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<document id="11919B462D82AE4BA87F349BA91A7215" ID-DOI="10.5852/ejt.2024.947.2635" ID-ISSN="2118-9773" ID-Zenodo-Dep="13643346" ID-ZooBank="5DC6E641-C928-48CC-A8D0-9144FD32DFB8" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725361599072" checkinUser="plazi" docAuthor="Makarikov, Arseny A. &amp; Stakheev, Valeriy V." docDate="2024" docId="200087D3F56AFFADFDB49316CA076D6D" docLanguage="en" docName="EJT.2024.947.268-288.pdf" docOrigin="European Journal of Taxonomy 947" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2635/12129" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Kontrimavichusia hobergi Makarikov &amp; Stakheev, 2024, sp. nov." docType="treatment" docUuid="8EBE9219-C6BA-4B85-8740-3A3E05F51581" docUuidSource="ZooBank" docVersion="5" lastPageNumber="282" masterDocId="DC39FFABF56DFFA3FFFE906BCF226B15" masterDocTitle="New species of Kontrimavichusia Makarikov &amp; Binkienė, 2022 (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae) from the North Caucasus" masterLastPageNumber="288" masterPageNumber="268" pageNumber="275" updateTime="1725885409113" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="89655A979E7CBFEEE98C03E367F68155">New species of Kontrimavichusia Makarikov &amp; Binkienė, 2022 (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae) from the North Caucasus</mods:title>
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<mods:namePart id="5ABF5410364DC1A445D0E631B32E31F5">Makarikov, Arseny A.</mods:namePart>
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<mods:affiliation id="B36FA2D6D5D35C60E78F30AD971A99CC">Institute of Systematics and Ecology of Animals, Siberian Branch, Russian Academy of Sciences, 11 Frunze Street, 630091 Novosibirsk, Russia.</mods:affiliation>
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<mods:affiliation id="05B33B092E39AF1ABA5DFE9275B4D0AA">The Southern Scientific Center of the Russian Academy of Sciences, 41 Chehova Street, 344006 Rostov-on-Don, Russia.</mods:affiliation>
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<heading id="F35E81A9F56AFFA4FDB49316CCCB688D" box="[586,1001,893,920]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="275" reason="2">
<taxonomicName id="6FA94D46F56AFFA4FDB49316CCAA688D" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[586,904,893,920]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="275" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F56AFFA4FDB49316CCAA688D" bold="true" box="[586,904,893,920]" italics="true" pageId="7" pageNumber="275">Kontrimavichusia hobergi</emphasis>
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<taxonomicNameLabel id="81EE57ACF56AFFA4FC719315CCCB688D" box="[911,1001,894,920]" pageId="7" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="30922A40F56AFFA4FD5E93AFCC2A68CB" box="[672,776,964,990]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="8.[189,232,1820,1846]" captionStartId-1="10.[189,232,1821,1847]" captionStartId-2="11.[189,232,1892,1918]" captionStartId-3="12.[190,233,1928,1954]" captionTargetBox-0="[227,1361,266,1775]" captionTargetBox-1="[189,1398,383,1772]" captionTargetBox-2="[219,1365,545,1848]" captionTargetBox-3="[190,1389,860,1876]" captionTargetId-0="figure-12@8.[227,1361,265,1783]" captionTargetId-1="figure-12@10.[189,1398,383,1781]" captionTargetId-2="figure-88@11.[216,1370,539,1855]" captionTargetId-3="figure-206@12.[189,1398,848,1889]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." captionText-1="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." captionText-2="Fig. 5. Maximum likelihood phylogenetic tree of Kontrimavichusia Makarikov &amp; Binkienė, 2022 and Hymenolepis Weinland, 1858 based on analysis of partial sequences of the 28S rRNA gene. Bootstrap support given for maximum likelihood analysis based on 1000 replicates. Bootstrap support values lower than 70% are not shown." captionText-3="Fig. 6. Maximum likelihood phylogenetic tree of species of Kontrimavichusia Makarikov &amp; Binkienė, 2022 based on analysis of partial sequences of the nad1 gen. Bootstrap support given for maximum likelihood analysis based on 1000 replicates. Bootstrap support values lower than 70% are not shown." figureDoi-0="http://doi.org/10.5281/zenodo.13643354" figureDoi-1="http://doi.org/10.5281/zenodo.13643356" figureDoi-2="http://doi.org/10.5281/zenodo.13643358" figureDoi-3="http://doi.org/10.5281/zenodo.13643360" httpUri-0="https://zenodo.org/record/13643354/files/figure.png" httpUri-1="https://zenodo.org/record/13643356/files/figure.png" httpUri-2="https://zenodo.org/record/13643358/files/figure.png" httpUri-3="https://zenodo.org/record/13643360/files/figure.png" pageId="7" pageNumber="275">Figs 3–6</figureCitation>
, 
<tableCitation id="E52B037EF56AFFA4FCEA93AFCCB668CB" box="[788,916,964,990]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="14.[189,259,265,291]" captionStartId-1="15.[184,254,547,575]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Table 1. Comparative morphometric data of species of Kontrimavichusia Makarikov &amp; Binkienė, 2022 (measurements in micrometres except where otherwise stated)." captionText-1="Table 2. Pairwise uncorrected genetic distances (below diagonal) and total nucleotide differences (above diagonal) in sequences of nad1 gen between specimens of Kontrimavichusia Makarikov &amp; Binkienė, 2022." httpUri-0="http://table.plazi.org/id/FCD6664DF563FFADFF439162CC8B6A53" httpUri-1="http://table.plazi.org/id/FCD6664DF562FFACFF469248CCBE6977" pageId="7" pageNumber="275" tableUuid-0="FCD6664DF563FFADFF439162CC8B6A53" tableUuid-1="FCD6664DF562FFACFF469248CCBE6977">Tables 1–2</tableCitation>
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<subSubSection id="E0B3654EF56AFFA4FF43946ECB7F6E37" pageId="7" pageNumber="275" type="diagnosis">
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<heading id="F35E81A9F56AFFA4FF43946ECE146F0A" bold="true" box="[189,310,1029,1055]" fontSize="11" level="3" pageId="7" pageNumber="275" reason="3">
<emphasis id="9ADDEAD7F56AFFA4FF43946ECE146F0A" bold="true" box="[189,310,1029,1055]" pageId="7" pageNumber="275">Diagnosis</emphasis>
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<taxonomicName id="6FA94D46F56AFFA4FF43945FCED26F5B" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[189,496,1076,1102]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="275" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F56AFFA4FF43945FCED26F5B" box="[189,496,1076,1102]" italics="true" pageId="7" pageNumber="275">Kontrimavichusia hobergi</emphasis>
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<taxonomicNameLabel id="81EE57ACF56AFFA4FE08945ECD726F5A" box="[502,592,1077,1103]" pageId="7" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
has morphological characters typical of the genus 
<taxonomicName id="6FA94D46F56AFFA4FB62945FCA4C6F5B" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" box="[1180,1390,1076,1102]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="275" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F56AFFA4FB62945FCA4C6F5B" box="[1180,1390,1076,1102]" italics="true" pageId="7" pageNumber="275">Kontrimavichusia</emphasis>
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, namely rhynchus armed with cricetoid-like hooks, apex of rostellum invaginable and blades of retracted hooks directed anteriorly, suckers armed with minute spines, ventral canals connected by irregularly spaced transverse anastomoses, copulatory part of vagina surrounded by circular musculature and covered externally by dense layer of intensely-stained cells, labyrinthine uterus extending beyond osmoregulatory canals into both lateral fields, situated dorsally to osmoregulatory canals and genital ducts and embryophore without polar filaments (
<bibRefCitation id="CC384B34F56AFFA4FD09956CCB726E37" author="Makarikov A. A. &amp; Binkiene R." box="[759,1104,1287,1314]" pageId="7" pageNumber="275" refId="ref11151" refString="Makarikov A. A. &amp; Binkiene R. 2022. Redescription and taxonomic position of Rodentolepis (sensu lato) asymmetrica (Janicki, 1904), with the erection of Kontrimavichusia n. g. (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae). Journal of Helminthology 96: E 63. https: // doi. org / 10.1017 / S 0022149 X 22000505" type="journal volume" year="2022">Makarikov &amp; Binkienė 2022</bibRefCitation>
).
</paragraph>
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<subSubSection id="E0B3654EF56AFFA4FF439522CB346D0B" pageId="7" pageNumber="275" type="etymology">
<paragraph id="A81636C5F56AFFA4FF439522CE666E76" blockId="7.[189,324,1353,1379]" box="[189,324,1353,1379]" pageId="7" pageNumber="275">
<heading id="F35E81A9F56AFFA4FF439522CE666E76" bold="true" box="[189,324,1353,1379]" fontSize="11" level="3" pageId="7" pageNumber="275" reason="3">
<emphasis id="9ADDEAD7F56AFFA4FF439522CE666E76" bold="true" box="[189,324,1353,1379]" pageId="7" pageNumber="275">Etymology</emphasis>
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This species has been named in honour of the outstanding parasitologist Dr Eric P. Hoberg in recognition of his seminal and critical studies of parasites of vertebrates, helminth systematics, biogeography, ecology, phylogeny and evolution. It is also recognition of his significant contribution to the advancement of parasitology and the preservation of important archival materials on helminths during his tenure as chief curator of the 
<collectingCountry id="D0BE7655F56AFFA4FE58966FCEF86D0B" box="[422,474,1540,1566]" name="United States of America" pageId="7" pageNumber="275">U.S.</collectingCountry>
National Parasite Collection from 1990 to 2014.
</paragraph>
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<emphasis id="9ADDEAD7F56AFFA4FF43962DCE4E6D4A" bold="true" box="[189,364,1605,1632]" pageId="7" pageNumber="275">Type material</emphasis>
</heading>
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<typeStatus id="77128867F56AFFA4FF14961FCE7E6D9B" box="[234,348,1652,1678]" pageId="7" pageNumber="275" type="holotype">Holotype</typeStatus>
</emphasis>
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<collectingCountry id="D0BE7655F56AFFA4FF4396FCCE076DA4" box="[189,293,1687,1713]" name="Russia" pageId="7" pageNumber="275">RUSSIA</collectingCountry>
• 
<collectingRegion id="6A6DF827F56AFFA4FEC496FCCDE16DA4" box="[314,707,1687,1713]" pageId="7" pageNumber="275">Republic of North Ossetia-Alania</collectingRegion>
, 
<collectingMunicipality id="4872ACBFF56AFFA4FD3296FCCC836DA4" box="[716,929,1687,1713]" pageId="7" pageNumber="275">Alagirsky District</collectingMunicipality>
, suburbs of 
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; 
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, 
<geoCoordinate id="CD9D5002F56AFFA4FF4396D2CE6F6DC0" box="[189,333,1721,1749]" degrees="43" direction="east" minutes="56" orientation="longitude" pageId="7" pageNumber="275" precision="15" seconds="03" value="43.934166">43º56′03″ E</geoCoordinate>
; 
<date id="DC171005F56AFFA4FEA796D0CECF6DC0" box="[345,493,1722,1749]" pageId="7" pageNumber="275" value="2017-07-27">
<collectingDate id="CC53E9EDF56AFFA4FEA796D0CECF6DC0" box="[345,493,1722,1749]" pageId="7" pageNumber="275" value="2017-07-27">27 Jul. 2017</collectingDate>
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; 
<collectorName id="055C5313F56AFFA4FE0696D0CD9B6DC1" box="[504,697,1722,1749]" pageId="7" pageNumber="275">A.A. Makarikov</collectorName>
leg.; 
<specimenCode id="F80F9EBEF56AFFA4FD0596D0CCFD6DC1" box="[763,991,1722,1749]" pageId="7" pageNumber="275">ISEA AM17-236#3</specimenCode>
(ex 379).
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<materialsCitation id="18C13C98F56AFFA4FF149697CCE46C2C" collectingDate="2017-07-27" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="RUSSIA" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-236#1" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<paragraph id="A81636C5F56AFFA4FF149697CE456C03" blockId="7.[188,1400,1788,1991]" box="[234,359,1788,1814]" pageId="7" pageNumber="275">
<emphasis id="9ADDEAD7F56AFFA4FF149697CE456C03" bold="true" box="[234,359,1788,1814]" pageId="7" pageNumber="275">
<typeStatus id="77128867F56AFFA4FF149697CE456C03" box="[234,359,1788,1814]" pageId="7" pageNumber="275" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="A81636C5F56AFFAAFF439774CD316A98" blockId="7.[188,1400,1788,1991]" lastBlockId="9.[188,1399,265,397]" lastPageId="9" lastPageNumber="277" pageId="7" pageNumber="275">
<collectingCountry id="D0BE7655F56AFFA4FF439774CE076C2C" box="[189,293,1823,1849]" name="Russia" pageId="7" pageNumber="275">RUSSIA</collectingCountry>
• 
<specimenCount id="BEAFFD4CF56AFFA4FEC19774CEB26C2F" box="[319,400,1823,1850]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FD1E9774CCE46C2C" box="[736,966,1823,1849]" pageId="7" pageNumber="275">ISEA AM17-236#1</specimenCode>
• 
<materialsCitation id="18C13C98F56AFFA4FC1C9774CD286C49" collectingDate="2017-07-27" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-236#2" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FC1C9774CB126C2F" box="[994,1072,1823,1850]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FF409729CE806C49" box="[190,418,1858,1884]" pageId="7" pageNumber="275">ISEA AM17-236#2</specimenCode>
(ex 378)
</materialsCitation>
• 
<materialsCitation id="18C13C98F56AFFA4FDDF9729CBB16C49" box="[545,1171,1858,1885]" collectingDate="2017-07-27" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-240#2" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FDDF9729CD4F6C48" box="[545,621,1858,1885]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FC519729CBB16C49" box="[943,1171,1858,1884]" pageId="7" pageNumber="275">ISEA AM17-240#2</specimenCode>
</materialsCitation>
• 
<materialsCitation id="18C13C98F56AFFA4FB559729CD926C95" collectingDate="2017-07-27" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-242" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FB559729CBD56C48" box="[1195,1271,1858,1885]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FE83970DCD676C95" box="[381,581,1894,1920]" pageId="7" pageNumber="275">ISEA AM17-242</specimenCode>
(ex 382)
</materialsCitation>
• 
<materialsCitation id="18C13C98F56AFFA4FD35970DCFD16CB6" collectingDate="2017-07-27" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-243#1" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FD35970DCC386C95" box="[715,794,1894,1920]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FB9A970DCA6B6C95" box="[1124,1353,1894,1920]" pageId="7" pageNumber="275">ISEA AM17-243#1</specimenCode>
(ex 377)
</materialsCitation>
• 
<materialsCitation id="18C13C98F56AFFA4FEF397E2CCAF6CB6" box="[269,909,1929,1955]" collectingDate="2017-07-27" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-243#2" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FEF397E2CE7E6CB6" box="[269,348,1929,1955]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FD5697E2CCAF6CB6" box="[680,909,1929,1955]" pageId="7" pageNumber="275">ISEA AM17-243#2</specimenCode>
</materialsCitation>
• 
<materialsCitation id="18C13C98F56AFFA4FC5997E2CEE86CD3" collectingDate="2017-07-27" collectionCode="AM" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="SEA AM17-243#3" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FC5997E2CCD46CB6" box="[935,1014,1929,1955]" count="1" pageId="7" pageNumber="275" type="generic">1 spec</specimenCount>
; same data as for holotype; 
<specimenCode id="F80F9EBEF56AFFA4FABF97E2CE7C6CD3" pageId="7" pageNumber="275">SEA AM17-243#3</specimenCode>
(ex 376)
</materialsCitation>
• 
<materialsCitation id="18C13C98F56AFFA4FE1897C7CA5A6CD3" box="[486,1400,1964,1991]" collectingDate="2017-07-25" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="7" pageNumber="275" specimenCode="ISEA AM17-206" specimenCount="1" stateProvince="North Ossetia" typeStatus="paratype">
<specimenCount id="BEAFFD4CF56AFFA4FE1897C7CD1C6CD2" box="[486,574,1964,1991]" count="1" pageId="7" pageNumber="275" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<date id="DC171005F56AFFA4FC6397C7CB166CD3" box="[925,1076,1964,1991]" pageId="7" pageNumber="275" value="2017-07-25">
<collectingDate id="CC53E9EDF56AFFA4FC6397C7CB166CD3" box="[925,1076,1964,1991]" pageId="7" pageNumber="275" value="2017-07-25">25 Jul. 2017</collectingDate>
</date>
; 
<specimenCode id="F80F9EBEF56AFFA4FBBC97C7CA296CD3" box="[1090,1291,1964,1990]" pageId="7" pageNumber="275">ISEA AM17-206</specimenCode>
(ex 477)
</materialsCitation>
• 
<materialsCitation id="18C13C98F564FFAAFF2E9162CC916A36" box="[208,947,265,291]" collectingDate="2017-07-25" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-211" specimenCount="1" stateProvince="North Ossetia">
<specimenCount id="BEAFFD4CF564FFAAFF2E9162CE056A36" box="[208,295,265,291]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<specimenCode id="F80F9EBEF564FFAAFD7E9162CC6A6A36" box="[640,840,265,291]" pageId="9" pageNumber="277">ISEA AM17-211</specimenCode>
(ex 474)
</materialsCitation>
• 
<materialsCitation id="18C13C98F564FFAAFC309162CEAB6A53" collectingDate="2017-07-25" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-213" specimenCount="1" stateProvince="North Ossetia">
<specimenCount id="BEAFFD4CF564FFAAFC309162CB076A36" box="[974,1061,265,291]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<specimenCode id="F80F9EBEF564FFAAFF439147CEAB6A53" box="[189,393,300,326]" pageId="9" pageNumber="277">ISEA AM17-213</specimenCode>
</materialsCitation>
• 
<materialsCitation id="18C13C98F564FFAAFE549147CA546A53" box="[426,1398,300,327]" collectingDate="2017-07-26" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-222#1" specimenCount="1" stateProvince="North Ossetia">
<specimenCount id="BEAFFD4CF564FFAAFE549147CD216A52" box="[426,515,300,327]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<date id="DC171005F564FFAAFC919147CB296A53" box="[879,1035,300,326]" pageId="9" pageNumber="277" value="2017-07-26">
<collectingDate id="CC53E9EDF564FFAAFC919147CB296A53" box="[879,1035,300,326]" pageId="9" pageNumber="277" value="2017-07-26">26 Jul. 2017</collectingDate>
</date>
; 
<specimenCode id="F80F9EBEF564FFAAFBE59147CA266A53" box="[1051,1284,300,326]" pageId="9" pageNumber="277">ISEA AM17-222#1</specimenCode>
(ex 368)
</materialsCitation>
• 
<materialsCitation id="18C13C98F564FFAAFF309124CC9D6A7C" box="[206,959,335,362]" collectingDate="2017-07-26" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-222#2" specimenCount="1" stateProvince="North Ossetia">
<specimenCount id="BEAFFD4CF564FFAAFF309124CE006A7F" box="[206,290,335,362]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<specimenCode id="F80F9EBEF564FFAAFD8D9124CC756A7C" box="[627,855,335,361]" pageId="9" pageNumber="277">ISEA AM17-222#2</specimenCode>
(ex 369)
</materialsCitation>
• 
<materialsCitation id="18C13C98F564FFAAFC269124CD2D6A98" collectingDate="2017-07-26" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-226#1" specimenCount="1" stateProvince="North Ossetia">
<specimenCount id="BEAFFD4CF564FFAAFC269124CB0E6A7F" box="[984,1068,335,362]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<specimenCode id="F80F9EBEF564FFAAFF429118CE836A99" box="[188,417,370,397]" pageId="9" pageNumber="277">ISEA AM17-226#1</specimenCode>
(ex 482)
</materialsCitation>
.
</paragraph>
</materialsCitation>
<caption id="FCD6664DF565FFABFF439777CD276CF2" ID-DOI="http://doi.org/10.5281/zenodo.13643354" ID-Zenodo-Dep="13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="8" pageNumber="276" startId="8.[189,232,1820,1846]" targetBox="[227,1361,266,1775]" targetPageId="8">
<paragraph id="A81636C5F565FFABFF439777CD276CF2" blockId="8.[189,1400,1820,2023]" pageId="8" pageNumber="276">
<emphasis id="9ADDEAD7F565FFABFF439777CE2F6C23" bold="true" box="[189,269,1820,1846]" pageId="8" pageNumber="276">Fig. 3.</emphasis>
<taxonomicName id="6FA94D46F565FFABFEE99777CD686C23" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[279,586,1820,1846]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="276" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F565FFABFEE99777CD686C23" box="[279,586,1820,1846]" italics="true" pageId="8" pageNumber="276">Kontrimavichusia hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF565FFABFDAD9776CD8D6C22" box="[595,687,1821,1847]" pageId="8" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9ADDEAD7F565FFABFD469777CDEE6C23" bold="true" box="[696,716,1820,1846]" pageId="8" pageNumber="276">A</emphasis>
. Paratype (ISEA AM17-242), scolex, dorso-ventral view. 
<emphasis id="9ADDEAD7F565FFABFF40972BCFF26C4F" bold="true" box="[190,208,1856,1882]" pageId="8" pageNumber="276">B</emphasis>
. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. 
<emphasis id="9ADDEAD7F565FFABFC609754CC936C4C" bold="true" box="[926,945,1855,1881]" pageId="8" pageNumber="276">C</emphasis>
. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). 
<emphasis id="9ADDEAD7F565FFABFE4997EDCEEE6CB5" bold="true" box="[439,460,1926,1952]" pageId="8" pageNumber="276">D</emphasis>
. Holotype, male mature proglottides, dorsal view. 
<emphasis id="9ADDEAD7F565FFABFBDA97EDCB156CB5" bold="true" box="[1060,1079,1926,1952]" pageId="8" pageNumber="276">E</emphasis>
. Holotype, hermaphroditic mature proglottis, dorsal view. 
<emphasis id="9ADDEAD7F565FFABFDCC97C2CD606CD6" bold="true" box="[562,578,1961,1987]" pageId="8" pageNumber="276">F</emphasis>
. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm.
</paragraph>
</caption>
<paragraph id="A81636C5F564FFAAFF4391DFCEDB6ADB" blockId="9.[189,505,436,462]" box="[189,505,436,462]" pageId="9" pageNumber="277">
<heading id="F35E81A9F564FFAAFF4391DFCEDB6ADB" bold="true" box="[189,505,436,462]" fontSize="11" level="3" pageId="9" pageNumber="277" reason="3">
<emphasis id="9ADDEAD7F564FFAAFF4391DFCEDB6ADB" bold="true" box="[189,505,436,462]" pageId="9" pageNumber="277">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A81636C5F564FFAAFF439188CC336934" blockId="9.[189,1398,482,545]" pageId="9" pageNumber="277">
<materialsCitation id="18C13C98F564FFAAFF439188CBE26AE8" box="[189,1216,482,509]" collectingDate="2017-07-26" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="RUSSIA" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-221#3" specimenCount="1" stateProvince="North Ossetia">
<collectingCountry id="D0BE7655F564FFAAFF439188CE076AE8" box="[189,293,483,509]" name="Russia" pageId="9" pageNumber="277">RUSSIA</collectingCountry>
• 
<specimenCount id="BEAFFD4CF564FFAAFEC39188CEB06AE8" box="[317,402,483,509]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for holotype; 
<date id="DC171005F564FFAAFD279189CC4E6AE8" box="[729,876,482,509]" pageId="9" pageNumber="277" value="2017-07-26">
<collectingDate id="CC53E9EDF564FFAAFD279189CC4E6AE8" box="[729,876,482,509]" pageId="9" pageNumber="277" value="2017-07-26">26 Jul. 2017</collectingDate>
</date>
; 
<specimenCode id="F80F9EBEF564FFAAFC879188CB7C6AE8" box="[889,1118,483,509]" pageId="9" pageNumber="277">ISEA AM17-221#3</specimenCode>
(ex455)
</materialsCitation>
• 
<materialsCitation id="18C13C98F564FFAAFB279188CC2E6934" collectingDate="2017-07-26" collectionCode="ISEA" collectorName="A. A. Makarikov" country="Russia" county="Republic" latitude="42.800556" location="Verkhniy Tsey" longLatPrecision="19" longitude="43.934166" municipality="Alagirsky District" pageId="9" pageNumber="277" specimenCode="ISEA AM17-221#4" specimenCount="1" stateProvince="North Ossetia">
<specimenCount id="BEAFFD4CF564FFAAFB279188CA0C6AE8" box="[1241,1326,483,509]" count="1" pageId="9" pageNumber="277" type="generic">1 spec.</specimenCount>
; same data as for preceding; 
<specimenCode id="F80F9EBEF564FFAAFE3C926DCD856935" box="[450,679,518,544]" pageId="9" pageNumber="277">ISEA AM17-221#4</specimenCode>
(ex456)
</materialsCitation>
.
</paragraph>
<paragraph id="A81636C5F564FFAAFF43922CCE146974" blockId="9.[189,310,583,609]" box="[189,310,583,609]" pageId="9" pageNumber="277">
<heading id="F35E81A9F564FFAAFF43922CCE146974" bold="true" box="[189,310,583,609]" fontSize="11" level="3" pageId="9" pageNumber="277" reason="3">
<emphasis id="9ADDEAD7F564FFAAFF43922CCE146974" bold="true" box="[189,310,583,609]" pageId="9" pageNumber="277">Type host</emphasis>
</heading>
</paragraph>
<paragraph id="A81636C5F564FFAAFF43921DCB796985" blockId="9.[189,1115,630,656]" box="[189,1115,630,656]" pageId="9" pageNumber="277">
<taxonomicName id="6FA94D46F564FFAAFF43921DCD996985" authority="(Shidlovsky, 1919)" baseAuthorityName="Shidlovsky" baseAuthorityYear="1919" box="[189,699,630,656]" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="277" phylum="Chordata" rank="species" species="daghestanicus">
<emphasis id="9ADDEAD7F564FFAAFF43921DCEF76985" box="[189,469,630,656]" italics="true" pageId="9" pageNumber="277">Microtus daghestanicus</emphasis>
(Shidlovsky, 1919)
</taxonomicName>
(
<taxonomicName id="6FA94D46F564FFAAFD35921DCC196985" authorityName="Bowdich" authorityYear="1821" box="[715,827,630,656]" class="Mammalia" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="277" phylum="Chordata" rank="order">Rodentia</taxonomicName>
: 
<taxonomicName id="6FA94D46F564FFAAFCB8921DCCE16985" authorityName="Fischer" authorityYear="1817" box="[838,963,630,656]" class="Mammalia" family="Cricetidae" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="277" phylum="Chordata" rank="family">Cricetidae</taxonomicName>
: Arvicoline).
</paragraph>
</subSubSection>
<subSubSection id="E0B3654EF564FFAEFF4392DCCC946A53" lastPageId="13" lastPageNumber="281" pageId="9" pageNumber="277" type="description">
<paragraph id="A81636C5F564FFAAFF4392DCCEED69C4" blockId="9.[189,463,695,721]" box="[189,463,695,721]" pageId="9" pageNumber="277">
<emphasis id="9ADDEAD7F564FFAAFF4392DCCE6C69C4" bold="true" box="[189,334,695,721]" pageId="9" pageNumber="277">Description</emphasis>
(
<figureCitation id="30922A40F564FFAAFEA092DCCEE469C4" box="[350,454,695,721]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1820,1846]" captionStartId-1="10.[189,232,1821,1847]" captionTargetBox-0="[227,1361,266,1775]" captionTargetBox-1="[189,1398,383,1772]" captionTargetId-0="figure-12@8.[227,1361,265,1783]" captionTargetId-1="figure-12@10.[189,1398,383,1781]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionText-0="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." captionText-1="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13643354" figureDoi-1="http://doi.org/10.5281/zenodo.13643356" httpUri-0="https://zenodo.org/record/13643354/files/figure.png" httpUri-1="https://zenodo.org/record/13643356/files/figure.png" pageId="9" pageNumber="277">Figs 3–4</figureCitation>
)
</paragraph>
<paragraph id="A81636C5F564FFAAFF43928DCEC16E05" blockId="9.[188,1401,742,1296]" pageId="9" pageNumber="277">
[Based on 11 stained and mounted specimens and 7 scoleces mounted in Berlese’s medium.] Fully developed strobila 65–78 (74; n = 7) mm long, with maximum width 1.7–2.1 (1.8; n = 5) mm at level gravid proglottides. Strobila flat, consisting of 400–480 craspedote proglottides. Scolex slightly compressed dorso-ventrally, 245–360 wide (301; n = 3), not clearly distinct from neck. Suckers small, thick-walled, rounded or oval, cup-shaped, 120–155 × 100–120 (138 × 109; n = 12), usually reaching lateral margins of scolex, armed with minute (less than 1 long) spines; spines covering entire sucker surface (
<figureCitation id="30922A40F564FFAAFEDB93D2CE8568C6" box="[293,423,953,979]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3A–B</figureCitation>
). Rostellar pouch 130–164 × 95–125 (148 × 112; n = 3), with muscular walls, its bottom not reaching level of posterior margin of suckers. Rostellum 105–161 × 50–70 (130 × 61; n = 3), sac-like, muscular, apex invaginable; when rostellar apparatus retracted, rostellar hooks with blades directed anteriorly (
<figureCitation id="30922A40F564FFAAFE4E9449CD2F6F28" box="[432,525,1058,1085]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3B</figureCitation>
). Rhynchus 60–73 long and 50–75 wide, with well-developed circular musculature, armed with single crown of 18–22 (n = 7) rostellar hooks of cricetoid-like 
<typeStatus id="77128867F564FFAAFB37942DCBDE6F75" box="[1225,1276,1094,1120]" pageId="9" pageNumber="277">type</typeStatus>
(
<figureCitation id="30922A40F564FFAAFAF2942DCA4B6F75" box="[1292,1385,1094,1120]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3C</figureCitation>
). Rostellar hooks with relatively short handle and straight blade; axis of blade situated to axis of guard at acute angle; guard narrow in anterior surface; handle and blade slightly shorter or equal in length with guard. Hook measurements: total length 20–24 (21.8; n = 39), handle 8–9.5 (8.7; n = 39), blade 7.8–9.4 (8.6; n = 39) and guard 8–10.6 (9; n = 39). Neck 200–240 wide (n = 3), approximately equal in width with scolex (
<figureCitation id="30922A40F564FFAAFEAB949DCEF46E05" box="[341,470,1270,1296]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3A–B</figureCitation>
).
</paragraph>
<paragraph id="A81636C5F564FFAAFF43955CCE736D14" blockId="9.[189,1399,1335,1537]" pageId="9" pageNumber="277">
Ventral osmoregulatory canals 40–110 (79; n = 36) wide, connected by irregularly spaced transverse anastomoses (present in up to 23% proglottides) (
<figureCitation id="30922A40F564FFAAFCE79531CC596E61" box="[793,891,1370,1396]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3D</figureCitation>
). Dorsal osmoregulatory canals very thin, 3–8 (5; n = 36) wide at level of hermaphroditic proglottides, usually situated directly dorsal (not shifted left or right) to ventral canals. Genital pores unilateral, dextral (
<figureCitation id="30922A40F564FFAAFC5995CACB016EAE" box="[935,1059,1441,1467]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3D–E</figureCitation>
). Genital ducts pass dorsally to both ventral and dorsal longitudinal osmoregulatory canals. Development of proglottides gradual, protandrous.
</paragraph>
<paragraph id="A81636C5F564FFA8FF439642CC386A53" blockId="9.[187,1400,1576,1991]" lastBlockId="11.[189,1398,265,326]" lastPageId="11" lastPageNumber="279" pageId="9" pageNumber="277">
Mature proglottides 180–240 × 970–1140 (210 × 1085; n = 29), transversely elongate, trapezoid (
<figureCitation id="30922A40F564FFAAFF399627CE6A6D73" box="[199,328,1612,1638]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3D–E</figureCitation>
). Testes 3, relatively large, almost equal in size, 135–210 × 95–150 (169 × 119; n = 36), round or oval, most often situated in triangle with flat angle (anterior antiporal testis shifted to lateral side of proglottis in relation to posterior antiporal testis) or, rarely, in triangle with right angle or in one row, poral testis separated from two antiporal testes by female gonads. Number and distribuition of testes constant, no variation in testes number observed, proglottis with three antiporal testes or two poral testes appear infrequent. Cirrus-sac relatively short, with thick muscular walls, club-shaped, 270–320 × 56–75 (296 × 66; n = 28). Antiporal part of cirrus-sac substantially crossing poral ventral longitudinal canal (
<figureCitation id="30922A40F564FFAAFF369729CE656C49" box="[200,327,1858,1884]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3E–F</figureCitation>
). Genital atrium simple, cup-shaped, opens laterally, approximately in middle of lateral proglottis margin. Cirrus large, 100–128 × 24–39 (112 × 31; n = 25), cylindrical, armed with very small (up to 1.0–1.5 long), needle-shaped spines (
<figureCitation id="30922A40F564FFAAFD4497E3CC356CB6" box="[698,791,1928,1955]" captionStart="Fig" captionStartId="10.[189,232,1821,1847]" captionTargetBox="[189,1398,383,1772]" captionTargetId="figure-12@10.[189,1398,383,1781]" captionTargetPageId="10" captionText="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="9" pageNumber="277">Fig. 4A</figureCitation>
). Internal seminal vesicle with circular musculature, ovoid, 130–180 × 52–70 (155 × 58; n = 28), occupying half of cirrus-sac length (
<figureCitation id="30922A40F564FFAAFB7D97C7CBDC6CD3" box="[1155,1278,1964,1990]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="9" pageNumber="277">Fig. 3E–F</figureCitation>
). External seminal vesicle, 105–190 × 82–120 (127 × 93; n = 28), round or oval, clearly distinguishable from vas deferens, normally smaller than seminal receptacle.
</paragraph>
<caption id="FCD6664DF567FFA9FF439776CC7D6CD0" ID-DOI="http://doi.org/10.5281/zenodo.13643356" ID-Zenodo-Dep="13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="10" pageNumber="278" startId="10.[189,232,1821,1847]" targetBox="[189,1398,383,1772]" targetPageId="10">
<paragraph id="A81636C5F567FFA9FF439776CC7D6CD0" blockId="10.[188,1399,1821,1989]" pageId="10" pageNumber="278">
<emphasis id="9ADDEAD7F567FFA9FF439776CE2E6C22" bold="true" box="[189,268,1821,1847]" pageId="10" pageNumber="278">Fig. 4.</emphasis>
<taxonomicName id="6FA94D46F567FFA9FEEA9776CD656C22" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[276,583,1821,1847]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="278" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F567FFA9FEEA9776CD656C22" box="[276,583,1821,1847]" italics="true" pageId="10" pageNumber="278">Kontrimavichusia hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF567FFA9FDB19775CD886C2D" box="[591,682,1822,1848]" pageId="10" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9ADDEAD7F567FFA9FD4F9776CDE46C22" bold="true" box="[689,710,1821,1847]" pageId="10" pageNumber="278">A</emphasis>
. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. 
<emphasis id="9ADDEAD7F567FFA9FEFF972ACE376C4E" bold="true" box="[257,277,1857,1883]" pageId="10" pageNumber="278">B</emphasis>
. Holotype (ISEA AM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. 
<emphasis id="9ADDEAD7F567FFA9FEAE970FCE476C6B" bold="true" box="[336,357,1892,1918]" pageId="10" pageNumber="278">C</emphasis>
. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. 
<emphasis id="9ADDEAD7F567FFA9FB58970FCB996C6B" bold="true" box="[1190,1211,1892,1918]" pageId="10" pageNumber="278">D</emphasis>
. Holotype, egg. 
<emphasis id="9ADDEAD7F567FFA9FF4297ECCFF26CB4" bold="true" box="[188,208,1927,1953]" pageId="10" pageNumber="278">E</emphasis>
. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm.
