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<document id="F101BE77E617784DF9E23E861DBF6E0B" ID-CLB-Dataset="39139" ID-DOI="10.11646/zootaxa.4107.4.2" ID-GBIF-Dataset="96189ebc-493c-44ef-9e46-1bf2af20c3e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="271360" ID-ZooBank="5CBAACB4-2A65-4766-8339-33D65C93FF9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462342958656" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2016" docId="597587E28F1AFFE6FF6CF906FB402DE1" docLanguage="en" docName="zootaxa.4107.4.2.pdf.imf" docOrigin="Zootaxa 4107 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cloeonini Newman 1853" docType="treatment" docVersion="12" lastPageNumber="493" masterDocId="A54CFF9A8F1BFFE4FFFBFFB7FFEF2C2E" masterDocTitle="A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae)" masterLastPageNumber="516" masterPageNumber="491" pageNumber="492" updateTime="1698662244020" updateUser="ExternalLinkService">
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<mods:title id="CEEF8AB8E9ADCDA9616E35E9E8F453D9">A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="EA96959DD805DE5B8FA6E0081A48F745">Kluge, Nikita J.</mods:namePart>
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<treatment id="597587E28F1AFFE6FF6CF906FB402DE1" ID-DOI="http://doi.org/10.5281/zenodo.5670313" ID-GBIF-Taxon="120620940" ID-Zenodo-Dep="5670313" LSID="urn:lsid:plazi:treatment:597587E28F1AFFE6FF6CF906FB402DE1" httpUri="http://treatment.plazi.org/id/597587E28F1AFFE6FF6CF906FB402DE1" lastPageId="2" lastPageNumber="493" pageId="1" pageNumber="492">
<subSubSection id="99C6657F8F1AFFE5FF6CF906FDFC2AE5" box="[151,531,1713,1739]" pageId="1" pageNumber="492" type="nomenclature">
<paragraph id="D16336F48F1AFFE5FF6CF906FDFC2AE5" blockId="1.[151,531,1713,1739]" box="[151,531,1713,1739]" pageId="1" pageNumber="492">
<heading id="8A2B81988F1AFFE5FF6CF906FDFC2AE5" bold="true" box="[151,531,1713,1739]" fontSize="11" level="1" pageId="1" pageNumber="492" reason="1">
<emphasis id="E3A8EAE68F1AFFE5FF6CF906FDFC2AE5" bold="true" box="[151,531,1713,1739]" pageId="1" pageNumber="492">
Tribe
<taxonomicName id="16DC4D778F1AFFE5FF19F906FDFC2AE5" box="[226,531,1713,1739]" pageId="1" pageNumber="492" rank="tribe" tribe="Cloeonini">Cloeonini Newman 1853</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F1AFFE5FF6CF946FD682B08" pageId="1" pageNumber="492" type="reference_group">
<paragraph id="D16336F48F1AFFE5FF6CF946FD892B29" blockId="1.[151,647,1777,1830]" box="[151,614,1777,1799]" pageId="1" pageNumber="492">
Branch “6”:
<bibRefCitation id="B54D4B058F1AFFE5FEECF946FDFA2B29" author="Kluge" box="[279,533,1777,1799]" pageId="1" pageNumber="492" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">Kluge &amp; Novikova 1992</bibRefCitation>
:
<figureCitation id="49E72A718F1AFFE5FDE4F946FD892B29" box="[543,614,1777,1799]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="1" pageNumber="492">Fig. 11</figureCitation>
</paragraph>
<paragraph id="D16336F48F1AFFE5FF6CF8A7FD682B08" blockId="1.[151,647,1777,1830]" box="[151,647,1808,1830]" pageId="1" pageNumber="492">
Hierarchical name:
<taxonomicName id="16DC4D778F1AFFE5FEA5F8A7FE422B08" ID-CoL="3R5Y" box="[350,429,1808,1830]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="492" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1:
<bibRefCitation id="B54D4B058F1AFFE5FE25F8A7FDB82B08" author="Kluge" box="[478,599,1808,1830]" pageId="1" pageNumber="492" refString="Kluge, N. J. (2011) Non-African representatives of the plesiomorphion Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2011">Kluge 2011</bibRefCitation>
: 362
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F1AFFE6FF6CF8FBFB402DE1" lastPageId="2" lastPageNumber="493" pageId="1" pageNumber="492" type="diagnosis">
<paragraph id="D16336F48F1AFFE5FF6CF8FBFADD2BDB" blockId="1.[151,1437,1868,2037]" pageId="1" pageNumber="492">
<emphasis id="E3A8EAE68F1AFFE5FF6CF8FBFE872B4D" bold="true" box="[151,360,1868,1893]" pageId="1" pageNumber="492">
Diagnosis.
