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<mods:title id="981B5BA820B309A71066B3EBBC75EC13">Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
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<mods:namePart id="74D9A056FFF4190800CB66A6F9798237">Pall-Gergely, Barna</mods:namePart>
<mods:affiliation id="E9EC1CDAFAF77425B8C00B2EF5168162">Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
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<mods:namePart id="5DFA4ACBD888000C7E9502E609877F1A">Sajan, Sheikh</mods:namePart>
<mods:affiliation id="4D1C96D5E8758A48B3029A148F47F1A7">Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India &amp; Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
<mods:nameIdentifier id="1711E1A7B27AADC0481037C0CE87036E" type="orcid">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
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<mods:namePart id="D2D4B36E1B96E1D140E19CE63EB71841">Tripathy, Basudev</mods:namePart>
<mods:affiliation id="508318B1C0365F42E36EDF3FC761C17D">Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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<mods:namePart id="9ECCFE594B6272323664429E63BE9671">Meng, Kaibaryer</mods:namePart>
<mods:affiliation id="0599BBFE1370B6C2CDCB052D3DA2474F">National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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<mods:namePart id="57DCAF4A26D7F40D92A76D633A798E36">Asami, Takahiro</mods:namePart>
<mods:affiliation id="49AAF9243716F7B379EBF86DE9665401">Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
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<mods:namePart id="8190FE01DA721A57DA1F5C24D5A8818A">Ablett, Jonathan D.</mods:namePart>
<mods:affiliation id="DF9131C194D0DD5625AFBAC835083311">Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
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<treatment id="3C5BDE9812C55B70BE2EA6701ED8F079" ID-GBIF-Taxon="168715134" LSID="urn:lsid:plazi:treatment:3C5BDE9812C55B70BE2EA6701ED8F079" httpUri="http://treatment.plazi.org/id/3C5BDE9812C55B70BE2EA6701ED8F079" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="F7E71AF6433F6CFAAFA71C8B2F752B49" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="765F9C77B2677EB44FCA08DC1BA1E968" pageId="0" pageNumber="1">
<taxonomicName id="674D24757FBE4890714EB2877680A90F" LSID="3C5BDE98-12C5-5B70-BE2E-A6701ED8F079" authority="Venmans, 1956" authorityName="Venmans" authorityYear="1956" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus eydouxi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="eydouxi" superFamily="Cyclophoroidea">Alycaeus eydouxi Venmans, 1956</taxonomicName>
<figureCitation id="68AB82A158050C94602C59B2D097D059" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Protoconch sculpture of Alycaeus Gray, 1850 species A Alycaeus eydouxi Venmans, 1956 (Cochinchina, coll. V. W. MacAndrew, NHMUK) B Alycaeus rolfbrandti Maassen, 2006 (MNHN-IM- 2012 - 27321) C Alycaeus conformis Fulton, 1902 (HNHM 99714, spec 3). Scale bars: 1 mm. All images: Barna Pall-Gergely." figureDoi="10.3897/zookeys.981.53583.figure2" httpUri="https://binary.pensoft.net/fig/468647" pageId="0" pageNumber="1">Figs 2A</figureCitation>
<figureCitation id="95B645CD25400E76736255663ED823FB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Type species of alycaeid genus-group taxa A Alycaeus eydouxi Venmans, 1956 (SMF 109290; type species of Alycaeus) B Chamalycaeus fruhstorferi (Moellendorff, 1897), lectotype (SMF 109481; type species of Chamalycaeus). Photographs: Barna Pall-Gergely (A) and Frank Walther (B)." figureDoi="10.3897/zookeys.981.53583.figure6" httpUri="https://binary.pensoft.net/fig/468651" pageId="0" pageNumber="1">, 6A</figureCitation>
<figureCitation id="C7AB4F0E3023CE31199198736872E909" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Living specimens of Alycaeus eydouxi Venmans, 1856. Th ủy Sơn (Water Mountain), Ngũ Hanh Sơn (Five Elements or Marble Mountains), Da Nang. 16 ° 0.254 ' N, 108 ° 15.756 ' E. Photograph: Junn Kitt Foon." figureDoi="10.3897/zookeys.981.53583.figure8" httpUri="https://binary.pensoft.net/fig/468653" pageId="0" pageNumber="1">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8B3575142C0A6A4BCB4216BF08BF628E" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="4A532AC8DF0D8CAFAEBC5DC7A732F93E" pageId="0" pageNumber="1">
<taxonomicName id="A5BCFAED57B0B4BAB87136191431328E" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma gibbum" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbum">Cyclostoma gibbum</taxonomicName>
Eydoux, 1838: 6, pl. 117, fig 1. (non
<taxonomicName id="B4CCA6F2478A067440E6B77B33B99EAD" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma gibbum" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbum">Cyclostoma gibbum</taxonomicName>
Draparnaud, 1805)
</paragraph>
<paragraph id="93AC465EED30BB7027EC65CEB732CC85" pageId="0" pageNumber="1">
<taxonomicName id="F1B429D0410EF1E4B877348A0EAB68FC" baseAuthorityName="Eydoux" baseAuthorityYear="1838" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus gibbus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbus" superFamily="Cyclophoroidea">Alycaeus gibbus</taxonomicName>
-
<bibRefCitation id="B8046784B4B130FACFD0780EC319A59B" author="Reeve, LA" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/8230573" refId="B192" refString="Reeve, LA, 1878. Monograph of the genus Alycaeus. In: Reeve LA (Ed.) Conchologica Iconica: or, Illustrations of the Shells of Molluscous Animals. Reeve and Co., London, 20: 13 pp. [+ 6 pls] https://www.biodiversitylibrary.org/page/8230573" title="Monograph of the genus Alycaeus. In: Reeve LA (Ed.) Conchologica Iconica: or, Illustrations of the Shells of Molluscous Animals. Reeve and Co., London, 20: 13 pp. [+ 6 pls]" url="https://www.biodiversitylibrary.org/page/8230573" year="1878">Reeve 1878</bibRefCitation>
: pl. 1, species 3.
</paragraph>
<paragraph id="8B2B41C26F31E0476B494C2A9929E83A" pageId="0" pageNumber="1">
<taxonomicName id="F2B991A455572404DA873DB0B1552210" baseAuthorityName="Eydoux" baseAuthorityYear="1838" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Alycaeus) gibbus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbus" subGenus="Alycaeus" superFamily="Cyclophoroidea">Alycaeus (Alycaeus) gibbus</taxonomicName>
-
<bibRefCitation id="AFD4EFC2A00181D8ADB0BBE50538A7F8" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
: 344-345.
</paragraph>
<paragraph id="85334EA4359D754D996096C50A2E1336" pageId="0" pageNumber="1">
<taxonomicName id="FFD00A6B1A0423BE0D00B7AB59FB3538" baseAuthorityName="Eydoux" baseAuthorityYear="1838" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Orthalycaeus) gibbus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbus" subGenus="Orthalycaeus" superFamily="Cyclophoroidea">Alycaeus (Orthalycaeus) gibbus</taxonomicName>
- Godwin-Austen 1914: 427, pl. 156, figs 5, 5a.
</paragraph>
<paragraph id="A38A09555833F5EE72D33ABBCA92D173" pageId="0" pageNumber="1">
<taxonomicName id="2A7BEED6798C83A4E4598CF456CDA451" authorityName="Venmans" authorityYear="1956" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus eydouxi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="eydouxi" superFamily="Cyclophoroidea">Alycaeus eydouxi</taxonomicName>
Venmans, 1956: 87, figs 6, 7 (radula). (nom. nov. pro
<taxonomicName id="275DFE87489D12403C3B21FA4510636B" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma gibbum" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbum">Cyclostoma gibbum</taxonomicName>
Eydoux, 1838, non
<taxonomicName id="017DED577C496B1646FDCE2BEA7DCE41" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma gibbum" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gibbum">Cyclostoma gibbum</taxonomicName>
Draparnaud, 1805)
</paragraph>
<paragraph id="0D8BC4D140528AD9B8762FB219EC58C9" pageId="0" pageNumber="1">
<taxonomicName id="A658606B4A75B4985B028D89C95FA863" authorityName="Venmans" authorityYear="1956" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus eydouxi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="eydouxi" superFamily="Cyclophoroidea">Alycaeus eydouxi</taxonomicName>
-
<bibRefCitation id="27A0B3F07AE28DA1B22F1D12B0828D2B" author="Egorov, RV" journalOrPublisher="Egorov, Moscow" pageId="0" pageNumber="1" refId="B45" refString="Egorov, RV, 2013. Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R. Egorov, Moscow" title="Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R." year="2013">Egorov 2013</bibRefCitation>
: fig. 58a;
<bibRefCitation id="B322DB21CB42216673A8F1AEECBC7024" DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken id="8E84E832D669A66A1EF386ED9F9C01F8" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 9-10, fig. 3A.