</paragraph>
</caption>
<paragraph id="A81636C5F566FFAFFF439105CCA56AA4" blockId="11.[187,1398,365,463]" lastBlockId="12.[189,1400,265,433]" lastPageId="12" lastPageNumber="280" pageId="11" pageNumber="279">
Ovary 280–396 (330; n = 25) wide, median, transversely elongate, fan-shaped, irregularly lobed, ventral to male genital organs, occupying less than half of median field, overlapping testes (
<figureCitation id="30922A40F566FFA8FB7891FACBFC6ABE" box="[1158,1246,401,427]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="11" pageNumber="279">Fig. 3E</figureCitation>
). Vitellarium 70–116 × 120–185 (85 × 145; n = 25), postovarian, slightly shifted to lateral side of proglottis, slightly lobed. Vagina tubular, clearly distinct from seminal receptacle; ventral to cirrus-sac. Copulatory part of vagina 94–120 × 8–32 (105 × 16; n = 25), shorter than cirrus-sac, thick-walled, surrounded by circular musculature and covered externally by dense layer of intensely stained cells; proximal part of vagina infundibular (
<figureCitation id="30922A40F561FFAFFE9C9119CEE26A98" box="[354,448,370,397]" captionStart="Fig" captionStartId="8.[189,232,1820,1846]" captionTargetBox="[227,1361,266,1775]" captionTargetId="figure-12@8.[227,1361,265,1783]" captionTargetPageId="8" captionText="Fig. 3. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-242), scolex, dorso-ventral view. B. Paratype (ISEA AM1 17-243#3), scolex, sub-lateral view. C. Holotype (ISEA AM17-236#3) (left) and paratype (ISEA AM 17-236#2) (centre, right), rostellar hooks in profile and frontal view (note narrow hook guard). D. Holotype, male mature proglottides, dorsal view. E. Holotype, hermaphroditic mature proglottis, dorsal view. F. Holotype, genital ducts, dorsal view. Scale bars: A–B, F = 100 µm; C = 10 µm; D–E = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.13643354" httpUri="https://zenodo.org/record/13643354/files/figure.png" pageId="12" pageNumber="280">Figs 3F</figureCitation>
, 
<figureCitation id="30922A40F561FFAFFE319118CED56A98" box="[463,503,371,397]" captionStart="Fig" captionStartId="10.[189,232,1821,1847]" captionTargetBox="[189,1398,383,1772]" captionTargetId="figure-12@10.[189,1398,383,1781]" captionTargetPageId="10" captionText="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="12" pageNumber="280">4A</figureCitation>
). Conductive part of vagina indistinct. Seminal receptacle relatively large, transversely elongate, 315–470 × 55–100 (387 × 75; n = 25).
</paragraph>
<caption id="FCD6664DF566FFA8FF43970FCD086CFD" ID-DOI="http://doi.org/10.5281/zenodo.13643358" ID-Zenodo-Dep="13643358" httpUri="https://zenodo.org/record/13643358/files/figure.png" pageId="11" pageNumber="279" startId="11.[189,232,1892,1918]" targetBox="[219,1365,545,1848]" targetPageId="11">
<paragraph id="A81636C5F566FFA8FF43970FCD086CFD" blockId="11.[189,1400,1892,2025]" pageId="11" pageNumber="279">
<emphasis id="9ADDEAD7F566FFA8FF43970FCE2F6C6A" bold="true" box="[189,269,1892,1919]" pageId="11" pageNumber="279">Fig. 5.</emphasis>
Maximum likelihood phylogenetic tree of 
<taxonomicName id="6FA94D46F566FFA8FCF3970ECA666C6A" authority="Makarikov &amp; Binkiene, 2022" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" box="[781,1348,1893,1919]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="279" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F566FFA8FCF3970ECCFD6C6A" box="[781,991,1893,1919]" italics="true" pageId="11" pageNumber="279">Kontrimavichusia</emphasis>
<bibRefCitation id="CC384B34F566FFA8FC19970ECA666C6A" author="Makarikov A. A. &amp; Binkiene R." box="[999,1348,1893,1919]" pageId="11" pageNumber="279" refId="ref11151" refString="Makarikov A. A. &amp; Binkiene R. 2022. Redescription and taxonomic position of Rodentolepis (sensu lato) asymmetrica (Janicki, 1904), with the erection of Kontrimavichusia n. g. (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae). Journal of Helminthology 96: E 63. https: // doi. org / 10.1017 / S 0022149 X 22000505" type="journal volume" year="2022">Makarikov &amp; Binkienė, 2022</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="6FA94D46F566FFA8FF4397E3CD3A6CB6" authority="Weinland, 1858" authorityName="Weinland" authorityYear="1858" box="[189,536,1928,1955]" class="Cestoda" family="Hymenolepididae" genus="Hymenolepis" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="279" phylum="Platyhelminthes" rank="genus">
<emphasis id="9ADDEAD7F566FFA8FF4397E3CE746CB7" box="[189,342,1928,1954]" italics="true" pageId="11" pageNumber="279">Hymenolepis</emphasis>
Weinland, 1858
</taxonomicName>
based on analysis of partial sequences of the 28S rRNA gene. Bootstrap support given for maximum likelihood analysis based on 1000 replicates. Bootstrap support values lower than 70% are not shown.
</paragraph>
</caption>
<paragraph id="A81636C5F561FFAEFF4391B3CC946A53" blockId="12.[188,1399,471,779]" lastBlockId="13.[189,1399,265,327]" lastPageId="13" lastPageNumber="281" pageId="12" pageNumber="280">
Uterus appears as perforated, transversely-elongate band, situated dorsally to testes, genital ducts and osmoregulatory canals and extending laterally beyond longitudinal osmoregulatory canals. With proglottis development, uterus forms numerous diverticula on ventral side and becomes labyrinthine in terminal postmature proglottides. Testes persist in postmature proglottides; cirrus-sac and vagina persist in gravid proglottides (
<figureCitation id="30922A40F561FFAFFE22920FCD1F696B" box="[476,573,612,639]" captionStart="Fig" captionStartId="10.[189,232,1821,1847]" captionTargetBox="[189,1398,383,1772]" captionTargetId="figure-12@10.[189,1398,383,1781]" captionTargetPageId="10" captionText="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="12" pageNumber="280">Fig. 4B</figureCitation>
). Gravid proglottides transversely elongate, 260–390 × 1690–2040 (325 × 1856; n = 20). Fully developed uterus labyrinthine, occupying entire median field, extending bilaterally, dorsally, beyond longitudinal osmoregulatory canals (
<figureCitation id="30922A40F561FFAFFC3792C1CB0A69D0" box="[969,1064,682,709]" captionStart="Fig" captionStartId="10.[189,232,1821,1847]" captionTargetBox="[189,1398,383,1772]" captionTargetId="figure-12@10.[189,1398,383,1781]" captionTargetPageId="10" captionText="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="12" pageNumber="280">Fig. 4C</figureCitation>
). Uterus contains numerous (up to 1500–1600) small eggs. Eggs 52–57 × 60–66, subspherical, with very thin outer coat (up to 0.7–1 thick); oncospheres 19–23 × 26–31 (
<figureCitation id="30922A40F561FFAFFD42929ACC36681E" box="[700,788,753,779]" captionStart="Fig" captionStartId="10.[189,232,1821,1847]" captionTargetBox="[189,1398,383,1772]" captionTargetId="figure-12@10.[189,1398,383,1781]" captionTargetPageId="10" captionText="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="12" pageNumber="280">Fig 4D</figureCitation>
). Embryophores very thin, 23–28 × 33–38, without polar filaments. Embryonic hooks small, antero-lateral hooks (9.8–10.7), much more robust than slender postero-lateral (9.8–10.5) and median (10.3–11) hooks (
<figureCitation id="30922A40F560FFAEFCB19147CC886A53" box="[847,938,300,327]" captionStart="Fig" captionStartId="10.[189,232,1821,1847]" captionTargetBox="[189,1398,383,1772]" captionTargetId="figure-12@10.[189,1398,383,1781]" captionTargetPageId="10" captionText="Fig. 4. Kontrimavichusia hobergi sp. nov. A. Paratype (ISEA AM17-236#1), cirrus and vagina, ventral view. B. Holotype (ISEAAM17-236#3), pregravid proglottis, showing appearance of uterine diverticula, dorsal view. C. Holotype, gravid proglottis, showing labyrinthine uterus, dorsal view. D. Holotype, egg. E. Holotype, embryonic hooks. Abbreviations: al = anterolateral; m = median; pl = postero-lateral. Scale bars: A = 50 µm; B–C = 300 µm; D = 20 µm; E = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13643356" httpUri="https://zenodo.org/record/13643356/files/figure.png" pageId="13" pageNumber="281">Fig. 4E</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="FCD6664DF561FFAFFF4097E3CA4B6CFD" ID-DOI="http://doi.org/10.5281/zenodo.13643360" ID-Zenodo-Dep="13643360" httpUri="https://zenodo.org/record/13643360/files/figure.png" pageId="12" pageNumber="280" startId="12.[190,233,1928,1954]" targetBox="[190,1389,860,1876]" targetPageId="12">
<paragraph id="A81636C5F561FFAFFF4097E3CA4B6CFD" blockId="12.[189,1398,1928,2025]" pageId="12" pageNumber="280">
<emphasis id="9ADDEAD7F561FFAFFF4097E3CE2F6CB7" bold="true" box="[190,269,1928,1954]" pageId="12" pageNumber="280">Fig. 6.</emphasis>
Maximum likelihood phylogenetic tree of species of 
<taxonomicName id="6FA94D46F561FFAFFC7B97E3CFDA6CD0" authority="Makarikov &amp; Binkiene, 2022" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="280" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F561FFAFFC7B97E3CB756CB7" box="[901,1111,1928,1954]" italics="true" pageId="12" pageNumber="280">Kontrimavichusia</emphasis>
<bibRefCitation id="CC384B34F561FFAFFBA097E3CFDA6CD0" author="Makarikov A. A. &amp; Binkiene R." pageId="12" pageNumber="280" refId="ref11151" refString="Makarikov A. A. &amp; Binkiene R. 2022. Redescription and taxonomic position of Rodentolepis (sensu lato) asymmetrica (Janicki, 1904), with the erection of Kontrimavichusia n. g. (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae). Journal of Helminthology 96: E 63. https: // doi. org / 10.1017 / S 0022149 X 22000505" type="journal volume" year="2022">Makarikov &amp; Binkienė, 2022</bibRefCitation>
</taxonomicName>
based on analysis of partial sequences of the nad1 gen. Bootstrap support given for maximum likelihood analysis based on 1000 replicates. Bootstrap support values lower than 70% are not shown.
</paragraph>
</caption>
<subSubSection id="E0B3654EF560FFAEFF439106CD9F6AA3" box="[189,701,365,438]" pageId="13" pageNumber="281" type="distribution">
<paragraph id="A81636C5F560FFAEFF439106CE746A92" blockId="13.[189,342,365,391]" box="[189,342,365,391]" pageId="13" pageNumber="281">
<heading id="F35E81A9F560FFAEFF439106CE746A92" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="13" pageNumber="281" reason="3">
<emphasis id="9ADDEAD7F560FFAEFF439106CE746A92" bold="true" box="[189,342,365,391]" pageId="13" pageNumber="281">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A81636C5F560FFAEFF4391F7CD9F6AA3" blockId="13.[189,701,412,438]" box="[189,701,412,438]" pageId="13" pageNumber="281">
<collectingCountry id="D0BE7655F560FFAEFF4391F7CE296AA3" box="[189,267,412,438]" name="Russia" pageId="13" pageNumber="281">Russia</collectingCountry>
(Republic of 
<collectingRegion id="6A6DF827F560FFAEFE5391F7CD6D6AA3" box="[429,591,412,438]" country="Russia" name="North Ossetia" pageId="13" pageNumber="281">North Ossetia</collectingRegion>
– 
<collectingRegion id="6A6DF827F560FFAEFDA091F7CD8E6AA3" box="[606,684,412,438]" country="Russia" name="North Ossetia" pageId="13" pageNumber="281">Alania</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="E0B3654EF560FFAEFF4391B5CD936893" pageId="13" pageNumber="281" type="discussion">
<paragraph id="A81636C5F560FFAEFF4391B5CE0D6AED" blockId="13.[189,303,478,504]" box="[189,303,478,504]" pageId="13" pageNumber="281">
<heading id="F35E81A9F560FFAEFF4391B5CE0D6AED" bold="true" box="[189,303,478,504]" fontSize="11" level="3" pageId="13" pageNumber="281" reason="3">
<emphasis id="9ADDEAD7F560FFAEFF4391B5CE0D6AED" bold="true" box="[189,303,478,504]" pageId="13" pageNumber="281">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="A81636C5F560FFAEFF439267CD936893" blockId="13.[188,1399,524,903]" pageId="13" pageNumber="281">
<taxonomicName id="6FA94D46F560FFAEFF439267CED66933" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[189,500,524,550]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F560FFAEFF439267CED66933" box="[189,500,524,550]" italics="true" pageId="13" pageNumber="281">Kontrimavichusia hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF560FFAEFDFE9266CD7C6932" box="[512,606,525,551]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
is readily distinguishable from 
<taxonomicName id="6FA94D46F560FFAEFC189266CB806933" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[998,1186,525,550]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFC189266CB806933" box="[998,1186,525,550]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
by the triangular arrangement of the testes; those in the 
<typeStatus id="77128867F560FFAEFD74925BCD9F695F" box="[650,701,560,586]" pageId="13" pageNumber="281">type</typeStatus>
species most often are situated in one row. The number of testes distinguishes the new species from 
<taxonomicName id="6FA94D46F560FFAEFD639238CC1B6979" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[669,825,594,620]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata" status="sp. nov.">
<emphasis id="9ADDEAD7F560FFAEFD639238CC1B6979" box="[669,825,594,620]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF560FFAEFCBE9238CCBF6978" box="[832,925,595,621]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
: the former has three 3 per proglottis, the latter has 4–6 per proglottis. The cirrus-sac in 
<taxonomicName id="6FA94D46F560FFAEFD28921DCC716985" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[726,851,630,656]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi">
<emphasis id="9ADDEAD7F560FFAEFD28921DCC716985" box="[726,851,630,656]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
substantially crosses the poral osmoregulatory canals; in contrast, the cirrus-sac of 
<taxonomicName id="6FA94D46F560FFAEFD9592F1CC0669A6" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[619,804,666,691]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFD9592F1CC0669A6" box="[619,804,666,691]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
overlaps or rarely crosses the ventral longitudinal canal. The distal end of the rostellar pouch does not attain the level of the posterior margins of the suckers in 
<taxonomicName id="6FA94D46F560FFAEFEC4928BCE9469EC" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[314,438,735,761]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi">
<emphasis id="9ADDEAD7F560FFAEFEC4928BCE9469EC" box="[314,438,735,761]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
, directly contrasting with the condition in 
<taxonomicName id="6FA94D46F560FFAEFC56928BCB4269EC" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[936,1120,736,761]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFC56928BCB4269EC" box="[936,1120,736,761]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
. In addition, specimens of 
<taxonomicName id="6FA94D46F560FFAEFF1E9368CE426809" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[224,352,770,796]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi">
<emphasis id="9ADDEAD7F560FFAEFF1E9368CE426809" box="[224,352,770,796]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="6FA94D46F560FFAEFE079368CD976809" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[505,693,771,796]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFE079368CD976809" box="[505,693,771,796]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
and 
<taxonomicName id="6FA94D46F560FFAEFD0A9368CCB66809" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[756,916,770,796]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata">
<emphasis id="9ADDEAD7F560FFAEFD0A9368CCB66809" box="[756,916,770,796]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
by distinctly smaller dimensions of the fully developed strobila. Further, the cirrus-sac is larger than in 
<taxonomicName id="6FA94D46F560FFAEFC35934DCBA4682A" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[971,1158,806,831]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFC35934DCBA4682A" box="[971,1158,806,831]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
but smaller than in 
<taxonomicName id="6FA94D46F560FFAEFF439321CE786876" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[189,346,841,867]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata">
<emphasis id="9ADDEAD7F560FFAEFF439321CE786876" box="[189,346,841,867]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
; the cirrus of the new species is smaller than in 
<taxonomicName id="6FA94D46F560FFAEFC6D9321CB0D6876" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[915,1071,841,867]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata">
<emphasis id="9ADDEAD7F560FFAEFC6D9321CB0D6876" box="[915,1071,841,867]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
. The ovary of 
<taxonomicName id="6FA94D46F560FFAEFB219321CA796876" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1247,1371,841,867]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi">
<emphasis id="9ADDEAD7F560FFAEFB219321CA796876" box="[1247,1371,841,867]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
is narrower than in both congeners (
<tableCitation id="E52B037EF560FFAEFDB69307CD826893" box="[584,672,876,902]" captionStart="Table 1" captionStartId="14.[189,259,265,291]" captionTargetPageId="14" captionText="Table 1. Comparative morphometric data of species of Kontrimavichusia Makarikov &amp; Binkienė, 2022 (measurements in micrometres except where otherwise stated)." httpUri="http://table.plazi.org/id/FCD6664DF563FFADFF439162CC8B6A53" pageId="13" pageNumber="281" tableUuid="FCD6664DF563FFADFF439162CC8B6A53">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E0B3654EF560FFADFF4393C5CA076D6D" lastPageId="14" lastPageNumber="282" pageId="13" pageNumber="281" type="description">
<paragraph id="A81636C5F560FFAEFF4393C5CD4A68DE" blockId="13.[189,616,942,971]" box="[189,616,942,971]" pageId="13" pageNumber="281">
<emphasis id="9ADDEAD7F560FFAEFF4393C5CD4A68DE" bold="true" box="[189,616,942,971]" italics="true" pageId="13" pageNumber="281">Molecular phylogenetic analysis</emphasis>
</paragraph>
<paragraph id="A81636C5F560FFAEFF439388CD4C6D25" blockId="13.[188,1400,995,1585]" pageId="13" pageNumber="281">
Phylogeny was based on the nuclear ribosomal 28S rRNA gene. The length of the alignment after trimming was 1042 nucleotides. Consistent with a previous study, the monophyly of each of the two clades corresponding to 
<taxonomicName id="6FA94D46F560FFAEFE1C9442CD966F56" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" box="[482,692,1065,1091]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F560FFAEFE1C9442CD966F56" box="[482,692,1065,1091]" italics="true" pageId="13" pageNumber="281">Kontrimavichusia</emphasis>
</taxonomicName>
and 
<taxonomicName id="6FA94D46F560FFAEFD0F9442CCA86F56" authorityName="Weinland" authorityYear="1858" box="[753,906,1065,1091]" class="Cestoda" family="Hymenolepididae" genus="Hymenolepis" kingdom="Animalia" order="Cyclophyllidea" pageId="13" pageNumber="281" phylum="Platyhelminthes" rank="genus">
<emphasis id="9ADDEAD7F560FFAEFD0F9442CCA86F56" box="[753,906,1065,1091]" italics="true" pageId="13" pageNumber="281">Hymenolepis</emphasis>
</taxonomicName>
is well supported (0.97 and 1.0 posterior probability, respectively) (
<bibRefCitation id="CC384B34F560FFAEFDFD9426CC566F72" author="Makarikov A. A. &amp; Binkiene R." box="[515,884,1101,1127]" pageId="13" pageNumber="281" refId="ref11151" refString="Makarikov A. A. &amp; Binkiene R. 2022. Redescription and taxonomic position of Rodentolepis (sensu lato) asymmetrica (Janicki, 1904), with the erection of Kontrimavichusia n. g. (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae). Journal of Helminthology 96: E 63. https: // doi. org / 10.1017 / S 0022149 X 22000505" type="journal volume" year="2022">Makarikov &amp; Binkienė 2022</bibRefCitation>
). Further, we demonstrated at least four independent phylogenetic lineages within the 
<taxonomicName id="6FA94D46F560FFAEFD1A941BCC946F9F" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" box="[740,950,1136,1162]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F560FFAEFD1A941BCC946F9F" box="[740,950,1136,1162]" italics="true" pageId="13" pageNumber="281">Kontrimavichusia</emphasis>
</taxonomicName>
clade including the two species from the North Caucasus described herein, with differences between all available sequences up to 1–11 bp (
<figureCitation id="30922A40F560FFAEFF3994DDCE2C6FC4" box="[199,270,1206,1233]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[219,1365,545,1848]" captionTargetId="figure-88@11.[216,1370,539,1855]" captionTargetPageId="11" captionText="Fig. 5. Maximum likelihood phylogenetic tree of Kontrimavichusia Makarikov &amp; Binkienė, 2022 and Hymenolepis Weinland, 1858 based on analysis of partial sequences of the 28S rRNA gene. Bootstrap support given for maximum likelihood analysis based on 1000 replicates. Bootstrap support values lower than 70% are not shown." figureDoi="http://doi.org/10.5281/zenodo.13643358" httpUri="https://zenodo.org/record/13643358/files/figure.png" pageId="13" pageNumber="281">Fig. 5</figureCitation>
). Current analyses strongly support the generic allocation of those species to the genus. One of those lineages is represented by specimens of 
<taxonomicName id="6FA94D46F560FFAEFD3394B1CCA46FE6" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[717,902,1242,1267]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFD3394B1CCA46FE6" box="[717,902,1242,1267]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
(sensu stricto) from Europe from 
<taxonomicName id="6FA94D46F560FFAEFAF294B1CECB6E02" authority="(Linnaeus, 1761)" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="agrestis">
<emphasis id="9ADDEAD7F560FFAEFAF294B1CE3B6E03" italics="true" pageId="13" pageNumber="281">Microtus agrestis</emphasis>
(Linnaeus, 1761)
</taxonomicName>
(GenBank: GU166232, HM138528, ON562540 and ON562542–ON562544) and 
<taxonomicName id="6FA94D46F560FFAEFF0D954ACD016E2F" authority="(Pallas, 1778)" baseAuthorityName="Pallas" baseAuthorityYear="1778" box="[243,547,1312,1338]" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="arvalis">
<emphasis id="9ADDEAD7F560FFAEFF0D954ACE4D6E2F" box="[243,367,1312,1338]" italics="true" pageId="13" pageNumber="281">M. arvalis</emphasis>
(Pallas, 1778)
</taxonomicName>
(GenBank: ON562541). The second lineage consists of species from the North Caucasus, namely 
<taxonomicName id="6FA94D46F560FFAEFDDE952FCDE26E48" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[544,704,1347,1373]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata" status="sp. nov.">
<emphasis id="9ADDEAD7F560FFAEFDDE952FCDE26E48" box="[544,704,1347,1373]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF560FFAEFD32952FCC096E4B" box="[716,811,1348,1374]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="6FA94D46F560FFAEFC95952FCCCE6E48" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[875,1004,1347,1373]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F560FFAEFC95952FCCCE6E48" box="[875,1004,1347,1373]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF560FFAEFC09952FCB746E4B" box="[1015,1110,1348,1374]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
including the sequence from 
<taxonomicName id="6FA94D46F560FFAEFEFF950CCE586E95" baseAuthorityName="Thomas" baseAuthorityYear="1906" box="[257,378,1383,1408]" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="majori">
<emphasis id="9ADDEAD7F560FFAEFEFF950CCE586E95" box="[257,378,1383,1408]" italics="true" pageId="13" pageNumber="281">M. majori</emphasis>
</taxonomicName>
from 
<collectingCountry id="D0BE7655F560FFAEFE34950DCD3C6E95" box="[458,542,1382,1408]" name="Turkey" pageId="13" pageNumber="281">Turkey</collectingCountry>
(GU166234) attributed to 
<taxonomicName id="6FA94D46F560FFAEFC96950CCB066E95" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[872,1060,1383,1408]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFC96950CCB066E95" box="[872,1060,1383,1408]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
(sensu lato) by 
<bibRefCitation id="CC384B34F560FFAEFB11950DCE746EB1" author="Haukisalmi V. &amp; Hardman L. M. &amp; Foronda P. &amp; Feliu C. &amp; Laakkonen J. &amp; Niemimaa J. &amp; Lehtonen J. T. &amp; Henttonen H." pageId="13" pageNumber="281" pagination="631 - 641" refId="ref10690" refString="Haukisalmi V., Hardman L. M., Foronda P., Feliu C., Laakkonen J., Niemimaa J., Lehtonen J. T. &amp; Henttonen H. 2010. Systematic relationships of hymenolepidid cestodes of rodents and shrews inferred from sequences of 28 S ribosomal RNA. Zoologica Scripta 39: 631 - 641. https: // doi. org / 10.1111 / j. 1463 - 6409.2010.00444. x" type="journal article" year="2010">
Haukisalmi 
<emphasis id="9ADDEAD7F560FFAEFF4395E1CFD86EB6" box="[189,250,1417,1443]" italics="true" pageId="13" pageNumber="281">et al.</emphasis>
(2010)
</bibRefCitation>
. It distinctly differs from the 
<taxonomicName id="6FA94D46F560FFAEFD4395E1CC5A6EB6" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[701,888,1418,1443]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFD4395E1CC5A6EB6" box="[701,888,1418,1443]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
lineage by 8–9 bp. However, interspecific differences based on 28S gene in the Caucasian cluster are not pronounced. The two remaining lineages represented by specimens from 
<taxonomicName id="6FA94D46F560FFAEFDB595BBCCD56EFF" authority="(Martins, 1842)" baseAuthorityName="Martins" baseAuthorityYear="1842" box="[587,1015,1488,1514]" class="Mammalia" family="Cricetidae" genus="Chionomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="nivalis">
<emphasis id="9ADDEAD7F560FFAEFDB595BBCC096EFF" box="[587,811,1488,1514]" italics="true" pageId="13" pageNumber="281">Chionomys nivalis</emphasis>
(Martins, 1842)
</taxonomicName>
from 
<collectingCountry id="D0BE7655F560FFAEFBB595BBCBB96EFF" box="[1099,1179,1488,1514]" name="France" pageId="13" pageNumber="281">France</collectingCountry>
(GU166231) and 
<taxonomicName id="6FA94D46F560FFAEFF43959FCDAB6D18" authority="(Martino, 1922)" baseAuthorityName="Martino" baseAuthorityYear="1922" box="[189,649,1523,1549]" class="Mammalia" family="Cricetidae" genus="Dinaromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="bogdanovi">
<emphasis id="9ADDEAD7F560FFAEFF43959FCEE16D18" box="[189,451,1523,1549]" italics="true" pageId="13" pageNumber="281">Dinaromys bogdanovi</emphasis>
(Martino, 1922)
</taxonomicName>
from 
<collectingCountry id="D0BE7655F560FFAEFD2A9598CC076D18" box="[724,805,1523,1549]" name="Bosnia and Herzegovina" pageId="13" pageNumber="281">Bosnia</collectingCountry>
(GU166233). Those differ from Caucasian linage by up to 11 bp and 7 bp respectively.
</paragraph>
<paragraph id="A81636C5F560FFAEFF439633CB396CBA" blockId="13.[188,1399,1624,1967]" pageId="13" pageNumber="281">
Phylogeny was based on the mitochondrial nad1 gen. The length of the alignment after trimming was 731 nucleotides. Until present, sequences of nad1 gen of 
<taxonomicName id="6FA94D46F560FFAEFCA89617CB2C6D80" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[854,1038,1660,1685]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFCA89617CB2C6D80" box="[854,1038,1660,1685]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
were missing in GenBank. We used for analysis the sequences of 
<taxonomicName id="6FA94D46F560FFAEFDA596F4CC316DAD" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[603,787,1695,1720]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFDA596F4CC316DAD" box="[603,787,1695,1720]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
(sensu stricto) from Eastern Europe (
<collectingCountry id="D0BE7655F560FFAEFB2F96F5CA616DAD" box="[1233,1347,1694,1720]" name="Lithuania" pageId="13" pageNumber="281">Lithuania</collectingCountry>
and the Republic of 
<collectingRegion id="6A6DF827F560FFAEFE7C96AACD0F6DCE" box="[386,557,1729,1755]" country="Russia" name="Bashkortostan" pageId="13" pageNumber="281">Bashkortostan</collectingRegion>
, 
<collectingCountry id="D0BE7655F560FFAEFDC596AACDA96DCE" box="[571,651,1729,1755]" name="Russia" pageId="13" pageNumber="281">Russia</collectingCountry>
) and two new species from the North Caucasus 
<taxonomicName id="6FA94D46F560FFAEFB2996A9CA546DCE" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1239,1398,1729,1755]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata" status="sp. nov.">
<emphasis id="9ADDEAD7F560FFAEFB2996A9CA546DCE" box="[1239,1398,1729,1755]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF560FFAEFF43968ECE376DEA" box="[189,277,1765,1791]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="6FA94D46F560FFAEFEB6968ECEE66DEB" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[328,452,1764,1790]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F560FFAEFEB6968ECEE66DEB" box="[328,452,1764,1790]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF560FFAEFE37968ECD006DEA" box="[457,546,1765,1791]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
that we obtained. Three well-supported lineages within 
<taxonomicName id="6FA94D46F560FFAEFB5C968FCA566DEB" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" box="[1186,1396,1764,1790]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F560FFAEFB5C968FCA566DEB" box="[1186,1396,1764,1790]" italics="true" pageId="13" pageNumber="281">Kontrimavichusia</emphasis>
</taxonomicName>
corresponded to morphologically recognized species (
<figureCitation id="30922A40F560FFAEFCBB9763CCAC6C37" box="[837,910,1800,1826]" captionStart="Fig" captionStartId="12.[190,233,1928,1954]" captionTargetBox="[190,1389,860,1876]" captionTargetId="figure-206@12.[189,1398,848,1889]" captionTargetPageId="12" captionText="Fig. 6. Maximum likelihood phylogenetic tree of species of Kontrimavichusia Makarikov &amp; Binkienė, 2022 based on analysis of partial sequences of the nad1 gen. Bootstrap support given for maximum likelihood analysis based on 1000 replicates. Bootstrap support values lower than 70% are not shown." figureDoi="http://doi.org/10.5281/zenodo.13643360" httpUri="https://zenodo.org/record/13643360/files/figure.png" pageId="13" pageNumber="281">Fig. 6</figureCitation>
). Unlike the 28S rRNA gene sequences, mitochondrial gene nad1 provided strong evidence for the description of 
<taxonomicName id="6FA94D46F560FFAEFBE29747CB9A6C50" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1052,1208,1835,1861]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata">
<emphasis id="9ADDEAD7F560FFAEFBE29747CB9A6C50" box="[1052,1208,1835,1861]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
and 
<taxonomicName id="6FA94D46F560FFAEFB0C9747CA4D6C50" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1266,1391,1835,1861]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi">
<emphasis id="9ADDEAD7F560FFAEFB0C9747CA4D6C50" box="[1266,1391,1835,1861]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
. The interspecific pairwise distances between lineages of 
<taxonomicName id="6FA94D46F560FFAEFCAC9724CCCC6C7D" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[850,1006,1870,1896]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="testiculata">
<emphasis id="9ADDEAD7F560FFAEFCAC9724CCCC6C7D" box="[850,1006,1870,1896]" italics="true" pageId="13" pageNumber="281">K. testiculata</emphasis>
</taxonomicName>
and 
<taxonomicName id="6FA94D46F560FFAEFBDB9724CB836C7D" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1061,1185,1870,1896]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="hobergi">
<emphasis id="9ADDEAD7F560FFAEFBDB9724CB836C7D" box="[1061,1185,1870,1896]" italics="true" pageId="13" pageNumber="281">K. hobergi</emphasis>
</taxonomicName>
vary within 4.76– 5.81% (33–40 bp). Also, both species distinctly differ from 
<taxonomicName id="6FA94D46F560FFAEFC739719CB676C9E" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[909,1093,1906,1931]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="281" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F560FFAEFC739719CB676C9E" box="[909,1093,1906,1931]" italics="true" pageId="13" pageNumber="281">K. asymmetrica</emphasis>
</taxonomicName>
(sensu stricto) by 13.36– 14.35% (84–92 bp) and 14.51–15.32% (91–98 bp) respectively (
<tableCitation id="E52B037EF560FFAEFC4F97FFCB2E6CBA" box="[945,1036,1940,1967]" captionStart="Table 2" captionStartId="15.[184,254,547,575]" captionTargetPageId="15" captionText="Table 2. Pairwise uncorrected genetic distances (below diagonal) and total nucleotide differences (above diagonal) in sequences of nad1 gen between specimens of Kontrimavichusia Makarikov &amp; Binkienė, 2022." httpUri="http://table.plazi.org/id/FCD6664DF562FFACFF469248CCBE6977" pageId="13" pageNumber="281" tableUuid="FCD6664DF562FFACFF469248CCBE6977">Table 2</tableCitation>
).
</paragraph>
<caption id="FCD6664DF563FFADFF439162CC8B6A53" ID-Table-UUID="FCD6664DF563FFADFF439162CC8B6A53" httpUri="http://table.plazi.org/id/FCD6664DF563FFADFF439162CC8B6A53" pageId="14" pageNumber="282" startId="14.[189,259,265,291]" targetBox="[195,1385,366,1257]" targetIsTable="true" targetPageId="14">
<paragraph id="A81636C5F563FFADFF439162CC8B6A53" blockId="14.[189,1398,265,327]" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFF439162CE3D6A36" bold="true" box="[189,287,265,291]" pageId="14" pageNumber="282">Table 1.</emphasis>
Comparative morphometric data of species of 
<taxonomicName id="6FA94D46F563FFADFCBC9162CA546A36" authority="Makarikov &amp; Binkiene, 2022" authorityName="Makarikov &amp; Binkiene" authorityYear="2022" box="[834,1398,265,291]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="genus">
<emphasis id="9ADDEAD7F563FFADFCBC9162CB346A36" box="[834,1046,265,291]" italics="true" pageId="14" pageNumber="282">Kontrimavichusia</emphasis>
<bibRefCitation id="CC384B34F563FFADFBE09162CA546A36" author="Makarikov A. A. &amp; Binkiene R." box="[1054,1398,265,291]" pageId="14" pageNumber="282" refId="ref11151" refString="Makarikov A. A. &amp; Binkiene R. 2022. Redescription and taxonomic position of Rodentolepis (sensu lato) asymmetrica (Janicki, 1904), with the erection of Kontrimavichusia n. g. (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae). Journal of Helminthology 96: E 63. https: // doi. org / 10.1017 / S 0022149 X 22000505" type="journal volume" year="2022">Makarikov &amp; Binkienė, 2022</bibRefCitation>
</taxonomicName>
(measurements in micrometres except where otherwise stated).
</paragraph>
</caption>
<paragraph id="A81636C5F563FFADFF3D9117CA2A6FDE" pageId="14" pageNumber="282">
<table id="DAA9C465F563005CFF3D9105CA4B6FFC" box="[195,1385,366,1257]" gridcols="4" gridrows="22" pageId="14" pageNumber="282">
<tr id="16993487F563005CFF3D9105CA4B6AB5" box="[195,1385,366,416]" gridrow="0" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D9105CEA86AB5" box="[195,394,366,416]" gridcol="0" gridrow="0" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFF3D9117CE286A87" bold="true" box="[195,266,380,402]" pageId="14" pageNumber="282">species</emphasis>
</th>
<th id="55485DFBF563005CFE599105CDDF6AB5" box="[423,765,366,416]" gridcol="1" gridrow="0" pageId="14" pageNumber="282">
<taxonomicName id="6FA94D46F563FFADFE469117CDCE6A84" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[440,748,380,402]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F563FFADFE469117CDCE6A84" bold="true" box="[440,748,380,402]" italics="true" pageId="14" pageNumber="282">Kontrimavichusia asymmetrica</emphasis>
</taxonomicName>
</th>
<th id="55485DFBF563005CFCEA9105CB6D6AB5" box="[788,1103,366,416]" gridcol="2" gridrow="0" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFCDD9105CCFA6AB5" bold="true" pageId="14" pageNumber="282">
<taxonomicName id="6FA94D46F563FFADFCDD9105CB1D6A91" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[803,1087,366,388]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="testiculata" status="sp. nov.">
<emphasis id="9ADDEAD7F563FFADFCDD9105CB1D6A91" bold="true" box="[803,1087,366,388]" italics="true" pageId="14" pageNumber="282">Kontrimavichusia testiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF563FFADFC7591E0CCFA6AB5" box="[907,984,395,416]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="55485DFBF563005CFB9B9105CA4B6AB5" box="[1125,1385,366,416]" gridcol="3" gridrow="0" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFB9B9105CA2F6AB5" bold="true" pageId="14" pageNumber="282">
<taxonomicName id="6FA94D46F563FFADFB9B9105CA4B6A91" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1125,1385,366,388]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F563FFADFB9B9105CA4B6A91" bold="true" box="[1125,1385,366,388]" italics="true" pageId="14" pageNumber="282">Kontrimavichusia hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF563FFADFB3F91E0CA2F6AB5" box="[1217,1293,395,416]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="16993487F563005CFF3D91C0CA4B6AD4" box="[195,1385,427,449]" gridrow="1" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D91C0CEA86AD4" box="[195,394,427,449]" gridcol="0" gridrow="1" pageId="14" pageNumber="282">source</th>
<td id="55485DFBF563005CFE5991C0CDDF6AD4" box="[423,765,427,449]" gridcol="1" gridrow="1" pageId="14" pageNumber="282">
<bibRefCitation id="CC384B34F563FFADFE3E91C0CDC76AD5" author="Makarikov A. A. &amp; Binkiene R." box="[448,741,427,449]" pageId="14" pageNumber="282" refId="ref11151" refString="Makarikov A. A. &amp; Binkiene R. 2022. Redescription and taxonomic position of Rodentolepis (sensu lato) asymmetrica (Janicki, 1904), with the erection of Kontrimavichusia n. g. (Eucestoda: Hymenolepididae) from arvicoline rodents (Rodentia: Cricetidae). Journal of Helminthology 96: E 63. https: // doi. org / 10.1017 / S 0022149 X 22000505" type="journal volume" year="2022">Makarikov &amp; Binkienė (2022)</bibRefCitation>
</td>
<td id="55485DFBF563005CFCEA91C0CB6D6AD4" box="[788,1103,427,449]" gridcol="2" gridrow="1" pageId="14" pageNumber="282">present study</td>
<td id="55485DFBF563005CFB9B91C0CA4B6AD4" box="[1125,1385,427,449]" gridcol="3" gridrow="1" pageId="14" pageNumber="282">present study</td>
</tr>
<tr id="16993487F563005CFF3D91A1CA4B6AE8" box="[195,1385,458,509]" gridrow="2" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D91A1CEA86AE8" box="[195,394,458,509]" gridcol="0" gridrow="2" pageId="14" pageNumber="282">host species</th>
<td id="55485DFBF563005CFE5991A1CDDF6AE8" box="[423,765,458,509]" gridcol="1" gridrow="2" pageId="14" pageNumber="282">
<taxonomicName id="6FA94D46F563FFADFE5991A0CDD56AF5" authority="(Linnaeus, 1761)" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[423,759,458,480]" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="agrestis">
<emphasis id="9ADDEAD7F563FFADFE5991A0CD6F6AF5" box="[423,589,459,480]" italics="true" pageId="14" pageNumber="282">Microtus agrestis</emphasis>
(Linnaeus, 1761)
</taxonomicName>
, 
<taxonomicName id="6FA94D46F563FFADFE25918DCDEB6AEF" authority="(Pallas, 1778)" baseAuthorityName="Pallas" baseAuthorityYear="1778" box="[475,713,485,507]" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="arvalis">
<emphasis id="9ADDEAD7F563FFADFE25918DCD1C6AEF" box="[475,574,485,507]" italics="true" pageId="14" pageNumber="282">M. arvalis</emphasis>
(Pallas, 1778)
</taxonomicName>
</td>
<td id="55485DFBF563005CFCEA91A1CB6D6AE8" box="[788,1103,458,509]" gridcol="2" gridrow="2" pageId="14" pageNumber="282">
<taxonomicName id="6FA94D46F563FFADFCEA91A0CB6D6AF5" authority="(Thomas, 1906)" baseAuthorityName="Thomas" baseAuthorityYear="1906" box="[788,1103,458,480]" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="majori">
<emphasis id="9ADDEAD7F563FFADFCEA91A0CC926AF5" box="[788,944,459,480]" italics="true" pageId="14" pageNumber="282">Microtus majori</emphasis>
(Thomas, 1906)
</taxonomicName>
</td>
<td id="55485DFBF563005CFB9B91A1CA4B6AE8" box="[1125,1385,458,509]" gridcol="3" gridrow="2" pageId="14" pageNumber="282">
<taxonomicName id="6FA94D46F563FFADFB8B91A0CA606AE9" authority="(Shidlovsky, 1919)" baseAuthorityName="Shidlovsky" baseAuthorityYear="1919" class="Mammalia" family="Cricetidae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="daghestanicus">
<emphasis id="9ADDEAD7F563FFADFB8B91A0CA7B6ACA" box="[1141,1369,458,480]" italics="true" pageId="14" pageNumber="282">Microtus daghestanicus</emphasis>
(Shidlovsky, 1919)
</taxonomicName>
</td>
</tr>
<tr id="16993487F563005CFF3D926DCA4B6908" box="[195,1385,518,541]" gridrow="3" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D926DCEA86908" box="[195,394,518,541]" gridcol="0" gridrow="3" pageId="14" pageNumber="282">strobila length (mm)</th>
<td id="55485DFBF563005CFE59926DCDDF6908" box="[423,765,518,541]" gridcol="1" gridrow="3" pageId="14" pageNumber="282">98–160 mm</td>
<td id="55485DFBF563005CFCEA926DCB6D6908" box="[788,1103,518,541]" gridcol="2" gridrow="3" pageId="14" pageNumber="282">72–128 mm</td>
<td id="55485DFBF563005CFB9B926DCA4B6908" box="[1125,1385,518,541]" gridcol="3" gridrow="3" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFB4C926DCA3E6909" bold="true" box="[1202,1308,518,540]" pageId="14" pageNumber="282">65–78 mm</emphasis>
</td>
</tr>
<tr id="16993487F563005CFF3D924DCA4B6928" box="[195,1385,550,573]" gridrow="4" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D924DCEA86928" box="[195,394,550,573]" gridcol="0" gridrow="4" pageId="14" pageNumber="282">strobila width (mm)</th>
<td id="55485DFBF563005CFE59924DCDDF6928" box="[423,765,550,573]" gridcol="1" gridrow="4" pageId="14" pageNumber="282">2.5–4.45 mm</td>
<td id="55485DFBF563005CFCEA924DCB6D6928" box="[788,1103,550,573]" gridcol="2" gridrow="4" pageId="14" pageNumber="282">2.6–3.4 mm</td>
<td id="55485DFBF563005CFB9B924DCA4B6928" box="[1125,1385,550,573]" gridcol="3" gridrow="4" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFB52924DCA00692E" bold="true" box="[1196,1314,550,571]" pageId="14" pageNumber="282">1.7–2.1 mm</emphasis>
</td>
</tr>
<tr id="16993487F563005CFF3D922DCA4B6949" box="[195,1385,582,604]" gridrow="5" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D922DCEA86949" box="[195,394,582,604]" gridcol="0" gridrow="5" pageId="14" pageNumber="282">scolex width</th>
<td id="55485DFBF563005CFE59922DCDDF6949" box="[423,765,582,604]" gridcol="1" gridrow="5" pageId="14" pageNumber="282">245–325</td>
<td id="55485DFBF563005CFCEA922DCB6D6949" box="[788,1103,582,604]" gridcol="2" gridrow="5" pageId="14" pageNumber="282">324–412</td>
<td id="55485DFBF563005CFB9B922DCA4B6949" box="[1125,1385,582,604]" gridcol="3" gridrow="5" pageId="14" pageNumber="282">245–360</td>
</tr>
<tr id="16993487F563005CFF3D920DCA4B6969" box="[195,1385,614,636]" gridrow="6" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D920DCEA86969" box="[195,394,614,636]" gridcol="0" gridrow="6" pageId="14" pageNumber="282">suckers size</th>
<td id="55485DFBF563005CFE59920DCDDF6969" box="[423,765,614,636]" gridcol="1" gridrow="6" pageId="14" pageNumber="282">120–144 × 105–125</td>
<td id="55485DFBF563005CFCEA920DCB6D6969" box="[788,1103,614,636]" gridcol="2" gridrow="6" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFCAF920DCB30696E" bold="true" box="[849,1042,614,635]" pageId="14" pageNumber="282">160–181 × 120–170</emphasis>
</td>
<td id="55485DFBF563005CFB9B920DCA4B6969" box="[1125,1385,614,636]" gridcol="3" gridrow="6" pageId="14" pageNumber="282">120–155 × 100–120</td>
</tr>
<tr id="16993487F563005CFF3D92EDCA4B6989" box="[195,1385,646,668]" gridrow="7" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D92EDCEA86989" box="[195,394,646,668]" gridcol="0" gridrow="7" pageId="14" pageNumber="282">rostellar pouch size</th>
<td id="55485DFBF563005CFE5992EDCDDF6989" box="[423,765,646,668]" gridcol="1" gridrow="7" pageId="14" pageNumber="282">168–185 × 83–115</td>
<td id="55485DFBF563005CFCEA92EDCB6D6989" box="[788,1103,646,668]" gridcol="2" gridrow="7" pageId="14" pageNumber="282">160–195 × 115–142</td>
<td id="55485DFBF563005CFB9B92EDCA4B6989" box="[1125,1385,646,668]" gridcol="3" gridrow="7" pageId="14" pageNumber="282">130–164 × 95–125</td>
</tr>
<tr id="16993487F563005CFF3D92CDCA4B69A9" box="[195,1385,678,700]" gridrow="8" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D92CDCEA869A9" box="[195,394,678,700]" gridcol="0" gridrow="8" pageId="14" pageNumber="282">rostellum size</th>
<td id="55485DFBF563005CFE5992CDCDDF69A9" box="[423,765,678,700]" gridcol="1" gridrow="8" pageId="14" pageNumber="282">113–135 × 42–81</td>
<td id="55485DFBF563005CFCEA92CDCB6D69A9" box="[788,1103,678,700]" gridcol="2" gridrow="8" pageId="14" pageNumber="282">115–153 × 50–88</td>
<td id="55485DFBF563005CFB9B92CDCA4B69A9" box="[1125,1385,678,700]" gridcol="3" gridrow="8" pageId="14" pageNumber="282">105–161 × 50–70</td>
</tr>
<tr id="16993487F563005CFF3D92AECA4B69ED" box="[195,1385,709,760]" gridrow="9" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D92AECEA869ED" box="[195,394,709,760]" gridcol="0" gridrow="9" pageId="14" pageNumber="282">rostellar hooks number</th>
<td id="55485DFBF563005CFE5992AECDDF69ED" box="[423,765,709,760]" gridcol="1" gridrow="9" pageId="14" pageNumber="282">18–23</td>
<td id="55485DFBF563005CFCEA92AECB6D69ED" box="[788,1103,709,760]" gridcol="2" gridrow="9" pageId="14" pageNumber="282">20–28</td>
<td id="55485DFBF563005CFB9B92AECA4B69ED" box="[1125,1385,709,760]" gridcol="3" gridrow="9" pageId="14" pageNumber="282">18–22</td>
</tr>
<tr id="16993487F563005CFF3D9369CA4B680D" box="[195,1385,770,792]" gridrow="10" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D9369CEA8680D" box="[195,394,770,792]" gridcol="0" gridrow="10" pageId="14" pageNumber="282">rostellar hooks size</th>
<td id="55485DFBF563005CFE599369CDDF680D" box="[423,765,770,792]" gridcol="1" gridrow="10" pageId="14" pageNumber="282">20–22.5</td>
<td id="55485DFBF563005CFCEA9369CB6D680D" box="[788,1103,770,792]" gridcol="2" gridrow="10" pageId="14" pageNumber="282">17.5–22</td>
<td id="55485DFBF563005CFB9B9369CA4B680D" box="[1125,1385,770,792]" gridcol="3" gridrow="10" pageId="14" pageNumber="282">20–24</td>
</tr>
<tr id="16993487F563005CFF3D9349CA4B682D" box="[195,1385,802,824]" gridrow="11" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D9349CEA8682D" box="[195,394,802,824]" gridcol="0" gridrow="11" pageId="14" pageNumber="282">testes size</th>
<td id="55485DFBF563005CFE599349CDDF682D" box="[423,765,802,824]" gridcol="1" gridrow="11" pageId="14" pageNumber="282">155–215 × 110–168</td>
<td id="55485DFBF563005CFCEA9349CB6D682D" box="[788,1103,802,824]" gridcol="2" gridrow="11" pageId="14" pageNumber="282">145–210 × 125–175</td>
<td id="55485DFBF563005CFB9B9349CA4B682D" box="[1125,1385,802,824]" gridcol="3" gridrow="11" pageId="14" pageNumber="282">135–210 × 95–150</td>
</tr>
<tr id="16993487F563005CFF3D932ACA4B684D" box="[195,1385,833,856]" gridrow="12" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D932ACEA8684D" box="[195,394,833,856]" gridcol="0" gridrow="12" pageId="14" pageNumber="282">cirrus-sac size</th>
<td id="55485DFBF563005CFE59932ACDDF684D" box="[423,765,833,856]" gridcol="1" gridrow="12" pageId="14" pageNumber="282">225–270 × 48–66</td>
<td id="55485DFBF563005CFCEA932ACB6D684D" box="[788,1103,833,856]" gridcol="2" gridrow="12" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFCA3932ACB246843" bold="true" box="[861,1030,833,854]" pageId="14" pageNumber="282">370–445 × 55–77</emphasis>
</td>
<td id="55485DFBF563005CFB9B932ACA4B684D" box="[1125,1385,833,856]" gridcol="3" gridrow="12" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFB6C932ACA1E6843" bold="true" box="[1170,1340,833,854]" pageId="14" pageNumber="282">270–320 × 56–75</emphasis>
</td>
</tr>
<tr id="16993487F563005CFF3D930ACA4B6862" box="[195,1385,865,887]" gridrow="13" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D930ACEA86862" box="[195,394,865,887]" gridcol="0" gridrow="13" pageId="14" pageNumber="282">cirrus size</th>
<td id="55485DFBF563005CFE59930ACDDF6862" box="[423,765,865,887]" gridcol="1" gridrow="13" pageId="14" pageNumber="282">110–166 × 25–36</td>
<td id="55485DFBF563005CFCEA930ACB6D6862" box="[788,1103,865,887]" gridcol="2" gridrow="13" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFCA3930ACB256863" bold="true" box="[861,1031,865,886]" pageId="14" pageNumber="282">152–200 × 30–42</emphasis>
</td>
<td id="55485DFBF563005CFB9B930ACA4B6862" box="[1125,1385,865,887]" gridcol="3" gridrow="13" pageId="14" pageNumber="282">100–128 × 24–39</td>
</tr>
<tr id="16993487F563005CFF3D93EACA4B68A6" box="[195,1385,897,947]" gridrow="14" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D93EACEA868A6" box="[195,394,897,947]" gridcol="0" gridrow="14" pageId="14" pageNumber="282">cirrus spines, presence</th>
<td id="55485DFBF563005CFE5993EACDDF68A6" box="[423,765,897,947]" gridcol="1" gridrow="14" pageId="14" pageNumber="282">armed</td>
<td id="55485DFBF563005CFCEA93EACB6D68A6" box="[788,1103,897,947]" gridcol="2" gridrow="14" pageId="14" pageNumber="282">armed</td>
<td id="55485DFBF563005CFB9B93EACA4B68A6" box="[1125,1385,897,947]" gridcol="3" gridrow="14" pageId="14" pageNumber="282">armed</td>
</tr>
<tr id="16993487F563005CFF3D93D5CA4B68E5" box="[195,1385,958,1008]" gridrow="15" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D93D5CEA868E5" box="[195,394,958,1008]" gridcol="0" gridrow="15" pageId="14" pageNumber="282">external seminal vesicle size</th>
<td id="55485DFBF563005CFE5993D5CDDF68E5" box="[423,765,958,1008]" gridcol="1" gridrow="15" pageId="14" pageNumber="282">75–152 × 70–105</td>
<td id="55485DFBF563005CFCEA93D5CB6D68E5" box="[788,1103,958,1008]" gridcol="2" gridrow="15" pageId="14" pageNumber="282">120–280 × 70–135</td>
<td id="55485DFBF563005CFB9B93D5CA4B68E5" box="[1125,1385,958,1008]" gridcol="3" gridrow="15" pageId="14" pageNumber="282">105–190 × 82–120</td>
</tr>
<tr id="16993487F563005CFF3D9391CA4B6F05" box="[195,1385,1018,1040]" gridrow="16" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D9391CEA86F05" box="[195,394,1018,1040]" gridcol="0" gridrow="16" pageId="14" pageNumber="282">ovary width</th>
<td id="55485DFBF563005CFE599391CDDF6F05" box="[423,765,1018,1040]" gridcol="1" gridrow="16" pageId="14" pageNumber="282">410–690</td>
<td id="55485DFBF563005CFCEA9391CB6D6F05" box="[788,1103,1018,1040]" gridcol="2" gridrow="16" pageId="14" pageNumber="282">380–530</td>
<td id="55485DFBF563005CFB9B9391CA4B6F05" box="[1125,1385,1018,1040]" gridcol="3" gridrow="16" pageId="14" pageNumber="282">
<emphasis id="9ADDEAD7F563FFADFB439391CA336F1A" bold="true" box="[1213,1297,1018,1039]" pageId="14" pageNumber="282">280–396</emphasis>
</td>
</tr>
<tr id="16993487F563005CFF3D9471CA4B6F25" box="[195,1385,1050,1072]" gridrow="17" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D9471CEA86F25" box="[195,394,1050,1072]" gridcol="0" gridrow="17" pageId="14" pageNumber="282">vitellarium size</th>
<td id="55485DFBF563005CFE599471CDDF6F25" box="[423,765,1050,1072]" gridcol="1" gridrow="17" pageId="14" pageNumber="282">70–105 × 192–276</td>
<td id="55485DFBF563005CFCEA9471CB6D6F25" box="[788,1103,1050,1072]" gridcol="2" gridrow="17" pageId="14" pageNumber="282">85–115 × 180–275</td>
<td id="55485DFBF563005CFB9B9471CA4B6F25" box="[1125,1385,1050,1072]" gridcol="3" gridrow="17" pageId="14" pageNumber="282">70–116 × 120–185</td>
</tr>
<tr id="16993487F563005CFF3D9451CA4B6F78" box="[195,1385,1082,1133]" gridrow="18" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D9451CEA86F78" box="[195,394,1082,1133]" gridcol="0" gridrow="18" pageId="14" pageNumber="282">seminal receptacle size</th>
<td id="55485DFBF563005CFE599451CDDF6F78" box="[423,765,1082,1133]" gridcol="1" gridrow="18" pageId="14" pageNumber="282">240–445 × 80–110</td>
<td id="55485DFBF563005CFCEA9451CB6D6F78" box="[788,1103,1082,1133]" gridcol="2" gridrow="18" pageId="14" pageNumber="282">375–550 × 80–110</td>
<td id="55485DFBF563005CFB9B9451CA4B6F78" box="[1125,1385,1082,1133]" gridcol="3" gridrow="18" pageId="14" pageNumber="282">315–470 × 55–100</td>
</tr>
<tr id="16993487F563005CFF3D941DCA4B6F99" box="[195,1385,1142,1164]" gridrow="19" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D941DCEA86F99" box="[195,394,1142,1164]" gridcol="0" gridrow="19" pageId="14" pageNumber="282">egg size</th>
<td id="55485DFBF563005CFE59941DCDDF6F99" box="[423,765,1142,1164]" gridcol="1" gridrow="19" pageId="14" pageNumber="282">42–50 × 47–54</td>
<td id="55485DFBF563005CFCEA941DCB6D6F99" box="[788,1103,1142,1164]" gridcol="2" gridrow="19" pageId="14" pageNumber="282">55–61 × 63–70</td>
<td id="55485DFBF563005CFB9B941DCA4B6F99" box="[1125,1385,1142,1164]" gridcol="3" gridrow="19" pageId="14" pageNumber="282">52–57 × 60–66</td>
</tr>
<tr id="16993487F563005CFF3D94FDCA4B6FB9" box="[195,1385,1174,1196]" gridrow="20" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D94FDCEA86FB9" box="[195,394,1174,1196]" gridcol="0" gridrow="20" pageId="14" pageNumber="282">oncosphere size</th>
<td id="55485DFBF563005CFE5994FDCDDF6FB9" box="[423,765,1174,1196]" gridcol="1" gridrow="20" pageId="14" pageNumber="282">18–21 × 20–25</td>
<td id="55485DFBF563005CFCEA94FDCB6D6FB9" box="[788,1103,1174,1196]" gridcol="2" gridrow="20" pageId="14" pageNumber="282">22–26 × 30–33</td>
<td id="55485DFBF563005CFB9B94FDCA4B6FB9" box="[1125,1385,1174,1196]" gridcol="3" gridrow="20" pageId="14" pageNumber="282">19–23 × 26–31</td>
</tr>
<tr id="16993487F563005CFF3D94DDCA4B6FFC" box="[195,1385,1206,1257]" gridrow="21" pageId="14" pageNumber="282">
<th id="55485DFBF563005CFF3D94DDCEA86FFC" box="[195,394,1206,1257]" gridcol="0" gridrow="21" pageId="14" pageNumber="282">embryonic hooks size</th>
<td id="55485DFBF563005CFE5994DDCDDF6FFC" box="[423,765,1206,1257]" gridcol="1" gridrow="21" pageId="14" pageNumber="282">9.0–10.5</td>
<td id="55485DFBF563005CFCEA94DDCB6D6FFC" box="[788,1103,1206,1257]" gridcol="2" gridrow="21" pageId="14" pageNumber="282">10.0–11.5</td>
<td id="55485DFBF563005CFB9B94DDCA4B6FFC" box="[1125,1385,1206,1257]" gridcol="3" gridrow="21" pageId="14" pageNumber="282">9.8–11</td>
</tr>
</table>
</paragraph>
<paragraph id="A81636C5F563FFADFF43956ECC3F6E20" blockId="14.[189,1373,1285,1333]" pageId="14" pageNumber="282">
<tableNote id="554F374BF563FFADFF43956ECC3F6E20" pageId="14" pageNumber="282" targetBox="[195,1385,366,1257]" targetPageId="14">
Note: the measurements of 
<taxonomicName id="6FA94D46F563FFADFE3B956ECDF96E0F" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[453,731,1285,1306]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="testiculata" status="sp. nov.">
<emphasis id="9ADDEAD7F563FFADFE3B956ECDF96E0F" box="[453,731,1285,1306]" italics="true" pageId="14" pageNumber="282">Kontrimavichusia testiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF563FFADFD1F956ECC0E6E0F" box="[737,812,1285,1306]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="6FA94D46F563FFADFCA7956DCC9D6E0F" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[857,959,1285,1307]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F563FFADFCA7956DCC9D6E0F" box="[857,959,1285,1307]" italics="true" pageId="14" pageNumber="282">K. hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF563FFADFC3B956ECB326E0F" box="[965,1040,1285,1306]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
highlighted in bold show the most remarkable differences between the described and new species.
</tableNote>
</paragraph>
<paragraph id="A81636C5F563FFADFF43950CCA076D6D" blockId="14.[189,1399,1383,1656]" pageId="14" pageNumber="282">
Among these, the level of intraspecific differences of the lineages based on nad1 gen has the following values: in the lineage of 
<taxonomicName id="6FA94D46F563FFADFE0A95E0CD936EB1" baseAuthorityName="Janicki" baseAuthorityYear="1904" box="[500,689,1419,1444]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="asymmetrica">
<emphasis id="9ADDEAD7F563FFADFE0A95E0CD936EB1" box="[500,689,1419,1444]" italics="true" pageId="14" pageNumber="282">K. asymmetrica</emphasis>
</taxonomicName>
(sensu stricto) the 
<specimenCount id="BEAFFD4CF563FFADFC5D95E1CB776EB1" box="[931,1109,1418,1444]" count="2" pageId="14" pageNumber="282" type="generic">two specimens</specimenCount>
from the same locality (Republic of Bashkortostan, Russsia) have no intraspecific variability, while the sequence originated from the relatively remote population (more than 
<quantity id="6F519B20F563FFADFCEF95BBCC5E6EFF" box="[785,892,1488,1514]" metricMagnitude="6" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="282" unit="km" value="2000.0">2000 km</quantity>
) in 
<collectingCountry id="D0BE7655F563FFADFC5395BBCB3F6EFF" box="[941,1053,1488,1514]" name="Lithuania" pageId="14" pageNumber="282">Lithuania</collectingCountry>
differs from these by 0.69% (5 bp) (
<tableCitation id="E52B037EF563FFADFEE79598CE566D1B" box="[281,372,1523,1550]" captionStart="Table 2" captionStartId="15.[184,254,547,575]" captionTargetPageId="15" captionText="Table 2. Pairwise uncorrected genetic distances (below diagonal) and total nucleotide differences (above diagonal) in sequences of nad1 gen between specimens of Kontrimavichusia Makarikov &amp; Binkienė, 2022." httpUri="http://table.plazi.org/id/FCD6664DF562FFACFF469248CCBE6977" pageId="14" pageNumber="282" tableUuid="FCD6664DF562FFACFF469248CCBE6977">Table 2</tableCitation>
); in 
<taxonomicName id="6FA94D46F563FFADFE50959FCD696D18" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[430,587,1523,1549]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="testiculata" status="sp. nov.">
<emphasis id="9ADDEAD7F563FFADFE50959FCD696D18" box="[430,587,1523,1549]" italics="true" pageId="14" pageNumber="282">K. testiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF563FFADFDAA959FCD936D1B" box="[596,689,1524,1550]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
from the Republic of Adygeya the intraspecific differences reached up to 0.97% (7 bp), while the sequences from Karachay-Cherkess Republic showed slightly higher values of intraspecific variability 1.11%–1.81% (8–13 bp); all sequences of 
<taxonomicName id="6FA94D46F563FFADFB6B9650CA316D41" authorityName="Makarikov &amp; Stakheev" authorityYear="2024" box="[1173,1299,1594,1620]" class="Mammalia" family="Cricetidae" genus="Kontrimavichusia" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="282" phylum="Chordata" rank="species" species="hobergi" status="sp. nov.">
<emphasis id="9ADDEAD7F563FFADFB6B9650CA316D41" box="[1173,1299,1594,1620]" italics="true" pageId="14" pageNumber="282">K. hobergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81EE57ACF563FFADFAE59651CA556D41" box="[1307,1399,1594,1620]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
originating from the same locality showed up to 0.14%–0.41% (1–3 bp) intraspecific variability.
</paragraph>
</subSubSection>
</treatment>
</document>