341 lines
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341 lines
40 KiB
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<document id="B35146875E55CC719D0DCE9DBD38F120" ID-CLB-Dataset="55712" ID-DOI="10.11646/zootaxa.5133.3.1" ID-GBIF-Dataset="dd1981ca-6b0b-42cf-839b-f1d3ba251c73" ID-ISSN="1175-5326" ID-Zenodo-Dep="6522449" ID-ZooBank="16A54EFD-028D-42FF-BD62-B8D35B7BA4DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651755885264" checkinUser="plazi" docAuthor="Makarkin, Vladimir N., Antell, Gwen S. & Archibald, S. Bruce" docDate="2022" docId="AD76878EFFE5FFEEE292A443CD957654" docLanguage="en" docName="zootaxa.5133.3.1.pdf" docOrigin="Zootaxa 5133 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cimbrochrysa Schluter 1982" docType="treatment" docVersion="6" lastPageNumber="313" masterDocId="514FFFF6FFEEFFE2E205A266CE40746A" masterDocTitle="A revision of Chrysopidae (Neuroptera) from the late Eocene Florissant Formation Colorado, with description of new species" masterLastPageNumber="345" masterPageNumber="301" pageNumber="312" updateTime="1699329839210" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F46D729B35D2E7BCE8C74D303C5B9BFA">A revision of Chrysopidae (Neuroptera) from the late Eocene Florissant Formation Colorado, with description of new species</mods:title>
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<mods:namePart id="87E68E1B744C58C2D05561DB68B87659">Makarkin, Vladimir N.</mods:namePart>
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<mods:affiliation id="2D31596512DEED0C028019F05D30BDD8">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia</mods:affiliation>
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<mods:namePart id="4470548AF8D724DA84D96B8E289C99F9">Antell, Gwen S.</mods:namePart>
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<mods:affiliation id="18392103AC7F8295022B960CDAB84F14">Department of Earth Sciences, University of Oxford, Oxford OX 1 3 AN, United Kingdom</mods:affiliation>
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<mods:name id="9D3ECA2CD9E328792A8D7235A26298C3" type="personal">
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<mods:namePart id="D35A2920811AF6E6182BAFABBD57C644">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="C9C4BDB6D03F5DB0BA8271DECE9ABC88">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada sba 48 @ sfu. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:title id="CF49E8BFDA94F70341FDA3D0772EC116">Zootaxa</mods:title>
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<mods:date id="BE9F5BFF31C41B488592EDC3EBBFD08D">2022</mods:date>
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Genus
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<taxonomicName id="E2DF4D1BFFE5FFE9E2EBA443CC127255" ID-CoL="3PX3" authority="Schluter, 1982" authorityName="Schluter" authorityYear="1982" box="[238,594,1573,1600]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
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<emphasis id="17ABEA8AFFE5FFE9E2EBA443CFD9722A" bold="true" box="[238,409,1573,1600]" italics="true" pageId="11" pageNumber="312">Cimbrochrysa</emphasis>
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<bibRefCitation id="414E4B69FFE5FFE9E3A4A443CC127255" author="Schluter, T." box="[417,594,1573,1599]" pageId="11" pageNumber="312" pagination="257 - 264" refId="ref25389" refString="Schluter, T. (1982) Cimbrochrysa moleriensis n. g. n. sp. und Hypochrysa hercyniensis n. sp., zwei fossile Chrysopidae-Arten (Insecta: Planipennia) aus dem europaischen Tertiar. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 5, 257 - 264. https: // doi. org / 10.1127 / njgpm / 1982 / 1982 / 257" type="journal article" year="1982">Schlüter, 1982</bibRefCitation>
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<paragraph id="25603698FFE5FFE9E292A40CCD5772D5" blockId="11.[151,1437,1642,1727]" pageId="11" pageNumber="312">
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<treatmentCitation id="A47E1089FFE5FFE9E292A40CCFB372E8" author="Schluter, T." box="[151,499,1642,1666]" page="258" pageId="11" pageNumber="312" year="1982">
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<taxonomicName id="E2DF4D1BFFE5FFE9E292A40CCFB372E8" ID-CoL="3PX3" authority="Schluter, 1982: 258" authorityName="Schluter" authorityPageNumber="258" authorityYear="1982" box="[151,499,1642,1666]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
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<emphasis id="17ABEA8AFFE5FFE9E292A40CCF6772E8" box="[151,295,1642,1666]" italics="true" pageId="11" pageNumber="312">Cimbrochrysa</emphasis>
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<bibRefCitation id="414E4B69FFE5FFE9E329A40CCFB372E8" author="Schluter, T." box="[300,499,1642,1666]" pageId="11" pageNumber="312" pagination="257 - 264" refId="ref25389" refString="Schluter, T. (1982) Cimbrochrysa moleriensis n. g. n. sp. und Hypochrysa hercyniensis n. sp., zwei fossile Chrysopidae-Arten (Insecta: Planipennia) aus dem europaischen Tertiar. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 5, 257 - 264. https: // doi. org / 10.1127 / njgpm / 1982 / 1982 / 257" type="journal article" year="1982">Schlüter, 1982: 258</bibRefCitation>
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; Schlüter 1984: 7;
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<treatmentCitation id="A47E1089FFE5FFE9E0A8A40CCDA472E8" author="Willmann, R. & Brooks, S. J." box="[685,996,1642,1666]" page="126" pageId="11" pageNumber="312" year="1991">
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<bibRefCitation id="414E4B69FFE5FFE9E0A8A40CCDA472E8" author="Willmann, R. & Brooks, S. J." box="[685,996,1642,1666]" pageId="11" pageNumber="312" pagination="125 - 135" refId="ref26306" refString="Willmann, R. & Brooks, S. J. (1991) Insekten aus der Fur-Formation von Danemark (Moler, ob. Paleozan / unt. Eozan?). 6. Chrysopidae (Neuroptera). Meyniana, 43, 125 - 135." type="journal article" year="1991">Willmann & Brooks 1991: 126</bibRefCitation>
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, 130;
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<treatmentCitation id="A47E1089FFE5FFE9E625A40CCAB072E8" author="Carpenter, F. M." box="[1056,1264,1642,1666]" page="347" pageId="11" pageNumber="312" year="1992">
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<bibRefCitation id="414E4B69FFE5FFE9E625A40CCAB072E8" author="Carpenter, F. M." box="[1056,1264,1642,1666]" pageId="11" pageNumber="312" pagination="1 - 655" refId="ref22887" refString="Carpenter, F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vols. 3 & 4. Geological Society of America, Boulder, Colorado & University of Kansas, Lawrence, Kansas, pp. 1 - 655." type="book chapter" year="1992">Carpenter 1992: 347</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFE5FFE9E6FCA40CCEAF72CA" author="Willmann, R." page="242" pageId="11" pageNumber="312" year="1993">
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<bibRefCitation id="414E4B69FFE5FFE9E6FCA40CCEAF72CA" author="Willmann, R." pageId="11" pageNumber="312" pagination="239 - 245" refId="ref26238" refString="Willmann, R. (1993) Insekten aus der Fur-Formation von Danemark (Moler, ob. Paleozan / unt. Eozan?). 8. Zwei neue Vertreter der Chrysopidae (Neuroptera). Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 4, 239 - 245. https: // doi. org / 10.1127 / njgpm / 1993 / 1993 / 239" type="journal article" year="1993">Willmann 1993: 242</bibRefCitation>
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, 244;
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<treatmentCitation id="A47E1089FFE5FFE9E334A4EECC7A72CA" author="Penalver, E. & Nel, A. & Martinex-Delclos, X." box="[305,570,1672,1696]" page="486" pageId="11" pageNumber="312" year="1995">
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<bibRefCitation id="414E4B69FFE5FFE9E334A4EECC7A72CA" author="Penalver, E. & Nel, A. & Martinex-Delclos, X." box="[305,570,1672,1696]" pageId="11" pageNumber="312" pagination="481 - 487" refId="ref25120" refString="Penalver, E., Nel, A. & Martinex-Delclos, X. (1995) New Nothochrysinae from the spanish Miocene (Neuroptera, Chrysopidae). Bulletin de la Societe Entomologique de France, 100, 481 - 487. https: // doi. org / 10.3406 / bsef. 1995.17194" type="journal article" year="1995">
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Peñalver
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<emphasis id="17ABEA8AFFE5FFE9E397A4EFCF8772CA" box="[402,455,1672,1696]" italics="true" pageId="11" pageNumber="312">et al.</emphasis>
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1995: 486
