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<mods:title id="AC2905DCD07F012F0134369AC349838D">A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae)</mods:title>
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<heading id="8A2B81988F1FFFE0FF6CF873FCDB2BF0" bold="true" box="[151,820,1988,2014]" fontSize="11" level="1" pageId="4" pageNumber="495" reason="1">
<taxonomicName id="16DC4D778F1FFFE0FF6CF873FCDB2BF0" ID-CoL="4MNVW" authority="Kimmins 1956" authorityName="Kimmins" authorityYear="1956" box="[151,820,1988,2014]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="495" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F1FFFE0FF6CF873FCDB2BF0" bold="true" box="[151,820,1988,2014]" pageId="4" pageNumber="495">
<emphasis id="E3A8EAE68F1FFFE0FF6CF873FEFC2BF0" bold="true" box="[151,275,1988,2014]" italics="true" pageId="4" pageNumber="495">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F1FFFE0FEDFF873FE592BF0" bold="true" box="[292,438,1988,2014]" italics="true" pageId="4" pageNumber="495">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F1FFFE0FE3CF873FD982BF0" bold="true" box="[455,631,1988,2014]" italics="true" pageId="4" pageNumber="495">cylindroculum</emphasis>
<bibRefCitation id="B54D4B058F1FFFE0FD85F873FCDB2BF0" author="Kimmins" box="[638,820,1988,2014]" pageId="4" pageNumber="495" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880." type="journal article" year="1956">Kimmins 1956</bibRefCitation>
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(
<figureCitation id="49E72A718F1EFFE1FF64FF20FEFF2C9E" box="[159,272,151,176]" captionStart-0="FIGURES 1 10" captionStart-1="FIGURES 11 20" captionStart-2="FIGURES 21 29" captionStart-3="FIGURES 30 32" captionStart-4="FIGURES 33 37" captionStart-5="FIGURES 38 42" captionStartId-0="6.[151,264,1903,1926]" captionStartId-1="7.[151,264,1612,1635]" captionStartId-2="8.[151,264,1932,1955]" captionStartId-3="9.[151,264,1845,1868]" captionStartId-4="10.[151,264,1855,1878]" captionStartId-5="11.[151,264,1844,1867]" captionTargetBox-0="[154,1436,277,1881]" captionTargetBox-1="[160,1415,207,1574]" captionTargetBox-2="[212,1367,193,1904]" captionTargetBox-3="[200,1361,197,1812]" captionTargetBox-4="[170,1416,193,1834]" captionTargetBox-5="[170,1416,193,1823]" captionTargetId-0="figure@6.[151,1436,265,1883]" captionTargetId-1="figure@7.[151,1436,193,1591]" captionTargetId-2="figure@8.[212,1367,193,1905]" captionTargetId-3="figure@9.[195,1372,193,1826]" captionTargetId-4="figure@10.[170,1417,193,1835]" captionTargetId-5="figure@11.[170,1417,193,1824]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionText-0="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." captionText-1="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." captionText-2="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." captionText-3="FIGURES 30 32. Procloeon (Oculogaster) cylindroculum from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." captionText-4="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." captionText-5="FIGURES 38 42. Procloeon (Oculogaster) cylindroculum, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl. l., lateral ocellus; ocl. m., median ocellus." httpUri-0="https://zenodo.org/record/271361/files/figure.png" httpUri-1="https://zenodo.org/record/271362/files/figure.png" httpUri-2="https://zenodo.org/record/271363/files/figure.png" httpUri-3="https://zenodo.org/record/271364/files/figure.png" httpUri-4="https://zenodo.org/record/271365/files/figure.png" httpUri-5="https://zenodo.org/record/271366/files/figure.png" pageId="5" pageNumber="496">Figs 142</figureCitation>
)
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<paragraph id="D16336F48F1EFFE1FF6CFF57FB012CD8" blockId="5.[151,1436,224,430]" box="[151,1262,224,246]" pageId="5" pageNumber="496">
<treatmentCitationGroup id="F1CC11DA8F1EFFE1FF6CFF57FB012CD8" box="[151,1262,224,246]" pageId="5" pageNumber="496">
?
<taxonomicName id="16DC4D778F1EFFE1FF59FF57FD8E2CD8" authority="Barnard 1932: 217" authorityName="Barnard" authorityPageNumber="217" authorityYear="1932" box="[162,609,224,246]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="africanum">
<emphasis id="E3A8EAE68F1EFFE1FF59FF57FE792CD8" box="[162,406,224,246]" italics="true" pageId="5" pageNumber="496">Austrocloeon africanum</emphasis>
:
<treatmentCitation id="507D10E58F1EFFE1FE5FFF57FD8E2CD8" author="Barnard" box="[420,609,224,246]" page="217" pageId="5" pageNumber="496" year="1932">
<bibRefCitation id="B54D4B058F1EFFE1FE5FFF57FDDD2CD8" author="Barnard" box="[420,562,224,246]" pageId="5" pageNumber="496" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. http: // dx. doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
: 217
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(♀ imago, larva) (non
<taxonomicName id="16DC4D778F1EFFE1FCB7FF57FB0D2CD8" authority="Esben-Petersen 1913" authorityName="Esben-Petersen" authorityYear="1913" box="[844,1250,224,246]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="africanum">
<emphasis id="E3A8EAE68F1EFFE1FCB7FF57FBEC2CD8" box="[844,1027,224,246]" italics="true" pageId="5" pageNumber="496">Cloeon africanum</emphasis>
<bibRefCitation id="B54D4B058F1EFFE1FBF1FF57FB0D2CD8" author="Esben-Petersen" box="[1034,1250,224,246]" pageId="5" pageNumber="496" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen 1913</bibRefCitation>
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</paragraph>
<paragraph id="D16336F48F1EFFE1FF6CFF49FCFA2D3A" blockId="5.[151,1436,224,430]" box="[151,789,254,276]" pageId="5" pageNumber="496">
<treatmentCitationGroup id="F1CC11DA8F1EFFE1FF6CFF49FCFA2D3A" box="[151,789,254,276]" pageId="5" pageNumber="496">
?
<taxonomicName id="16DC4D778F1EFFE1FF59FF49FDBB2D3A" ID-CoL="4MNVQ" authority="Kimmins 1956: 866" authorityName="Kimmins" authorityPageNumber="866" authorityYear="1956" box="[162,596,254,276]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="africanum">
<emphasis id="E3A8EAE68F1EFFE1FF59FF49FE962D3A" box="[162,377,254,276]" italics="true" pageId="5" pageNumber="496">Procloeon africanum</emphasis>
:
<treatmentCitation id="507D10E58F1EFFE1FE7DFF49FDBB2D3A" author="Kimmins" box="[390,596,254,276]" page="866" pageId="5" pageNumber="496" year="1956">
<bibRefCitation id="B54D4B058F1EFFE1FE7DFF49FDCE2D3A" author="Kimmins" box="[390,545,254,276]" pageId="5" pageNumber="496" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880." type="journal article" year="1956">Kimmins 1956</bibRefCitation>
: 866
</treatmentCitation>
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;
<treatmentCitation id="507D10E58F1EFFE1FDA5FF49FCFE2D3A" author="Gillies" box="[606,785,254,276]" page="247" pageId="5" pageNumber="496" year="1997">
<bibRefCitation id="B54D4B058F1EFFE1FDA5FF49FD312D3A" author="Gillies" box="[606,734,254,276]" pageId="5" pageNumber="496" refString="Gillies, M. T. (1997) A new species of Procloeon Bengtsson from the forest zone of West Africa (Ephemeroptera, Baetidae). Entomologist's Monthly Magazine, 133, 247 - 250." type="journal article" year="1997">Gillies 1997</bibRefCitation>
: 247
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;
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</paragraph>
<paragraph id="D16336F48F1EFFE1FF6CFEAAFD4C2D7C" blockId="5.[151,1436,224,430]" pageId="5" pageNumber="496">
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<emphasis id="E3A8EAE68F1EFFE1FF6CFEAAFE782D1D" box="[151,407,285,307]" italics="true" pageId="5" pageNumber="496">Procloeon cylindroculum</emphasis>
<treatmentCitation id="507D10E58F1EFFE1FE64FEAAFD852D1D" author="Kimmins" box="[415,618,285,307]" page="865" pageId="5" pageNumber="496" year="1956">
<bibRefCitation id="B54D4B058F1EFFE1FE64FEAAFDD52D1D" author="Kimmins" box="[415,570,285,307]" pageId="5" pageNumber="496" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880." type="journal article" year="1956">Kimmins 1956</bibRefCitation>
: 865
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(♂ imago);
<treatmentCitation id="507D10E58F1EFFE1FD11FEAAFC592D1C" author="Kimmins" box="[746,950,285,307]" page="341" pageId="5" pageNumber="496" year="1960">
<bibRefCitation id="B54D4B058F1EFFE1FD11FEAAFC6A2D1D" author="Kimmins" box="[746,901,285,307]" pageId="5" pageNumber="496" refString="Kimmins, D. E. (1960) Notes on East African Ephemeroptera with descriptions of new species. Bulletin of the British Museum (Natural History), Entomology, 9 (6), 337 - 355. http: // dx. doi. org / 10.5962 / bhl. part. 27555" type="journal article" year="1960">Kimmins 1960</bibRefCitation>
: 341
</treatmentCitation>
(♀ imago, ♂ and ♀ subimago);
<bibRefCitation id="B54D4B058F1EFFE1FAFFFEAAFA732D1C" author="Kimmins" box="[1284,1436,285,307]" pageId="5" pageNumber="496" refString="Kimmins, D. E. (1971) A list of the type-specimens of Ephemeroptera in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 25 (7), 307 - 324. http: // dx. doi. org / 10.5962 / bhl. part. 19678" type="journal article" year="1971">Kimmins 1971</bibRefCitation>
(
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);
<treatmentCitation id="507D10E58F1EFFE1FEC6FE8BFE022D7F" author="Gillies" box="[317,493,316,338]" page="247" pageId="5" pageNumber="496" year="1997">
<bibRefCitation id="B54D4B058F1EFFE1FEC6FE8BFE552D7C" author="Gillies" box="[317,442,316,338]" pageId="5" pageNumber="496" refString="Gillies, M. T. (1997) A new species of Procloeon Bengtsson from the forest zone of West Africa (Ephemeroptera, Baetidae). Entomologist's Monthly Magazine, 133, 247 - 250." type="journal article" year="1997">Gillies 1997</bibRefCitation>
: 247
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(♀ imago, larva);
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<paragraph id="D16336F48F1EFFE1FF6CFEEDFEF52DA1" blockId="5.[151,1436,224,430]" pageId="5" pageNumber="496">
<treatmentCitationGroup id="F1CC11DA8F1EFFE1FF6CFEEDFEF52DA1" pageId="5" pageNumber="496">
<taxonomicName id="16DC4D778F1EFFE1FF6CFEEDFDBB2D5E" authority="Demoulin 1970: 55" authorityName="Demoulin" authorityPageNumber="55" authorityYear="1970" box="[151,596,346,369]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F1EFFE1FF6CFEEDFE962D5E" box="[151,377,346,368]" italics="true" pageId="5" pageNumber="496">Cloeon cylindroculum</emphasis>
:
<treatmentCitation id="507D10E58F1EFFE1FE72FEECFDBB2D5E" author="Demoulin" box="[393,596,346,369]" page="55" pageId="5" pageNumber="496" year="1970">
<bibRefCitation id="B54D4B058F1EFFE1FE72FEECFDC32D5E" author="Demoulin" box="[393,556,347,369]" pageId="5" pageNumber="496" refString="Demoulin, G. (1970) Ephemeroptera des faunes ethiopienne et malgache. In: Hanstrom, Bertil, Per Brinck &amp; Gustaf Rudebeck (Eds.), South African Animal Life, 14, pp. 24 - 170. [Results of the Lunds University Expedition in 1950 - 1951]" type="journal article" year="1970">Demoulin 1970</bibRefCitation>
: 55
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;
<treatmentCitation id="507D10E58F1EFFE1FD9AFEECFCFA2D5E" author="Gillies" box="[609,789,346,369]" page="155" pageId="5" pageNumber="496" year="1979">
<bibRefCitation id="B54D4B058F1EFFE1FD9AFEECFD0B2D5E" author="Gillies" box="[609,740,347,369]" pageId="5" pageNumber="496" refString="Gillies, M. T. (1979) Polymorphism in the turbinate eyes of an African Cloeon (Ephemeroptera: Baetidae). Entomologist's Monthly Magazine, 114, 155." type="journal article" year="1979">Gillies 1979</bibRefCitation>
: 155
</treatmentCitation>
(♂ imago);
<treatmentCitation id="507D10E58F1EFFE1FC63FEECFBA32D5E" author="Gillies" box="[920,1100,346,369]" page="145" pageId="5" pageNumber="496" year="1980">
<bibRefCitation id="B54D4B058F1EFFE1FC63FEECFBF52D5E" author="Gillies" box="[920,1050,347,369]" pageId="5" pageNumber="496" refString="Gillies, M. T. (1980) An introduction to the study of Cloeon Leach (Baetidae, Ephemeroptera) in West Africa. Bulletin de l'Institute Francaise d'Afrique Noire, Serie A, 42 (1), 135 - 156." type="journal article" year="1980">Gillies 1980</bibRefCitation>
: 145
</treatmentCitation>
(♂ and ♀ imago, larva);
<treatmentCitation id="507D10E58F1EFFE1FAA3FEECFEF92DA1" author="Gillies" page="8" pageId="5" pageNumber="496" year="1985">
<bibRefCitation id="B54D4B058F1EFFE1FAA3FEECFF122DA1" author="Gillies" pageId="5" pageNumber="496" refString="Gillies, M. T. (1985) A preliminary account of the East African species of Cloeon Leach and Rhithrocloeon gen. n. (Ephemeroptera). Aquatic Insects, 7 (1), 1 - 17. http: // dx. doi. org / 10.1080 / 01650428509361193" type="journal article" year="1985">Gillies 1985</bibRefCitation>
: 8
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="D16336F48F1EFFE1FF6CFE2FFCB02D80" blockId="5.[151,1436,224,430]" box="[151,863,408,430]" pageId="5" pageNumber="496">
<treatmentCitationGroup id="F1CC11DA8F1EFFE1FF6CFE2FFCB02D80" box="[151,863,408,430]" pageId="5" pageNumber="496">
?
<taxonomicName id="16DC4D778F1EFFE1FF59FE2FFDCE2D80" authority="Gillies 1997: 248" authorityName="Gillies" authorityPageNumber="248" authorityYear="1997" box="[162,545,408,430]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="sylvicola">
<emphasis id="E3A8EAE68F1EFFE1FF59FE2FFE842D80" box="[162,363,408,430]" italics="true" pageId="5" pageNumber="496">Procloeon sylvicola</emphasis>
<treatmentCitation id="507D10E58F1EFFE1FE8AFE2FFDCE2D80" author="Gillies" box="[369,545,408,430]" page="248" pageId="5" pageNumber="496" year="1997">
<bibRefCitation id="B54D4B058F1EFFE1FE8AFE2FFE1D2D80" author="Gillies" box="[369,498,408,430]" pageId="5" pageNumber="496" refString="Gillies, M. T. (1997) A new species of Procloeon Bengtsson from the forest zone of West Africa (Ephemeroptera, Baetidae). Entomologist's Monthly Magazine, 133, 247 - 250." type="journal article" year="1997">Gillies 1997</bibRefCitation>
: 248
</treatmentCitation>
</taxonomicName>
(♀ imago, ♂ subimago, larva).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F1EFFE9FF6CFE6DFB9D2D83" lastPageId="13" lastPageNumber="504" pageId="5" pageNumber="496" type="materials_examined">
<paragraph id="D16336F48F1EFFE1FF6CFE6DFEA02EE5" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="496">
<emphasis id="E3A8EAE68F1EFFE1FF6CFE6DFE6F2DDD" bold="true" box="[151,384,474,499]" pageId="5" pageNumber="496">Material examined.</emphasis>
<collectingCountry id="A9CB76648F1EFFE1FE7DFE6CFE182DDD" box="[390,503,475,499]" name="Zambia" pageId="5" pageNumber="496">ZAMBIA</collectingCountry>
, River Luangwa near bridge,
<date id="A56210348F1EFFE1FCBBFE6CFC0C2DDD" box="[832,995,475,499]" pageId="5" pageNumber="496" value="2014-08-02" valueMax="2014-08-07" valueMin="2014-08-02">27.VIII.2014</date>
, coll. N. Kluge &amp; L. Sheyko: 3 L-S-I♂, 1 L/S♂, 3 L-S♀; River Zambezi above Victoria Falls,
<date id="A56210348F1EFFE1FCF8FE49FC2D2E38" box="[771,962,510,534]" pageId="5" pageNumber="496" value="2014-08-25" valueMax="2014-08-31" valueMin="2014-08-25">2531.VIII.2014</date>
, coll. N. Kluge &amp; L. Sheyko:
<date id="A56210348F1EFFE1FAE0FE48FAD22E38" box="[1307,1341,511,535]" pageId="5" pageNumber="496">1 I</date>
♀ (filled with embryos),
<date id="A56210348F1EFFE1FEB3FD95FE972E14" box="[328,376,546,571]" pageId="5" pageNumber="496" value="2015-01">15 I</date>
♀ (empty, without embryos).
<collectingCountry id="A9CB76648F1EFFE1FD39FD94FCAF2E15" box="[706,832,547,571]" name="Uganda" pageId="5" pageNumber="496">UGANDA</collectingCountry>
: River Victoria Nile at Bujagali Falls,
<date id="A56210348F1EFFE1FB0BFD95FA772E15" box="[1264,1432,546,571]" pageId="5" pageNumber="496" value="2007-07-06" valueMax="2007-07-10" valueMin="2007-07-06">610.VII.2007</date>
, coll. N. Kluge: 1 L-S-I♀, 1 L♀, 1 L/S♂, 2 S-I♀; Lake Victoria in Jinja,
<date id="A56210348F1EFFE1FC30FDF1FB9C2E70" box="[971,1139,582,606]" pageId="5" pageNumber="496" value="2007-07-06" valueMax="2007-07-10" valueMin="2007-07-06">610.VII.2007</date>
, coll. N. Kluge: 147 I♂, 5 S♂,
<date id="A56210348F1EFFE1FF37FDDCFF002EAC" box="[204,239,619,643]" pageId="5" pageNumber="496">3 I</date>
♀ (filled with embryos), 13 S♀; Entebbe, airport,
<date id="A56210348F1EFFE1FCD6FDDDFC4E2EAC" box="[813,929,618,642]" pageId="5" pageNumber="496" value="2007-09-05">5.IX.2007</date>
, coll. N. Kluge:
<date id="A56210348F1EFFE1FB99FDDCFB6C2EAC" box="[1122,1155,619,643]" pageId="5" pageNumber="496">2 I</date>
♂, 2 S-I♀.
<collectingCountry id="A9CB76648F1EFFE1FAFDFDDCFABD2EAD" box="[1286,1362,619,643]" name="Serbia" pageId="5" pageNumber="496">MALI</collectingCountry>
: River Bafing near Manantali, August
<date id="A56210348F1EFFE1FDF8FD39FD442E89" box="[515,683,654,679]" pageId="5" pageNumber="496" value="1990-02">February 1990</date>
,
<date id="A56210348F1EFFE1FD43FD38FC8C2E89" box="[696,867,654,679]" pageId="5" pageNumber="496" value="1991" valueMax="1992">1991 and 1992</date>
, coll. W. Tobias &amp; D. Tobias: 113 I♀ (partly with embryos), 1 S♂.
</paragraph>
<paragraph id="D16336F48F1EFFE1FF3CFD61FE5C2FC5" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="496">
<emphasis id="E3A8EAE68F1EFFE1FF3CFD61FE522EC1" bold="true" box="[199,445,726,751]" pageId="5" pageNumber="496">
Descriptions.
<emphasis id="E3A8EAE68F1EFFE1FE8AFD60FE5A2EC0" bold="true" box="[369,437,727,750]" italics="true" pageId="5" pageNumber="496">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION. Head with diffusive brownish and colorless areas. Pronotum and mesonotum ocher or brownish with diffusive lighter and darker areas; cuticle of fore protoptera either nearly unicolor, or with darker brown stripes along convex veins and stripes lighter than background along concave veins (
<figureCitation id="49E72A718F1EFFE1FEB6FCF5FE492F75" box="[333,422,834,859]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Fig. 26</figureCitation>
). Metanotum and thoracic pleura with diffusive colorless and brownish areas, thoracic sterna colorless. Legs colorless, with diffusive brown bands in distal part of femur, in proximal part of tibia and in proximal part of tarsus. Abdominal terga with variable maculation; abdominal sterna lighter than terga, either unicolor, or with diffusive lighter and darker areas (
<figureCitation id="49E72A718F1EFFE1FD21FC19FCB22FE9" box="[730,861,942,967]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Figs 2325</figureCitation>
). Caudalii ocher with posterior margin of each 4-th segment brown (
<figureCitation id="49E72A718F1EFFE1FEA9FC65FE4A2FC5" box="[338,421,978,1003]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F1EFFE1FF3CFC40FCB12855" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="496">
HYPODERMAL COLORATION. Mature larva with markings characteristic for winged stages: mature male larva with reddish coloration on posterior part of abdominal tergum VII and terga VIIIIX; middle terga either without hypodermal markings, or with pair of reddish spots on abdominal tergum IV (
<figureCitation id="49E72A718F1EFFE1FBADFB89FB472879" box="[1110,1192,1086,1111]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="5" pageNumber="496">Fig. 35</figureCitation>
); mature female larva with pair of curved stripes along thorax and abdomen (
<figureCitation id="49E72A718F1EFFE1FD06FBD5FCBE2855" box="[765,849,1122,1147]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="5" pageNumber="496">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F1EFFE1FF3CFB30FE8A2B65" blockId="5.[151,1437,474,2011]" pageId="5" pageNumber="496">
<collectionCode id="B7CDAE318F1EFFE1FF3CFB30FECF28B1" box="[199,288,1159,1183]" pageId="5" pageNumber="496">SHAPE</collectionCode>
AND SETATION. Mandibles with incisor and kinetodontium fused more than to middle, with ventral denticle well-developed; incisor of right mandible with 2nd denticle shortest; right prostheca apically with 36 short parallel processes (
<figureCitation id="49E72A718F1EFFE1FE52FB79FDF328C9" box="[425,540,1230,1255]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="5" pageNumber="496">Figs 810</figureCitation>
). Maxillary palp 2-segmented (
<figureCitation id="49E72A718F1EFFE1FC83FB79FC2328C9" box="[888,972,1230,1255]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Fig. 28</figureCitation>
). Labium with glossae and paraglossae of subequal size, palp with median angle moderately projected (as in
<figureCitation id="49E72A718F1EFFE1FC7EFB45FBE82925" box="[901,1031,1266,1291]" captionStart="FIGURES 43 48" captionStartId="16.[151,264,1836,1859]" captionTargetBox="[170,1408,196,1806]" captionTargetId="figure@16.[170,1417,193,1816]" captionTargetPageId="16" captionText="FIGURES 43 48. Procloeon (Oculogaster) album sp. n., larva. 43, labrum, 44, 45, apices of left and right mandibles; 46, maxilla, dorsal view; 47, half of labium, ventral view; 48, the same, dorsal view. Abbreviations: ds 1, ds 2, ds 3, dentisetae; in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271367/files/figure.png" pageId="5" pageNumber="496">Figs 4748</figureCitation>
). Femora with very small spine-like setae on outer and inner sides, with 2 (rarely 3) larger pointed spine-like setae on outer side near apex (
<figureCitation id="49E72A718F1EFFE1FAC1FAA1FA7F2901" box="[1338,1424,1302,1327]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="5" pageNumber="496">Fig. 20</figureCitation>
). Claws moderately long, with two rows of minute denticles in proximal part and two rows of larger denticles reaching mid-length of claw (
<figureCitation id="49E72A718F1EFFE1FE13FAE9FDD22959" box="[488,573,1374,1399]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="5" pageNumber="496">Fig. 19</figureCitation>
).
<collectingCountry id="A9CB76648F1EFFE1FDA8FAE9FD642959" box="[595,651,1374,1399]" name="India" pageId="5" pageNumber="496">Hind</collectingCountry>
protoptera completely absent. Lateral spines on abdominal segments VIX, occasionally also on segments IV and III, with numbers varying on abdominal segments IIIIX as follows: III 01, IV 01,
<date id="A56210348F1EFFE1FEABFA10FE762991" box="[336,409,1447,1471]" pageId="5" pageNumber="496">V 24</date>
,
<date id="A56210348F1EFFE1FE5EFA10FE192991" box="[421,502,1447,1471]" pageId="5" pageNumber="496">VI 34</date>
,
<date id="A56210348F1EFFE1FDF9FA10FDB22991" box="[514,605,1446,1471]" pageId="5" pageNumber="496">VII 35</date>
,
<date id="A56210348F1EFFE1FD92FA10FD222991" box="[617,717,1446,1471]" pageId="5" pageNumber="496">VIII 57</date>
,
<date id="A56210348F1EFFE1FD22FA10FCC02991" box="[729,815,1446,1471]" pageId="5" pageNumber="496">IX 57.</date>
Posterolateral spines present on segments IIIX, being minute on anterior segments. Posterior margin of abdominal tergum I with few small sparse spine-like pointed denticles; posterior margins of terga IIX with larger spine-like pointed denticles of unequal length; tergum X with sublateral denticles longest (
<figureCitation id="49E72A718F1EFFE1FE25F9A5FDDB2A05" box="[478,564,1554,1579]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Fig. 21</figureCitation>
; as in
<figureCitation id="49E72A718F1EFFE1FD78F9A5FD382A05" box="[643,727,1554,1579]" captionStart="FIGURES 59 64" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[187,1393,193,1915]" captionTargetId="figure@18.[187,1394,193,1916]" captionTargetPageId="18" captionText="FIGURES 59 64. Procloeon (Oculogaster) album sp. n., larva. 59, paracercus and cercus (with subimaginal cercus inside); 60, distal part of cercus; 61, posterior margin of abdominal tergum X; 62, fragment of abdominal tergum III; 63, fragment of abdominal sternum VI; 64, tergalius VI (arrow shows additional middle rib) (59, 60, holotype)." httpUri="https://zenodo.org/record/271369/files/figure.png" pageId="5" pageNumber="496">Fig. 61</figureCitation>
). Abdominal terga with short scales in wide semilunar or Wshaped non-operculated sockets, with sparse simple fine setae (similar to
<figureCitation id="49E72A718F1EFFE1FC29F981FBC92A61" box="[978,1062,1590,1615]" captionStart="FIGURES 59 64" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[187,1393,193,1915]" captionTargetId="figure@18.[187,1394,193,1916]" captionTargetPageId="18" captionText="FIGURES 59 64. Procloeon (Oculogaster) album sp. n., larva. 59, paracercus and cercus (with subimaginal cercus inside); 60, distal part of cercus; 61, posterior margin of abdominal tergum X; 62, fragment of abdominal tergum III; 63, fragment of abdominal sternum VI; 64, tergalius VI (arrow shows additional middle rib) (59, 60, holotype)." httpUri="https://zenodo.org/record/271369/files/figure.png" pageId="5" pageNumber="496">Fig. 62</figureCitation>
). Posterior margins of abdominal sterna IV smooth, posterior margins of sterna VIIX and paraprocts with spine-like pointed denticles of unequal length. Abdominal sterna with sparse scales and sparse fine setae; in lateral areas of sterna both simple and bifid fine setae vary from small to very long, located sparsely and irregularly, not forming rows or groups (as in
<figureCitation id="49E72A718F1EFFE1FAC6F915FA602A95" box="[1341,1423,1698,1723]" captionStart="FIGURES 59 64" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[187,1393,193,1915]" captionTargetId="figure@18.[187,1394,193,1916]" captionTargetPageId="18" captionText="FIGURES 59 64. Procloeon (Oculogaster) album sp. n., larva. 59, paracercus and cercus (with subimaginal cercus inside); 60, distal part of cercus; 61, posterior margin of abdominal tergum X; 62, fragment of abdominal tergum III; 63, fragment of abdominal sternum VI; 64, tergalius VI (arrow shows additional middle rib) (59, 60, holotype)." httpUri="https://zenodo.org/record/271369/files/figure.png" pageId="5" pageNumber="496">Fig. 63</figureCitation>
). Tergalius I with costal rib very short and lacking denticles; tergalii IIVI with 25 denticles on apex of costal rib, tergalius VII with 410 denticles on costal rib; dorsal lamella present either on tergalii IVI (
<figureCitation id="49E72A718F1EFFE1FB2BF95DFAD72B2D" box="[1232,1336,1770,1795]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="5" pageNumber="496">Figs 17</figureCitation>
), or IV (
<figureCitation id="49E72A718F1EFFE1FF64F8B9FECD2B09" box="[159,290,1806,1831]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="5" pageNumber="496">Figs 1117</figureCitation>
). In distal part of cercus spine on lateral side of each segment subequal or 1.5 times longer than next segment (
<figureCitation id="49E72A718F1EFFE1FEFFF885FEB72B65" box="[260,344,1842,1867]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F1EFFE2FF3CF8E1FD0D2CFB" blockId="5.[151,1437,474,2011]" lastBlockId="6.[151,1264,151,213]" lastPageId="6" lastPageNumber="497" pageId="5" pageNumber="496">
<emphasis id="E3A8EAE68F1EFFE1FF3CF8E1FEAD2B41" bold="true" box="[199,322,1878,1903]" italics="true" pageId="5" pageNumber="496">Subimago.</emphasis>
CUTICULAR COLORATION. Head colorless. Pronotum with diffusive light brownish and colorless areas. Mesonotum light brownish with sutures darker brown, with large colorless areas occupying anterior part of submedioscutum and posterior scutal protuberance (
<figureCitation id="49E72A718F1EFFE1FC5CF829FC112B99" box="[935,1022,1950,1975]" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="5" pageNumber="496">Fig. 27</figureCitation>
). Thoracic pleura at most colorless, with some sclerites light brownish (
<figureCitation id="49E72A718F1EFFE1FDC3F875FD602BF5" box="[568,655,1986,2011]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="5" pageNumber="496">Fig. 37</figureCitation>
). Legs nearly colorless. Abdomen very light brownish with pair of diffusive darker triangular maculae adjacent to posterior margin of each tergum. Caudalii colorless. HYPODERMAL COLORATION.
<collectingCountry id="A9CB76648F1DFFE2FDAAFF0BFD9F2CFA" box="[593,624,188,212]" name="American Samoa" pageId="6" pageNumber="497">As</collectingCountry>
in imago.
</paragraph>
<caption id="85A3667C8F1DFFE2FF6CF8D8FBB32BED" httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="6" pageNumber="497" targetBox="[154,1436,277,1881]" targetPageId="6">
<paragraph id="D16336F48F1DFFE2FF6CF8D8FBB32BED" blockId="6.[151,1436,1903,1987]" pageId="6" pageNumber="497">
<emphasis id="E3A8EAE68F1DFFE2FF6CF8D8FEA62BAB" bold="true" box="[151,329,1903,1926]" pageId="6" pageNumber="497">FIGURES 110.</emphasis>
<taxonomicName id="16DC4D778F1DFFE2FEA9F8C7FD042BA8" box="[338,747,1904,1926]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="497" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F1DFFE2FEA9F8C7FE542BA8" box="[338,443,1904,1926]" italics="true" pageId="6" pageNumber="497">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F1DFFE2FE37F8C7FDA62BA8" box="[460,585,1904,1926]" italics="true" pageId="6" pageNumber="497">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F1DFFE2FDA1F8C7FD042BA8" box="[602,747,1904,1926]" italics="true" pageId="6" pageNumber="497">cylindroculum</emphasis>
</taxonomicName>
, larva. 17, tergalii of IVII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in1, in2, in3, in4, denticles of incisor; in-V, ventral denticle of incisor; kd1, kd2, kd3, denticles of kinetodontium.
</paragraph>
</caption>
<caption id="85A3667C8F1CFFE3FF6CF9FBFC862A8E" httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="7" pageNumber="498" targetBox="[160,1415,207,1574]" targetPageId="7">
<paragraph id="D16336F48F1CFFE3FF6CF9FBFC862A8E" blockId="7.[151,1436,1612,1696]" pageId="7" pageNumber="498">
<emphasis id="E3A8EAE68F1CFFE3FF6CF9FBFEBB2A4C" bold="true" box="[151,340,1612,1635]" pageId="7" pageNumber="498">FIGURES 1120.</emphasis>
<taxonomicName id="16DC4D778F1CFFE3FEA0F9FBFD1E2A4C" box="[347,753,1612,1634]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="498" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F1CFFE3FEA0F9FBFE2B2A4C" box="[347,452,1612,1634]" italics="true" pageId="7" pageNumber="498">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F1CFFE3FE28F9FBFDBF2A4C" box="[467,592,1612,1634]" italics="true" pageId="7" pageNumber="498">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F1CFFE3FD9BF9FBFD1E2A4C" box="[608,753,1612,1634]" italics="true" pageId="7" pageNumber="498">cylindroculum</emphasis>
</taxonomicName>
, larva. 1117, tergalii of IVII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus).
</paragraph>
</caption>
<paragraph id="D16336F48F1CFFE3FF3CF97AFAEA2B27" blockId="7.[151,1436,1740,2017]" pageId="7" pageNumber="498">TEXTURE. On fore leg of male and female 1st tarsal segment covered mainly with microtrichiae (as tibia), at apex with pointed microlepides; all other tarsal segments entirely covered with pointed microlepides.</paragraph>
<paragraph id="D16336F48F1CFFE8FF3CF8A2FE8A2DDA" blockId="7.[151,1436,1740,2017]" lastBlockId="12.[151,1437,151,2013]" lastPageId="12" lastPageNumber="503" pageId="7" pageNumber="498">
<emphasis id="E3A8EAE68F1CFFE3FF3CF8A2FEB82B03" bold="true" box="[199,343,1812,1837]" italics="true" pageId="7" pageNumber="498">Imago, male</emphasis>
(
<figureCitation id="49E72A718F1CFFE3FE9DF8A3FE062B03" box="[358,489,1812,1837]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="7" pageNumber="498">Figs 3334</figureCitation>
). Head brown with ocher. Turbinate eyes cylindrical or widened at apex; color varies from orange to dark red. Thorax brown; sterna somewhat lighter than terga; membranes of pleura either ocher, or white. Legs pale ocher or whitish. In reared specimen from
<collectingCountry id="A9CB76648F1CFFE3FCC8F8EBFC642B5B" box="[819,907,1884,1909]" name="Zambia" pageId="7" pageNumber="498">Zambia</collectingCountry>
, ratio femur / tibia / tarsal segments on fore leg 100:140:5:72:44:23:16, on middle and hind leg 85:80:35:12:3:14. Middle and hind legs without apical spine of 1st+2nd tarsomere, with single apical spine on primary 3rd tarsomere. Fore wing widest at proximal part; membrane colorless, veins light brown; pterostigma with one oblique vein, no other veins in costal field (among more than
<specimenCount id="C7DAFD7D8F17FFE8FEE9FF2FFE402C9E" box="[274,431,151,176]" pageId="12" pageNumber="503" type="generic">300 specimen</specimenCount>
examined, only
<specimenCount id="C7DAFD7D8F17FFE8FD94FF2FFD142C9E" box="[623,763,151,176]" pageId="12" pageNumber="503" type="generic">4 specimens</specimenCount>
have 2 or 1½ veins in pterostigma of one wing). Marginal intercalary present at least in each 2nd space, namely in spaces located behind convex vein; in spaces located behind concave veins marginal intercalary either absent (
<figureCitation id="49E72A718F17FFE8FCDEFF68FC902CD6" box="[805,895,223,248]" captionStart="FIGURES 30 32" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[200,1361,197,1812]" captionTargetId="figure@9.[195,1372,193,1826]" captionTargetPageId="9" captionText="FIGURES 30 32. Procloeon (Oculogaster) cylindroculum from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271364/files/figure.png" pageId="12" pageNumber="503">Figs 32</figureCitation>
,
<figureCitation id="49E72A718F17FFE8FC77FF57FC442CD6" box="[908,939,224,248]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">33</figureCitation>
), or present.
<collectingCountry id="A9CB76648F17FFE8FBBFFF68FB932CD6" box="[1092,1148,223,248]" name="India" pageId="12" pageNumber="503">Hind</collectingCountry>
wing absent. Abdominal terga IVI translucent, whitish, either unicolor, or with posterior margins slightly darkened; tergum IV either as others (
<figureCitation id="49E72A718F17FFE8FF11FE90FED22D6E" box="[234,317,295,320]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 34</figureCitation>
), or with pair of red or brown spots (
<figureCitation id="49E72A718F17FFE8FD2DFE90FCC62D6E" box="[726,809,295,320]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 33</figureCitation>
), or with unpaired transverse red or brown spot. Tergum VII with anterior part translucent and whitish as previous segments, posterior part brown; terga VIIIX brown. Sterna IVII translucent, whitish, either unicolor, or with posterior margin darkened by brown. Sterna VIII, IX, styligers and gonostyli ocher-brownish. Median projection of penis semicircular (
<figureCitation id="49E72A718F17FFE8FBBCFE23FB702D82" box="[1095,1183,404,429]" captionStart="FIGURES 30 32" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[200,1361,197,1812]" captionTargetId="figure@9.[195,1372,193,1826]" captionTargetPageId="9" captionText="FIGURES 30 32. Procloeon (Oculogaster) cylindroculum from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271364/files/figure.png" pageId="12" pageNumber="503">Fig. 30</figureCitation>
). Cerci either entirely colorless (
<figureCitation id="49E72A718F17FFE8FEF5FE00FE882DFE" box="[270,359,439,464]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 34</figureCitation>
), or with dark bands formed by brown hypodermal pigmentation in distal part of each 4th segment (
<figureCitation id="49E72A718F17FFE8FEFFFE6BFEB72DDA" box="[260,344,476,501]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 33</figureCitation>
).
</paragraph>
<caption id="85A3667C8F13FFECFF6CF83BFDE02BCE" httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="8" pageNumber="499" targetBox="[212,1367,193,1904]" targetPageId="8">
<paragraph id="D16336F48F13FFECFF6CF83BFDE02BCE" blockId="8.[151,1436,1932,2016]" pageId="8" pageNumber="499">
<emphasis id="E3A8EAE68F13FFECFF6CF83BFEBA2B8C" bold="true" box="[151,341,1932,1955]" pageId="8" pageNumber="499">FIGURES 2129.</emphasis>
<taxonomicName id="16DC4D778F13FFECFEA7F83AFD1C2B8D" box="[348,755,1933,1955]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="499" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F13FFECFEA7F83AFE2A2B8D" box="[348,453,1933,1955]" italics="true" pageId="8" pageNumber="499">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F13FFECFE2EF83AFDBD2B8D" box="[469,594,1933,1955]" italics="true" pageId="8" pageNumber="499">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F13FFECFD99F83AFD1C2B8D" box="[610,755,1933,1955]" italics="true" pageId="8" pageNumber="499">cylindroculum</emphasis>
</taxonomicName>
, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 2325, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum.
</paragraph>
</caption>
<caption id="85A3667C8F12FFEDFF6CF882FB232BCB" httpUri="https://zenodo.org/record/271364/files/figure.png" pageId="9" pageNumber="500" targetBox="[200,1361,197,1812]" targetPageId="9">
<paragraph id="D16336F48F12FFEDFF6CF882FB232BCB" blockId="9.[151,1436,1845,2021]" pageId="9" pageNumber="500">
<emphasis id="E3A8EAE68F12FFEDFF6CF882FEBB2B65" bold="true" box="[151,340,1845,1868]" pageId="9" pageNumber="500">FIGURES 3032.</emphasis>
<taxonomicName id="16DC4D778F12FFEDFEA6F882FD1A2B65" box="[349,757,1845,1867]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="500" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F12FFEDFEA6F882FE292B65" box="[349,454,1845,1867]" italics="true" pageId="9" pageNumber="500">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F12FFEDFE2DF882FDBC2B65" box="[470,595,1845,1867]" italics="true" pageId="9" pageNumber="500">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F12FFEDFD9FF882FD1A2B65" box="[612,757,1845,1867]" italics="true" pageId="9" pageNumber="500">cylindroculum</emphasis>
</taxonomicName>
from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i.e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger.
</paragraph>
</caption>
<caption id="85A3667C8F11FFEEFF6CF888FB6D2BBD" httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="10" pageNumber="501" targetBox="[170,1416,193,1834]" targetPageId="10">
<paragraph id="D16336F48F11FFEEFF6CF888FB6D2BBD" blockId="10.[151,1435,1855,1939]" pageId="10" pageNumber="501">
<emphasis id="E3A8EAE68F11FFEEFF6CF888FEB92B7B" bold="true" box="[151,342,1855,1878]" pageId="10" pageNumber="501">FIGURES 3337.</emphasis>
<taxonomicName id="16DC4D778F11FFEEFEA5F8F7FD192B78" box="[350,758,1856,1878]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="501" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F11FFEEFEA5F8F7FE282B78" box="[350,455,1856,1878]" italics="true" pageId="10" pageNumber="501">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F11FFEEFE2CF8F7FDBB2B78" box="[471,596,1856,1878]" italics="true" pageId="10" pageNumber="501">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F11FFEEFD9EF8F7FD192B78" box="[613,758,1856,1878]" italics="true" pageId="10" pageNumber="501">cylindroculum</emphasis>
</taxonomicName>
. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 3537, specimens from Zambia; 34, specimen from Uganda).
