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<document id="6704756FE288CB0C02F2F3138041E4DD" ID-CLB-Dataset="299407" ID-DOI="10.5281/zenodo.11067242" ID-GBIF-Dataset="36a87cae-beb8-49c4-a561-a9e9a73afdf1" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067242" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052119798" checkinUser="felipe" docAuthor="Maddison, David R." docDate="2023" docId="03C82573441EA56A88FE1EAE0C76FB07" docLanguage="en" docName="ProcCalifAcadSci.67.19.433-450.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (19)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Bembidion elescarabajo Maddison 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="445" masterDocId="FFF15D0B4414A566892F1D770F4AFFB4" masterDocTitle="A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae)" masterLastPageNumber="450" masterPageNumber="433" pageNumber="443" updateTime="1719951180206" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1F14D155E4A13B740351387562235DC6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4A3E89CFA239DF6E004CD2D1D330AC13">
<mods:title id="405848A913C9A6E351434F2635394CD5">A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="EBFECBF0DD46B379D9BA77D53256195F" type="personal">
<mods:role id="AD344B5BBB325B60424C27E0EF928301">
<mods:roleTerm id="9B0A2BFF1634788CF25D3E30B4FCD708">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA4BDD35C7F54BDE6EB31F74B1059F10">Maddison, David R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3D6E7AC1E46FFED26348CA4F8963E8F1">text</mods:typeOfResource>
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<mods:title id="2985B20CC3FEFAC9C3273D9CBE233B83">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="2865380B638C2F7E7D6200AC34E3CD49">
<mods:date id="75EFD725EB7DC4B590BFA16C5401002B">2023</mods:date>
<mods:detail id="AD16E9D738054CDEA492D23A180A2D14" type="pubDate">
<mods:number id="B047E8C0D1D700AC280FC428DA63B458">2023-02-15</mods:number>
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<mods:detail id="9AE4D096370FCB21F9F5599DEB578EDF" type="volume">
<mods:number id="1B9F4A93AF606E5AE22C510FCCAC6AC6">67</mods:number>
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<mods:number id="A350FA0DFCC29A04DB9D03EA0BD832AE">19</mods:number>
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<mods:classification id="C51A2CF5A362D0689E2C12B7C20A44E0">journal article</mods:classification>
<mods:identifier id="4F3EB0C73A932E7F37B6A77451EF65AC" type="CLB-Dataset">299407</mods:identifier>
<mods:identifier id="789B27771A564E5C30861A2C34916782" type="DOI">10.5281/zenodo.11067242</mods:identifier>
<mods:identifier id="77E35EF603617FB26C9EEB39E5499B4A" type="GBIF-Dataset">36a87cae-beb8-49c4-a561-a9e9a73afdf1</mods:identifier>
<mods:identifier id="EA1DA2DAFACF753928A03F21382B99D0" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="CB66324528ED0DE5E2209C1F4F38DB7D" type="Zenodo-Dep">11067242</mods:identifier>
</mods:mods>
<treatment id="03C82573441EA56A88FE1EAE0C76FB07" ID-DOI="http://doi.org/10.5281/zenodo.12627931" ID-GBIF-Taxon="234411868" ID-Zenodo-Dep="12627931" LSID="urn:lsid:plazi:treatment:03C82573441EA56A88FE1EAE0C76FB07" httpUri="http://treatment.plazi.org/id/03C82573441EA56A88FE1EAE0C76FB07" lastPageId="12" lastPageNumber="445" pageId="10" pageNumber="443">
<subSubSection id="C37BC7EE441EA56C88FE1EAE0C75FC44" box="[465,831,985,1009]" pageId="10" pageNumber="443" type="nomenclature">
<paragraph id="8BDE9465441EA56C88FE1EAE0C75FC44" blockId="10.[123,1174,985,1725]" box="[465,831,985,1009]" pageId="10" pageNumber="443">
<heading id="D0962309441EA56C88FE1EAE0C75FC44" box="[465,831,985,1009]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="443" reason="4">
<emphasis id="B9154877441EA56C88FE1EAE0C75FC44" bold="true" box="[465,831,985,1009]" pageId="10" pageNumber="443">
<taxonomicName id="4C61EFE6441EA56C88FE1EAE0D91FC45" authority="Maddison, 2023" authorityName="Maddison" authorityYear="2023" box="[465,731,985,1009]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="443" phylum="Arthropoda" rank="species" species="elescarabajo" status="sp. nov.">
<emphasis id="B9154877441EA56C88FE1EAE0D91FC45" bold="true" box="[465,731,985,1009]" italics="true" pageId="10" pageNumber="443">Bembidion elescarabajo</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A226F50C441EA56C8BC61EAE0C75FC44" box="[745,831,985,1008]" pageId="10" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE441EA56C8B221E8E0C49FBBB" box="[525,771,1017,1039]" pageId="10" pageNumber="443" type="description">
<paragraph id="8BDE9465441EA56C8B221E8E0C49FBBB" blockId="10.[123,1174,985,1725]" box="[525,771,1017,1039]" pageId="10" pageNumber="443">
(
<figureCitation id="135A88E0441EA56C8B3B1E8E0DCBFBBB" box="[532,641,1017,1039]" captionStart="FIGURE" captionStartId="1.[155,167,1563,1582]" captionTargetBox="[156,1171,245,1547]" captionTargetId="figure-14@1.[120,1176,216,1551]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Geocosmius species. (A) Bembidion “Chiapas”, brachypterous form, voucher V101274. (B) B. “Chiapas”, fully winged form, voucher V101273. (C) B. elescarabajo, paratype, voucher V101491. (D) B. franiae, specimen UASM371160. Note: B. nahuala (not shown) is extremely similar externally to B. “Chiapas”. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067244" httpUri="https://zenodo.org/record/11067244/files/figure.png" pageId="10" pageNumber="443">Figures 1C</figureCitation>
,
<figureCitation id="135A88E0441EA56C8BA31E8E0D9EFBBB" box="[652,724,1017,1039]" captionStart="FIGURE" captionStartId="11.[155,167,1212,1231]" captionTargetBox="[120,1164,236,1178]" captionTargetId="figure-170@11.[120,1176,216,1200]" captionTargetPageId="11" captionText="FIGURE 5. Aedeagi of Geocosmius. (A) Bembidion “Chiapas”, voucher DNA5317. (B) B. “Chiapas”, voucher DNA4882.Arrow indicates the prominent ridge characteristic of this species. (C) B. elescarabajo, voucher V101490.Arrow indicates the less curved sclerite margin characteristic of this species. (D) B. elescarabajo, voucher V101489. (E) B. nahuala voucher DNA5033. (F) B. nahuala voucher DNA5034. (G) B. nahuala, holotype. (H) B. franiae, voucher DNA4623. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067252" httpUri="https://zenodo.org/record/11067252/files/figure.png" pageId="10" pageNumber="443">5C, 5D</figureCitation>
,
<figureCitation id="135A88E0441EA56C8BF01E8E0DB1FBBB" box="[735,763,1017,1039]" captionStart="FIGURE" captionStartId="12.[539,551,932,951]" captionTargetBox="[504,1176,216,914]" captionTargetId="figure-386@12.[504,1176,216,914]" captionTargetPageId="12" captionText="FIGURE 6. Microsculpture of Geocosmius. (A) Bembidion franiae, vouch- er DNA4623, (B) B. elescarabajo, voucher V101491. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12627937" httpUri="https://zenodo.org/record/12627937/files/figure.png" pageId="10" pageNumber="443">6B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE441EA56C898419680CE0F9AF" pageId="10" pageNumber="443" type="materials_examined">
<paragraph id="8BDE9465441EA56C898419680CEBFB03" blockId="10.[123,1174,985,1725]" pageId="10" pageNumber="443">
<materialsCitation id="3B099E38441EA56C898419680CEBFB03" ID-GBIF-Occurrence="4902825306" collectingDate="1974-08-25" collectionCode="UASM" collectorName="H. Frania &amp; G. E. Ball" country="Mexico" elevation="2040" location="Oaxaca Rte" pageId="10" pageNumber="443" specimenCode="UASM# 410000" specimenCount="1" stateProvince="Oaxaca Rte" typeStatus="holotype">
<emphasis id="B9154877441EA56C898419680E59FB82" bold="true" box="[171,275,1055,1078]" pageId="10" pageNumber="443">
<typeStatus id="54DA2AC7441EA56C898419680E59FB82" box="[171,275,1055,1078]" pageId="10" pageNumber="443" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D675FEC441EA56C883619690E05FB82" box="[281,335,1054,1078]" pageId="10" pageNumber="443" type="male">male</specimenCount>
, in
<collectionCode id="ED700CA0441EA56C885A19690E8FFB82" box="[373,453,1054,1078]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM</collectionCode>
, labeled “
<collectingCountry id="F376D4F5441EA56C8B0319690D26FB82" box="[556,620,1054,1078]" name="Mexico" pageId="10" pageNumber="443">MEX</collectingCountry>
.
<collectingRegion id="49A55A87441EA56C8B5919690DBEFB82" box="[630,756,1054,1078]" pageId="10" pageNumber="443">Oaxaca Rte</collectingRegion>
.
<locationDeviation id="9A6C8D47441EA56C8BD119690B7FFB82" box="[766,1077,1054,1078]" pageId="10" pageNumber="443">
<quantity id="4C993980441EA56C8BD119690C3DFB82" box="[766,887,1054,1078]" metricMagnitude="5" metricUnit="m" metricValue="2.8208581632000005" pageId="10" pageNumber="443" unit="mi" value="175.28">175. 28 mi.</quantity>
s. Valle Nacional
</locationDeviation>
.
<quantity id="4C993980441EA56C8D1119690BD8FB82" box="[1086,1170,1054,1078]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="10" pageNumber="443" unit="m" value="2040.0">
<elevation id="004C7356441EA56C8D1119690BD8FB82" box="[1086,1170,1054,1078]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="10" pageNumber="443" unit="m" value="2040.0">2040 m</elevation>
</quantity>
.
<date id="FFDFB2A5441EA56C895419490E52FBE2" box="[123,280,1086,1110]" pageId="10" pageNumber="443" value="1974-08-25">
<collectingDate id="EF9B4B4D441EA56C895419490E52FBE2" box="[123,280,1086,1110]" pageId="10" pageNumber="443" value="1974-08-25">Aug. 25, 1974</collectingDate>
</date>
” [handwritten], “MIDDLE AMER. EXP. 1974,
<collectorName id="2694F1B3441EA56C8A0D19480CC3FBE2" box="[802,905,1086,1110]" pageId="10" pageNumber="443">H. Frania</collectorName>
&amp;
<collectorName id="2694F1B3441EA56C8A8119490B59FBE2" box="[942,1043,1086,1110]" pageId="10" pageNumber="443">G.E. Ball</collectorName>
collectors”, “David R.
<taxonomicName id="4C61EFE6441EA56C89C119280E13FBC3" authority="Maddison, 2023" authorityName="Maddison" authorityYear="2023" box="[238,345,1119,1143]" pageId="10" pageNumber="441" rank="subGenus" status="subgen. nov." subGenus="Maddison">Maddison</taxonomicName>
V101490 Voucher Specimen”, “HOLOTYPE
<taxonomicName id="4C61EFE6441EA56C8A6319280FDEFB23" authority="David R." authorityName="David R." class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="443" phylum="Arthropoda" rank="species" species="elescarabajo">Bembidion elescarabajo David R.</taxonomicName>
Maddison [partly handwritten, on red paper]”, “
<specimenCode id="DBC73C1E441EA56C8B9D19080C26FB23" box="[690,876,1151,1175]" collectionCode="UASM" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM# 410000</specimenCode>
” [printed on white paper]. Genitalia mounted in Euparal in between coverslips pinned with specimen.
</materialsCitation>
</paragraph>
<paragraph id="8BDE9465441EA56C898419B70DFEFA6F" blockId="10.[123,1174,985,1725]" pageId="10" pageNumber="443">
<materialsCitation id="3B099E38441EA56C898419B70CFBFB4C" ID-GBIF-Occurrence="4902825307" collectionCode="UASM, CTVR" collectorName="George E. Ball &amp; D. R. Whitehead" country="Mexico" location="Oaxaca" pageId="10" pageNumber="443" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Oaxaca" typeStatus="paratype">
<emphasis id="B9154877441EA56C898419B70E57FB63" bold="true" box="[171,285,1216,1239]" pageId="10" pageNumber="443">
<typeStatus id="54DA2AC7441EA56C898419B70E57FB63" box="[171,285,1216,1239]" pageId="10" pageNumber="443" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D675FEC441EA56C880219B70ECEFB6C" box="[301,388,1216,1240]" count="6" pageId="10" pageNumber="443" type="male">6 males</specimenCount>
,
<specimenCount id="9D675FEC441EA56C88A019B70EB6FB6C" box="[399,508,1216,1240]" count="6" pageId="10" pageNumber="443" type="female">6 females</specimenCount>
). “
<collectingCountry id="F376D4F5441EA56C8B3619B70D3AFB6C" box="[537,624,1216,1240]" name="Mexico" pageId="10" pageNumber="443">Mexico</collectingCountry>
:
<collectingRegion id="49A55A87441EA56C8B5419B70D84FB6C" box="[635,718,1216,1240]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="443">Oaxaca</collectingRegion>
.
<locationDeviation id="9A6C8D47441EA56C8BF519B70CA5FB6C" box="[730,1007,1216,1240]" pageId="10" pageNumber="443">
<quantity id="4C993980441EA56C8BF519B70C66FB6C" box="[730,812,1216,1240]" metricMagnitude="4" metricUnit="m" metricValue="4.425696" pageId="10" pageNumber="443" unit="mi" value="27.5">27.5 mi</quantity>
s. Valle Nacional
</locationDeviation>
. 5600’
<date id="FFDFB2A5441EA56C8D1219B70BC4FB6C" box="[1085,1166,1216,1240]" pageId="10" pageNumber="443" value="2015-08">VIII.15</date>
- 16.65”, “
<collectorName id="2694F1B3441EA56C89F519970E31FB4C" box="[218,379,1248,1272]" pageId="10" pageNumber="443">George E. Ball</collectorName>
and
<collectorName id="2694F1B3441EA56C889F19960D17FB4C" box="[432,605,1248,1272]" pageId="10" pageNumber="443">D.R. Whitehead</collectorName>
collectors” (2:
<collectionCode id="ED700CA0441EA56C8A2C19970C19FB4C" box="[771,851,1248,1272]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM</collectionCode>
,
<collectionCode id="ED700CA0441EA56C8A7119970CE0FB4C" box="[862,938,1248,1272]" pageId="10" pageNumber="443">CTVR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B099E38441EA56C8A9419970DC1FA8D" ID-GBIF-Occurrence="4902825301" collectionCode="UASM, UNAM, OSAC, NHMUK, NMNH" collectorName="George E. Ball &amp; D. R. Whitehead" country="Mexico" isTailEnumeration="true" location="Oaxaca" pageId="10" pageNumber="443" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
“
<collectingCountry id="F376D4F5441EA56C8AEA19970B7EFB4C" box="[965,1076,1248,1272]" name="Mexico" pageId="10" pageNumber="443">MEXICO</collectingCountry>
:
<collectingRegion id="49A55A87441EA56C8D1019970BD8FB4C" box="[1087,1170,1248,1272]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="443">Oaxaca</collectingRegion>
.
<quantity id="4C993980441EA56C895418770F85FAAC" box="[123,207,1280,1304]" metricMagnitude="4" metricUnit="m" metricValue="4.425696" pageId="10" pageNumber="443" unit="mi" value="27.5">27.5 mi</quantity>
s. Valle Nacional. 5600’
<date id="FFDFB2A5441EA56C88C418770D45FAAC" box="[491,527,1280,1304]" pageId="10" pageNumber="443">V.4</date>
-5.1966”, “
<collectorName id="2694F1B3441EA56C8BA818770C64FAAC" box="[647,814,1280,1304]" pageId="10" pageNumber="443">George E. Ball</collectorName>
and
<collectorName id="2694F1B3441EA56C8A4618760B52FAAC" box="[873,1048,1280,1304]" pageId="10" pageNumber="443">D.R. Whitehead</collectorName>
collectors” (5:
<collectionCode id="ED700CA0441EA56C898D18560FB8FA8D" box="[162,242,1313,1337]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM</collectionCode>
,
<collectionCode id="ED700CA0441EA56C89D018560E19FA8D" box="[255,339,1313,1337]" collectionName="Mexico, Mexico D.F., Universidad Nacional Autonoma de Mexico" country="Mexico" name="Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="443">UNAM</collectionCode>
,
<collectionCode id="ED700CA0441EA56C884E18560EE1FA8D" box="[353,427,1313,1337]" collectionName="USA, Oregon, Corvallis, Oregon State University" country="USA" name="Oregon State Arthropod Collection" pageId="10" pageNumber="443" type="University or college">OSAC</collectionCode>
,
<collectionCode id="ED700CA0441EA56C889718550D6AFA8D" box="[440,544,1314,1337]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="443" type="Museum">NHMUK</collectionCode>
,
<collectionCode id="ED700CA0441EA56C8B0218560DC9FA8D" box="[557,643,1313,1337]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="443" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B099E38441EA56C8BB718560EFFFACD" ID-GBIF-Occurrence="4902825305" collectingDate="1977-04-27" collectionCode="UASM" collectorName="J. S. Ashe &amp; H. E. Frania &amp; D. Shpeley" country="Mexico" elevation="2010" location="Oaxaca" municipality="Valle Nacional" pageId="10" pageNumber="443" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
“
<collectingCountry id="F376D4F5441EA56C8B8E18560C47FA8D" box="[673,781,1313,1337]" name="Mexico" pageId="10" pageNumber="443">MEXICO</collectingCountry>
<collectingRegion id="49A55A87441EA56C8A3918560C2CFA8D" box="[790,870,1313,1337]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="443">Oaxaca</collectingRegion>
<locationDeviation id="9A6C8D47441EA56C8A5F18560BDBFA8D" box="[880,1169,1313,1337]" pageId="10" pageNumber="443">
<quantity id="4C993980441EA56C8A5F18560C80FA8D" box="[880,970,1313,1337]" metricMagnitude="4" metricUnit="m" metricValue="3.62" pageId="10" pageNumber="443" unit="km" value="36.2">36.2 km</quantity>
s. Valle Nacional
</locationDeviation>
, Rte. 175 cloud forest ca.
<quantity id="4C993980441EA56C88AA18360E91FAED" box="[389,475,1345,1369]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="10" pageNumber="443" unit="m" value="2010.0">
<elevation id="004C7356441EA56C88AA18360E91FAED" box="[389,475,1345,1369]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="10" pageNumber="443" unit="m" value="2010.0">2010 m</elevation>
</quantity>
.,
<date id="FFDFB2A5441EA56C88C718360DCDFAED" box="[488,647,1345,1369]" pageId="10" pageNumber="443" value="1977-04-27">April 27, 1977</date>
”, “MEXICAN EXP. 1977
<collectorName id="2694F1B3441EA56C8A8B18360B42FAED" box="[932,1032,1345,1369]" pageId="10" pageNumber="443">J.S. Ashe</collectorName>
,
<collectorName id="2694F1B3441EA56C8D3C18350BDBFAED" box="[1043,1169,1345,1369]" pageId="10" pageNumber="443">H.E. Frania</collectorName>
,
<collectorName id="2694F1B3441EA56C895418150FBBFACD" box="[123,241,1377,1401]" pageId="10" pageNumber="443">D. Shpeley</collectorName>
coll.” (1:
<collectionCode id="ED700CA0441EA56C887318160EE4FACD" box="[348,430,1377,1401]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B099E38441EA56C889018160B73FA2E" ID-GBIF-Occurrence="4902825303" collectingDate="1974-08-28" collectionCode="UASM" collectorName="H. Frania &amp; G. E. Ball" country="Mexico" elevation="2040" location="Rte." municipality="Rte." pageId="10" pageNumber="443" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
“
<collectingCountry id="F376D4F5441EA56C88E618160D43FACD" box="[457,521,1377,1401]" name="Mexico" pageId="10" pageNumber="443">MEX</collectingCountry>
.