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFE5FFE9E043A4EECD4E72CA" author="Nel, A. & Delclos, X. & Hutin, A." box="[582,782,1672,1696]" page="67" pageId="11" pageNumber="312" year="2005">
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<bibRefCitation id="414E4B69FFE5FFE9E043A4EECD4E72CA" author="Nel, A. & Delclos, X. & Hutin, A." box="[582,782,1672,1696]" pageId="11" pageNumber="312" pagination="29 - 68" refId="ref24759" refString="Nel, A., Delclos, X. & Hutin, A. (2005) Mesozoic chrysopid-like Planippenia: a phylogenetic approach (Insecta: Neuroptera). Annales de la Societe Entomologique de France, 41, 29 - 68. https: // doi. org / 10.1080 / 00379271.2005.10697440" type="journal article" year="2005">
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Nel
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<emphasis id="17ABEA8AFFE5FFE9E076A4EFCCE872CA" box="[627,680,1672,1696]" italics="true" pageId="11" pageNumber="312">et al.</emphasis>
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2005: 67
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFE5FFE9E11FA4EECA3872CA" author="Makarkin, V. N. & Archibald, S. B." box="[794,1144,1672,1696]" page="127" pageId="11" pageNumber="312" year="2013">
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<bibRefCitation id="414E4B69FFE5FFE9E11FA4EECA3872CA" author="Makarkin, V. N. & Archibald, S. B." box="[794,1144,1672,1696]" pageId="11" pageNumber="312" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald 2013: 127</bibRefCitation>
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;
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<treatmentCitation id="A47E1089FFE5FFE9E686A4EECBD872CA" author="Archibald, S. B. & Makarkin, V. N. & Greenwood, D. R. & Gunnell, G. F." box="[1155,1432,1672,1696]" page="191" pageId="11" pageNumber="312" year="2014">
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<bibRefCitation id="414E4B69FFE5FFE9E686A4EECBD872CA" author="Archibald, S. B. & Makarkin, V. N. & Greenwood, D. R. & Gunnell, G. F." box="[1155,1432,1672,1696]" pageId="11" pageNumber="312" pagination="185 - 191" refId="ref22595" refString="Archibald, S. B., Makarkin, V. N., Greenwood, D. R. & Gunnell, G. F. (2014) The Red Queen and Court Jester in green lacewing evolution: bat predation and global climate change. Palaios, 29, 185 - 191. https: // doi. org / 10.2110 / palo. 2013.089" type="journal article" year="2014">
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Archibald
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<emphasis id="17ABEA8AFFE5FFE9E6F5A4EFCB5F72CA" box="[1264,1311,1672,1696]" italics="true" pageId="11" pageNumber="312">et al</emphasis>
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. 2014: 191
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;
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<treatmentCitation id="A47E1089FFE5FFE9E2C2A4C1CF3F72D5" author="Makarkin V. N." box="[199,383,1703,1727]" page="4" pageId="11" pageNumber="312" year="2014">
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<bibRefCitation id="414E4B69FFE5FFE9E2C2A4C1CF3F72D5" author="Makarkin V. N." box="[199,383,1703,1727]" pageId="11" pageNumber="312" pagination="1 - 7" refId="ref24254" refString="Makarkin V. N. (2014) A new fossil green lacewing (Neuroptera: Chrysopidae) from the Eocene Tadushi Formation, eastern Sikhote-Alin. Far Eastern Entomologist, 272, 1 - 7." type="journal article" year="2014">Makarkin 2014: 4</bibRefCitation>
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;
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<treatmentCitation id="A47E1089FFE5FFE9E38CA4C1CC9F72D5" author="Archibald, S. B. & Makarkin, V. N." box="[393,735,1703,1727]" page="365" pageId="11" pageNumber="312" year="2015">
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<bibRefCitation id="414E4B69FFE5FFE9E38CA4C1CC9F72D5" author="Archibald, S. B. & Makarkin, V. N." box="[393,735,1703,1727]" pageId="11" pageNumber="312" pagination="359 - 369" refId="ref22418" refString="Archibald, S. B. & Makarkin, V. N. (2015) A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the Early Eocene of Driftwood Canyon, British Columbia, Canada. Canadian Entomologist, 147, 359 - 369. https: // doi. org / 10.4039 / tce. 2014.53" type="journal article" year="2015">Archibald & Makarkin 2015: 365</bibRefCitation>
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, 366.