<emphasis id="E3A8EAE68F1AFFE5FEE6F8FBFE872B4D" bold="true" box="[285,360,1868,1891]" italics="true" pageId="1" pageNumber="492">Larva.</emphasis>
</emphasis>
Labial palp with 3rd segment truncate and 2nd segment non-widened. Patella-tibial suture equally developed on all legs, often interrupted on outer side and continued by row of fine setae (
<figureCitation id="49E72A718F1AFFE5FB37F8D8FACD2BA6" box="[1228,1314,1903,1928]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="1" pageNumber="492">Figs 20</figureCitation>
,
<figureCitation id="49E72A718F1AFFE5FAD5F8D8FAA22BA6" box="[1326,1357,1903,1928]" captionStart="FIGURES 49 58" captionStartId="17.[151,264,1666,1689]" captionTargetBox="[162,1426,196,1637]" captionTargetId="figure@17.[151,1436,187,1653]" captionTargetPageId="17" captionText="FIGURES 49 58. Procloeon (Oculogaster) album sp. n., larva. 49 55, tergalii I VII; 56, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus); 57, claw; 58, paraproct (49 52, 54 55, 58, holotype)." httpUri="https://zenodo.org/record/271368/files/figure.png" pageId="1" pageNumber="492">56</figureCitation>
;
<bibRefCitation id="B54D4B058F1AFFE5FAA3F8D8FE8B2B82" author="Kluge" pageId="1" pageNumber="492" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">Kluge &amp; Novikova 1992</bibRefCitation>
:
<figureCitation id="49E72A718F1AFFE5FE94F823FE5D2B82" box="[367,434,1940,1965]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="1" pageNumber="492">Fig. 1</figureCitation>
: 1, 2). Claws slightly bent, symmetric, either with two equal rows of denticles, or without denticles. At least abdominal segment IX with denticles on lateral sides. Tergalii able to rhythmical respiratory movements. Tergalii without anal rib, anal margin with or without flap forming additional dorsal lamella.
</paragraph>
<paragraph id="D16336F48F19FFE6FF3CFF2FFBAA2CFB" blockId="2.[151,1437,151,464]" pageId="2" pageNumber="493">
<emphasis id="E3A8EAE68F19FFE6FF3CFF2FFC912C9E" bold="true" box="[199,894,151,176]" italics="true" pageId="2" pageNumber="493">Pose of developing subimaginal gonostyli in mature larva</emphasis>
of the «
<taxonomicName id="16DC4D778F19FFE6FC13FF2FFBD62C9E" box="[1000,1081,152,176]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F19FFE6FC13FF2FFBD62C9E" box="[1000,1081,152,176]" italics="true" pageId="2" pageNumber="493">Cloeon</emphasis>
</taxonomicName>
-
<typeStatus id="0E6788568F19FFE6FBBAFF2FFB992C9E" box="[1089,1142,152,176]" pageId="2" pageNumber="493">type</typeStatus>
», i.e. with 2nd segments directed laterally and 3rd segments bent medially (
<bibRefCitation id="B54D4B058F19FFE6FD35FF0BFBD82CFB" author="Kluge" box="[718,1079,188,213]" pageId="2" pageNumber="493" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">
Kluge &amp; Novikova 1992:
<figureCitation id="49E72A718F19FFE6FC0FFF0BFBD82CFB" box="[1012,1079,188,213]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1903,1926]" captionTargetBox="[154,1436,277,1881]" captionTargetId="figure@6.[151,1436,265,1883]" captionTargetPageId="6" captionText="FIGURES 1 10. Procloeon (Oculogaster) cylindroculum, larva. 1 7, tergalii of I VII pairs (specimen from Uganda); 8, 9, apices of left and right mandibles, dorsal view; 10, apex of right mandible, apical view. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/271361/files/figure.png" pageId="2" pageNumber="493">Fig. 5</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="D16336F48F19FFE6FF3CFF68FABD2D83" blockId="2.[151,1437,151,464]" pageId="2" pageNumber="493">
<emphasis id="E3A8EAE68F19FFE6FF3CFF68FE9D2CD9" bold="true" box="[199,370,223,248]" italics="true" pageId="2" pageNumber="493">Winged stages.</emphasis>
Fore wing with no more than one intercalary vein in each space.
<collectingCountry id="A9CB76648F19FFE6FBB6FF68FB6A2CD6" box="[1101,1157,223,248]" name="India" pageId="2" pageNumber="493">Hind</collectingCountry>
wing either absent, or of the «
<taxonomicName id="16DC4D778F19FFE6FF35FEB3FE8D2D32" box="[206,354,260,284]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F19FFE6FF35FEB3FE8D2D32" box="[206,354,260,284]" italics="true" pageId="2" pageNumber="493">Centroptilum</emphasis>
</taxonomicName>
-
<typeStatus id="0E6788568F19FFE6FE91FEB2FE702D33" box="[362,415,261,285]" pageId="2" pageNumber="493">type</typeStatus>
», i.e. narrow, with 2 longitudinal veins and always with hooked costal process. Postsubalar sclerite (well visible on subimaginal exuviae) has posterior-dorsal process narrow, with concave dorsal margin (
<figureCitation id="49E72A718F19FFE6FF64FEFBFF1D2D4A" box="[159,242,332,357]" captionStart="FIGURES 33 37" captionStartId="10.[151,264,1855,1878]" captionTargetBox="[170,1416,193,1834]" captionTargetId="figure@10.[170,1417,193,1835]" captionTargetPageId="10" captionText="FIGURES 33 37. Procloeon (Oculogaster) cylindroculum. 33, 34, male imagoes; 35, subimaginal abdomen extracted from mature male larva; 36, subimaginal abdomen extracted from mature female larva; 37, subimaginal exuviae of postsubalar sclerite and lateropostnotal crest (33, 35 37, specimens from Zambia; 34, specimen from Uganda)." httpUri="https://zenodo.org/record/271365/files/figure.png" pageId="2" pageNumber="493">Fig. 37</figureCitation>
). Subimaginal tarsi of all legs covered by pointed microlepides, without blunt microlepides. In male imago apices of gonovectes fused with penial bridge; penial bridge medially with unpaired sclerotized projection (
<figureCitation id="49E72A718F19FFE6FABAFED8FA782DA6" box="[1345,1431,367,392]" captionStart="FIGURES 30 32" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[200,1361,197,1812]" captionTargetId="figure@9.[195,1372,193,1826]" captionTargetPageId="9" captionText="FIGURES 30 32. Procloeon (Oculogaster) cylindroculum from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271364/files/figure.png" pageId="2" pageNumber="493">Figs 30</figureCitation>
,
<figureCitation id="49E72A718F19FFE6FF6CFE23FF582D83" box="[151,183,404,429]" captionStart="FIGURES 65 67" captionStartId="19.[151,264,1843,1866]" captionTargetBox="[163,1414,196,1810]" captionTargetId="figure@19.[163,1423,193,1822]" captionTargetPageId="19" captionText="FIGURES 65 67. Procloeon (Oculogaster) album sp. n. 65, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 66, genitals of male subimago; 67, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271370/files/figure.png" pageId="2" pageNumber="493">65</figureCitation>
). Gonostyli with 2nd segment widened apically, 3rd (terminal) segment small and petiolate (
<figureCitation id="49E72A718F19FFE6FB39FE23FAF52D82" box="[1218,1306,404,429]" captionStart="FIGURES 30 32" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[200,1361,197,1812]" captionTargetId="figure@9.[195,1372,193,1826]" captionTargetPageId="9" captionText="FIGURES 30 32. Procloeon (Oculogaster) cylindroculum from Zambia. 30, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 31, exuviae of subimaginal genitals; 32, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271364/files/figure.png" pageId="2" pageNumber="493">Figs 30</figureCitation>
,
<figureCitation id="49E72A718F19FFE6FADEFE23FAAA2D83" box="[1317,1349,404,429]" captionStart="FIGURES 65 67" captionStartId="19.[151,264,1843,1866]" captionTargetBox="[163,1414,196,1810]" captionTargetId="figure@19.[163,1423,193,1822]" captionTargetPageId="19" captionText="FIGURES 65 67. Procloeon (Oculogaster) album sp. n. 65, genitals of male imago (at left half gonovectal muscles shown by interrupted lines, hidden part of penis shown by interrupted line and dotted, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles not shown; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). 66, genitals of male subimago; 67, wing. Abbreviations: 1, 2, 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/271370/files/figure.png" pageId="2" pageNumber="493">65</figureCitation>
).
</paragraph>
<paragraph id="D16336F48F19FFE6FF3CFE00FB402DE1" blockId="2.[151,1437,151,464]" box="[199,1199,439,464]" pageId="2" pageNumber="493">
<emphasis id="E3A8EAE68F19FFE6FF3CFE00FE892DFE" bold="true" box="[199,358,439,464]" pageId="2" pageNumber="493">Composition.</emphasis>
Includes genera
<taxonomicName id="16DC4D778F19FFE6FDDFFE0FFD9A2DFE" box="[548,629,440,464]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F19FFE6FDDFFE0FFD9A2DFE" box="[548,629,440,464]" italics="true" pageId="2" pageNumber="493">Cloeon</emphasis>
</taxonomicName>
,
<taxonomicName id="16DC4D778F19FFE6FD7FFE0FFCFF2DFE" box="[644,784,440,464]" class="Insecta" family="Baetidae" genus="Similicloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F19FFE6FD7FFE0FFCFF2DFE" box="[644,784,440,464]" italics="true" pageId="2" pageNumber="493">Similicloeon</emphasis>
</taxonomicName>
,
<taxonomicName id="16DC4D778F19FFE6FCE6FE0FFC112DFE" box="[797,1022,440,464]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F19FFE6FCE6FE0FFC112DFE" box="[797,1022,440,464]" italics="true" pageId="2" pageNumber="493">Pseudocentroptilum</emphasis>
</taxonomicName>
and
<taxonomicName id="16DC4D778F19FFE6FBCFFE0FFB472DFE" box="[1076,1192,440,464]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F19FFE6FBCFFE0FFB472DFE" box="[1076,1192,440,464]" italics="true" pageId="2" pageNumber="493">Procloeon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>