<collectingRegion id="49A55A87441EA56C8B3B18160D2EFACD" box="[532,612,1377,1401]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="443">Oaxaca</collectingRegion>
<collectingMunicipality id="6BBA0E1F441EA56C8B4518150DDCFACD" box="[618,662,1378,1401]" pageId="10" pageNumber="443">Rte.</collectingMunicipality>
<locationDeviation id="9A6C8D47441EA56C8BB318160C9EFACD" box="[668,980,1377,1401]" pageId="10" pageNumber="443">
<quantity id="4C993980441EA56C8BB318160C5CFACD" box="[668,790,1377,1401]" metricMagnitude="5" metricUnit="m" metricValue="2.8208581632000005" pageId="10" pageNumber="443" unit="mi" value="175.28">175. 28 mi.</quantity>
s. Valle Nacional
</locationDeviation>
.
<quantity id="4C993980441EA56C8AF218160B7BFACD" box="[989,1073,1377,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="10" pageNumber="443" unit="m" value="2040.0">
<elevation id="004C7356441EA56C8AF218160B7BFACD" box="[989,1073,1377,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="10" pageNumber="443" unit="m" value="2040.0">2040 m</elevation>
</quantity>
.
<date id="FFDFB2A5441EA56C8D1618160FFEFA2E" pageId="10" pageNumber="443" value="1974-08-28">Aug. 28, 1974</date>
”, “MIDDLE AMER. EXP. 1974,
<collectorName id="2694F1B3441EA56C8B1D18F40DD0FA2E" box="[562,666,1410,1434]" pageId="10" pageNumber="443">H. Frania</collectorName>
&amp;
<collectorName id="2694F1B3441EA56C8BEA18F50C67FA2E" box="[709,813,1410,1434]" pageId="10" pageNumber="443">G.E. Ball</collectorName>
collectors” (1:
<collectionCode id="ED700CA0441EA56C8ACF18F50B78FA2E" box="[992,1074,1410,1434]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B099E38441EA56C8D6718F50DFBFA6F" ID-GBIF-Occurrence="4902825304" collectingDate="1975-07-22" collectionCode="UASM, UNAM, OSAC" collectorName="H. Frania &amp; G. E. Ball" country="Mexico" elevation="175" location="Oaxaca" municipality="Istlan de Juarez" pageId="10" pageNumber="443" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
“
<collectingCountry id="F376D4F5441EA56C8D7D18F50BD8FA2E" box="[1106,1170,1410,1434]" name="Mexico" pageId="10" pageNumber="443">MEX</collectingCountry>
.
<collectingRegion id="49A55A87441EA56C895418D50F81FA0E" box="[123,203,1442,1466]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="443">Oaxaca</collectingRegion>
<quantity id="4C993980441EA56C89FC18D50E64FA0E" box="[211,302,1442,1466]" metricMagnitude="4" metricUnit="m" metricValue="5.64879744" pageId="10" pageNumber="443" unit="mi" value="35.1">35.1 mi.</quantity>
n. Istlan de Juarez, Rte.
<quantity id="4C993980441EA56C8B1B18D50DF2FA0E" box="[564,696,1442,1466]" metricMagnitude="2" metricUnit="m" metricValue="1.75205" pageId="10" pageNumber="443" unit="m" value="175.205">175. 2050m</quantity>
,
<date id="FFDFB2A5441EA56C8BEB18D50C0BFA0E" box="[708,833,1442,1466]" pageId="10" pageNumber="443" value="1975-07-22">
<collectingDate id="EF9B4B4D441EA56C8BEB18D50C0BFA0E" box="[708,833,1442,1466]" pageId="10" pageNumber="443" value="1975-07-22">22.07.1975</collectingDate>
</date>
”, “MEX. EXP. 1975,
<collectorName id="2694F1B3441EA56C8D0018D40BDFFA0E" box="[1071,1173,1442,1466]" pageId="10" pageNumber="443">H. Frania</collectorName>
&amp;
<collectorName id="2694F1B3441EA56C89B818B40FB1FA6F" box="[151,251,1475,1499]" pageId="10" pageNumber="443">G.E. Ball</collectorName>
collectors” (3:
<collectionCode id="ED700CA0441EA56C888C18B40EB9FA6F" box="[419,499,1475,1499]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="10" pageNumber="443">UASM</collectionCode>
,
<collectionCode id="ED700CA0441EA56C88D118B40D18FA6F" box="[510,594,1475,1499]" collectionName="Mexico, Mexico D.F., Universidad Nacional Autonoma de Mexico" country="Mexico" name="Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="443">UNAM</collectionCode>
,
<collectionCode id="ED700CA0441EA56C8B7218B40DE3FA6F" box="[605,681,1475,1499]" collectionName="USA, Oregon, Corvallis, Oregon State University" country="USA" name="Oregon State Arthropod Collection" pageId="10" pageNumber="443" type="University or college">OSAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDE9465441EA56C898418930CE0F9AF" blockId="10.[123,1174,985,1725]" pageId="10" pageNumber="443">
<emphasis id="B9154877441EA56C898418930E08FA4F" bold="true" box="[171,322,1508,1531]" pageId="10" pageNumber="443">Type locality.</emphasis>
<materialsCitation id="3B099E38441EA56C886618940CECF9AF" ID-GBIF-Occurrence="4902825310" country="Mexico" latitude="17.594" location="Oaxaca" longLatPrecision="76" longitude="-96.427" pageId="10" pageNumber="443" specimenCount="1" stateProvince="Oaxaca">
<collectingCountry id="F376D4F5441EA56C886618940ED5FA4F" box="[329,415,1507,1531]" name="Mexico" pageId="10" pageNumber="443">México</collectingCountry>
:
<collectingRegion id="49A55A87441EA56C888518940EB4FA4F" box="[426,510,1507,1531]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="443">Oaxaca</collectingRegion>
:
<locationDeviation id="9A6C8D47441EA56C8B2618940E9FF9AF" pageId="10" pageNumber="443">
<quantity id="4C993980441EA56C8B2618940D2FFA4F" box="[521,613,1507,1531]" metricMagnitude="4" metricUnit="m" metricValue="4.5061632" pageId="10" pageNumber="443" unit="mi" value="28.0">28 miles</quantity>
(
<quantity id="4C993980441EA56C8B5B18940DF1FA4F" box="[628,699,1507,1531]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="443" unit="km" value="45.0">45 km</quantity>
) south of Valle Nacional (San Juan Bautista Valle Nacional) along route 175
</locationDeviation>
; this is approximately
<geoCoordinate id="EE55F2A2441EA56C8BE61B740C7AF9AF" box="[713,816,1539,1563]" degrees="17.594" direction="north" orientation="latitude" pageId="10" pageNumber="443" precision="55" value="17.594">17.594°N</geoCoordinate>
<geoCoordinate id="EE55F2A2441EA56C8A171B740CECF9AF" box="[824,934,1539,1563]" degrees="96.427" direction="west" orientation="longitude" pageId="10" pageNumber="443" precision="55" value="-96.427">96.427°W</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE441EA56C89841B520FF7F909" pageId="10" pageNumber="443" type="etymology">
<paragraph id="8BDE9465441EA56C89841B520FF7F909" blockId="10.[123,1174,985,1725]" pageId="10" pageNumber="443">
<emphasis id="B9154877441EA56C89841B520EB5F988" bold="true" box="[171,511,1573,1596]" pageId="10" pageNumber="443">Derivation of specific epithet</emphasis>
. This species is named in honor of George Ball’s three
<emphasis id="B9154877441EA56C8D511B530FBCF9E8" italics="true" pageId="10" pageNumber="443">El Escarabajo</emphasis>
campers. George and his colleagues used those trusty vehicles to explore the regions of
<collectingCountry id="F376D4F5441EA56C89B81B120FA2F9C9" box="[151,232,1637,1661]" name="Mexico" pageId="10" pageNumber="443">México</collectingCountry>
inhabited by
<taxonomicName id="4C61EFE6441EA56C88571B120EB0F9C8" authorityName="Maddison" authorityYear="2023" box="[376,506,1637,1660]" pageId="10" pageNumber="443" rank="subGenus" subGenus="Geocosmius">
<emphasis id="B9154877441EA56C88571B120EB0F9C8" box="[376,506,1637,1660]" italics="true" pageId="10" pageNumber="443">Geocosmius</emphasis>
</taxonomicName>
, and on these journeys most known specimens of the subgenus were collected, including the
<typeStatus id="54DA2AC7441EA56C889E1BF10E95F92A" box="[433,479,1670,1694]" pageId="10" pageNumber="443">type</typeStatus>
series of this species. The name is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF1EC4ED441FA56D89B419CB0E37FA82" ID-DOI="http://doi.org/10.5281/zenodo.11067252" ID-Zenodo-Dep="11067252" httpUri="https://zenodo.org/record/11067252/files/figure.png" pageId="11" pageNumber="444" startId="11.[155,167,1212,1231]" targetBox="[120,1164,236,1178]" targetPageId="11" targetType="figure">
<paragraph id="8BDE9465441FA56D89B419CB0E37FA82" blockId="11.[123,1175,1212,1334]" pageId="11" pageNumber="444">
<smallCapsWord id="8D3802B9441FA56D89B419CB0F92FB7A" baselines="1226,1226" box="[155,216,1212,1231]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="11" pageNumber="444">FIGURE</smallCapsWord>
5. Aedeagi of
<taxonomicName id="4C61EFE6441FA56D885F19CB0E92FB7B" authorityName="Maddison" authorityYear="2023" box="[368,472,1212,1231]" pageId="11" pageNumber="443" rank="subGenus" subGenus="Geocosmius">
<emphasis id="B9154877441FA56D885F19CB0E92FB7B" box="[368,472,1212,1231]" italics="true" pageId="11" pageNumber="444">Geocosmius</emphasis>
</taxonomicName>
. (A)
<taxonomicName id="4C61EFE6441FA56D8B3B19CB0D3AFB7B" authorityName="Latreille" authorityYear="1802" box="[532,624,1212,1231]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9154877441FA56D8B3B19CB0D3AFB7B" box="[532,624,1212,1231]" italics="true" pageId="11" pageNumber="444">Bembidion</emphasis>
</taxonomicName>
“Chiapas”, voucher DNA5317. (B)
<emphasis id="B9154877441FA56D8AE619CA0C9CFB7B" box="[969,982,1213,1231]" italics="true" pageId="11" pageNumber="444">B</emphasis>
. “Chiapas”, voucher DNA4882.Arrow indicates the prominent ridge characteristic of this species. (C)
<taxonomicName id="4C61EFE6441FA56D8A0219A10CF8FB5C" box="[813,946,1237,1256]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="species" species="elescarabajo">
<emphasis id="B9154877441FA56D8A0219A10CF8FB5C" box="[813,946,1237,1256]" italics="true" pageId="11" pageNumber="444">B. elescarabajo</emphasis>
</taxonomicName>
, voucher V101490.Arrow indicates the less curved sclerite margin characteristic of this species. (D)
<taxonomicName id="4C61EFE6441FA56D8A6219870C96FAB6" box="[845,988,1263,1282]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="species" species="elescarabajo">
<emphasis id="B9154877441FA56D8A6219870C96FAB6" box="[845,988,1263,1282]" italics="true" pageId="11" pageNumber="444">B. elescarabajo</emphasis>
</taxonomicName>
, voucher V101489. (E)
<taxonomicName id="4C61EFE6441FA56D89B3187E0FB1FAA8" authorityName="Erwin" authorityYear="1982" box="[156,251,1289,1308]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="species" species="nahuala">
<emphasis id="B9154877441FA56D89B3187E0FB1FAA8" box="[156,251,1289,1308]" italics="true" pageId="11" pageNumber="444">B. nahuala</emphasis>
</taxonomicName>
voucher DNA5033. (F)
<taxonomicName id="4C61EFE6441FA56D88FE187E0D7AFAA8" authorityName="Erwin" authorityYear="1982" box="[465,560,1289,1308]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="species" species="nahuala">
<emphasis id="B9154877441FA56D88FE187E0D7AFAA8" box="[465,560,1289,1308]" italics="true" pageId="11" pageNumber="444">B. nahuala</emphasis>
</taxonomicName>
voucher DNA5034. (G)
<taxonomicName id="4C61EFE6441FA56D8A25187E0C23FAA8" authorityName="Erwin" authorityYear="1982" box="[778,873,1289,1308]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="species" species="nahuala">
<emphasis id="B9154877441FA56D8A25187E0C23FAA8" box="[778,873,1289,1308]" italics="true" pageId="11" pageNumber="444">B. nahuala</emphasis>
</taxonomicName>
, holotype. (H)
<taxonomicName id="4C61EFE6441FA56D8AC0187E0B0FFAA8" authorityName="Erwin" authorityYear="1982" box="[1007,1093,1289,1308]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="444" phylum="Arthropoda" rank="species" species="franiae">
<emphasis id="B9154877441FA56D8AC0187E0B0FFAA8" box="[1007,1093,1289,1308]" italics="true" pageId="11" pageNumber="444">B. franiae</emphasis>
</taxonomicName>
, voucher DNA4623. Scale bar 100 µm.
</paragraph>
</caption>
<subSubSection id="C37BC7EE441FA56A898418390E66FD67" lastPageId="12" lastPageNumber="445" pageId="11" pageNumber="444" type="description">
<paragraph id="8BDE9465441FA56A898418390E94FDC7" blockId="11.[123,1174,1358,1702]" lastBlockId="12.[123,478,219,915]" lastPageId="12" lastPageNumber="445" pageId="11" pageNumber="444">
<emphasis id="B9154877441FA56D898418390E65FAD1" bold="true" box="[171,303,1358,1381]" pageId="11" pageNumber="444">Description</emphasis>
. Body length
<quantity id="4C993980441FA56D88EA18390D29FAD1" box="[453,611,1358,1382]" metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.9" metricValueMin="3.25" pageId="11" pageNumber="444" unit="mm" value="3.575" valueMax="3.9" valueMin="3.25">3.25–3.90 mm</quantity>
. Body brown, with head and pronotum slightly darker than elytra. Legs uniform in color, testaceous or slightly infuscated; antennomere 1 testaceous, antennomeres 2 and 3 with distal half slightly infuscated, remaining antennomeres slightly infuscated; penultimate maxillary palpomere infuscated, antepenultimate testaceous. Mentum with anterior lateral regions large, triangular; medial tooth triangular with rounded tip; frontal furrows weakly defined, somewhat rugose; eyes not prominent; prothorax with greatest width in front of middle; sides sinuate, with sides in front of hind angle straight; hind angle approximately right; posterolateral carina evident, moderately long, and straight; posterior region of pronotum slightly rugose with poorly defined punctures. Elytra with lateral bead not prolonged medially at shoulder; inner three striae well-defined around ed3; striae 4 and 5 faint, even anteriorly, with striae 6 and 7 extremely faint; only first stria evident to apex, with other striae more or less absent from apical fourth. Microsculpture absent from dorsal surface of head at center and from pronotal disc; evident in both sexes over entire surface of elytra, nearly isodiametric (Fig. 6B). Pronotum with two lateral setae on each side; elytron with two setae in third stria. Aedeagus (Fig. 5C, D) similar to members of the
<emphasis id="B91548774418A56A88A91C8C0E97FDA7" box="[390,477,507,531]" italics="true" pageId="12" pageNumber="445">nahuala</emphasis>
group (Fig. 5A, B, E, F, G), but with ventral margin of the large central sclerite less abruptly bent.
</paragraph>
<paragraph id="8BDE94654418A56A89841F0B0E66FD67" blockId="12.[123,478,219,915]" pageId="12" pageNumber="445">
<emphasis id="B91548774418A56A89841F0B0E9DFD27" bold="true" box="[171,471,636,659]" pageId="12" pageNumber="445">Additional characteristics</emphasis>
. All thirteen specimens examined lack hind wings.
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE4418A56A89841FAB0B6BFBC7" pageId="12" pageNumber="445" type="distribution">
<paragraph id="8BDE94654418A56A89841FAB0FBCFBA6" blockId="12.[123,478,219,915]" lastBlockId="12.[123,1174,987,1203]" pageId="12" pageNumber="445">
<emphasis id="B91548774418A56A89841FAB0E9CFD47" bold="true" box="[171,470,732,755]" pageId="12" pageNumber="445">Geographic distribution</emphasis>
. Known only from four closely proximate localities in Oaxaca along route 175 between San Juan Bautista Valle Nacional and Ixtlán de Juárez, between about 17.604°N 96.381°W and 17.594°N 96.460°W (Fig. 2, star). These localities are at most 9 km apart.
</paragraph>
<caption id="DF1EC4ED4418A56A8B341ED30B26FC64" ID-DOI="http://doi.org/10.5281/zenodo.12627937" ID-Zenodo-Dep="12627937" httpUri="https://zenodo.org/record/12627937/files/figure.png" pageId="12" pageNumber="445" startId="12.[539,551,932,951]" targetBox="[504,1176,216,914]" targetPageId="12">
<paragraph id="8BDE94654418A56A8B341ED30B26FC64" blockId="12.[507,1173,931,976]" pageId="12" pageNumber="445">
FIGURE 6. Microsculpture of
<emphasis id="B91548774418A56A8A391ED40C34FC02" box="[790,894,931,950]" italics="true" pageId="12" pageNumber="445">Geocosmius</emphasis>
. (A)
<emphasis id="B91548774418A56A8A811ED40B1FFC02" box="[942,1109,931,950]" italics="true" pageId="12" pageNumber="445">Bembidion franiae,</emphasis>
voucher DNA4623, (B)
<emphasis id="B91548774418A56A8BB81ECA0C54FC64" box="[663,798,957,976]" italics="true" pageId="12" pageNumber="445">B. elescarabajo</emphasis>
, voucher V101491. Scale bar 100 µm.
</paragraph>
</caption>
<paragraph id="8BDE94654418A56A8984196B0B6BFBC7" blockId="12.[123,1174,987,1203]" pageId="12" pageNumber="445">
<emphasis id="B91548774418A56A8984196B0DF9FB87" bold="true" box="[171,691,1052,1075]" pageId="12" pageNumber="445">Geographic relationships with other species.</emphasis>
One specimen of a
<emphasis id="B91548774418A56A8AB0196C0CBCFB87" box="[927,1014,1051,1075]" italics="true" pageId="12" pageNumber="445">nahuala</emphasis>
-group species (perhaps
<emphasis id="B91548774418A56A89F1194C0E18FBE7" box="[222,338,1083,1107]" italics="true" pageId="12" pageNumber="445">Bembidion</emphasis>
“Chiapas”) was collected at the same place and time as five of the paratypes of
<emphasis id="B91548774418A56A89B7192B0E0BFBC7" box="[152,321,1115,1139]" italics="true" pageId="12" pageNumber="445">B. elescarabajo</emphasis>
: “Mexico: Oaxaca. 27.5 mi s. Valle Nacional. 5600’ VIII.15-16.65”.
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE4418A56A8984190B0C76FB07" pageId="12" pageNumber="445" type="biology_ecology">
<paragraph id="8BDE94654418A56A8984190B0C76FB07" blockId="12.[123,1174,987,1203]" pageId="12" pageNumber="445">
<emphasis id="B91548774418A56A8984190B0E48FB27" bold="true" box="[171,258,1148,1171]" pageId="12" pageNumber="445">Habitat</emphasis>
. Based upon imagery in Google Earth, and the annotation “cloud forest” on the label of the 1977 specimen, the known localities are all in cloud forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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