</paragraph>
</subSubSection>
<subSubSection id="8F37535CC65FF0F0C7ACB4D5E6A03094" pageId="0" pageNumber="1" type="type locality">
<paragraph id="4A893FD84A8BB28E6D5BECFB898648CD" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="A2366F574F2BC6C7A6ADD48D7CF68F4C" pageId="0" pageNumber="1">
&quot;que dans les grottes
<normalizedToken id="D70A7E89AC9156D556B10DDA932700D0" originalValue="formées">formees</normalizedToken>
dans
<normalizedToken id="0341B36A852AA0730517C88C83AA926B" originalValue="l’intérieur">l'interieur</normalizedToken>
des montagnes de marbre qui
<normalizedToken id="135F44EA942C6E316C5B170BB63DFBD5" originalValue="s’élèvent">s'elevent</normalizedToken>
au milieu de la plaine
<normalizedToken id="65A95B450CD3DDE63A13942F99CFC093" originalValue="oú">ou</normalizedToken>
est
<normalizedToken id="66262F03274F5FF73DD03CD949FC7E31" originalValue="bâtie">batie</normalizedToken>
la ville de Turanne, en Cochinchine&quot;.
</paragraph>
</subSubSection>
<subSubSection id="EF664B80648C33ED7AF517A1909F063F" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="67ED1AF79765EEB37701C9995B383669" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="91D0DB39E5D8D17A3D23E5DFFDC98252" pageId="0" pageNumber="1">
Annam, Touranne, leg.
<normalizedToken id="BE87A7EFEA7F3B1DC8D2E2AE8B8804AE" originalValue="Frühstorfer">Fruehstorfer</normalizedToken>
, coll.
<normalizedToken id="7C6CBA591A7E7D78AC76AFA5007442A5" originalValue="Möllendorff">Moellendorff</normalizedToken>
, SMF 109290 (5 shells); Same data, NHMW 43182 (4 shells); Cochinchina, coll. V.W. MacAndrew, NHMUK.
</paragraph>
</subSubSection>
<subSubSection id="B8B0D57195144C865AEF311CF21BCAEC" pageId="0" pageNumber="1" type="remarks">
<paragraph id="4B6977D4BFEC495E2583DA842841F332" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="C770EB3F6804F3E7D088CF2B7C744C57" pageId="0" pageNumber="1">Protoconch matte, without spiral lines; R1 fine, dense, rather regular ribs with weak spiral striation; R2 long, with dense, lamellae-like, elevated ribs, which are most elevated closer to the suture. Below the ribs, the microtunnels are visible as narrow light bands between the darker, thicker stripes (visible where there are weathered areas of the shell).</paragraph>
<paragraph id="3EB553212C5F1C128F574B6FBBC8B896" pageId="0" pageNumber="1">
<bibRefCitation id="9416A3E5897EAB3152F1D8E703645EBD" author="Habe, T" journalOrPublisher="Nature and life in southeast Asia" pageId="0" pageNumber="1" pagination="111 - 128" refId="B75" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and life in southeast Asia 4: 111 - 128" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe (1965)</bibRefCitation>
reported &quot;
<emphasis id="41AD826E30583839A2A4917C43401CF3" italics="true" pageId="0" pageNumber="1">Dioryx gibbus</emphasis>
(Reeve)&quot; from &quot;Kao Phlong, north or Sara Buri, Central Thailand&quot;, without publishing a picture. This almost certainly refers to a different species.
</paragraph>
<caption id="887C071D3D6C9B4886C944D1D1DB13BD" doi="10.3897/zookeys.981.53583.figure8" httpUri="https://binary.pensoft.net/fig/468653" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph id="DB85AD5B92CE1E95209F327812A48562" pageId="0" pageNumber="1">
<emphasis id="5A5D761BF7A15C804B8D61EA49DD76D4" bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Living specimens of
<taxonomicName id="776566D25137743C785B37F13813F38E" authorityName="Venmans" authorityYear="1956" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus eydouxi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="eydouxi" superFamily="Cyclophoroidea">
<emphasis id="C57C7668ECB99FE6A98E668E97DF3DA2" italics="true" pageId="0" pageNumber="1">Alycaeus eydouxi</emphasis>
</taxonomicName>
Venmans, 1856. Thủy
<normalizedToken id="1A445DA2D0A10A4AD383BA71EC0F6CD0" originalValue="Sơn">Sơn</normalizedToken>
(Water Mountain),
<normalizedToken id="62B2372FD4646B983C87652B6128C7E5" originalValue="Ngũ">Ngũ</normalizedToken>
<normalizedToken id="9B80A1F38FBD00B7D7E8E7724563ABF4" originalValue="Hành">Hanh</normalizedToken>
<normalizedToken id="30423ED72B2855FBF8660E318C20471F" originalValue="Sơn">Sơn</normalizedToken>
(Five Elements or Marble Mountains), Da Nang. 16°0.254'N, 108°15.756'E. Photograph: Junn Kitt Foon.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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