</paragraph>
</caption>
<caption id="85A3667C8F10FFEFFF6CF883FDB32BE8" httpUri="https://zenodo.org/record/271366/files/figure.png" pageId="11" pageNumber="502" targetBox="[170,1416,193,1823]" targetPageId="11">
<paragraph id="D16336F48F10FFEFFF6CF883FDB32BE8" blockId="11.[151,1436,1844,1990]" pageId="11" pageNumber="502">
<emphasis id="E3A8EAE68F10FFEFFF6CF883FEB92B64" bold="true" box="[151,342,1844,1867]" pageId="11" pageNumber="502">FIGURES 3842.</emphasis>
<taxonomicName id="16DC4D778F10FFEFFEA5F882FD182B65" box="[350,759,1845,1867]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="502" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="E3A8EAE68F10FFEFFEA5F882FE282B65" box="[350,455,1845,1867]" italics="true" pageId="11" pageNumber="502">Procloeon</emphasis>
(
<emphasis id="E3A8EAE68F10FFEFFE2CF882FDBB2B65" box="[471,596,1845,1867]" italics="true" pageId="11" pageNumber="502">Oculogaster</emphasis>
)
<emphasis id="E3A8EAE68F10FFEFFD9DF882FD182B65" box="[614,759,1845,1867]" italics="true" pageId="11" pageNumber="502">cylindroculum</emphasis>
</taxonomicName>
, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl.l., lateral ocellus; ocl.m., median ocellus.
</paragraph>
</caption>
<paragraph id="D16336F48F17FFE8FF3CFDB7FA8F2F3B" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CFDB7FE802E36" bold="true" box="[199,367,511,536]" italics="true" pageId="12" pageNumber="503">Imago, female</emphasis>
(
<figureCitation id="49E72A718F17FFE8FE7BFE48FE362E36" box="[384,473,511,536]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Figs 36</figureCitation>
,
<figureCitation id="49E72A718F17FFE8FE1DFDB7FDDF2E36" box="[486,560,512,536]" captionStart="FIGURES 38 42" captionStartId="11.[151,264,1844,1867]" captionTargetBox="[170,1416,193,1823]" captionTargetId="figure@11.[170,1417,193,1824]" captionTargetPageId="11" captionText="FIGURES 38 42. Procloeon (Oculogaster) cylindroculum, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl. l., lateral ocellus; ocl. m., median ocellus." httpUri="https://zenodo.org/record/271366/files/figure.png" pageId="12" pageNumber="503">3842</figureCitation>
). Head relatively wide, distance between eyes exceeds eye length; eyes only slightly elevated above head surface. Head, thorax and abdomen ocher with contrasting red, purple, brown or/and black hypodermal strip of composite form, stretching along lateral side of body (along thoracic pleura and along lateral parts of abdominal terga) from prothorax to tip of abdomen. In fresh specimens dorsal margin of this stripe can be bordered by contrasting white stripe (
<figureCitation id="49E72A718F17FFE8FD6BFD38FD092E86" box="[656,742,655,680]" captionStart="FIGURES 38 42" captionStartId="11.[151,264,1844,1867]" captionTargetBox="[170,1416,193,1823]" captionTargetId="figure@11.[170,1417,193,1824]" captionTargetPageId="11" captionText="FIGURES 38 42. Procloeon (Oculogaster) cylindroculum, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl. l., lateral ocellus; ocl. m., median ocellus." httpUri="https://zenodo.org/record/271366/files/figure.png" pageId="12" pageNumber="503">Fig. 40</figureCitation>
); on posterior margins of abdominal terga IV and VIII lateral stripes turned medially (
<figureCitation id="49E72A718F17FFE8FE5FFD03FE132EE3" box="[420,508,692,717]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Figs 36</figureCitation>
). Mesonotum either unicolor ocher, or with contrasting pigmented stripes on lateral areas (
<figureCitation id="49E72A718F17FFE8FF1BFD60FEDA2EDE" box="[224,309,727,752]" captionStart="FIGURES 38 42" captionStartId="11.[151,264,1844,1867]" captionTargetBox="[170,1416,193,1823]" captionTargetId="figure@11.[170,1417,193,1824]" captionTargetPageId="11" captionText="FIGURES 38 42. Procloeon (Oculogaster) cylindroculum, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl. l., lateral ocellus; ocl. m., median ocellus." httpUri="https://zenodo.org/record/271366/files/figure.png" pageId="12" pageNumber="503">Fig. 40</figureCitation>
). All legs with single apical spine on primary 3rd tarsomere. Wings as in male. Cerci colorless with contrasting dark brown or black bands formed by hypodermal pigmentation of each 4th or each 2nd segment.
</paragraph>
<paragraph id="D16336F48F17FFE8FF3CFC97FB2A2F72" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CFC97FF122F19" bold="true" box="[199,253,800,823]" italics="true" pageId="12" pageNumber="503">Egg.</emphasis>
Without rigid chorion; embryos are developed inside female imago, so that mature female imago looks maculated because of black eyes and ocelli of embryos visible through its integument (
<figureCitation id="49E72A718F17FFE8FB9FFCF3FB572F72" box="[1124,1208,836,861]" captionStart="FIGURES 38 42" captionStartId="11.[151,264,1844,1867]" captionTargetBox="[170,1416,193,1823]" captionTargetId="figure@11.[170,1417,193,1824]" captionTargetPageId="11" captionText="FIGURES 38 42. Procloeon (Oculogaster) cylindroculum, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl. l., lateral ocellus; ocl. m., median ocellus." httpUri="https://zenodo.org/record/271366/files/figure.png" pageId="12" pageNumber="503">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F17FFE8FF3CFCD0FCB22F51" blockId="12.[151,1437,151,2013]" box="[199,861,871,896]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CFCD0FEA12FAE" bold="true" box="[199,334,871,896]" pageId="12" pageNumber="503">Dimension.</emphasis>
Fore wing length of male and female
<quantity id="16249B118F17FFE8FD02FCD0FCB62F51" box="[761,857,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="12" pageNumber="503" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
.
</paragraph>
<paragraph id="D16336F48F17FFE8FF3CFC3BFE3E283E" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CFC3BFD5C2F8B" bold="true" box="[199,691,908,933]" pageId="12" pageNumber="503">
Variability.
<emphasis id="E3A8EAE68F17FFE8FEAFFC3BFD5C2F8B" bold="true" box="[340,691,908,933]" italics="true" pageId="12" pageNumber="503">Turbinate eyes of male imago.</emphasis>
</emphasis>
<bibRefCitation id="B54D4B058F17FFE8FD40FC3BFCB72F8A" author="Gillies" box="[699,856,908,933]" pageId="12" pageNumber="503" refString="Gillies, M. T. (1979) Polymorphism in the turbinate eyes of an African Cloeon (Ephemeroptera: Baetidae). Entomologist's Monthly Magazine, 114, 155." type="journal article" year="1979">Gillies (1979)</bibRefCitation>
reported variability in shape of turbinate eyes of
<taxonomicName id="16DC4D778F17FFE8FA7CFC3AFED72FE6" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F17FFE8FA7CFC3AFED72FE6" italics="true" pageId="12" pageNumber="503">P. cylindroculum</emphasis>
</taxonomicName>
, which in some populations are mainly tall and cylindrical (height to width about 1.5:1), and in others mainly broad (height to width about 1:1). Most specimens examined by me have intermediate shape of turbinate eyes (
<figureCitation id="49E72A718F17FFE8FEB9FC40FE2A283E" box="[322,453,1015,1040]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Figs 3334</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F17FFE8FF3CFBABFB6828EB" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CFBABFE3A281B" bold="true" box="[199,469,1052,1077]" italics="true" pageId="12" pageNumber="503">Marginal intercalaries.</emphasis>
In the original description wing on
<taxonomicName id="16DC4D778F17FFE8FC84FBAAFBD2281A" box="[895,1085,1052,1076]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F17FFE8FC84FBAAFBD2281A" box="[895,1085,1052,1076]" italics="true" pageId="12" pageNumber="503">P. cylindroculum</emphasis>
</taxonomicName>
is figured as having marginal intercalaries in each space (
<bibRefCitation id="B54D4B058F17FFE8FE37FB88FD262876" author="Kimmins" box="[460,713,1087,1112]" pageId="12" pageNumber="503" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880." type="journal article" year="1956">
Kimmins 1956:
<figureCitation id="49E72A718F17FFE8FD7EFB88FD262876" box="[645,713,1087,1112]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="12" pageNumber="503">Fig. 2</figureCitation>
</bibRefCitation>
). Gillies, based on reexamination of
<typeStatus id="0E6788568F17FFE8FB93FB88FB252876" box="[1128,1226,1087,1112]" pageId="12" pageNumber="503" type="holotype">holotype</typeStatus>
and
<typeStatus id="0E6788568F17FFE8FAFAFBF7FA822876" box="[1281,1389,1088,1112]" pageId="12" pageNumber="503" type="paratype">paratypes</typeStatus>
and examination of other material, reported that actually marginal intercalaries are present in alternated spaces (
<bibRefCitation id="B54D4B058F17FFE8FAABFBD3FEE4288E" author="Gillies" pageId="12" pageNumber="503" refString="Gillies, M. T. (1980) An introduction to the study of Cloeon Leach (Baetidae, Ephemeroptera) in West Africa. Bulletin de l'Institute Francaise d'Afrique Noire, Serie A, 42 (1), 135 - 156." type="journal article" year="1980">Gillies 1980: 147</bibRefCitation>
). Specimens examined by me, have marginal intercalaries at least in spaces located behind convex veins; spaces behind concave veins are either empty (
<figureCitation id="49E72A718F17FFE8FD5FFB1BFD1328EA" box="[676,764,1196,1221]" captionStart="FIGURES 30 32" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[200,1361,197,1812]" captionTargetId="figure@9.[195,1372,193,1826]" captionTargetPageId="9" captionText="FIGURES 30 32. Procloeon (Oculogaster) cylindroculum from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271364/files/figure.png" pageId="12" pageNumber="503">Figs 32</figureCitation>
), or some of them have intercalary.
</paragraph>
<paragraph id="D16336F48F17FFE8FF3CFB78FB6D2A7E" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CFB78FD4C28C6" bold="true" box="[199,675,1231,1256]" italics="true" pageId="12" pageNumber="503">Coloration of male imago and subimago.</emphasis>
According to the original description of
<taxonomicName id="16DC4D778F17FFE8FB8FFB66FADE28C6" box="[1140,1329,1232,1256]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F17FFE8FB8FFB66FADE28C6" box="[1140,1329,1232,1256]" italics="true" pageId="12" pageNumber="503">P. cylindroculum</emphasis>
</taxonomicName>
based on male imagoes from
<collectingCountry id="A9CB76648F17FFE8FE8EFB43FE252923" box="[373,458,1268,1293]" name="Malawi" pageId="12" pageNumber="503">Malawi</collectingCountry>
(
<collectingCountry id="A9CB76648F17FFE8FE22FB43FDBA2923" box="[473,597,1268,1293]" name="Malawi" pageId="12" pageNumber="503">Nyasaland</collectingCountry>
), abdominal tergum IV has no pair of reddish spots, and cerci have no dark bands (
<bibRefCitation id="B54D4B058F17FFE8FF17FAA0FE71291E" author="Kimmins" box="[236,414,1303,1328]" pageId="12" pageNumber="503" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880." type="journal article" year="1956">Kimmins 1956</bibRefCitation>
). According to the additional description based on material from Entebbe and Jinja in
<collectingCountry id="A9CB76648F17FFE8FF6CFA8BFF1E297B" box="[151,241,1340,1365]" name="Uganda" pageId="12" pageNumber="503">Uganda</collectingCountry>
, most male specimens have the same color, but some specimens have pair of reddish brown spots on tergum IV and annulated cerci (
<bibRefCitation id="B54D4B058F17FFE8FE03FAE8FD4B2956" author="Kimmins" box="[504,676,1375,1400]" pageId="12" pageNumber="503" refString="Kimmins, D. E. (1960) Notes on East African Ephemeroptera with descriptions of new species. Bulletin of the British Museum (Natural History), Entomology, 9 (6), 337 - 355. http: // dx. doi. org / 10.5962 / bhl. part. 27555" type="journal article" year="1960">Kimmins 1960</bibRefCitation>
). Among
<specimenCount id="C7DAFD7D8F17FFE8FCE9FAE8FC582956" box="[786,951,1375,1400]" pageId="12" pageNumber="503" type="generic">152 specimens</specimenCount>
from the same locality (Jinja) examined by me, most specimens have no spots on tergum IV (
<figureCitation id="49E72A718F17FFE8FD25FA33FCDB29B2" box="[734,820,1412,1437]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 34</figureCitation>
), and
<specimenCount id="C7DAFD7D8F17FFE8FC85FA33FBF729B3" box="[894,1048,1412,1437]" pageId="12" pageNumber="503" type="generic">23 specimens</specimenCount>
have pair of reddish or brownish spots, contrasting in some specimens and poorly expressed in others; all preserved cerci have no dark bands (
<figureCitation id="49E72A718F17FFE8FA94FA10FF5829CA" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 34</figureCitation>
). Among
<specimenCount id="C7DAFD7D8F17FFE8FED7FA7BFE6929CB" box="[300,390,1484,1509]" pageId="12" pageNumber="503" type="male">4 males</specimenCount>
collected in
<collectingCountry id="A9CB76648F17FFE8FDDAFA7BFD9829CB" box="[545,631,1484,1509]" name="Zambia" pageId="12" pageNumber="503">Zambia</collectingCountry>
(three imagoes reared from larvae and one subimago extracted from larva),
<specimenCount id="C7DAFD7D8F17FFE8FF1CFA47FE9D2A26" box="[231,370,1519,1544]" pageId="12" pageNumber="503" type="generic">3 specimens</specimenCount>
have a pair of well-expressed spots on abdominal tergum IV (
<figureCitation id="49E72A718F17FFE8FBC3FA58FB532A26" box="[1080,1212,1519,1544]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Figs 33, 35</figureCitation>
), and one specimen has less distinct unpaired transverse macula on the same place; cerci of all four specimens have dark bands (
<figureCitation id="49E72A718F17FFE8FA94F9A3FF592A7E" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="12" pageNumber="503">Fig. 33</figureCitation>
), which are less contrasting than in females collected in
<collectingCountry id="A9CB76648F17FFE8FCD5F980FC6A2A7E" box="[814,901,1591,1616]" name="Uganda" pageId="12" pageNumber="503">Uganda</collectingCountry>
and
<collectingCountry id="A9CB76648F17FFE8FC47F980FBFE2A7E" box="[956,1041,1591,1616]" name="Zambia" pageId="12" pageNumber="503">Zambia</collectingCountry>
(
<figureCitation id="49E72A718F17FFE8FBDAF980FB9A2A7E" box="[1057,1141,1591,1616]" captionStart="FIGURES 38 42" captionStartId="11.[151,264,1844,1867]" captionTargetBox="[170,1416,193,1823]" captionTargetId="figure@11.[170,1417,193,1824]" captionTargetPageId="11" captionText="FIGURES 38 42. Procloeon (Oculogaster) cylindroculum, female imago. 38, head and prothorax in Canadian balsam with embryos visible inside; 39, enlarged embryo with focus on median ocellus and mandibles; 40, mature female imago free of embryos; 41, abdomen of female imago with embryos (black eyes and ocelli visible through integument); 42, embryo extracted from female imago (38, 39, 41, 42, from Uganda; 40, from Zambezi). Abbreviations: md, mandibles; oc, future facetted eye; ocl. l., lateral ocellus; ocl. m., median ocellus." httpUri="https://zenodo.org/record/271366/files/figure.png" pageId="12" pageNumber="503">Fig. 40</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F17FFE8FF3CF9EBFAF32ACE" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CF9EBFD562A5B" bold="true" box="[199,697,1628,1653]" italics="true" pageId="12" pageNumber="503">Coloration of female imago and subimago.</emphasis>
<bibRefCitation id="B54D4B058F17FFE8FD3AF9EBFCB12A5A" author="Gillies" box="[705,862,1628,1653]" pageId="12" pageNumber="503" refString="Gillies, M. T. (1980) An introduction to the study of Cloeon Leach (Baetidae, Ephemeroptera) in West Africa. Bulletin de l'Institute Francaise d'Afrique Noire, Serie A, 42 (1), 135 - 156." type="journal article" year="1980">Gillies (1980)</bibRefCitation>
reported variability in coloration of freshly caught females, some of which have no red lateral lines on thorax and abdomen. Among specimens examined by me, all females collected in
<collectingCountry id="A9CB76648F17FFE8FE77F913FE0C2A93" box="[396,483,1700,1725]" name="Uganda" pageId="12" pageNumber="503">Uganda</collectingCountry>
and
<collectingCountry id="A9CB76648F17FFE8FDD8F913FD942A93" box="[547,635,1700,1725]" name="Zambia" pageId="12" pageNumber="503">Zambia</collectingCountry>
, have these lines. Females without this coloration are found among specimens from
<collectingCountry id="A9CB76648F17FFE8FEABF970FE672ACE" box="[336,392,1735,1760]" name="Serbia" pageId="12" pageNumber="503">Mali</collectingCountry>
; fixation of these specimens was not well, and their initial coloration is unknown.
</paragraph>
<paragraph id="D16336F48F17FFE9FF3CF95BFB9D2D83" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1436,151,429]" lastPageId="13" lastPageNumber="504" pageId="12" pageNumber="503">
<emphasis id="E3A8EAE68F17FFE8FF3CF95BFE972B2B" bold="true" box="[199,376,1772,1797]" italics="true" pageId="12" pageNumber="503">Larval tergalii.</emphasis>
All three examined larvae from Victoria Nile have tergalii large, greatly widened, dorsal lamella being present on tergalii IVI (
<figureCitation id="49E72A718F17FFE8FD8BF8B8FD332B06" box="[624,732,1807,1832]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="12" pageNumber="503">Figs 17</figureCitation>
). Among seven examined larvae from Luangwa, only five specimens have tergalii preserved; their tergalii are relatively small, with small dorsal lamella on tergalii IV only (
<figureCitation id="49E72A718F17FFE8FF64F8E0FECB2B5E" box="[159,292,1879,1904]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="12" pageNumber="503">Figs 1117</figureCitation>
). Larvae described by
<bibRefCitation id="B54D4B058F17FFE8FDD4F8E0FD312B5E" author="Barnard" box="[559,734,1879,1904]" pageId="12" pageNumber="503" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. http: // dx. doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard (1932)</bibRefCitation>
under the name “
<taxonomicName id="16DC4D778F17FFE8FC48F8EFFB2B2B5E" box="[947,1220,1880,1904]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="africanum">
<emphasis id="E3A8EAE68F17FFE8FC48F8EFFB2B2B5E" box="[947,1220,1880,1904]" italics="true" pageId="12" pageNumber="503">Austrocloeon africanum</emphasis>
</taxonomicName>
”, according to his description, have dorsal lamella either on tergalii IV, or on tergalii IVI. On the drawing made by
<bibRefCitation id="B54D4B058F17FFE8FAFBF8CBFF1B2B96" author="Gillies" pageId="12" pageNumber="503" refString="Gillies, M. T. (1980) An introduction to the study of Cloeon Leach (Baetidae, Ephemeroptera) in West Africa. Bulletin de l'Institute Francaise d'Afrique Noire, Serie A, 42 (1), 135 - 156." type="journal article" year="1980">
Gillies (1980:
<figureCitation id="49E72A718F17FFE8FF6CF817FF032B96" box="[151,236,1952,1977]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="12" pageNumber="503">Fig. 20</figureCitation>
)
</bibRefCitation>
tergalii of
<taxonomicName id="16DC4D778F17FFE8FE85F817FD932B96" box="[382,636,1952,1976]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F17FFE8FE85F817FD932B96" box="[382,636,1952,1976]" italics="true" pageId="12" pageNumber="503">Cloeon cylindroculum</emphasis>
</taxonomicName>
are wide, tergalii IVI with dorsal lamella; tergalius VII is figured wrongly, probably being either overturned, or damaged. For the newly described species
<taxonomicName id="16DC4D778F17FFE9FB5AF873FEDB2C9E" authority="Gillies (1997)" authorityName="Gillies" authorityYear="1997" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" lastPageId="13" lastPageNumber="504" order="Ephemeroptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="silvicola">
<emphasis id="E3A8EAE68F17FFE8FB5AF873FA6D2BF2" box="[1185,1410,1988,2012]" italics="true" pageId="12" pageNumber="503">Procloeon silvicola</emphasis>
,
<bibRefCitation id="B54D4B058F16FFE9FF6CFF20FEDB2C9E" author="Gillies" box="[151,308,151,176]" pageId="13" pageNumber="504" refString="Gillies, M. T. (1997) A new species of Procloeon Bengtsson from the forest zone of West Africa (Ephemeroptera, Baetidae). Entomologist's Monthly Magazine, 133, 247 - 250." type="journal article" year="1997">Gillies (1997)</bibRefCitation>
</taxonomicName>
reported presence of dorsal lamellae on tergalii IIV only; this conclusion was based on one larval skin and was not suggested as a character separating
<emphasis id="E3A8EAE68F16FFE9FD1CFF0AFCB12CFA" box="[743,862,188,212]" italics="true" pageId="13" pageNumber="504">
<taxonomicName id="16DC4D778F16FFE9FD1CFF0AFCBF2CFA" box="[743,848,188,212]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="subSpecies" species="sil" subSpecies="vicol">P. sil vicol</taxonomicName>
a
</emphasis>
from
<taxonomicName id="16DC4D778F16FFE9FC5EFF0AFB8E2CFA" box="[933,1121,188,212]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F16FFE9FC5EFF0AFB8E2CFA" box="[933,1121,188,212]" italics="true" pageId="13" pageNumber="504">P. cylindroculum</emphasis>
</taxonomicName>
. In the same paper tergalius III of
<taxonomicName id="16DC4D778F16FFE9FF18FF57FE452CD6" box="[227,426,224,248]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="E3A8EAE68F16FFE9FF18FF57FE452CD6" box="[227,426,224,248]" italics="true" pageId="13" pageNumber="504">C. cylindroculum</emphasis>
</taxonomicName>
is figured as narrow and having small dorsal lamella (
<bibRefCitation id="B54D4B058F16FFE9FBBAFF68FAC02CD6" author="Gillies" box="[1089,1327,223,248]" pageId="13" pageNumber="504" refString="Gillies, M. T. (1985) A preliminary account of the East African species of Cloeon Leach and Rhithrocloeon gen. n. (Ephemeroptera). Aquatic Insects, 7 (1), 1 - 17. http: // dx. doi. org / 10.1080 / 01650428509361193" type="journal article" year="1985">
Gillies 1985:
<figureCitation id="49E72A718F16FFE9FB1DFF68FAC02CD6" box="[1254,1327,223,248]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="13" pageNumber="504">Fig. 2</figureCitation>
</bibRefCitation>
). Similar variations of size and shape of tergalii, and presence/absence of dorsal lamella on tergalii of posterior pairs, are usual for other species of
<taxonomicName id="16DC4D778F16FFE9FE39FE9FFDD92D6E" box="[450,566,296,320]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F16FFE9FE39FE9FFDD92D6E" box="[450,566,296,320]" italics="true" pageId="13" pageNumber="504">Procloeon</emphasis>
</taxonomicName>
, for which long series of larvae were examined. For example, in
<taxonomicName id="16DC4D778F16FFE9FAD3FE9FFDF02D4A" authority="Kluge &amp; Novikova 1992" authorityName="Kluge &amp; Novikova" authorityYear="1992" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="macronyx">
<emphasis id="E3A8EAE68F16FFE9FAD3FE9FFEEB2D4A" italics="true" pageId="13" pageNumber="504">Procloeon macronyx</emphasis>
<bibRefCitation id="B54D4B058F16FFE9FEF0FEFBFDF02D4A" author="Kluge" box="[267,543,332,357]" pageId="13" pageNumber="504" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">Kluge &amp; Novikova 1992</bibRefCitation>
</taxonomicName>
dorsal lamella can be larger or smaller, present or absent on tergalius VI (
<bibRefCitation id="B54D4B058F16FFE9FAADFEFBFE082DA6" author="Kluge" pageId="13" pageNumber="504" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">
Kluge &amp; Novikova 1992:
<figureCitation id="49E72A718F16FFE9FE8FFED8FE082DA6" box="[372,487,367,392]" captionStart-0="FIGURES 1 10" captionStart-1="FIGURES 11 20" captionStartId-0="6.[151,264,1903,1926]" captionStartId-1="7.[151,264,1612,1635]" captionTargetBox-0="[154,1436,277,1881]" captionTargetBox-1="[160,1415,207,1574]" captionTargetId-0="figure@6.[151,1436,265,1883]" captionTargetId-1="figure@7.[151,1436,193,1591]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." captionText-1="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri-0="https://zenodo.org/record/271361/files/figure.png" httpUri-1="https://zenodo.org/record/271362/files/figure.png" pageId="13" pageNumber="504">Figs 115</figureCitation>
</bibRefCitation>
); in
<taxonomicName id="16DC4D778F16FFE9FDE7FEC6FC142DA6" authority="Kluge &amp; Novikova 1992" authorityName="Kluge &amp; Novikova" authorityYear="1992" box="[540,1019,367,392]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="heterophyllum">
<emphasis id="E3A8EAE68F16FFE9FDE7FEC6FD362DA6" box="[540,729,368,392]" italics="true" pageId="13" pageNumber="504">P. heterophyllum</emphasis>
<bibRefCitation id="B54D4B058F16FFE9FD19FED8FC142DA6" author="Kluge" box="[738,1019,367,392]" pageId="13" pageNumber="504" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">Kluge &amp; Novikova 1992</bibRefCitation>
</taxonomicName>
dorsal lamellae vary from absent on all tergalii to present on tergalii IV (
<specimenCount id="C7DAFD7D8F16FFE9FDC3FE23FD422D83" box="[568,685,404,429]" pageId="13" pageNumber="504" type="larva">110 larvae</specimenCount>
from one small stream were examined).
</paragraph>
</subSubSection>
</treatment>
</document>