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<document id="F122DC50CABE2FD1D7AF7C8A82E7F0C2" ID-DOI="10.24199/j.mmv.2006.63.17" ID-ISSN="1447-2554" ID-Zenodo-Dep="12210410" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718991714234" checkinUser="felipe" docAuthor="Sandford, Andrew C. &amp; Holloway, David J." docDate="2006" docId="9C6887D7FF813F0B6525F92FABA3F8FD" docLanguage="en" docName="MemMusVictoria.63.2.215-255.pdf" docOrigin="Memoirs of Museum Victoria 63 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-63-issue-2-2006/pages-215-255/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Ananaspis , Campbell 1967" docType="treatment" docVersion="2" lastPageNumber="224" masterDocId="6051FFAFFF893F02660FFF86AF6EFF93" masterDocTitle="Early Silurian phacopide trilobites from central Victoria, Australia" masterLastPageNumber="255" masterPageNumber="215" pageNumber="223" updateTime="1719002137797" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D1E8AE56A65F1E54CBF1FA8F9852AE62">Early Silurian phacopide trilobites from central Victoria, Australia</mods:title>
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<mods:namePart id="E61A2900E9CD3EB6B33CA3E2AB9190A4">Sandford, Andrew C.</mods:namePart>
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<mods:namePart id="CA72D869A3A54014856E71C7DF0E7CC7">Holloway, David J.</mods:namePart>
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<mods:title id="797EDF8EFF9AA24F048679933FD6C53C">Memoirs of Museum Victoria</mods:title>
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<mods:date id="80A06330386604C636E479D475200F23">2006</mods:date>
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<treatment id="9C6887D7FF813F0B6525F92FABA3F8FD" LSID="urn:lsid:plazi:treatment:9C6887D7FF813F0B6525F92FABA3F8FD" httpUri="http://treatment.plazi.org/id/9C6887D7FF813F0B6525F92FABA3F8FD" lastPageId="9" lastPageNumber="224" pageId="8" pageNumber="223">
<subSubSection id="5CDB654AFF813F0A6525F92FAB51F92C" box="[810,1087,1705,1727]" pageId="8" pageNumber="223" type="nomenclature">
<paragraph id="147E36C1FF813F0A6525F92FAB51F92C" blockId="8.[810,1087,1705,1727]" box="[810,1087,1705,1727]" pageId="8" pageNumber="223">
<heading id="4F3681ADFF813F0A6525F92FAB51F92C" box="[810,1087,1705,1727]" fontSize="9" level="2" pageId="8" pageNumber="223" reason="5">
<taxonomicName id="D3C14D42FF813F0A6525F92FAB51F92C" authority="Campbell, 1967" authorityName=", Campbell" authorityYear="1967" box="[810,1087,1705,1727]" class="Trilobita" family="Phacopidae" genus="Ananaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="8" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF813F0A6525F92FACF7F92D" bold="true" box="[810,921,1705,1726]" italics="true" pageId="8" pageNumber="223">Ananaspis</emphasis>
<bibRefCitation id="70504B30FF813F0A6590F92FAB51F92C" author="Campbell, K. S. W." box="[927,1087,1705,1727]" pageId="8" pageNumber="223" pagination="1 - 68" refId="ref29079" refString="Campbell, K. S. W. 1967. Trilobites of the Henryhouse Formation (Silurian) in Oklahoma. Bulletin of the Oklahoma Geological Survey 115: 1 - 68." type="journal article" year="1967">Campbell, 1967</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CDB654AFF813F0B6525F950ABA3F8FD" lastPageId="9" lastPageNumber="224" pageId="8" pageNumber="223" type="materials_examined">
<paragraph id="147E36C1FF813F0A6525F950ACCBF8B6" blockId="8.[810,1441,1749,1829]" pageId="8" pageNumber="223">
<materialsCitation id="A4A93C9CFF813F0A6525F950ACCFF8B6" country="Czech Republic" location="Kopanina Formation" municipality="Kolednik" pageId="8" pageNumber="223" specimenCount="1" typeStatus="holotype">
<emphasis id="26B5EAD3FF813F0A6525F950ACD6F978" box="[810,952,1750,1771]" italics="true" pageId="8" pageNumber="223">
<typeStatus id="CB7A8863FF813F0A6525F950AC34F978" box="[810,858,1750,1771]" pageId="8" pageNumber="223">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="D3C14D42FF813F0A65CBF953AA40F978" authority="Barrande, 1846" authorityName="Barrande" authorityYear="1846" box="[964,1326,1749,1771]" class="Trilobita" family="Phacopidae" genus="Phacops" kingdom="Animalia" order="Phacopida" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="fecundus">
<emphasis id="26B5EAD3FF813F0A65CBF953ABEBF979" box="[964,1157,1749,1770]" italics="true" pageId="8" pageNumber="223">Phacops fecundus</emphasis>
<bibRefCitation id="70504B30FF813F0A629DF953AA40F978" author="Barrande, J." box="[1170,1326,1749,1771]" pageId="8" pageNumber="223" refId="ref28892" refString="Barrande, J. 1846. Notice preliminaire sur le Systeme silurien et les trilobites de Boheme. C. L. Hirschfeld: Leipzig. vi + 97 pp." type="book" year="1846">Barrande, 1846</bibRefCitation>
</taxonomicName>
from the
<location id="111E601AFF813F0A6525F974AC94F89B" LSID="urn:lsid:plazi:treatment:9C6887D7FF813F0B6525F92FABA3F8FD:111E601AFF813F0A6525F974AC94F89B" box="[810,1018,1778,1800]" country="Czech Republic" municipality="Kolednik" name="Kopanina Formation" pageId="8" pageNumber="223">Kopanina Formation</location>
(Ludlow),
<collectingMunicipality id="F41AACBBFF813F0A626AF974ABA9F89B" box="[1125,1223,1778,1800]" pageId="8" pageNumber="223">Koledník</collectingMunicipality>
,
<collectingCountry id="6CD67651FF813F0A62C0F974AA43F89B" box="[1231,1325,1778,1800]" name="Czech Republic" pageId="8" pageNumber="223">Bohemia</collectingCountry>
, by original designation
</materialsCitation>
.
</paragraph>
<paragraph id="147E36C1FF813F0B6525F8BAAD87F895" blockId="8.[810,1442,1852,1903]" lastBlockId="9.[146,777,1357,1798]" lastPageId="9" lastPageNumber="224" pageId="8" pageNumber="223">
<emphasis id="26B5EAD3FF813F0A6525F8BAACE3F8C2" box="[810,909,1852,1873]" italics="true" pageId="8" pageNumber="223">Diagnosis</emphasis>
. Glabella moderately to strongly convex (sag., tr.), vertical or slightly overhanging anteriorly in dorsal view. Composite lobe expanding strongly forward, maximum width twice width at L1 or a little more. L2 commonly significantly shorter (exsag.) than L3. Eye of moderate to large size, lower edge situated in or slightly above lateral border furrow anteriorly and distant from lateral border furrow posteriorly, visual surface lacking strongly raised sclera (may be slightly thickened dorsally). Palpebral area as high as or higher than palpebral lobe. Fixigenal portion of lateral border furrow deep and continuous with posterior border furrow. Vincular furrow shallow to moderately impressed medially, commonly rather weakly notched laterally. Glabellar sculpture of bimodal tubercles lacking perforations and with superimposed and interspersed granules, doublure finely and densely granulate. Hypostome with short (sag.) posterior border having 3 angular points on margin. Pygidial pleural furrows deep and wide (exsag.), interpleural furrows distinct.
</paragraph>
<caption id="40BE6649FF803F0B669CFB46ADFAFB7E" ID-DOI="http://doi.org/10.5281/zenodo.12210424" ID-Zenodo-Dep="12210424" httpUri="https://zenodo.org/record/12210424/files/figure.png" pageId="9" pageNumber="224" startId="9.[147,207,1216,1235]" targetBox="[267,1328,229,1200]" targetPageId="9" targetType="figure">
<paragraph id="147E36C1FF803F0B669CFB46ADFAFB7E" blockId="9.[147,1449,1216,1262]" pageId="9" pageNumber="224">
Figure 5. AD,
<taxonomicName id="D3C14D42FF803F0B6710FB46AEEBFB40" authorityName=", Ramskold and Werdelin" authorityYear="1991" box="[287,389,1216,1235]" class="Trilobita" family="Phacopidae" genus="Acernaspis" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B6710FB46AEEBFB40" box="[287,389,1216,1235]" italics="true" pageId="9" pageNumber="224">Acernaspis</emphasis>
</taxonomicName>
? sp., NMV P139804, cephalon, × 4, from PL385, Costerfield. EG,
<taxonomicName id="D3C14D42FF803F0B65E9FB46AB21FB40" authorityName="Hawle and Corda" authorityYear="1847" box="[998,1103,1216,1235]" class="Trilobita" family="Phacopidae" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="family">Phacopidae</taxonomicName>
gen.indet.3, NMV P139353, enrolled exoskeleton from PL1369, Springfield; E, × 4.0; F, × 4.5.
</paragraph>
</caption>
<paragraph id="147E36C1FF803F0B669DF89BABA3F8FD" blockId="9.[146,777,1821,1903]" lastBlockId="9.[814,1446,1357,1903]" pageId="9" pageNumber="224">
<emphasis id="26B5EAD3FF803F0B669DF89BAF9EF8A1" box="[146,240,1821,1842]" italics="true" pageId="9" pageNumber="224">Remarks.</emphasis>
This diagnosis distinguishes
<taxonomicName id="D3C14D42FF803F0B6403F89BAD1DF8A1" authorityName=", Campbell" authorityYear="1967" box="[524,627,1821,1842]" class="Trilobita" family="Phacopidae" genus="Ananaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B6403F89BAD1DF8A1" box="[524,627,1821,1842]" italics="true" pageId="9" pageNumber="224">Ananaspis</emphasis>
</taxonomicName>
from a number of closely related genera recognised since the work of
<bibRefCitation id="70504B30FF803F0B64A7F8BDAFB9F8FC" author="Campbell, K. S. W." pageId="9" pageNumber="224" pagination="1 - 68" refId="ref29079" refString="Campbell, K. S. W. 1967. Trilobites of the Henryhouse Formation (Silurian) in Oklahoma. Bulletin of the Oklahoma Geological Survey 115: 1 - 68." type="journal article" year="1967">Campbell (1967)</bibRefCitation>
.
<taxonomicName id="D3C14D42FF803F0B66ECF8DFAE9EF8FD" authority="Sandford, 2002" authorityName="Sandford" authorityYear="2002" box="[227,496,1881,1903]" class="Trilobita" family="Phacopidae" genus="Echidnops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B66ECF8DFAE22F8FD" box="[227,332,1881,1902]" italics="true" pageId="9" pageNumber="224">Echidnops</emphasis>
<bibRefCitation id="70504B30FF803F0B675BF8DFAE9EF8FD" author="Sandford, A. C." box="[340,496,1881,1903]" pageId="9" pageNumber="224" pagination="1 - 31" refId="ref31816" refString="Sandford, A. C. 2002. Systematics, biostratigraphy and palaeoenvironments of Echidnops, a new genus of trilobite from the late Silurian - Early Devonian of south-eastern Australia: Phacopidae of Victoria, Part 1. Memoirs of the Association of Australasian Palaeontologists 27: 1 - 31." type="journal article" year="2002">Sandford, 2002</bibRefCitation>
</taxonomicName>
is the closest to
<taxonomicName id="D3C14D42FF803F0B64AEF8DFAC66F8FD" authorityName=", Campbell" authorityYear="1967" box="[673,776,1881,1902]" class="Trilobita" family="Phacopidae" genus="Ananaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B64AEF8DFAC66F8FD" box="[673,776,1881,1902]" italics="true" pageId="9" pageNumber="224">Ananaspis</emphasis>
</taxonomicName>
but differs not only in having characteristic occipital and thoracic axial spines but also a much more deeply incised vincular furrow medially.
<taxonomicName id="D3C14D42FF803F0B6594FA01AB82FA0F" authority="Maksimova, 1972" authorityName="Maksimova" authorityYear="1972" box="[923,1260,1415,1437]" class="Trilobita" family="Phacopidae" genus="Paciphacops" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B6594FA01AB75FA0F" box="[923,1051,1415,1436]" italics="true" pageId="9" pageNumber="224">Paciphacops</emphasis>
<bibRefCitation id="70504B30FF803F0B6222FA01AB82FA0F" author="Maksimova, Z. A." box="[1069,1260,1415,1437]" pageId="9" pageNumber="224" pagination="78 - 83" refId="ref31052" refString="Maksimova, Z. A. 1972. Novye devonskie trilobity Phacopoidea. Paleontologicheskii Zhurnal (1972): 88 - 94. [English translation published 1972 in Paleontological Journal 6: 78 - 83.]" type="journal article" year="1972">Maksimova, 1972</bibRefCitation>
</taxonomicName>
is distinguished especially by the strongly raised sclera on the visual surface and the perforate glabellar tubercles, but in addition L1 is more depressed medially, the glabella is more raised and subquadrate in anterior profile, and the vincular furrow is deep medially.
<taxonomicName id="D3C14D42FF803F0B6521F99FAB28F9BD" authority="Chlupac, 1972" authorityName="Chlupac" authorityYear="1972" box="[814,1094,1561,1583]" class="Trilobita" family="Phacopidae" genus="Lochkovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B6521F99FACC9F9BD" box="[814,935,1561,1582]" italics="true" pageId="9" pageNumber="224">Lochkovella</emphasis>
<bibRefCitation id="70504B30FF803F0B65BEF99FAB28F9BD" author="Chlupac, I." box="[945,1094,1561,1583]" pageId="9" pageNumber="224" pagination="395 - 401" refId="ref29366" refString="Chlupac, I. 1972. New Silurian and Lower Devonian phacopid trilobites from the Barrandian area (Czechoslovakia). Casopis pro Mineralogii a Geologii 17: 395 - 401." type="journal article" year="1972">Chlupáč, 1972</bibRefCitation>
</taxonomicName>
(see also
<bibRefCitation id="70504B30FF803F0B62BAF99FAA3DF9BD" author="Sandford, A. C." box="[1205,1363,1561,1583]" pageId="9" pageNumber="224" pagination="209 - 222" refId="ref31918" refString="Sandford, A. C. 2004. A revision of Lochkovella Chlupac, 1972 (Arthropoda: Trilobita) and a species from Lilydale and Tyers. Phacopidae of Victoria, Part 5. Proceedings of the Royal Society of Victoria 116: 209 - 222." type="journal article" year="2004">Sandford, 2004</bibRefCitation>
) differs from
<taxonomicName id="D3C14D42FF803F0B656BF9B0ACA5F9D8" authorityName=", Campbell" authorityYear="1967" box="[868,971,1590,1611]" class="Trilobita" family="Phacopidae" genus="Ananaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B656BF9B0ACA5F9D8" box="[868,971,1590,1611]" italics="true" pageId="9" pageNumber="224">Ananaspis</emphasis>
</taxonomicName>
in that the cephalic tuberculation is finer and of more uniform size, the eye is situated very low on the cheek with its lower edge indenting the lateral cephalic border, the hypostome has a long (sag.) posterior border with five marginal denticles, the pygidial interpleural furrows are not as deep, and the pygidial granulation is coarser. Similar distinctions can be made with
<taxonomicName id="D3C14D42FF803F0B65AAF962ABACF969" authority="Erben, 1952" authorityName="Erben" authorityYear="1952" box="[933,1218,1764,1786]" class="Trilobita" family="Phacopidae" genus="Nephranomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B65AAF962AB57F96A" box="[933,1081,1764,1785]" italics="true" pageId="9" pageNumber="224">Nephranomma</emphasis>
<bibRefCitation id="70504B30FF803F0B624BF962ABACF969" author="Erben, H. K." box="[1092,1218,1764,1786]" pageId="9" pageNumber="224" pagination="150 - 362" refId="ref29810" refString="Erben, H. K. 1952. Trilobiten aus dem alteren Hercyn (Unterdevon) des Unterharzes. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 94: 150 - 362, pls 17 - 24." type="journal article" year="1952">Erben, 1952</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="70504B30FF803F0B62F0F962AAF3F969" author="Sandford, A. C." box="[1279,1437,1764,1786]" pageId="9" pageNumber="224" pagination="309 - 329" refId="ref31868" refString="Sandford, A. C. 2003. A revision of Nephranomma Erben, 1952 (Trilobita: Phacopidae), with new species from the Lower Devonian of Victoria, Australia: Phacopidae of Victoria, Part 2. Special Papers in Palaeontology 70: 309 - 329." type="journal article" year="2003">Sandford, 2003</bibRefCitation>
) which further differs from
<taxonomicName id="D3C14D42FF803F0B625EF884ABD6F884" authorityName=", Campbell" authorityYear="1967" box="[1105,1208,1794,1815]" class="Trilobita" family="Phacopidae" genus="Ananaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B625EF884ABD6F884" box="[1105,1208,1794,1815]" italics="true" pageId="9" pageNumber="224">Ananaspis</emphasis>
</taxonomicName>
in lacking a vincular furrow medially and in having a distinctive scaly sculpture on the cephalic doublure. See the remarks on
<taxonomicName id="D3C14D42FF803F0B62EBF8BBAA74F8C1" authorityName="Sandford &amp; Holloway" authorityYear="2006" box="[1252,1306,1853,1874]" class="Trilobita" family="Phacopidae" genus="Ivops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="26B5EAD3FF803F0B62EBF8BBAA74F8C1" box="[1252,1306,1853,1874]" italics="true" pageId="9" pageNumber="224">Ivops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D8657A8FF803F0B632DF8BAAAEEF8C2" box="[1314,1408,1852,1873]" pageId="9" pageNumber="224" rank="genus">gen. nov.</taxonomicNameLabel>
for comparison of that genus with
<taxonomicName id="D3C14D42FF803F0B626FF8DFABA9F8FD" authorityName=", Campbell" authorityYear="1967" box="[1120,1223,1881,1902]" class="Trilobita" family="Phacopidae" genus="Ananaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="26B5EAD3FF803F0B626FF8DFABA9F8FD" box="[1120,1223,1881,1902]" italics="true" pageId="9" pageNumber="224">Ananaspis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>