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</paragraph>
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<emphasis id="17ABEA8AFFE5FFE9E292A48BCF72736D" bold="true" box="[151,306,1773,1799]" pageId="11" pageNumber="312">
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<typeStatus id="FA64883AFFE5FFE9E292A48BCE92736D" box="[151,210,1773,1799]" pageId="11" pageNumber="312">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="E2DF4D1BFFE5FFE9E33CA48BCD4D736D" ID-CoL="VC2D" authority="Schluter, 1982" authorityName="Schluter" authorityYear="1982" box="[313,781,1773,1799]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="moleriensis">
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<emphasis id="17ABEA8AFFE5FFE9E33CA48BCC20736D" box="[313,608,1773,1799]" italics="true" pageId="11" pageNumber="312">Cimbrochrysa moleriensis</emphasis>
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<bibRefCitation id="414E4B69FFE5FFE9E062A48BCD4D736D" author="Schluter, T." box="[615,781,1773,1799]" pageId="11" pageNumber="312" pagination="257 - 264" refId="ref25389" refString="Schluter, T. (1982) Cimbrochrysa moleriensis n. g. n. sp. und Hypochrysa hercyniensis n. sp., zwei fossile Chrysopidae-Arten (Insecta: Planipennia) aus dem europaischen Tertiar. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 5, 257 - 264. https: // doi. org / 10.1127 / njgpm / 1982 / 1982 / 257" type="journal article" year="1982">Schlüter, 1982</bibRefCitation>
|
|
</taxonomicName>
|
|
, by original designation.
|
|
</paragraph>
|
|
<paragraph id="25603698FFE5FFEEE2C2A577CA297493" blockId="11.[151,1436,1773,2016]" lastBlockId="12.[151,1437,150,574]" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">
|
|
<emphasis id="17ABEA8AFFE5FFE9E2C2A577CFF67341" bold="true" box="[199,438,1809,1835]" pageId="11" pageNumber="312">Emended diagnosis.</emphasis>
|
|
May be distinguished from other genera of
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E1A2A577CA187341" authorityName="Navas" authorityYear="1910" box="[935,1112,1809,1835]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="subFamily" subFamily="Nothochrysinae">Nothochrysinae</taxonomicName>
|
|
by a combination of the following. Forewing: (1)
|
|
<emphasis id="17ABEA8AFFE5FFE9E392A550CFF37325" box="[407,435,1846,1871]" italics="true" pageId="11" pageNumber="312">im</emphasis>
|
|
broad, pentagonal [
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E090A553CD0F7325" ID-CoL="7N4Z" authorityName="Wilmann" authorityYear="1993" box="[661,847,1845,1871]" class="Insecta" family="Chrysopidae" genus="Stephenbrooksia" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E090A553CD0F7325" box="[661,847,1845,1871]" italics="true" pageId="11" pageNumber="312">Stephenbrooksia</emphasis>
|
|
</taxonomicName>
|
|
: very long, narrow;
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E63DA553CAA07325" ID-CoL="ZW2" authorityName="Adams" authorityYear="1967" box="[1080,1248,1845,1871]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E63DA553CAA07325" box="[1080,1248,1845,1871]" italics="true" pageId="11" pageNumber="312">Archaeochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E6EAA553CBD57325" ID-CoL="496V" authorityName="Adams" authorityYear="1967" box="[1263,1429,1845,1871]" class="Insecta" family="Chrysopidae" genus="Dyspetochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E6EAA553CBD57325" box="[1263,1429,1845,1871]" italics="true" pageId="11" pageNumber="312">Dyspetochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E292A53FCF9F731E" ID-CoL="5FLP" authority="Carpenter, 1935" authorityName="Carpenter" authorityYear="1935" box="[151,479,1881,1908]" class="Insecta" family="Chrysopidae" genus="Lithochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E292A53FCF5D7319" box="[151,285,1881,1907]" italics="true" pageId="11" pageNumber="312">Lithochrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFE5FFE9E323A53FCF9F731E" author="Carpenter, F. M." box="[294,479,1881,1908]" pageId="11" pageNumber="312" pagination="259 - 271" refId="ref22778" refString="Carpenter, F. M. (1935) Tertiary insects of the family Chrysopidae. Journal of Paleontology, 9, 259 - 271." type="journal article" year="1935">Carpenter, 1935</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E3E8A53FCD767319" ID-CoL="63K25" authority="Scudder, 1883" authorityName="Scudder" authorityYear="1883" box="[493,822,1881,1907]" class="Insecta" family="Chrysopidae" genus="Palaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E3E8A53FCCC77319" box="[493,647,1881,1907]" italics="true" pageId="11" pageNumber="312">Palaeochrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFE5FFE9E095A53FCD767319" author="Scudder, S. H." box="[656,822,1881,1907]" pageId="11" pageNumber="312" pagination="271 - 293" refId="ref25580" refString="Scudder, S. H. (1883) The tertiary Lake Basin at Florissant, Colo., between South and Hayden Parks. Annual Report of the United States Geological and Geographical Survey of the Territories, 12, 271 - 293. https: // doi. org / 10.3133 / 70159191" type="journal article" year="1883">Scudder, 1883</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E141A53FCA3B7319" ID-CoL="7YYW" authority="Scudder, 1883" authorityName="Scudder" authorityYear="1883" box="[836,1147,1881,1907]" class="Insecta" family="Chrysopidae" genus="Tribochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E141A53FCD8B7319" box="[836,971,1881,1907]" italics="true" pageId="11" pageNumber="312">Tribochrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFE5FFE9E1D1A53FCA3B7319" author="Scudder, S. H." box="[980,1147,1881,1907]" pageId="11" pageNumber="312" pagination="271 - 293" refId="ref25580" refString="Scudder, S. H. (1883) The tertiary Lake Basin at Florissant, Colo., between South and Hayden Parks. Annual Report of the United States Geological and Geographical Survey of the Territories, 12, 271 - 293. https: // doi. org / 10.3133 / 70159191" type="journal article" year="1883">Scudder, 1883</bibRefCitation>
|
|
</taxonomicName>
|
|
: elongate]; (2) 2m-cu located approximately in middle of
|
|
<emphasis id="17ABEA8AFFE5FFE9E013A518CC7273FD" box="[534,562,1918,1943]" italics="true" pageId="11" pageNumber="312">im</emphasis>
|
|
[
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E046A51BCA0E73FD" ID-CoL="6ZLP" authority="Makarkin & Archibald, 2013" authorityName="Makarkin & Archibald" authorityYear="2013" box="[579,1102,1917,1943]" class="Insecta" family="Chrysopidae" genus="Pseudochrysopa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E046A51BCCBC73FD" box="[579,764,1917,1943]" italics="true" pageId="11" pageNumber="312">Pseudochrysopa</emphasis>
|
|
<bibRefCitation id="414E4B69FFE5FFE9E101A51BCA0E73FD" author="Makarkin, V. N. & Archibald, S. B." box="[772,1102,1917,1943]" pageId="11" pageNumber="312" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E65FA51BCB4073FD" ID-CoL="496V" authorityName="Adams" authorityYear="1967" box="[1114,1280,1917,1943]" class="Insecta" family="Chrysopidae" genus="Dyspetochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E65FA51BCB4073FD" box="[1114,1280,1917,1943]" italics="true" pageId="11" pageNumber="312">Dyspetochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E70AA51BCBD573FD" ID-CoL="5FLP" authorityName="Carpenter" authorityYear="1935" box="[1295,1429,1917,1943]" class="Insecta" family="Chrysopidae" genus="Lithochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E70AA51BCBD573FD" box="[1295,1429,1917,1943]" italics="true" pageId="11" pageNumber="312">Lithochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E292A5C7CF7173D1" ID-CoL="63K25" authorityName="Scudder" authorityYear="1883" box="[151,305,1953,1979]" class="Insecta" family="Chrysopidae" genus="Palaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E292A5C7CF7173D1" box="[151,305,1953,1979]" italics="true" pageId="11" pageNumber="312">Palaeochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E344A5C7CF8873D1" ID-CoL="7YYW" authorityName="Scudder" authorityYear="1883" box="[321,456,1953,1979]" class="Insecta" family="Chrysopidae" genus="Tribochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E344A5C7CF8873D1" box="[321,456,1953,1979]" italics="true" pageId="11" pageNumber="312">Tribochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E3DDA5C7CC2773D1" ID-CoL="653R" authorityName="McLachlan" authorityYear="1868" box="[472,615,1953,1979]" class="Insecta" family="Chrysopidae" genus="Nothochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E3DDA5C7CC2773D1" box="[472,615,1953,1979]" italics="true" pageId="11" pageNumber="312">Nothochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E072A5C7CD6A73D1" ID-CoL="63Q5S" box="[631,810,1953,1979]" class="Insecta" family="Chrysopidae" genus="Pronothochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E072A5C7CD6A73D1" box="[631,810,1953,1979]" italics="true" pageId="11" pageNumber="312">Pronothochrysa</emphasis>
