210 lines
29 KiB
XML
210 lines
29 KiB
XML
<document id="A74BD94B11D9687FCE41C36F455FDED6" ID-CLB-Dataset="7175" ID-DOI="10.5252/geodiversitas2021v43a9" ID-GBIF-Dataset="04426a8e-c179-4af0-8e23-d72908e8b4cc" ID-ISSN="1638-9395" ID-Zenodo-Dep="4720776" ID-ZooBank="urn:lsid:zoobank.org:pub:11D0D852-39D7-449C-9EB3-C3D804114556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445770811" checkinUser="felipe" docAuthor="Georgalis, Georgios L., Čerňanský, Andrej & Klembara, Jozef" docDate="2021" docId="03A1633BFFBCFFE3316BFA4BF8AD3C03" docLanguage="en" docName="Geodiversitas.43.9.219-293.pdf" docOrigin="Geodiversitas 43 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Melanosaurini Sullivan 1979" docType="treatment" docVersion="6" lastPageNumber="250" masterDocId="FF981B43FFA3FFC1303EFFB8FA5E380D" masterDocTitle="Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material" masterLastPageNumber="293" masterPageNumber="219" pageNumber="248" updateTime="1698937615995" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="653760E95CAA2F03D563D6F7F47A5BDB">
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<mods:title id="5F2B15434C9BFF7F3F43D3745CAEE322">Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material</mods:title>
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<mods:name id="608A1DC0C9C08C1B0ADD632148795238" type="personal">
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<mods:namePart id="B58D76B213E21747EF6E9C82B9817C5E">Georgalis, Georgios L.</mods:namePart>
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<mods:affiliation id="D80031FBA4192C939C24059377E4AA3A">Palaeontological Institute and Museum, University of Zurich, Karl Schmid-Strasse 4, 8006 Zurich (Switzerland) and Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia) and Department of Earth Sciences, University of Torino, Via Valperga Caluso 35, 10125 Turin (Italy)</mods:affiliation>
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<mods:nameIdentifier id="36966686B12B84B0F443DD6147C9BC7F" type="email">georgios.georgalis@pim.uzh.ch</mods:nameIdentifier>
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<mods:name id="CB2CA629FF9F159B114F7D5574925AA2" type="personal">
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<mods:roleTerm id="FFAE2987195592D736D97F38C884942D">Author</mods:roleTerm>
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<mods:namePart id="68AA7382372AF90F508748EC6A40215C">Čerňanský, Andrej</mods:namePart>
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<mods:affiliation id="D5D551EF406083878B1DD236E6EFABC0">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:namePart id="E58F1D23D9E58ECA0F91FACAF8966A42">Klembara, Jozef</mods:namePart>
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<mods:affiliation id="94E24EA05639F433AB329225DD8796D3">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:title id="E495999FED81BB8C5953E67D26561E1F">Geodiversitas</mods:title>
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<mods:part id="B286471A0EEDA0761FA458129A9B3616">
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<mods:date id="1017CE5F2561411136709FC5076289C8">2021</mods:date>
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<mods:number id="F73E813A593C1012BB6DCC544A836020">2021-04-22</mods:number>
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<treatment id="03A1633BFFBCFFE3316BFA4BF8AD3C03" ID-DOI="http://doi.org/10.5281/zenodo.4721443" ID-GBIF-Taxon="181601094" ID-Zenodo-Dep="4721443" LSID="urn:lsid:plazi:treatment:03A1633BFFBCFFE3316BFA4BF8AD3C03" httpUri="http://treatment.plazi.org/id/03A1633BFFBCFFE3316BFA4BF8AD3C03" lastPageId="34" lastPageNumber="250" pageId="31" pageNumber="248">
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<subSection id="E287CBC6FFBCFFDE316BFA4BFF1D3FE6" pageId="31" pageNumber="248" type="multiple">
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<subSubSection id="C31281A6FFBCFFDE316BFA4BF8563E20" pageId="31" pageNumber="248" type="nomenclature">
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<paragraph id="8BB7D22DFFBCFFDE316BFA4BF8563E20" blockId="31.[341,567,1523,1581]" pageId="31" pageNumber="248">
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<taxonomicName id="4C08A9AEFFBCFFDE316BFA4BFBAF3E00" authorityName="Sullivan" authorityYear="1979" box="[341,497,1523,1549]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="248" phylum="Chordata" rank="tribe" tribe="Melanosaurini">Melanosaurini</taxonomicName>
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indet. (
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<figureCitation id="1333CEA8FFBCFFDE31B2F9AAFBA13E20" box="[396,511,1554,1581]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="37.[132,143,1752,1769]" captionStartId-1="38.[132,143,1613,1630]" captionStartId-2="39.[132,143,1201,1218]" captionTargetBox-0="[137,1448,216,1711]" captionTargetBox-1="[140,1451,215,1571]" captionTargetBox-2="[132,1455,211,1163]" captionTargetId-0="figure-152@37.[132,1455,213,1720]" captionTargetId-1="figure-199@38.[132,1455,203,1575]" captionTargetId-2="figure-447@39.[132,1455,211,1163]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionText-0="FIG. 30. — Melanosaurini indet. Photographs of presacral vertebra NHMW 2019/0094/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views. Scale bar: 2 mm." captionText-1="FIG. 31. — Melanosaurini indet.A-E, photographs of presacral vertebra NHMW 2019/0094/0002 in anterior (A), dorsal (B), right lateral (C), posterior (D), and ventral (E) views; F, G, photographs of presacral vertebra NHMW 2019/0094/0003 in posterior (F) and ventral (G) views. Scale bar: 2 mm." captionText-2="FIG. 32. — Melanosaurini indet. Photographs of sacral vertebra NHMW 2019/0094/0004 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720869" figureDoi-1="http://doi.org/10.5281/zenodo.4720871" figureDoi-2="http://doi.org/10.5281/zenodo.4720873" httpUri-0="https://zenodo.org/record/4720869/files/figure.png" httpUri-1="https://zenodo.org/record/4720871/files/figure.png" httpUri-2="https://zenodo.org/record/4720873/files/figure.png" pageId="31" pageNumber="248">Figs 30-32</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31281A6FFBCFFDE30BAF9F4F9563E94" pageId="31" pageNumber="248" type="materials_examined">
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<paragraph id="8BB7D22DFFBCFFDE30BAF9F4F9563E94" blockId="31.[132,776,1611,1689]" pageId="31" pageNumber="248">
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REFERRED SPECIMENS. —
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<materialsCitation id="3B60D870FFBCFFDE31A4F9F4F9563E94" ID-GBIF-Occurrence="3091185311" pageId="31" pageNumber="248">
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Four presacral vertebrae (
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<collectionCode id="ED194AE8FFBCFFDE3294F9F4F9583E69" box="[682,774,1612,1636]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="248" type="Museum">NHMW</collectionCode>
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2019/0094/0001-
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<collectionCode id="ED194AE8FFBCFFDE316EF9DEFBF23E70" box="[336,428,1638,1661]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="248" type="Museum">NHMW</collectionCode>
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2019/0094/0003 and
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<collectionCode id="ED194AE8FFBCFFDE3295F9DEF9583E70" box="[683,774,1638,1661]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="248" type="Museum">NHMW</collectionCode>
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2019/0094/0005); one sacral vertebra (
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<collectionCode id="ED194AE8FFBCFFDE31C8F939F8103E94" box="[502,590,1665,1689]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="248" type="Museum">NHMW</collectionCode>
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2019/0094/0004).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31281A6FFBCFFDE30BAF96AFF1D3FE6" pageId="31" pageNumber="248" type="description">
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<paragraph id="8BB7D22DFFBCFFDE30BAF96AFB4B3EE7" blockId="31.[130,775,1746,2028]" box="[132,277,1746,1772]" pageId="31" pageNumber="248">DESCRIPTION</paragraph>
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<paragraph id="8BB7D22DFFBCFFDE30BAF94AFB8F3F01" blockId="31.[130,775,1746,2028]" box="[132,465,1778,1804]" pageId="31" pageNumber="248">
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<emphasis id="B97C0E3FFFBCFFDE30BAF94AFB8F3F01" box="[132,465,1778,1804]" italics="true" pageId="31" pageNumber="248">
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Presacral vertebrae (
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<figureCitation id="1333CEA8FFBCFFDE316DF94BFBC03F01" box="[339,414,1778,1804]" captionStart="FIG" captionStartId="37.[132,143,1752,1769]" captionTargetBox="[137,1448,216,1711]" captionTargetId="figure-152@37.[132,1455,213,1720]" captionTargetPageId="37" captionText="FIG. 30. — Melanosaurini indet. Photographs of presacral vertebra NHMW 2019/0094/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720869" httpUri="https://zenodo.org/record/4720869/files/figure.png" pageId="31" pageNumber="248">Figs 30</figureCitation>
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;
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<figureCitation id="1333CEA8FFBCFFDE3192F94AFB963F01" box="[428,456,1778,1804]" captionStart="FIG" captionStartId="38.[132,143,1613,1630]" captionTargetBox="[140,1451,215,1571]" captionTargetId="figure-199@38.[132,1455,203,1575]" captionTargetPageId="38" captionText="FIG. 31. — Melanosaurini indet.A-E, photographs of presacral vertebra NHMW 2019/0094/0002 in anterior (A), dorsal (B), right lateral (C), posterior (D), and ventral (E) views; F, G, photographs of presacral vertebra NHMW 2019/0094/0003 in posterior (F) and ventral (G) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720871" httpUri="https://zenodo.org/record/4720871/files/figure.png" pageId="31" pageNumber="248">31</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="8BB7D22DFFBCFFDE30BAF8AAFF1D3FE6" blockId="31.[130,775,1746,2028]" lastBlockId="31.[810,1458,1523,2028]" pageId="31" pageNumber="248">
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In all specimens, the centrum is significantly anteriorly widened (
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<figureCitation id="1333CEA8FFBCFFDE30B3F889FA873F41" box="[141,217,1841,1868]" captionStart="FIG" captionStartId="37.[132,143,1752,1769]" captionTargetBox="[137,1448,216,1711]" captionTargetId="figure-152@37.[132,1455,213,1720]" captionTargetPageId="37" captionText="FIG. 30. — Melanosaurini indet. Photographs of presacral vertebra NHMW 2019/0094/0001 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720869" httpUri="https://zenodo.org/record/4720869/files/figure.png" pageId="31" pageNumber="248">Figs 30</figureCitation>
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;
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<figureCitation id="1333CEA8FFBCFFDE30D6F88AFB5D3F41" box="[232,259,1842,1868]" captionStart="FIG" captionStartId="38.[132,143,1613,1630]" captionTargetBox="[140,1451,215,1571]" captionTargetId="figure-199@38.[132,1455,203,1575]" captionTargetPageId="38" captionText="FIG. 31. — Melanosaurini indet.A-E, photographs of presacral vertebra NHMW 2019/0094/0002 in anterior (A), dorsal (B), right lateral (C), posterior (D), and ventral (E) views; F, G, photographs of presacral vertebra NHMW 2019/0094/0003 in posterior (F) and ventral (G) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720871" httpUri="https://zenodo.org/record/4720871/files/figure.png" pageId="31" pageNumber="248">31</figureCitation>
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). Their size varies, with centrum lengths ranging between 6 and
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<quantity id="4CF07FC8FFBCFFDE3119F8EAFB233F61" box="[295,381,1874,1900]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="31" pageNumber="248" unit="mm" value="9.4">9.4 mm</quantity>
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(
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<figureCitation id="1333CEA8FFBCFFDE31B5F8E9F8593F61" box="[395,519,1873,1900]" captionStart="APPENDIX" captionStartId="75.[427,439,219,236]" captionTargetPageId="75" captionText="APPENDIX 1. — Centrum lengths of all lizard vertebrae from the NHMW Quercy collection." pageId="31" pageNumber="248">Appendix 1</figureCitation>
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). There is a distinct and rather wide groove in the ventral surface of the centrum, originating anteriorly almost at the level of the cotyle and terminating posteriorly at around the level of the condyle, being almost uniform in wideness across its length. The prezygapophyses are strongly dorsally inclined in anterior view. Both cotyle and condyle are strongly dorsoventrally compressed. The neural spine, when preserved, develops mostly at the posterior half of the neural arch, however, its base extends anteriorly in the shape of a narrow longitudinal ridge until the anterior most edge of the neural arch. The height of the neural spine varies, being either high (NHMW 2019/0094/0002) or rather short (NHMW 2019/0094/0001).In
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<specimenCount id="9D0E19A4FFBCFFDE3438F96AFED53EE1" box="[1030,1163,1746,1772]" pageId="31" pageNumber="248" type="generic">one specimen</specimenCount>
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(NHMW 2019/0094/0001), the posterior edge of the neural spine (as seen in dorsal view) is bifurcated. The postzygapophyses are large and extend much laterally in dorsal view. The neural canal is relatively large and triangular in shape. The shape of the neural arch in posterior view varies, apparently depending on the exact position of the vertebra in the column; as such it can be either depressed (e.g., NHMW 2019/0094/0005 and NHMW 2019/0094/0003) or relatively vaulted (e.g., NHMW 2019/0094/0002).
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</paragraph>
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</subSubSection>
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</subSection>
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<caption id="DF7782A5FF83FFE130BAFA1DFFEF3DBB" ID-DOI="http://doi.org/10.5281/zenodo.4720851" ID-Zenodo-Dep="4720851" box="[132,1457,1445,1462]" httpUri="https://zenodo.org/record/4720851/files/figure.png" pageId="32" pageNumber="249" startId="32.[132,143,1445,1462]" targetBox="[132,1455,209,1411]" targetPageId="32">
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<paragraph id="8BB7D22DFF83FFE130BAFA1DFFEF3DBB" blockId="32.[132,1457,1445,1462]" box="[132,1457,1445,1462]" pageId="32" pageNumber="249">
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FIG. 25. —
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<taxonomicName id="4C08A9AEFF83FFE130E4FA1DF8483DBB" authority="(Filhol, 1882)" baseAuthorityName="Filhol" baseAuthorityYear="1882" box="[218,534,1445,1462]" class="Reptilia" family="Anguidae" genus="Paraplacosauriops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="249" phylum="Chordata" rank="species" species="quercyi">
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<emphasis id="B97C0E3FFF83FFE130E4FA1DFBF33DBB" box="[218,429,1445,1462]" italics="true" pageId="32" pageNumber="249">Paraplacosauriops quercyi</emphasis>
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(Filhol,1882)
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</taxonomicName>
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.Photographs of right dentary NHMW 2019/0049/0001 in lateral (
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<emphasis id="B97C0E3FFF83FFE1342DFA1DFE413DBB" bold="true" box="[1043,1055,1445,1462]" pageId="32" pageNumber="249">A</emphasis>
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),medial (
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<emphasis id="B97C0E3FFF83FFE13454FA1DFE293DBB" bold="true" box="[1130,1143,1445,1462]" pageId="32" pageNumber="249">B</emphasis>
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),and dorsal (
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<emphasis id="B97C0E3FFF83FFE134E1FA1DFEB23DBB" bold="true" box="[1247,1260,1445,1462]" pageId="32" pageNumber="249">C</emphasis>
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) views.Scale bar:5 mm.
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</paragraph>
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</caption>
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<subSection id="E287CBC6FF83FFE330BAFA4BF8AD3C03" lastPageId="34" lastPageNumber="251" pageId="32" pageNumber="249" type="multiple">
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<paragraph id="8BB7D22DFF83FFE130BAFA4BF8FE3E00" blockId="32.[132,672,1523,1549]" box="[132,672,1523,1549]" pageId="32" pageNumber="249">
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<heading id="D0FF6541FF83FFE130BAFA4BF8FE3E00" box="[132,672,1523,1549]" fontSize="11" level="2" pageId="32" pageNumber="249" reason="8">
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<emphasis id="B97C0E3FFF83FFE130BAFA4BF8FE3E00" box="[132,672,1523,1549]" italics="true" pageId="32" pageNumber="249">
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Sacral vertebra NHMW 2019/0094/0004 (
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<figureCitation id="1333CEA8FF83FFE13272FA4BF8C93E00" box="[588,663,1523,1549]" captionStart="FIG" captionStartId="39.[132,143,1201,1218]" captionTargetBox="[132,1455,211,1163]" captionTargetId="figure-447@39.[132,1455,211,1163]" captionTargetPageId="39" captionText="FIG. 32. — Melanosaurini indet. Photographs of sacral vertebra NHMW 2019/0094/0004 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720873" httpUri="https://zenodo.org/record/4720873/files/figure.png" pageId="32" pageNumber="249">Fig. 32</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB7D22DFF83FFE130BAF9AAFBC53FE6" blockId="32.[130,776,1554,2027]" pageId="32" pageNumber="249">
|
||
The sacral vertebra NHMW 2019/0094/0004 is rather similar to the above described presacral ones, especially at the degree of the anterior widening of its centrum, the dorsally inclined prezygapophyses, and the much dorsoventrally compressed cotyle and condyle (
|
||
<figureCitation id="1333CEA8FF83FFE13148F92AFB973EA1" box="[374,457,1682,1708]" captionStart="FIG" captionStartId="39.[132,143,1201,1218]" captionTargetBox="[132,1455,211,1163]" captionTargetId="figure-447@39.[132,1455,211,1163]" captionTargetPageId="39" captionText="FIG. 32. — Melanosaurini indet. Photographs of sacral vertebra NHMW 2019/0094/0004 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720873" httpUri="https://zenodo.org/record/4720873/files/figure.png" pageId="32" pageNumber="249">Fig. 32</figureCitation>
|
||
). The prezygapophyses are robust. The postzygapophyses are short and do not extend significantly laterally. The neural spine is high, and is mostly developed and augmenting in height in the posterior half of the neural arch. The neural arch is moderately vaulted in posterior view. Subcentral foramina are present.Two distinct foramina, each situated between each prezygapophysis, are present above the cotyle, a structure herein defined as “anocotylar” foramina (see Remarks below). Interestingly also, this specimen is pierced by distinct foramina in the dorsal surface of its neural arch.
|
||
</paragraph>
|
||
<subSubSection id="C31281A6FF83FFE33313FA4BF8AD3C03" lastPageId="34" lastPageNumber="251" pageId="32" pageNumber="249" type="discussion">
|
||
<paragraph id="8BB7D22DFF83FFE13313FA4BF9CF3E06" blockId="32.[811,1456,1523,2027]" box="[813,913,1523,1549]" pageId="32" pageNumber="249">REMARKS</paragraph>
|
||
<paragraph id="8BB7D22DFF83FFE13313F9AAFEAF3F66" blockId="32.[811,1456,1523,2027]" pageId="32" pageNumber="249">
|
||
The referral of this vertebral material to
|
||
<taxonomicName id="4C08A9AEFF83FFE134FAF9AAFF023E21" authorityName="Sullivan" authorityYear="1979" box="[1220,1372,1554,1580]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="249" phylum="Chordata" rank="tribe" tribe="Melanosaurini">Melanosaurini</taxonomicName>
|
||
is made primarily on the basis of the much anteriorly widened centrum than in other glyptosaurines, similar to that observed for the North American
|
||
<taxonomicName id="4C08A9AEFF83FFE133E1F9CAFF2B3E81" authority="Gilmore, 1928" authorityName="Gilmore" authorityYear="1928" box="[991,1397,1650,1676]" class="Reptilia" family="Anguidae" genus="Melanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="249" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B97C0E3FFF83FFE133E1F9CAFE913E81" box="[991,1231,1650,1676]" italics="true" pageId="32" pageNumber="249">Melanosaurus maximus</emphasis>
|
||
<bibRefCitation id="EF99AFDCFF83FFE134EBF9CAFF2B3E81" author="GILMORE C. W." box="[1237,1397,1650,1676]" pageId="32" pageNumber="249" pagination="1 - 197" refId="ref36362" refString="GILMORE C. W. 1928. - The fossil lizards of North America. Memoirs of the National Academy of Sciences 11: 1 - 197." type="journal article" year="1928">Gilmore, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
, and, to a lesser degree,
|
||
<taxonomicName id="4C08A9AEFF83FFE133DEF92AFECC3EA1" authorityName="Auge & Sullivan" authorityYear="2006" box="[992,1170,1682,1708]" class="Reptilia" family="Anguidae" genus="Paraplacosauriops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="249" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C0E3FFF83FFE133DEF92AFECC3EA1" box="[992,1170,1682,1708]" italics="true" pageId="32" pageNumber="249">Paraplacosauriops</emphasis>
|
||
</taxonomicName>
|
||
from the Eocene of Europe (see figures in
|
||
<bibRefCitation id="EF99AFDCFF83FFE13387F90AFE123EC1" author="GILMORE C. W." box="[953,1100,1714,1740]" pageId="32" pageNumber="249" pagination="1 - 197" refId="ref36362" refString="GILMORE C. W. 1928. - The fossil lizards of North America. Memoirs of the National Academy of Sciences 11: 1 - 197." type="journal article" year="1928">Gilmore 1928</bibRefCitation>
|
||
and Augé 2003, 2005). One other important difference between NHMW 2019/0094/0001 (but not the other melanosaurine vertebrae from our collection) and the above ones referred to
|
||
<taxonomicName id="4C08A9AEFF83FFE1344BF8AAFEB33F21" authorityName="Gervais" authorityYear="1848" box="[1141,1261,1810,1836]" class="Reptilia" family="Anguidae" genus="Placosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="249" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C0E3FFF83FFE1344BF8AAFEB33F21" box="[1141,1261,1810,1836]" italics="true" pageId="32" pageNumber="249">Placosaurus</emphasis>
|
||
</taxonomicName>
|
||
is that the former possess much more massive postzygapophyses that extend more prominently laterally in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="8BB7D22DFF83FFE1337DF8CAFFEE3FE6" blockId="32.[811,1456,1523,2027]" pageId="32" pageNumber="249">
|
||
A plausible taxonomic scenario could be that these specimens pertain to
|
||
<taxonomicName id="4C08A9AEFF83FFE1341AF829FF653FA6" baseAuthorityName="Filhol" baseAuthorityYear="1882" box="[1060,1339,1937,1963]" class="Reptilia" family="Anguidae" genus="Paraplacosauriops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="249" phylum="Chordata" rank="species" species="quercyi">
|
||
<emphasis id="B97C0E3FFF83FFE1341AF829FF653FA6" box="[1060,1339,1937,1963]" italics="true" pageId="32" pageNumber="249">Paraplacosauriops quercyi</emphasis>
|
||
</taxonomicName>
|
||
described above from cranial material, however, on the absence of articulated specimens and the imprecisely known locality
|
||
</paragraph>
|
||
<paragraph id="8BB7D22DFF82FFE03131F91CFBF33EB5" blockId="33.[271,429,1700,1720]" box="[271,429,1700,1720]" pageId="33" pageNumber="250">Meckel’s groove</paragraph>
|
||
<caption id="DF7782A5FF82FFE030BAF956FBF83F1B" ID-DOI="http://doi.org/10.5281/zenodo.4720853" ID-Zenodo-Dep="4720853" httpUri="https://zenodo.org/record/4720853/files/figure.png" pageId="33" pageNumber="250" startId="33.[132,143,1774,1791]" targetBox="[151,1434,217,1700]" targetPageId="33">
|
||
<paragraph id="8BB7D22DFF82FFE030BAF956FBF83F1B" blockId="33.[132,1455,1774,1814]" pageId="33" pageNumber="250">
|
||
FIG. 26. —
|
||
<taxonomicName id="4C08A9AEFF82FFE030DEF956F8713EF2" authority="(Filhol, 1882)" baseAuthorityName="Filhol" baseAuthorityYear="1882" box="[224,559,1774,1791]" class="Reptilia" family="Anguidae" genus="Paraplacosauriops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="250" phylum="Chordata" rank="species" species="quercyi">
|
||
<emphasis id="B97C0E3FFF82FFE030DEF956FBE23EF2" box="[224,444,1774,1791]" italics="true" pageId="33" pageNumber="250">Paraplacosauriops quercyi</emphasis>
|
||
(Filhol, 1882)
|
||
</taxonomicName>
|
||
. Virtual 3D models of right dentary NHMW 2019/0049/0001 in lateral (
|
||
<emphasis id="B97C0E3FFF82FFE03446F956FEDB3EF2" bold="true" box="[1144,1157,1774,1791]" pageId="33" pageNumber="250">A</emphasis>
|
||
), dorsal (
|
||
<emphasis id="B97C0E3FFF82FFE034ECF956FE813EF2" bold="true" box="[1234,1247,1774,1791]" pageId="33" pageNumber="250">B</emphasis>
|
||
), medial (
|
||
<emphasis id="B97C0E3FFF82FFE0350FF956FF613EF2" bold="true" box="[1329,1343,1774,1791]" pageId="33" pageNumber="250">C</emphasis>
|
||
), and ventromedial (
|
||
<emphasis id="B97C0E3FFF82FFE030F8F8BDFA8A3F1B" bold="true" box="[198,212,1797,1814]" pageId="33" pageNumber="250">D</emphasis>
|
||
) views. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB7D22DFF82FFE030BAF8E9FBC93FA1" blockId="33.[132,775,1873,2028]" pageId="33" pageNumber="250">data (including the fact that the material was probably collected from different localities), we refrain from referring them to the same taxon.</paragraph>
|
||
<paragraph id="8BB7D22DFF82FFE330A5F809F8AD3C03" blockId="33.[132,775,1873,2028]" lastBlockId="34.[131,776,757,1039]" lastPageId="34" lastPageNumber="251" pageId="33" pageNumber="250">
|
||
The presence of two distinct foramina above the cotyle of the sacral vertebra, a feature also prominent in several presacral vertebrae of
|
||
<taxonomicName id="4C08A9AEFF82FFE03419F8E9FE9F3F66" authorityName="Zittel" authorityYear="1887" box="[1063,1217,1873,1899]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="250" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C0E3FFF82FFE03419F8E9FE9F3F66" box="[1063,1217,1873,1899]" italics="true" pageId="33" pageNumber="250">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
(see below), is interesting. We acknowledge the presence of these structures in large-sized vertebrae of extant specimens of the anguid
|
||
<taxonomicName id="4C08A9AEFF82FFE03313F809FE153FC6" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" box="[813,1099,1969,1996]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="250" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C0E3FFF82FFE03313F809F9C73FC6" box="[813,921,1969,1995]" italics="true" pageId="33" pageNumber="250">Pseudopus</emphasis>
|
||
<bibRefCitation id="EF99AFDCFF82FFE0339DF80AFE153FC6" author="MERREM B." box="[931,1099,1969,1996]" pageId="33" pageNumber="250" refId="ref38229" refString="MERREM B. 1820. - Versuch eines systems der Amphibien, Vol. 8. J. C. Krieger, Marburg, 191 p. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem, 1820</bibRefCitation>
|
||
</taxonomicName>
|
||
, and the varanid
|
||
<taxonomicName id="4C08A9AEFF82FFE03527F80AF9FB3FE6" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="250" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C0E3FFF82FFE03527F80AFF2E3FC6" box="[1305,1392,1970,1995]" italics="true" pageId="33" pageNumber="250">Varanus</emphasis>
|
||
<bibRefCitation id="EF99AFDCFF82FFE03549F80AF9FB3FE6" author="MERREM B." pageId="33" pageNumber="250" refId="ref38229" refString="MERREM B. 1820. - Versuch eines systems der Amphibien, Vol. 8. J. C. Krieger, Marburg, 191 p. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem, 1820</bibRefCitation>
|
||
</taxonomicName>
|
||
. We consider that their presence is widespread in large-sized anguimorphs and is apparently correlated with large size; we define these structures as “anocotylar foramina”, from the Greek words “ἄνω” (“ano”), meaning “above” and “κότυλος” (“cotylos”), meaning “cup”, in a similar trend of the term “paracotylar foramina”, which applies in snake vertebrae terminology. The potential taxonomic utility of anocotylar foramina needs to be further investigated in the light of detailed quantitative analyses on extant forms, as well as articulated fossil specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |