRefCitation id="432582C0AD1C5F76FE05E05D387F3F3A" author="Yu D. S. K. &amp; van Achterberg C. &amp; Horstmann K." box="[435,603,612,638]" pageId="101" pageNumber="102" refId="ref80895" refString="Yu D. S. K., van Achterberg C. &amp; Horstmann K. 2016. Taxapad 2016, Ichneumonoidea 2015. Database on flash-drive. Nepean, Ontario." type="book" year="2016">
Yu
<emphasis id="15C02323AD1C5F76FE6DE05C38323F3A" box="[475,534,612,638]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
2016
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. —
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<bibRefCitation id="432582C0AD1C5F76FD3DE05D39DB3F3B" author="Jasso-Martinez J. M. &amp; Belokobylskij S. A. &amp; Zaldivar-Riveron A." box="[651,1023,612,639]" pageId="101" pageNumber="102" pagination="164 - 171" refId="ref79845" refString="Jasso-Martinez J. M., Belokobylskij S. A. &amp; Zaldivar-Riveron A. 2019. Molecular phylogenetics and evolution of generic diagnostic morphological features in the doryctine wasp tribe Rhaconotini (Hymenoptera: Braconidae). Zoologischer Anzeiger 279: 164 - 171. https: // doi. org / 10.1016 / j. jcz. 2019.02.002" type="journal article" year="2019">
Jasso-Martínez
<emphasis id="15C02323AD1C5F76FCF2E05C395B3F3A" box="[836,895,612,638]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
2019: 165
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<typeStatus id="F80F4193AD1C5F76FF0BE09F3ADE3F84" box="[189,250,678,704]" pageId="101" pageNumber="102">Type</typeStatus>
species
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<taxonomicName id="E0B484B2AD1C5F76FF0BE0EC39533FAB" authority="van Achterberg &amp; Shaw, 2010" authorityName="van Achterberg &amp; Shaw" authorityYear="2010" box="[189,887,725,751]" class="Insecta" family="Braconidae" genus="Pseudorhaconotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="enervatus">
<emphasis id="15C02323AD1C5F76FF0BE0EC382B3FAB" box="[189,527,725,751]" italics="true" pageId="101" pageNumber="102">Pseudorhaconotus enervatus</emphasis>
<bibRefCitation id="432582C0AD1C5F76FDA0E0EC39533FAB" author="van Achterberg C. &amp; Shaw M. R." box="[534,887,725,751]" pageId="101" pageNumber="102" pagination="1 - 5" refId="ref80765" refString="van Achterberg C. &amp; Shaw M. R. 2010. Pseudorhaconotus enervatus, a new genus and species from Spain (Hymenoptera: Braconidae: Doryctinae). Zoologische Mededelingen 84 (1 - 9): 1 - 5." type="journal article" year="2010">van Achterberg &amp; Shaw, 2010</bibRefCitation>
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, by monotypy and original designation (
<bibRefCitation id="432582C0AD1C5F76FAFCE0EC3BCA3E57" author="van Achterberg C. &amp; Shaw M. R." pageId="101" pageNumber="102" pagination="1 - 5" refId="ref80765" refString="van Achterberg C. &amp; Shaw M. R. 2010. Pseudorhaconotus enervatus, a new genus and species from Spain (Hymenoptera: Braconidae: Doryctinae). Zoologische Mededelingen 84 (1 - 9): 1 - 5." type="journal article" year="2010">van Achterberg &amp; Shaw 2010</bibRefCitation>
).
</paragraph>
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<emphasis id="15C02323AD1C5F76FF0BE1023B6A3E11" bold="true" box="[189,334,827,853]" pageId="101" pageNumber="102">Description</emphasis>
(
<figureCitation id="BF8FE3B4AD1C5F76FEE8E1023BC03E11" box="[350,484,827,853]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="103.[189,232,1685,1711]" captionStartId-1="104.[189,232,1965,1991]" captionTargetBox-0="[189,1390,269,1646]" captionTargetBox-1="[216,1372,267,1912]" captionTargetId-0="figure-12@103.[186,1401,262,1650]" captionTargetId-1="figure-14@104.[213,1375,262,1929]" captionTargetPageId-0="103" captionTargetPageId-1="104" captionText-0="Fig. 38. Pseudorhaconotus enervatus van Achterberg &amp; Shaw, 2010, ♀, holotype, NMS. A. Habitus, dorsal view. B. Head, dorsal view. C. Head, front view. D. Head, lateral view. E. Basal segments of antenna. F. Mesosoma, dorsal view. G. Mesosoma, lateral view. H. Hind leg." captionText-1="Fig. 39. Pseudorhaconotus enervatus van Achterberg &amp; Shaw, 2010, ♀, holotype, NMS. A. Wings. B. Metasoma, dorsal view. C. Metasoma and hind leg, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4649775" figureDoi-1="http://doi.org/10.5281/zenodo.4649777" httpUri-0="https://zenodo.org/record/4649775/files/figure.png" httpUri-1="https://zenodo.org/record/4649777/files/figure.png" pageId="101" pageNumber="102">Figs 3839</figureCitation>
)
</paragraph>
<paragraph id="270BFF31AD1C5F76FF0BE15339E53918" blockId="101.[189,1399,874,1116]" pageId="101" pageNumber="102">HEAD. Head weakly transverse. Ocelli arranged in triangle with base larger than its sides. Frons flat and without median keel. Eyes with sparse and short setae. Occipital carina distinct dorsally, obliterated below at short distance and not fused with hypostomal carina. Malar suture absent. Clypeal suture distinct and complete. Clypeus with short and fine ventral flange. Hypoclypeal depression round. Postgenal bridge narrow. Palps relatively short; maxillary palps 6-segmented, labial palps 4-segmented; third segment of labial palps not shortened. Scapus rather wide and long, without apical lobe. First flagellar segment subcylindrical, almost straight, almost as long as second segment.</paragraph>
<paragraph id="270BFF31AD1C5F76FF0BE6BA3BD43830" blockId="101.[189,1399,1155,1396]" pageId="101" pageNumber="102">MESOSOMA. Pronotum dorsally flat, pronotal keel distinct. Mesonotum not highly, gently roundly elevated above pronotum, entirely finely reticulate-granulate. Notauli entirely shallow and fine. Prescutellar depression rather short, with median carina. Scuto-scutellar suture present and distinct. Metanotal tooth almost absent. Sternaulus (precoxal furrow) shallow, long, sculptured. Prepectal carina distinct and complete. Postpectal carina absent. Propodeum with areas mostly delineated by distinct carinae in basal half and fine carinae in apical half, areola narrow; lateral tubercles absent; propodeal bridge absent; spiracles small and round.</paragraph>
<paragraph id="270BFF31AD1C5F76FF0BE7A539AD38BA" blockId="101.[189,1399,1436,1535]" pageId="101" pageNumber="102">LEGS. Fore and middle tibiae with slender several spines arranged in single row. Hind coxa with distinct basoventral corner and basoventral tooth. All femora with fine dorsal protuberances. Hind basitarsus almost 0.5 times as long as secondfifth segments combined.</paragraph>
<paragraph id="270BFF31AD1C5F76FF0BE41F3EC23A5C" blockId="101.[189,1399,1574,1816]" pageId="101" pageNumber="102">WINGS. Pterostigma of fore wing narrow; radial vein (r) arising from middle of pterostigma. Radial (marginal) cell rather distinctly shortened. Second radiomedial vein (r-m) absent. Recurrent vein (mcu) weakly postfurcal or almost interstitial. Nervulus (cu-a) postfurcal. Discoidal (discal) cell long petiolate. Parallel vein (CU1a) interstitial. Brachial (subdiscal) cell closed at level of recurrent vein (m-cu). Hind wing with three hamuli. Nervellus (cu-a) present. Medial (basal) cell narrow, but weakly widened posteriorly. Submedial (subbasal) cell rather short; first abscissa of mediocubital vein (M+CU) 0.6 times as long as second abscissa (1-M). Recurrent vein (m-cu) present, but transparent.</paragraph>
<paragraph id="270BFF31AD1C5F75FF0BE5063E253CC9" blockId="101.[189,1399,1855,2025]" lastBlockId="102.[189,1399,265,397]" lastPageId="102" lastPageNumber="103" pageId="101" pageNumber="102">METASOMA. First metasomal tergite not petiolate, relatively long and wide, moveably fused with second tergite. Acrosternite 0.150.20 times as long as first tergite, its apical margin situated distinctly before level of spiracles; dorsope of first tergite absent; spiracular tubercles small, dorsal carinae distinct, complete and strongly convergent basally; basolateral lobes absent. Second suture deep, almost straight, without lateral bends. Second tergite often with fine subbasal transverse furrow, with apical narrow area delineated anteriorly by deep and rather wide furrow. Present five visible tergites. Secondsixth tergites with separate laterotergites. Fourth and fifth tergite with distinct and crenulate subbasal transverse furrows. Fifth tergite of female enlarged, larger than previous tergite, almost concealed by the succeeding segments, entirely sculptured. Ovipositor much shorter than metasoma.</paragraph>
</subSubSection>
<subSubSection id="6FAEACBAAD1F5F75FF0BE3953ADB3F7F" pageId="102" pageNumber="103" type="diagnosis">
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<heading id="7C43485DAD1F5F75FF0BE3953B123C82" bold="true" box="[189,310,428,454]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="15C02323AD1F5F75FF0BE3953B123C82" bold="true" box="[189,310,428,454]" pageId="102" pageNumber="103">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="270BFF31AD1F5F75FF0BE3E23ADB3F7F" blockId="102.[189,1398,475,571]" pageId="102" pageNumber="103">
This genus is morphologically similar to
<taxonomicName id="E0B484B2AD1F5F75FD0FE3E239C73CB2" authority="Ruthe, 1854" authorityName="Ruthe" authorityYear="1854" box="[697,995,475,502]" class="Insecta" family="Braconidae" genus="Rhaconotus" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="15C02323AD1F5F75FD0FE3E239603CB1" box="[697,836,475,501]" italics="true" pageId="102" pageNumber="103">Rhaconotus</emphasis>
<bibRefCitation id="432582C0AD1F5F75FCF9E3E239C73CB2" author="Ruthe" box="[847,995,475,502]" pageId="102" pageNumber="103" pagination="343 - 355" refId="ref80392" refString="Ruthe, 1854. Beitrage zur Kenntnis der Braconiden. Stettiner Entomologische Zeitung 15: 343 - 355." type="journal article" year="1854">Ruthe, 1854</bibRefCitation>
</taxonomicName>
but differs from the latter genus mainly by the missing second radiomedial vein (r-m) and second radiomedial (submarginal) cell of fore wing.
</paragraph>
</subSubSection>
<subSubSection id="6FAEACBAAD1F5F75FF0BE06239A53FE0" box="[189,897,603,677]" pageId="102" pageNumber="103" type="description">
<paragraph id="270BFF31AD1F5F75FF0BE0623B793F31" blockId="102.[189,349,603,629]" box="[189,349,603,629]" pageId="102" pageNumber="103">
<heading id="7C43485DAD1F5F75FF0BE0623B793F31" bold="true" box="[189,349,603,629]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="15C02323AD1F5F75FF0BE0623B793F31" bold="true" box="[189,349,603,629]" pageId="102" pageNumber="103">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="270BFF31AD1F5F75FF0BE0B339A53FE0" blockId="102.[189,897,650,677]" box="[189,897,650,677]" pageId="102" pageNumber="103">
<taxonomicName id="E0B484B2AD1F5F75FF0BE0B339583FE0" authority="van Achterberg &amp; Shaw, 2010" authorityName="van Achterberg &amp; Shaw" authorityYear="2010" box="[189,892,650,677]" class="Insecta" family="Braconidae" genus="Pseudorhaconotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="enervatus">
<emphasis id="15C02323AD1F5F75FF0BE0B338343FE0" box="[189,528,650,676]" italics="true" pageId="102" pageNumber="103">Pseudorhaconotus enervatus</emphasis>
<bibRefCitation id="432582C0AD1F5F75FDAEE0B239583FE0" author="van Achterberg C. &amp; Shaw M. R." box="[536,892,650,677]" pageId="102" pageNumber="103" pagination="1 - 5" refId="ref80765" refString="van Achterberg C. &amp; Shaw M. R. 2010. Pseudorhaconotus enervatus, a new genus and species from Spain (Hymenoptera: Braconidae: Doryctinae). Zoologische Mededelingen 84 (1 - 9): 1 - 5." type="journal article" year="2010">van Achterberg &amp; Shaw, 2010</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="6FAEACBAAD1F5F75FF0BE0FD3B1D3E48" box="[189,313,708,780]" pageId="102" pageNumber="103" type="biology_ecology">
<paragraph id="270BFF31AD1F5F75FF0BE0FD3B273F9A" blockId="102.[189,259,708,734]" box="[189,259,708,734]" pageId="102" pageNumber="103">
<heading id="7C43485DAD1F5F75FF0BE0FD3B273F9A" bold="true" box="[189,259,708,734]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="15C02323AD1F5F75FF0BE0FD3B273F9A" bold="true" box="[189,259,708,734]" pageId="102" pageNumber="103">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="270BFF31AD1F5F75FF0BE0CB3B1D3E48" blockId="102.[189,313,754,780]" box="[189,313,754,780]" pageId="102" pageNumber="103">Unknown.</paragraph>
</subSubSection>
<subSubSection id="6FAEACBAAD1F5F75FF0BE1153BBC3E31" box="[189,408,812,885]" pageId="102" pageNumber="103" type="distribution">
<paragraph id="270BFF31AD1F5F75FF0BE1153B723E02" blockId="102.[189,342,812,838]" box="[189,342,812,838]" pageId="102" pageNumber="103">
<heading id="7C43485DAD1F5F75FF0BE1153B723E02" bold="true" box="[189,342,812,838]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="15C02323AD1F5F75FF0BE1153B723E02" bold="true" box="[189,342,812,838]" pageId="102" pageNumber="103">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="270BFF31AD1F5F75FF0BE1623BBC3E31" blockId="102.[189,408,859,885]" box="[189,408,859,885]" pageId="102" pageNumber="103">Palaearctic region.</paragraph>
</subSubSection>
<subSubSection id="6FAEACBAAD1F5F75FF0BE1AC3F1F39F5" pageId="102" pageNumber="103" type="discussion">
<paragraph id="270BFF31AD1F5F75FF0BE1AC3B0B3EEB" blockId="102.[189,303,917,943]" box="[189,303,917,943]" pageId="102" pageNumber="103">
<heading id="7C43485DAD1F5F75FF0BE1AC3B0B3EEB" bold="true" box="[189,303,917,943]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="15C02323AD1F5F75FF0BE1AC3B0B3EEB" bold="true" box="[189,303,917,943]" pageId="102" pageNumber="103">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="270BFF31AD1F5F75FF0BE1FA3F1F39F5" blockId="102.[189,1399,963,1201]" pageId="102" pageNumber="103">
<taxonomicName id="E0B484B2AD1F5F75FF0BE1FA3BB33E99" authorityName="van Achterberg &amp; Shaw" authorityYear="2010" box="[189,407,963,989]" class="Insecta" family="Braconidae" genus="Pseudorhaconotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="15C02323AD1F5F75FF0BE1FA3BB33E99" box="[189,407,963,989]" italics="true" pageId="102" pageNumber="103">Pseudorhaconotus</emphasis>
</taxonomicName>
is a morphologically distinctive monotypic genus that was originally described from southern
<collectingCountry id="5FA3BFA1AD1F5F75FE9EE1DE3B4F3945" box="[296,363,999,1025]" name="Spain" pageId="102" pageNumber="103">Spain</collectingCountry>
(
<bibRefCitation id="432582C0AD1F5F75FECFE1DE38FC3945" author="van Achterberg C. &amp; Shaw M. R." box="[377,728,999,1025]" pageId="102" pageNumber="103" pagination="1 - 5" refId="ref80765" refString="van Achterberg C. &amp; Shaw M. R. 2010. Pseudorhaconotus enervatus, a new genus and species from Spain (Hymenoptera: Braconidae: Doryctinae). Zoologische Mededelingen 84 (1 - 9): 1 - 5." type="journal article" year="2010">van Achterberg &amp; Shaw 2010</bibRefCitation>
). This is one of the two
<taxonomicName id="E0B484B2AD1F5F75FC5EE1DE3E533945" authorityName="Fahringer" authorityYear="1928" box="[1000,1143,999,1025]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="tribe" tribe="Rhaconotini">Rhaconotini</taxonomicName>
genera (together with Malagasy
<taxonomicName id="E0B484B2AD1F5F75FE81E63338D23961" authority="Belokobylskij, 2005" authorityName="Belokobylskij" authorityYear="2005" box="[311,758,1034,1061]" class="Insecta" family="Braconidae" genus="Grangerdoryctes" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="15C02323AD1F5F75FE81E6333BDA3960" box="[311,510,1034,1060]" italics="true" pageId="102" pageNumber="103">Grangerdoryctes</emphasis>
Belokobylskij, 2005
</taxonomicName>
) that does not have the second radiomedial vein (r-m) and second radiomedial (submarginal) cell. The position of this genus within
<taxonomicName id="E0B484B2AD1F5F75FBE6E6143EFB3903" authorityName="Fahringer" authorityYear="1928" box="[1104,1247,1069,1095]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="tribe" tribe="Rhaconotini">Rhaconotini</taxonomicName>
is supported by its enlarged fifth metasomal tergite, which almost covers the following segments, the separated laterotergites on the fifth metasomal tergite, interstitial position of parallel vein (CU1a) in fore wing, postfurcal position of recurrent vein (m-cu), and densely granulate-reticulate head and mesosoma.
</paragraph>
</subSubSection>
</treatment>
</document>