|
|
</taxonomicName>
|
|
: in proximal part of
|
|
<emphasis id="17ABEA8AFFE5FFE9E61EA5C4CA7773D1" box="[1051,1079,1954,1979]" italics="true" pageId="11" pageNumber="312">im</emphasis>
|
|
;
|
|
<emphasis id="17ABEA8AFFE5FFE9E642A5C7CA8E73D1" box="[1095,1230,1953,1979]" italics="true" pageId="11" pageNumber="312">Kimachrysa</emphasis>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E6DBA5C7CB2473D1" ID-CoL="6QK5" authorityName="Adams" authorityYear="1957" box="[1246,1380,1953,1979]" class="Insecta" family="Chrysopidae" genus="Pimachrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E6DBA5C7CB2473D1" box="[1246,1380,1953,1979]" italics="true" pageId="11" pageNumber="312">Pimachrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E771A5C4CF4473B5" ID-CoL="6CZK" class="Insecta" family="Chrysopidae" genus="Pamochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E771A5C4CF4473B5" italics="true" pageId="11" pageNumber="312">Pamochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E313A5A3CFDD73B5" ID-CoL="52NJ" authorityName="Hagen" authorityYear="1966" box="[278,413,1989,2015]" class="Insecta" family="Chrysopidae" genus="Hypochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E313A5A3CFDD73B5" box="[278,413,1989,2015]" italics="true" pageId="11" pageNumber="312">Hypochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE5FFE9E3AAA5A3CC1473B5" ID-CoL="34G9" authorityName="Adams & Penny" authorityYear="1992" box="[431,596,1989,2015]" class="Insecta" family="Chrysopidae" genus="Asthenochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFE9E3AAA5A3CC1473B5" box="[431,596,1989,2015]" italics="true" pageId="11" pageNumber="312">Asthenochrysa</emphasis>
|
|
</taxonomicName>
|
|
: proximad
|
|
<emphasis id="17ABEA8AFFE5FFE9E0D8A5A0CCB973B5" box="[733,761,1990,2015]" italics="true" pageId="11" pageNumber="312">im</emphasis>
|
|
]; (3) 2m-cu located distad proximal branch of CuA [
|
|
<taxonomicName id="E2DF4D1BFFE5FFEEE76AA5A0CEB074DB" ID-CoL="62CN7" authorityName="Makarkin" authorityYear="2014" class="Insecta" family="Chrysopidae" genus="Asiachrysa" kingdom="Animalia" lastPageId="12" lastPageNumber="313" order="Neuroptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE5FFEEE76AA5A0CEB074DB" italics="true" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">Asiachrysa</emphasis>
|
|
</taxonomicName>
|
|
: proximad]; (4) CuP once forked [
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE06BA2F1CCB774DB" ID-CoL="3ZRR" authorityName="Wilmann" authorityYear="1993" box="[622,759,151,177]" class="Insecta" family="Chrysopidae" genus="Danochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE06BA2F1CCB774DB" box="[622,759,151,177]" italics="true" pageId="12" pageNumber="313">Danochrysa</emphasis>
|
|
</taxonomicName>
|
|
: twice forked]; (5) two gradate series in radial space [
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE74DA2F1CF5A74BF" ID-CoL="67H3" authorityName="Makarkin & Archibald" authorityYear="2013" class="Insecta" family="Chrysopidae" genus="Okanaganochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE74DA2F1CF5A74BF" italics="true" pageId="12" pageNumber="313">Okanaganochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE32CA2DDCFF874BF" ID-CoL="82CV" authorityName="Kimmins" authorityYear="1952" box="[297,440,187,213]" class="Insecta" family="Chrysopidae" genus="Triplochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE32CA2DDCFF874BF" box="[297,440,187,213]" italics="true" pageId="12" pageNumber="313">Triplochrysa</emphasis>
|
|
</taxonomicName>
|
|
: three;
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE014A2DDCCE674BF" ID-CoL="62RB6" authorityName="Esben-Petersen" authorityYear="1917" box="[529,678,187,213]" class="Insecta" family="Chrysopidae" genus="Dictyochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE014A2DDCCE674BF" box="[529,678,187,213]" italics="true" pageId="12" pageNumber="313">Dictyochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE0B3A2DDCD1B74BF" ID-CoL="NYH" authorityName="Makarkin & Archibald" authorityYear="2013" box="[694,859,187,213]" class="Insecta" family="Chrysopidae" genus="Adamsochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE0B3A2DDCD1B74BF" box="[694,859,187,213]" italics="true" pageId="12" pageNumber="313">Adamsochrysa</emphasis>
|
|
</taxonomicName>
|
|
: four or more]. Hind wing: (6) MA fused with RP proximally for long distance [
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE3E3A2B9CCCE7493" ID-CoL="ZW2" authorityName="Adams" authorityYear="1967" box="[486,654,223,249]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE3E3A2B9CCCE7493" box="[486,654,223,249]" italics="true" pageId="12" pageNumber="313">Archaeochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE099A2B9CD167493" ID-CoL="7N4Z" authorityName="Wilmann" authorityYear="1993" box="[668,854,223,249]" class="Insecta" family="Chrysopidae" genus="Stephenbrooksia" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE099A2B9CD167493" box="[668,854,223,249]" italics="true" pageId="12" pageNumber="313">Stephenbrooksia</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE161A2B9CDAE7493" ID-CoL="5CX8" authorityName="Adams & Penny" authorityYear="1992" box="[868,1006,223,249]" class="Insecta" family="Chrysopidae" genus="Leptochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE161A2B9CDAE7493" box="[868,1006,223,249]" italics="true" pageId="12" pageNumber="313">Leptochrysa</emphasis>
|
|
</taxonomicName>
|
|
: separate].
|
|
</paragraph>
|
|
<paragraph id="25603698FFE2FFEEE2C2A364CDAF752B" blockId="12.[151,1437,150,574]" pageId="12" pageNumber="313">
|
|
<emphasis id="17ABEA8AFFE2FFEEE2C2A364CFCC7576" bold="true" box="[199,396,258,284]" pageId="12" pageNumber="313">Species included</emphasis>
|
|
.
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE39FA365CC7B7577" ID-CoL="VC2D" authorityName="Schluter" authorityYear="1982" box="[410,571,259,285]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="moleriensis">
|
|
<emphasis id="17ABEA8AFFE2FFEEE39FA365CC7B7577" box="[410,571,259,285]" italics="true" pageId="12" pageNumber="313">C. moleriensis</emphasis>
|
|
</taxonomicName>
|
|
(earliest Eocene of the Fur Formation,
|
|
<collectingCountry id="5DC87608FFE2FFEEE1F6A365CA217574" box="[1011,1121,259,286]" name="Denmark" pageId="12" pageNumber="313">Denmark</collectingCountry>
|
|
);
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE676A365CB4C7576" authority="Makarkin & Antell & Archibald, 2022" authorityName="Makarkin & Antell & Archibald" authorityYear="2022" box="[1139,1292,259,285]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="americana" status="sp. nov.">
|
|
<emphasis id="17ABEA8AFFE2FFEEE676A365CB4C7576" box="[1139,1292,259,285]" italics="true" pageId="12" pageNumber="313">C. americana</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="0C9857F1FFE2FFEEE717A365CB2D7577" box="[1298,1389,259,285]" pageId="12" pageNumber="313" rank="species">
|
|
<emphasis id="17ABEA8AFFE2FFEEE717A365CB267577" bold="true" box="[1298,1382,259,285]" pageId="12" pageNumber="313">sp. nov</emphasis>
|
|
.
|
|
</taxonomicNameLabel>
|
|
and
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE292A341CEBB752A" authority="Makarkin & Antell & Archibald, 2022" authorityName="Makarkin & Antell & Archibald" authorityYear="2022" box="[151,251,295,321]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
|
|
<emphasis id="17ABEA8AFFE2FFEEE292A341CEBB752A" box="[151,251,295,321]" italics="true" pageId="12" pageNumber="313">C. major</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="0C9857F1FFE2FFEEE307A341CF1D752B" box="[258,349,295,321]" pageId="12" pageNumber="313" rank="species">
|
|
<emphasis id="17ABEA8AFFE2FFEEE307A341CF16752B" bold="true" box="[258,342,295,321]" pageId="12" pageNumber="313">sp. nov</emphasis>
|
|
.
|
|
</taxonomicNameLabel>
|
|
(both from the late Eocene of Florissant,
|
|
<collectingRegion id="E71BF87AFFE2FFEEE135A341CDDB752B" box="[816,923,295,321]" country="United States of America" name="Colorado" pageId="12" pageNumber="313">Colorado</collectingRegion>
|
|
,
|
|
<collectingCountry id="5DC87608FFE2FFEEE1A3A341CDA3752B" box="[934,995,295,321]" name="United States of America" pageId="12" pageNumber="313">USA</collectingCountry>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="25603698FFE2FFEEE2C2A32CCAFE750F" blockId="12.[151,1437,150,574]" box="[199,1214,330,357]" pageId="12" pageNumber="313">
|
|
<emphasis id="17ABEA8AFFE2FFEEE2C2A32CCF12750E" bold="true" box="[199,338,330,356]" pageId="12" pageNumber="313">Occurrence</emphasis>
|
|
. The Eocene: early Ypresian of Europe and late Priabonian of North America.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6DC56513FFE2FFEEE2C2A309CD957654" pageId="12" pageNumber="313" type="discussion">
|
|
<paragraph id="25603698FFE2FFEEE2C2A309CB6C75C7" blockId="12.[151,1437,150,574]" pageId="12" pageNumber="313">
|
|
<emphasis id="17ABEA8AFFE2FFEEE2C2A309CF7B75E3" bold="true" box="[199,315,367,393]" pageId="12" pageNumber="313">Remarks.</emphasis>
|
|
We could not examine the
|
|
<typeStatus id="FA64883AFFE2FFEEE06FA309CC8C75E3" box="[618,716,367,393]" pageId="12" pageNumber="313" type="holotype">holotype</typeStatus>
|
|
of
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE0F4A309CDD275E3" authorityName="Schluter" authorityYear="1982" box="[753,914,367,393]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="moleriensis">
|
|
<emphasis id="17ABEA8AFFE2FFEEE0F4A309CDD275E3" box="[753,914,367,393]" italics="true" pageId="12" pageNumber="313">C. moleriensis</emphasis>
|
|
</taxonomicName>
|
|
which is apparently missing from the Fur Museum (
|
|
<collectingCountry id="5DC87608FFE2FFEEE2E4A3F5CF0D75C4" box="[225,333,403,430]" name="Denmark" pageId="12" pageNumber="313">Denmark</collectingCountry>
|
|
) where it was deposited according to the original description (Bo Schultz, pers. comm.).
|
|
</paragraph>
|
|
<paragraph id="25603698FFE2FFEEE2C2A3D1CD957654" blockId="12.[151,1437,150,574]" pageId="12" pageNumber="313">
|
|
The forewing venation of
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE3F1A3D1CCD375BB" authorityName="Schluter" authorityYear="1982" box="[500,659,439,465]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE3F1A3D1CCD375BB" box="[500,659,439,465]" italics="true" pageId="12" pageNumber="313">Cimbrochrysa</emphasis>
|
|
</taxonomicName>
|
|
is most similar to that of
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE1C5A3D1CA7C75BB" authorityName="Makarkin" authorityYear="2014" box="[960,1084,439,465]" class="Insecta" family="Chrysopidae" genus="Asiachrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE1C5A3D1CA7C75BB" box="[960,1084,439,465]" italics="true" pageId="12" pageNumber="313">Asiachrysa</emphasis>
|
|
</taxonomicName>
|
|
from the Eocene Tadushi Formation of the Russian Far East. These genera differ only in small details and are probably closely related. Green lacewings with venation similar to
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE021A399CC837673" authorityName="Schluter" authorityYear="1982" box="[548,707,511,537]" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE021A399CC837673" box="[548,707,511,537]" italics="true" pageId="12" pageNumber="313">Cimbrochrysa</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="E2DF4D1BFFE2FFEEE0FEA399CD377673" authorityName="Makarkin" authorityYear="2014" box="[763,887,511,537]" class="Insecta" family="Chrysopidae" genus="Asiachrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFE2FFEEE0FEA399CD377673" box="[763,887,511,537]" italics="true" pageId="12" pageNumber="313">Asiachrysa</emphasis>
|
|
</taxonomicName>
|
|
were therefore distributed in Eocene throughout the Holarctic (
|
|
<emphasis id="17ABEA8AFFE2FFEEE33CA045CF147656" box="[313,340,547,572]" italics="true" pageId="12" pageNumber="313">i.e</emphasis>
|
|
., Europe, Far Eastern Asia, and western North America).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |