treatments-xml/data/03/A1/63/03A1633BFFA9FFD2339EFC2CFE233D80.xml

<document id="EC3A837B96EA6245FCF37A5C9F433D18" ID-CLB-Dataset="7175" ID-DOI="10.5252/geodiversitas2021v43a9" ID-GBIF-Dataset="04426a8e-c179-4af0-8e23-d72908e8b4cc" ID-ISSN="1638-9395" ID-Zenodo-Dep="4720776" ID-ZooBank="urn:lsid:zoobank.org:pub:11D0D852-39D7-449C-9EB3-C3D804114556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445770811" checkinUser="felipe" docAuthor="Georgalis, Georgios L., Čerňanský, Andrej &amp; Klembara, Jozef" docDate="2021" docId="03A1633BFFA9FFD2339EFC2CFE233D80" docLanguage="en" docName="Geodiversitas.43.9.219-293.pdf" docOrigin="Geodiversitas 43 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pseudeumeces kyrillomethodicus Georgalis &amp; Čerňanský &amp; Klembara 2021, n. sp." docType="treatment" docUuid="FE001956-70BD-4493-8FD4-2F15E9A4778D" docUuidSource="ZooBank" docVersion="6" lastPageNumber="236" masterDocId="FF981B43FFA3FFC1303EFFB8FA5E380D" masterDocTitle="Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material" masterLastPageNumber="293" masterPageNumber="219" pageNumber="227" updateTime="1698937615995" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="4CE58FBE42112E6EF9BB1F960DD36E5B">Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material</mods:title>
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<mods:namePart id="91A4A8FA3C38EE6E39D4C7CD5D0497A3">Georgalis, Georgios L.</mods:namePart>
<mods:affiliation id="881397EED894BF80B9A92D2E49D83495">Palaeontological Institute and Museum, University of Zurich, Karl Schmid-Strasse 4, 8006 Zurich (Switzerland) and Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia) and Department of Earth Sciences, University of Torino, Via Valperga Caluso 35, 10125 Turin (Italy)</mods:affiliation>
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<mods:namePart id="6F303BB13BAC6E50808CCA9D4892A7DB">Čerňanský, Andrej</mods:namePart>
<mods:affiliation id="D4182DBB731495E57D9EBE210A572A52">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:namePart id="B18B8AFEA2A329761256A3CD5CA9ED37">Klembara, Jozef</mods:namePart>
<mods:affiliation id="6E2F23747903D05870056C278388547B">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<taxonomicName id="4C08A9AEFFA9FFCB339EFC2CFEA43BA2" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[928,1274,916,943]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="227" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFA9FFCB339EFC2CFEA43BA2" bold="true" box="[928,1274,916,943]" italics="true" pageId="10" pageNumber="227">Pseudeumeces kyrillomethodicus</emphasis>
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<taxonomicNameLabel id="A24FB344FFA9FFCB353FFC2DFF623BA2" box="[1281,1340,917,943]" pageId="10" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="1333CEA8FFA9FFCB3405FC0DFEFE3BC2" box="[1083,1184,949,975]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStart-8="FIG" captionStartId-0="11.[132,143,1985,2002]" captionStartId-1="12.[132,143,1804,1821]" captionStartId-2="13.[132,143,1410,1427]" captionStartId-3="14.[132,143,1565,1582]" captionStartId-4="15.[132,143,1985,2002]" captionStartId-5="16.[132,143,1821,1838]" captionStartId-6="17.[132,143,1376,1393]" captionStartId-7="18.[813,823,1078,1095]" captionStartId-8="19.[132,143,1114,1131]" captionTargetBox-0="[136,1448,215,1866]" captionTargetBox-1="[148,1456,187,1773]" captionTargetBox-2="[137,1450,213,1368]" captionTargetBox-3="[241,1346,215,1523]" captionTargetBox-4="[132,1455,177,1931]" captionTargetBox-5="[204,1408,177,1791]" captionTargetBox-6="[132,1455,198,1349]" captionTargetBox-7="[819,1449,215,1036]" captionTargetBox-8="[141,1447,215,1070]" captionTargetId-0="figure-16@11.[132,1455,211,1874]" captionTargetId-1="figure-91@12.[272,1280,231,1767]" captionTargetId-2="figure-360@13.[699,1262,746,1279]" captionTargetId-3="figure-212@14.[241,1346,215,1523]" captionTargetId-4="figure-16@15.[132,1455,215,1878]" captionTargetId-5="figure-73@16.[294,1323,209,1787]" captionTargetId-6="figure-309@17.[132,1455,198,1349]" captionTargetId-7="figure-833@18.[813,1456,211,1050]" captionTargetId-8="figure-428@19.[117,1483,215,1080]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="17" captionTargetPageId-7="18" captionTargetPageId-8="19" captionText-0="FIG. 6. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), posterior (B), medial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-1="FIG. 7. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-2="FIG. 8. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the holotype left dentary NHMW 2019/0051/0001 in medial (A) and dorsal (B) views. Scale bar: 1 mm." captionText-3="FIG. 9. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the holotype left dentary NHMW 2019/0051/0001: horizontal, with a special emphasis on the anterior and mid-section of the tooth row (A), horizonal, upper-more section with a special emphasis on the mid- and posterior section of the tooth row (B), and sagittal (C). Note the preserved tooth position showing the tooth number of 15. Scale bars: 2 mm." captionText-4="FIG. 10. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the paratype right dentary NHMW 2019/0051/0002 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-5="FIG. 11. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the paratype right dentary NHMW 2019/0051/0002 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-6="FIG. 12. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the paratype right dentary NHMW 2019/0051/0002 in medial (A) and dorsal (B) view. Scale bar: 1 mm." captionText-7="FIG. 13. — Pseudeumeces kyrillomethodicus n. sp. Close up photograph of the most robust tooth of the paratype right dentary NHMW 2019/0051/0002 in medial view. Note the prominent striations. Scale bar: 1 mm." captionText-8="FIG. 14. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the paratype right dentary NHMW 2019/0051/0002: horizontal (A) and sagittal (B). Note that the posterior section of the tooth row is missing. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720796" figureDoi-1="http://doi.org/10.5281/zenodo.4721436" figureDoi-2="http://doi.org/10.5281/zenodo.4720800" figureDoi-3="http://doi.org/10.5281/zenodo.4720802" figureDoi-4="http://doi.org/10.5281/zenodo.4720807" figureDoi-5="http://doi.org/10.5281/zenodo.4721441" figureDoi-6="http://doi.org/10.5281/zenodo.4720813" figureDoi-7="http://doi.org/10.5281/zenodo.4720815" figureDoi-8="http://doi.org/10.5281/zenodo.4720819" httpUri-0="https://zenodo.org/record/4720796/files/figure.png" httpUri-1="https://zenodo.org/record/4721436/files/figure.png" httpUri-2="https://zenodo.org/record/4720800/files/figure.png" httpUri-3="https://zenodo.org/record/4720802/files/figure.png" httpUri-4="https://zenodo.org/record/4720807/files/figure.png" httpUri-5="https://zenodo.org/record/4721441/files/figure.png" httpUri-6="https://zenodo.org/record/4720813/files/figure.png" httpUri-7="https://zenodo.org/record/4720815/files/figure.png" httpUri-8="https://zenodo.org/record/4720819/files/figure.png" pageId="10" pageNumber="227">Figs 6-14</figureCitation>
)
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BB7D22DFFA9FFCB3313FB8BFF333C46" blockId="10.[811,1457,1075,2026]" box="[813,1389,1075,1099]" pageId="10" pageNumber="227">
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<typeStatus id="54B36C8FFFA9FFCB3313FB8BF9C93C44" box="[813,919,1075,1098]" pageId="10" pageNumber="227" type="holotype">HOLOTYPE</typeStatus>
. — A left dentary (
<specimenCode id="DBAE7A56FFA9FFCB3468FB8BFEB53C46" box="[1110,1259,1075,1099]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="227" type="Museum">NHMW 2019</specimenCode>
/0051/0001).
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<materialsCitation id="3B60D870FFA9FFCB3313FBE5FF2C3C79" ID-GBIF-Occurrence="3091185316" box="[813,1394,1116,1141]" pageId="10" pageNumber="227" typeStatus="paratype">
<typeStatus id="54B36C8FFFA9FFCB3313FBE5F9D23C7E" box="[813,908,1117,1140]" pageId="10" pageNumber="227" type="paratype">PARATYPE</typeStatus>
. — A right dentary (
<collectionCode id="ED194AE8FFA9FFCB3465FBE5FEEA3C78" box="[1115,1204,1117,1141]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="227" type="Museum">NHMW</collectionCode>
2019/0051/0002).
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<subSubSection id="C31281A6FFA9FFCB3313FB3FFE8E3D83" pageId="10" pageNumber="227" type="etymology">
<paragraph id="8BB7D22DFFA9FFCB3313FB3FFE8E3D83" blockId="10.[811,1457,1075,2026]" pageId="10" pageNumber="227">
ETYMOLOGY. — The new species epithet “ 
<taxonomicName id="4C08A9AEFFA9FFCB348AFB3EFF0C3C93" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[1204,1362,1158,1182]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="227" phylum="Chordata" rank="species" species="kyrillomethodicus">
<emphasis id="B97C0E3FFFA9FFCB348AFB3EFF0C3C93" box="[1204,1362,1158,1182]" italics="true" pageId="10" pageNumber="227">kyrillomethodicus</emphasis>
</taxonomicName>
” honours Kyrillos (or Cyril; Greek: Κύριλλος) and Methodios (or Methodius; Greek: Μεθόδιος), the two Byzantine brothers from Thessaloniki that were sent by the Byzantine Emperor to the area of Bratislava (city of Dovina [currently Devin]) in 863 AD, where they created the first scripture for the people of Great 
<collectingRegion id="49CC1CCFFFA9FFCB34F9FAB3FF423D2E" box="[1223,1308,1291,1315]" country="Czech Republic" name="Jihomoravsky" pageId="10" pageNumber="227">Moravia</collectingRegion>
. The name alludes to the fact that one of us (GLG) came from Thessaloniki to Bratislava (during which project also this paper was written) and generally to our nice collaboration among the three of us, Greek and Slovak scientists. Gender is masculine.
</paragraph>
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<subSubSection id="C31281A6FFA9FFCB3313FA18FE1E3DE0" pageId="10" pageNumber="227" type="materials_examined">
<paragraph id="8BB7D22DFFA9FFCB3313FA18FE1E3DE0" blockId="10.[811,1457,1075,2026]" pageId="10" pageNumber="227">
<collectionCode id="ED194AE8FFA9FFCB3313FA18F99C3DBB" box="[813,962,1440,1463]" pageId="10" pageNumber="227">TYPE LOCALITY</collectionCode>
. — 
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Imprecisely known locality, Phosphorites du Quercy, Department of Lot orTarn-et-Garonne, Occitanie, southern 
<collectingCountry id="F31F92BDFFA9FFCB3315FA6EF9313DE0" box="[811,879,1494,1517]" name="France" pageId="10" pageNumber="227">France</collectingCountry>
; probably Oligocene.
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</paragraph>
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<subSubSection id="C31281A6FFA9FFCB3313FA47FF773FAB" pageId="10" pageNumber="227" type="diagnosis">
<paragraph id="8BB7D22DFFA9FFCB3313FA47FF773FAB" blockId="10.[811,1457,1075,2026]" pageId="10" pageNumber="227">
DIAGNOSIS. — A medium sized lacertid, morphologically similar to 
<taxonomicName id="4C08A9AEFFA9FFCB3315F9A1FE4C3E3C" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[811,1042,1561,1585]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="227" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFA9FFCB3315F9A1FE4C3E3C" box="[811,1042,1561,1585]" italics="true" pageId="10" pageNumber="227">Pseudeumeces cadurcensis</emphasis>
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, but differing from it in the following combination of features: 1) the last posterior three or four dentary teeth are reduced (instead of only the last one or two posterior teeth being reduced); 2) the dentary is short and massive rather than anteroposteriorly long relative to its dorsoventral height (the overall morphology of dentary in 
<taxonomicName id="4C08A9AEFFA9FFCB33EBF927FEE33EBA" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[981,1213,1695,1719]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="227" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFA9FFCB33EBF927FEE33EBA" box="[981,1213,1695,1719]" italics="true" pageId="10" pageNumber="227">Pseudeumeces cadurcensis</emphasis>
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has narrower appearance relative to that in 
<taxonomicName id="4C08A9AEFFA9FFCB33EAF901FEA93EDC" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[980,1271,1721,1745]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="227" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFA9FFCB33EAF901FEA93EDC" box="[980,1271,1721,1745]" italics="true" pageId="10" pageNumber="227">Pseudeumeces kyrillomethodicus</emphasis>
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<taxonomicNameLabel id="A24FB344FFA9FFCB34C3F902FF6D3EDC" box="[1277,1331,1721,1745]" pageId="10" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
); 3) the facet for the anterolateral process of the coronoid reaches around the level of the 4 
<superScript id="7C7D7F65FFA9FFCB334AF957F9DC3F0D" attach="left" box="[884,898,1775,1792]" fontSize="7" pageId="10" pageNumber="227">th</superScript>
tooth position (counted from posterior) rather than reaching the level of the posteriormost tooth position (in some cases, the level between the last and penultimate tooth positions); 4) the facet for anteromedial process of the coronoid reaches the level of the 3rd tooth position (counted from posterior) rather than only the level of the posteriormost tooth position; and 5) the dorsal elevation of the posterior section of the dentary is more pronounced.
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FFA9FFCB3313F801F9293FE7" pageId="10" pageNumber="227" type="distribution">
<paragraph id="8BB7D22DFFA9FFCB3313F801F9293FE7" blockId="10.[811,1457,1075,2026]" pageId="10" pageNumber="227">
DISTRIBUTION. — Oligocene, Phosphorites du Quercy, southern 
<collectingCountry id="F31F92BDFFA9FFCB3310F86BF92C3FE7" box="[814,882,2003,2026]" name="France" pageId="10" pageNumber="227">France</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF7782A5FFA8FFCA30BAF879FBFC3FE4" ID-DOI="http://doi.org/10.5281/zenodo.4720796" ID-Zenodo-Dep="4720796" httpUri="https://zenodo.org/record/4720796/files/figure.png" pageId="11" pageNumber="228" startId="11.[132,143,1985,2002]" targetBox="[136,1448,215,1866]" targetPageId="11">
<paragraph id="8BB7D22DFFA8FFCA30BAF879FBFC3FE4" blockId="11.[132,1455,1985,2025]" pageId="11" pageNumber="228">
FIG. 6. — 
<taxonomicName id="4C08A9AEFFA8FFCA30EBF879FBB23FDF" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[213,492,1985,2002]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="228" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFA8FFCA30EBF879FBB23FDF" box="[213,492,1985,2002]" italics="true" pageId="11" pageNumber="228">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFA8FFCA31CCF879F87D3FDF" box="[498,547,1985,2002]" pageId="11" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
Photographs of the holotype left dentary NHMW 2019/0051/0001 in lateral (
<emphasis id="B97C0E3FFFA8FFCA349CF879FEF13FDF" bold="true" box="[1186,1199,1985,2002]" pageId="11" pageNumber="228">A</emphasis>
), posterior (
<emphasis id="B97C0E3FFFA8FFCA352DF879FF7E3FDF" bold="true" box="[1299,1312,1985,2002]" pageId="11" pageNumber="228">B</emphasis>
), medial (
<emphasis id="B97C0E3FFFA8FFCA354CF879FFDE3FDF" bold="true" box="[1394,1408,1985,2002]" pageId="11" pageNumber="228">C</emphasis>
), and dorsal (
<emphasis id="B97C0E3FFFA8FFCA30FCF860FA8E3FE4" bold="true" box="[194,208,2008,2025]" pageId="11" pageNumber="228">D</emphasis>
) views. Scale bar: 2 mm.
</paragraph>
</caption>
<caption id="DF7782A5FFAFFFCD30BAF8B4FBB73F39" ID-DOI="http://doi.org/10.5281/zenodo.4721436" ID-Zenodo-Dep="4721436" httpUri="https://zenodo.org/record/4721436/files/figure.png" pageId="12" pageNumber="229" startId="12.[132,143,1804,1821]" targetBox="[148,1456,187,1773]" targetPageId="12">
<paragraph id="8BB7D22DFFAFFFCD30BAF8B4FBB73F39" blockId="12.[132,1455,1804,1844]" pageId="12" pageNumber="229">
FIG. 7. — 
<taxonomicName id="4C08A9AEFFAFFFCD30EBF8B4FBB23F10" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[213,492,1804,1821]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="229" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFAFFFCD30EBF8B4FBB23F10" box="[213,492,1804,1821]" italics="true" pageId="12" pageNumber="229">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFAFFFCD31CFF8B4F87C3F10" box="[497,546,1804,1821]" pageId="12" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
Virtual 3D models of the holotype left dentary NHMW 2019/0051/0001 in lateral (
<emphasis id="B97C0E3FFFAFFFCD34F6F8B4FE8B3F10" bold="true" box="[1224,1237,1804,1821]" pageId="12" pageNumber="229">A</emphasis>
), medial (
<emphasis id="B97C0E3FFFAFFFCD3518F8B4FF6D3F10" bold="true" box="[1318,1331,1804,1821]" pageId="12" pageNumber="229">B</emphasis>
), ventromedial (
<emphasis id="B97C0E3FFFAFFFCD30B7F89BFAC93F39" bold="true" box="[137,151,1827,1844]" pageId="12" pageNumber="229">C</emphasis>
), and dorsal (
<emphasis id="B97C0E3FFFAFFFCD3136F89BFB483F39" bold="true" box="[264,278,1827,1844]" pageId="12" pageNumber="229">D</emphasis>
) views. Scale bar: 2 mm.
</paragraph>
</caption>
<subSubSection id="C31281A6FFAFFFD030BAF8CAFAB33F01" lastPageId="17" lastPageNumber="234" pageId="12" pageNumber="229" type="description">
<paragraph id="8BB7D22DFFAFFFCD30BAF8CAFB4B3F87" blockId="12.[132,775,1906,2028]" box="[132,277,1906,1932]" pageId="12" pageNumber="229">DESCRIPTION</paragraph>
<paragraph id="8BB7D22DFFAFFFCD30BAF829F8323FA1" blockId="12.[132,775,1906,2028]" box="[132,620,1937,1964]" pageId="12" pageNumber="229">
<emphasis id="B97C0E3FFFAFFFCD30BAF829F8323FA1" box="[132,620,1937,1964]" italics="true" pageId="12" pageNumber="229">
<typeStatus id="54B36C8FFFAFFFCD30BAF829FA813FA6" box="[132,223,1937,1963]" pageId="12" pageNumber="229" type="holotype">Holotype</typeStatus>
<specimenCode id="DBAE7A56FFAFFFCD30D8F82AFB203FA6" box="[230,382,1937,1963]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="229" type="Museum">NHMW 2019</specimenCode>
/0051/0001 (
<figureCitation id="1333CEA8FFAFFFCD3230F82AF83D3FA6" box="[526,611,1937,1963]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="11.[132,143,1985,2002]" captionStartId-1="12.[132,143,1804,1821]" captionStartId-2="13.[132,143,1410,1427]" captionStartId-3="14.[132,143,1565,1582]" captionTargetBox-0="[136,1448,215,1866]" captionTargetBox-1="[148,1456,187,1773]" captionTargetBox-2="[137,1450,213,1368]" captionTargetBox-3="[241,1346,215,1523]" captionTargetId-0="figure-16@11.[132,1455,211,1874]" captionTargetId-1="figure-91@12.[272,1280,231,1767]" captionTargetId-2="figure-360@13.[699,1262,746,1279]" captionTargetId-3="figure-212@14.[241,1346,215,1523]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIG. 6. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), posterior (B), medial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-1="FIG. 7. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-2="FIG. 8. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the holotype left dentary NHMW 2019/0051/0001 in medial (A) and dorsal (B) views. Scale bar: 1 mm." captionText-3="FIG. 9. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the holotype left dentary NHMW 2019/0051/0001: horizontal, with a special emphasis on the anterior and mid-section of the tooth row (A), horizonal, upper-more section with a special emphasis on the mid- and posterior section of the tooth row (B), and sagittal (C). Note the preserved tooth position showing the tooth number of 15. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720796" figureDoi-1="http://doi.org/10.5281/zenodo.4721436" figureDoi-2="http://doi.org/10.5281/zenodo.4720800" figureDoi-3="http://doi.org/10.5281/zenodo.4720802" httpUri-0="https://zenodo.org/record/4720796/files/figure.png" httpUri-1="https://zenodo.org/record/4721436/files/figure.png" httpUri-2="https://zenodo.org/record/4720800/files/figure.png" httpUri-3="https://zenodo.org/record/4720802/files/figure.png" pageId="12" pageNumber="229">Figs 6-9</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BB7D22DFFAFFFCD30BAF809F9593FE1" blockId="12.[132,775,1906,2028]" pageId="12" pageNumber="229">
The 
<typeStatus id="54B36C8FFFAFFFCD308BF809FB4A3FC6" box="[181,276,1969,1995]" pageId="12" pageNumber="229" type="holotype">holotype</typeStatus>
left dentary NHMW 2019/0051/0001 is the most complete specimen (
<figureCitation id="1333CEA8FFAFFFCD31A0F869FBAA3FE6" box="[414,500,2001,2028]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="11.[132,143,1985,2002]" captionStartId-1="12.[132,143,1804,1821]" captionStartId-2="13.[132,143,1410,1427]" captionStartId-3="14.[132,143,1565,1582]" captionTargetBox-0="[136,1448,215,1866]" captionTargetBox-1="[148,1456,187,1773]" captionTargetBox-2="[137,1450,213,1368]" captionTargetBox-3="[241,1346,215,1523]" captionTargetId-0="figure-16@11.[132,1455,211,1874]" captionTargetId-1="figure-91@12.[272,1280,231,1767]" captionTargetId-2="figure-360@13.[699,1262,746,1279]" captionTargetId-3="figure-212@14.[241,1346,215,1523]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIG. 6. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), posterior (B), medial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-1="FIG. 7. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-2="FIG. 8. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the holotype left dentary NHMW 2019/0051/0001 in medial (A) and dorsal (B) views. Scale bar: 1 mm." captionText-3="FIG. 9. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the holotype left dentary NHMW 2019/0051/0001: horizontal, with a special emphasis on the anterior and mid-section of the tooth row (A), horizonal, upper-more section with a special emphasis on the mid- and posterior section of the tooth row (B), and sagittal (C). Note the preserved tooth position showing the tooth number of 15. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720796" figureDoi-1="http://doi.org/10.5281/zenodo.4721436" figureDoi-2="http://doi.org/10.5281/zenodo.4720800" figureDoi-3="http://doi.org/10.5281/zenodo.4720802" httpUri-0="https://zenodo.org/record/4720796/files/figure.png" httpUri-1="https://zenodo.org/record/4721436/files/figure.png" httpUri-2="https://zenodo.org/record/4720800/files/figure.png" httpUri-3="https://zenodo.org/record/4720802/files/figure.png" pageId="12" pageNumber="229">Figs 6-9</figureCitation>
). The specimen measures
</paragraph>
<paragraph id="8BB7D22DFFAFFFCC3312F8CAFF613F86" blockId="12.[811,1457,1905,2028]" lastBlockId="13.[811,1458,1523,2028]" lastPageId="13" lastPageNumber="230" pageId="12" pageNumber="229">
a total length of 
<quantity id="4CF07FC8FFAFFFCD33D2F8C9FE1D3F81" box="[1004,1091,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="229" unit="mm" value="16.0">16 mm</quantity>
. It is short, massively built, and ventrally deep. In cross-section, the dentary is C-shaped. In medial aspect, the dentary has a dorsally concave appearance and it gradually widens posteriorly. The CT scan reveals that the 
<typeStatus id="54B36C8FFFAEFFCC3095FA4BFB573E00" box="[171,265,1523,1549]" pageId="13" pageNumber="230" type="holotype">holotype</typeStatus>
dentary bears 15 tooth positions (see 
<figureCitation id="1333CEA8FFAEFFCC32AEFA4BF8923E00" box="[656,716,1523,1549]" captionStart="FIG" captionStartId="14.[132,143,1565,1582]" captionTargetBox="[241,1346,215,1523]" captionTargetId="figure-212@14.[241,1346,215,1523]" captionTargetPageId="14" captionText="FIG. 9. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the holotype left dentary NHMW 2019/0051/0001: horizontal, with a special emphasis on the anterior and mid-section of the tooth row (A), horizonal, upper-more section with a special emphasis on the mid- and posterior section of the tooth row (B), and sagittal (C). Note the preserved tooth position showing the tooth number of 15. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720802" httpUri="https://zenodo.org/record/4720802/files/figure.png" pageId="13" pageNumber="230">Fig. 9</figureCitation>
). On the medial side, the deep Meckel’s groove is fully open and distinctly broad posteriorly. It gradually narrows anteriorly, but it is still relatively wide even in this region. It disappears posterior to the large symphyseal region. CT images show that it continues inside to the bone further anteriorly only as a small internal canal. The alveolar foramen is small, being located at the level between the 4 
<superScript id="7C7D7F65FFAEFFCC31BFF96BFBCF3EE9" attach="left" box="[385,401,1747,1764]" fontSize="7" pageId="13" pageNumber="230">th</superScript>
and 5 
<superScript id="7C7D7F65FFAEFFCC31E6F96BFBB63EE9" attach="left" box="[472,488,1747,1764]" fontSize="7" pageId="13" pageNumber="230">th</superScript>
tooth positions (counting from the posterior tooth; in other words, at the level of 11 
<superScript id="7C7D7F65FFAEFFCC32C9F94BF9593F09" attach="none" box="[759,775,1779,1796]" fontSize="7" pageId="13" pageNumber="230">th</superScript>
and 12 
<superScript id="7C7D7F65FFAEFFCC30F3F8ABFA833F29" attach="left" box="[205,221,1811,1828]" fontSize="7" pageId="13" pageNumber="230">th</superScript>
tooth position counted from anterior). Further anteriorly, the alveolar canal is separated from the Meckel’s groove by the intramandibular septum, which is completely fused to the bone. Meckel’s groove is roofed dorsally by the subdental shelf, which bears a facet for the splenial on its ventral side. This facet reaches the level of around the 5th tooth position (counted from anterior) and its anterior end is well stepped. Dorsally, the subdental shelf bears the sulcus dentalis. The whole shelf gradually narrows posteriorly in medial aspect. The posterodorsal portion of the dentary is inclined dorsally, being well elevated. This inclination starts at the level of the 4th tooth position (counted from posterior). Posterior to the tooth row, there is a long and robust coronoid process. It reaches highly above the level of the largest tooth crowns. On its dorsomedial side, it bears a facet for the anteromedial process of the coronoid. It reaches the level of the 3 
<superScript id="7C7D7F65FFAEFFCC3562F94BFF323F09" attach="left" box="[1372,1388,1779,1796]" fontSize="7" pageId="13" pageNumber="230">rd</superScript>
tooth position (counted from posterior). The posteroventral portion of the dentary is preserved. This region is slightly curved medially, forming a short angular process. Nevertheless, this region is also slightly weathered in this specimen.
</paragraph>
<caption id="DF7782A5FFAEFFCC30BAFA3AFBFC3DA7" ID-DOI="http://doi.org/10.5281/zenodo.4720800" ID-Zenodo-Dep="4720800" httpUri="https://zenodo.org/record/4720800/files/figure.png" pageId="13" pageNumber="230" startId="13.[132,143,1410,1427]" targetBox="[137,1450,213,1368]" targetPageId="13">
<paragraph id="8BB7D22DFFAEFFCC30BAFA3AFBFC3DA7" blockId="13.[132,1455,1410,1450]" pageId="13" pageNumber="230">
FIG. 8. — 
<taxonomicName id="4C08A9AEFFAEFFCC30EAFA3AFBB73D9E" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[212,489,1410,1427]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="230" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFAEFFCC30EAFA3AFBB73D9E" box="[212,489,1410,1427]" italics="true" pageId="13" pageNumber="230">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFAEFFCC31D0FA3AF8433D9E" box="[494,541,1410,1427]" pageId="13" pageNumber="230" rank="species">n. sp.</taxonomicNameLabel>
Close up photographs of posterior teeth of the holotype left dentary NHMW 2019/0051/0001 in medial (
<emphasis id="B97C0E3FFFAEFFCC3547FA3AFFD83D9E" bold="true" box="[1401,1414,1410,1427]" pageId="13" pageNumber="230">A</emphasis>
) and dorsal (
<emphasis id="B97C0E3FFFAEFFCC30FCFA21FA913DA7" bold="true" box="[194,207,1433,1450]" pageId="13" pageNumber="230">B</emphasis>
) views. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFFAEFFCF337DF829F95B3F00" blockId="13.[811,1458,1523,2028]" lastBlockId="14.[130,775,1682,1805]" lastPageId="14" lastPageNumber="231" pageId="13" pageNumber="230">
The otherwise smooth lateral surface of the dentary is pierced by five labial foramina (
<figureCitation id="1333CEA8FFAEFFCC341FF809FE2C3FC6" box="[1057,1138,1969,1995]" captionStart="FIG" captionStartId="11.[132,143,1985,2002]" captionTargetBox="[136,1448,215,1866]" captionTargetId="figure-16@11.[132,1455,211,1874]" captionTargetPageId="11" captionText="FIG. 6. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), posterior (B), medial (C), and dorsal (D) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720796" httpUri="https://zenodo.org/record/4720796/files/figure.png" pageId="13" pageNumber="230">Figs 6A</figureCitation>
; 
<figureCitation id="1333CEA8FFAEFFCC3445F80AFEC03FC6" box="[1147,1182,1969,1996]" captionStart="FIG" captionStartId="12.[132,143,1804,1821]" captionTargetBox="[148,1456,187,1773]" captionTargetId="figure-91@12.[272,1280,231,1767]" captionTargetPageId="12" captionText="FIG. 7. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the holotype left dentary NHMW 2019/0051/0001 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4721436" httpUri="https://zenodo.org/record/4721436/files/figure.png" pageId="13" pageNumber="230">7A</figureCitation>
). The posteriormost one is located at the level of around the 5th tooth position (counted from posterior). In the posterodorsal region of the dentary, the wedge shaped facet for the anterolateral process of coronoid is present, being, however, very shallow. It appears to reach the level of the 4 
<superScript id="7C7D7F65FFADFFCF3106F94BFB163F09" attach="left" box="[312,328,1779,1796]" fontSize="7" pageId="14" pageNumber="231">th</superScript>
tooth position (counted from posterior).
</paragraph>
<caption id="DF7782A5FFADFFCF30BAF9A5F9ED3E56" ID-DOI="http://doi.org/10.5281/zenodo.4720802" ID-Zenodo-Dep="4720802" httpUri="https://zenodo.org/record/4720802/files/figure.png" pageId="14" pageNumber="231" startId="14.[132,143,1565,1582]" targetBox="[241,1346,215,1523]" targetPageId="14">
<paragraph id="8BB7D22DFFADFFCF30BAF9A5F9ED3E56" blockId="14.[132,1455,1565,1627]" pageId="14" pageNumber="231">
FIG. 9. — 
<taxonomicName id="4C08A9AEFFADFFCF30EBF9A5FBB53E23" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[213,491,1565,1582]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="231" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFADFFCF30EBF9A5FBB53E23" box="[213,491,1565,1582]" italics="true" pageId="14" pageNumber="231">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFADFFCF31CFF9A5F87F3E23" box="[497,545,1565,1582]" pageId="14" pageNumber="231" rank="species">n. sp.</taxonomicNameLabel>
Virtual slices of the holotype left dentary NHMW 2019/0051/0001: horizontal, with a special emphasis on the anterior and mid-section of the tooth row (
<emphasis id="B97C0E3FFFADFFCF31DAF98CFBAF3E48" bold="true" box="[484,497,1588,1605]" pageId="14" pageNumber="231">A</emphasis>
), horizonal, upper-more section with a special emphasis on the mid- and posterior section of the tooth row (
<emphasis id="B97C0E3FFFADFFCF354DF98CFFDE3E48" bold="true" box="[1395,1408,1588,1605]" pageId="14" pageNumber="231">B</emphasis>
), and sagittal (
<emphasis id="B97C0E3FFFADFFCF30F4F9F2FA863E56" bold="true" box="[202,216,1610,1627]" pageId="14" pageNumber="231">C</emphasis>
). Note the preserved tooth position showing the tooth number of 15. Scale bars: 2 mm.
</paragraph>
</caption>
<subSection id="E287CBC6FFADFFD130BAF88AFE2F3FE1" lastPageId="16" lastPageNumber="233" pageId="14" pageNumber="231" type="multiple">
<paragraph id="8BB7D22DFFADFFCF30BAF88AF8DB3F41" blockId="14.[132,645,1842,1868]" box="[132,645,1842,1868]" pageId="14" pageNumber="231">
<heading id="D0FF6541FFADFFCF30BAF88AF8DB3F41" box="[132,645,1842,1868]" fontSize="11" level="2" pageId="14" pageNumber="231" reason="8">
<emphasis id="B97C0E3FFFADFFCF30BAF88AF8DB3F41" box="[132,645,1842,1868]" italics="true" pageId="14" pageNumber="231">
<typeStatus id="54B36C8FFFADFFCF30BAF88AFA823F46" box="[132,220,1842,1867]" pageId="14" pageNumber="231" type="paratype">Paratype</typeStatus>
NHMW 2019/0051/0002 (
<figureCitation id="1333CEA8FFADFFCF3235F88AF8223F41" box="[523,636,1842,1868]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStartId-0="15.[132,143,1985,2002]" captionStartId-1="16.[132,143,1821,1838]" captionStartId-2="17.[132,143,1376,1393]" captionStartId-3="18.[813,823,1078,1095]" captionStartId-4="19.[132,143,1114,1131]" captionTargetBox-0="[132,1455,177,1931]" captionTargetBox-1="[204,1408,177,1791]" captionTargetBox-2="[132,1455,198,1349]" captionTargetBox-3="[819,1449,215,1036]" captionTargetBox-4="[141,1447,215,1070]" captionTargetId-0="figure-16@15.[132,1455,215,1878]" captionTargetId-1="figure-73@16.[294,1323,209,1787]" captionTargetId-2="figure-309@17.[132,1455,198,1349]" captionTargetId-3="figure-833@18.[813,1456,211,1050]" captionTargetId-4="figure-428@19.[117,1483,215,1080]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionText-0="FIG. 10. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the paratype right dentary NHMW 2019/0051/0002 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-1="FIG. 11. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the paratype right dentary NHMW 2019/0051/0002 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-2="FIG. 12. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the paratype right dentary NHMW 2019/0051/0002 in medial (A) and dorsal (B) view. Scale bar: 1 mm." captionText-3="FIG. 13. — Pseudeumeces kyrillomethodicus n. sp. Close up photograph of the most robust tooth of the paratype right dentary NHMW 2019/0051/0002 in medial view. Note the prominent striations. Scale bar: 1 mm." captionText-4="FIG. 14. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the paratype right dentary NHMW 2019/0051/0002: horizontal (A) and sagittal (B). Note that the posterior section of the tooth row is missing. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720807" figureDoi-1="http://doi.org/10.5281/zenodo.4721441" figureDoi-2="http://doi.org/10.5281/zenodo.4720813" figureDoi-3="http://doi.org/10.5281/zenodo.4720815" figureDoi-4="http://doi.org/10.5281/zenodo.4720819" httpUri-0="https://zenodo.org/record/4720807/files/figure.png" httpUri-1="https://zenodo.org/record/4721441/files/figure.png" httpUri-2="https://zenodo.org/record/4720813/files/figure.png" httpUri-3="https://zenodo.org/record/4720815/files/figure.png" httpUri-4="https://zenodo.org/record/4720819/files/figure.png" pageId="14" pageNumber="231">Figs 10-14</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7D22DFFADFFCF30BAF8E9F9593FE6" blockId="14.[132,777,1873,2028]" pageId="14" pageNumber="231">
Besides the 
<typeStatus id="54B36C8FFFADFFCF3134F8E9FB353F66" box="[266,363,1873,1899]" pageId="14" pageNumber="231" type="holotype">holotype</typeStatus>
described above, only a single other dentary, the 
<typeStatus id="54B36C8FFFADFFCF3129F8CAFB293F81" box="[279,375,1906,1932]" pageId="14" pageNumber="231" type="paratype">paratype</typeStatus>
NHMW 2019/0051/0002, can be confidently referred to 
<taxonomicName id="4C08A9AEFFADFFCF31AFF829F8B73FA6" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[401,745,1937,1963]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="231" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFADFFCF31AFF829F8B73FA6" box="[401,745,1937,1963]" italics="true" pageId="14" pageNumber="231">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFADFFCF32CDF82AFAFA3FC1" pageId="14" pageNumber="231" rank="species">n. sp.</taxonomicNameLabel>
This is rather similar to the 
<typeStatus id="54B36C8FFFADFFCF31C4F809F83D3FC6" box="[506,611,1969,1995]" pageId="14" pageNumber="231" type="holotype">holotype</typeStatus>
. The 
<typeStatus id="54B36C8FFFADFFCF329BF80AF9593FC1" box="[677,775,1970,1996]" pageId="14" pageNumber="231" type="paratype">paratype</typeStatus>
NHMW 2019/0051/0002 is larger and more robust than
</paragraph>
<paragraph id="8BB7D22DFFADFFCF3310F92AFFEE3FE1" blockId="14.[812,1457,1682,2028]" pageId="14" pageNumber="231">
the 
<typeStatus id="54B36C8FFFADFFCF3366F92AF9E73EA1" box="[856,953,1682,1708]" pageId="14" pageNumber="231" type="holotype">holotype</typeStatus>
NHMW 2019/0051/0001 (
<figureCitation id="1333CEA8FFADFFCF34C6F92AFF333EA1" box="[1272,1389,1682,1709]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStartId-0="15.[132,143,1985,2002]" captionStartId-1="16.[132,143,1821,1838]" captionStartId-2="17.[132,143,1376,1393]" captionStartId-3="18.[813,823,1078,1095]" captionStartId-4="19.[132,143,1114,1131]" captionTargetBox-0="[132,1455,177,1931]" captionTargetBox-1="[204,1408,177,1791]" captionTargetBox-2="[132,1455,198,1349]" captionTargetBox-3="[819,1449,215,1036]" captionTargetBox-4="[141,1447,215,1070]" captionTargetId-0="figure-16@15.[132,1455,215,1878]" captionTargetId-1="figure-73@16.[294,1323,209,1787]" captionTargetId-2="figure-309@17.[132,1455,198,1349]" captionTargetId-3="figure-833@18.[813,1456,211,1050]" captionTargetId-4="figure-428@19.[117,1483,215,1080]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionText-0="FIG. 10. — Pseudeumeces kyrillomethodicus n. sp. Photographs of the paratype right dentary NHMW 2019/0051/0002 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-1="FIG. 11. — Pseudeumeces kyrillomethodicus n. sp. Virtual 3D models of the paratype right dentary NHMW 2019/0051/0002 in lateral (A), medial (B), ventromedial (C), and dorsal (D) views. Scale bar: 2 mm." captionText-2="FIG. 12. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the paratype right dentary NHMW 2019/0051/0002 in medial (A) and dorsal (B) view. Scale bar: 1 mm." captionText-3="FIG. 13. — Pseudeumeces kyrillomethodicus n. sp. Close up photograph of the most robust tooth of the paratype right dentary NHMW 2019/0051/0002 in medial view. Note the prominent striations. Scale bar: 1 mm." captionText-4="FIG. 14. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the paratype right dentary NHMW 2019/0051/0002: horizontal (A) and sagittal (B). Note that the posterior section of the tooth row is missing. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720807" figureDoi-1="http://doi.org/10.5281/zenodo.4721441" figureDoi-2="http://doi.org/10.5281/zenodo.4720813" figureDoi-3="http://doi.org/10.5281/zenodo.4720815" figureDoi-4="http://doi.org/10.5281/zenodo.4720819" httpUri-0="https://zenodo.org/record/4720807/files/figure.png" httpUri-1="https://zenodo.org/record/4721441/files/figure.png" httpUri-2="https://zenodo.org/record/4720813/files/figure.png" httpUri-3="https://zenodo.org/record/4720815/files/figure.png" httpUri-4="https://zenodo.org/record/4720819/files/figure.png" pageId="14" pageNumber="231">Figs 10-14</figureCitation>
). The posteroventral portion of the dentary that is preserved in the 
<typeStatus id="54B36C8FFFADFFCF3367F96AF9E43EE1" box="[857,954,1746,1772]" pageId="14" pageNumber="231" type="holotype">holotype</typeStatus>
is not preserved here and only 12 tooth positions can be therefore observed in the 
<typeStatus id="54B36C8FFFADFFCF34DAF94AFF1D3F01" box="[1252,1347,1778,1804]" pageId="14" pageNumber="231" type="paratype">paratype</typeStatus>
NHMW 2019/0051/0002 (see 
<figureCitation id="1333CEA8FFADFFCF3416F8AAFE2B3F21" box="[1064,1141,1810,1836]" captionStart="FIG" captionStartId="19.[132,143,1114,1131]" captionTargetBox="[141,1447,215,1070]" captionTargetId="figure-428@19.[117,1483,215,1080]" captionTargetPageId="19" captionText="FIG. 14. — Pseudeumeces kyrillomethodicus n. sp. Virtual slices of the paratype right dentary NHMW 2019/0051/0002: horizontal (A) and sagittal (B). Note that the posterior section of the tooth row is missing. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720819" httpUri="https://zenodo.org/record/4720819/files/figure.png" pageId="14" pageNumber="231">Fig. 14</figureCitation>
). The facet for coronoid on the lateral side is more developed in the 
<typeStatus id="54B36C8FFFADFFCF34C8F88AFF0A3F41" box="[1270,1364,1842,1868]" pageId="14" pageNumber="231" type="paratype">paratype</typeStatus>
dentary if compared to the 
<typeStatus id="54B36C8FFFADFFCF33C1F8E9FE3A3F66" box="[1023,1124,1873,1899]" pageId="14" pageNumber="231" type="holotype">holotype</typeStatus>
. When the specimen NHMW 2019/0051/0002 was originally complete and in the case it possessed 15 tooth positions as the 
<typeStatus id="54B36C8FFFADFFCF34DEF829FF1B3FA6" box="[1248,1349,1937,1963]" pageId="14" pageNumber="231" type="holotype">holotype</typeStatus>
, then the facet for the anterolateral process of the coronoid reached the level of the 5th tooth position (counted from posterior).
</paragraph>
<caption id="DF7782A5FFACFFCE30BAF879FB993FE4" ID-DOI="http://doi.org/10.5281/zenodo.4720807" ID-Zenodo-Dep="4720807" httpUri="https://zenodo.org/record/4720807/files/figure.png" pageId="15" pageNumber="232" startId="15.[132,143,1985,2002]" targetBox="[132,1455,177,1931]" targetPageId="15">
<paragraph id="8BB7D22DFFACFFCE30BAF879FB993FE4" blockId="15.[132,1456,1985,2025]" pageId="15" pageNumber="232">
FIG. 10. — 
<taxonomicName id="4C08A9AEFFACFFCE30E0F879FBAD3FDF" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[222,499,1985,2002]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="232" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFACFFCE30E0F879FBAD3FDF" box="[222,499,1985,2002]" italics="true" pageId="15" pageNumber="232">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFACFFCE31C6F879F8793FDF" box="[504,551,1985,2002]" pageId="15" pageNumber="232" rank="species">n. sp.</taxonomicNameLabel>
Photographs of the paratype right dentary NHMW 2019/0051/0002 in lateral (
<emphasis id="B97C0E3FFFACFFCE3497F879FEE83FDF" bold="true" box="[1193,1206,1985,2002]" pageId="15" pageNumber="232">A</emphasis>
), medial (
<emphasis id="B97C0E3FFFACFFCE3538F879FF4D3FDF" bold="true" box="[1286,1299,1985,2002]" pageId="15" pageNumber="232">B</emphasis>
), ventromedial (
<emphasis id="B97C0E3FFFACFFCE35A9F879FFFB3FDF" bold="true" box="[1431,1445,1985,2002]" pageId="15" pageNumber="232">C</emphasis>
), and dorsal (
<emphasis id="B97C0E3FFFACFFCE30D8F860FAAA3FE4" bold="true" box="[230,244,2008,2025]" pageId="15" pageNumber="232">D</emphasis>
) views. Scale bar: 2 mm.
</paragraph>
</caption>
<caption id="DF7782A5FFB3FFD130BAF8A5FBB73F48" ID-DOI="http://doi.org/10.5281/zenodo.4721441" ID-Zenodo-Dep="4721441" httpUri="https://zenodo.org/record/4721441/files/figure.png" pageId="16" pageNumber="233" startId="16.[132,143,1821,1838]" targetBox="[204,1408,177,1791]" targetPageId="16">
<paragraph id="8BB7D22DFFB3FFD130BAF8A5FBB73F48" blockId="16.[132,1455,1821,1861]" pageId="16" pageNumber="233">
FIG. 11. — 
<taxonomicName id="4C08A9AEFFB3FFD130E3F8A5FBAD3F23" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[221,499,1821,1838]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="233" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB3FFD130E3F8A5FBAD3F23" box="[221,499,1821,1838]" italics="true" pageId="16" pageNumber="233">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB3FFD131C8F8A5F87B3F23" box="[502,549,1821,1838]" pageId="16" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
Virtual 3D models of the paratype right dentary NHMW 2019/0051/0002 in lateral (
<emphasis id="B97C0E3FFFB3FFD134F2F8A5FE873F23" bold="true" box="[1228,1241,1821,1838]" pageId="16" pageNumber="233">A</emphasis>
), medial (
<emphasis id="B97C0E3FFFB3FFD13516F8A5FF6B3F23" bold="true" box="[1320,1333,1821,1838]" pageId="16" pageNumber="233">B</emphasis>
), ventromedial (
<emphasis id="B97C0E3FFFB3FFD130B7F88CFAC93F48" bold="true" box="[137,151,1844,1861]" pageId="16" pageNumber="233">C</emphasis>
), and dorsal (
<emphasis id="B97C0E3FFFB3FFD13136F88CFB483F48" bold="true" box="[264,278,1844,1861]" pageId="16" pageNumber="233">D</emphasis>
) views. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFFB3FFD130BAF829F9593FE6" blockId="16.[132,775,1937,2028]" pageId="16" pageNumber="233">
If the tooth count was only 14, the coronoid reached the level of the 4th tooth position, the condition being identical to the 
<typeStatus id="54B36C8FFFB3FFD130C9F869FB023FE6" box="[247,348,2001,2027]" pageId="16" pageNumber="233" type="holotype">holotype</typeStatus>
. However, the difference between this
</paragraph>
<paragraph id="8BB7D22DFFB3FFD13311F829FE2F3FE1" blockId="16.[812,1457,1937,2028]" pageId="16" pageNumber="233">
specimen and the 
<typeStatus id="54B36C8FFFB3FFD133C3F829FE003FA6" box="[1021,1118,1937,1963]" pageId="16" pageNumber="233" type="holotype">holotype</typeStatus>
is the position of the alveolar foramen, being located at the level of the 9th tooth position here (counted from anterior).
</paragraph>
</subSection>
<caption id="DF7782A5FFB2FFD030BAFAD8FBE33D85" ID-DOI="http://doi.org/10.5281/zenodo.4720813" ID-Zenodo-Dep="4720813" httpUri="https://zenodo.org/record/4720813/files/figure.png" pageId="17" pageNumber="234" startId="17.[132,143,1376,1393]" targetBox="[132,1455,198,1349]" targetPageId="17">
<paragraph id="8BB7D22DFFB2FFD030BAFAD8FBE33D85" blockId="17.[132,1455,1376,1416]" pageId="17" pageNumber="234">
FIG. 12. — 
<taxonomicName id="4C08A9AEFFB2FFD030E1FAD8FBAB3D7C" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[223,501,1376,1393]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB2FFD030E1FAD8FBAB3D7C" box="[223,501,1376,1393]" italics="true" pageId="17" pageNumber="234">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB2FFD031C5FAD8F8753D7C" box="[507,555,1376,1393]" pageId="17" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
Close up photographs of posterior teeth of the paratype right dentary NHMW 2019/0051/0002 in medial (
<emphasis id="B97C0E3FFFB2FFD035A3FAD8FFF43D7C" bold="true" box="[1437,1450,1376,1393]" pageId="17" pageNumber="234">A</emphasis>
) and dorsal (
<emphasis id="B97C0E3FFFB2FFD030D8FACFFAAD3D85" bold="true" box="[230,243,1399,1416]" pageId="17" pageNumber="234">B</emphasis>
) view. Scale bar: 1 mm.
</paragraph>
</caption>
<subSection id="E287CBC6FFB2FFD030BAFA6CFAB33F01" pageId="17" pageNumber="234" type="multiple">
<paragraph id="8BB7D22DFFB2FFD030BAFA6CFAB73DE0" blockId="17.[132,233,1492,1517]" box="[132,233,1492,1517]" pageId="17" pageNumber="234">
<heading id="D0FF6541FFB2FFD030BAFA6CFAB73DE0" box="[132,233,1492,1517]" fontSize="11" level="2" pageId="17" pageNumber="234" reason="8">
<emphasis id="B97C0E3FFFB2FFD030BAFA6CFAB73DE0" box="[132,233,1492,1517]" italics="true" pageId="17" pageNumber="234">Dentition</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7D22DFFB2FFD030BAFA4BFAB33F01" blockId="17.[130,777,1523,1804]" pageId="17" pageNumber="234">
The dentition is pleurodont and strongly heterodont. The teeth are closely spaced. While the tooth size increases posteriorly (except for the last three or four teeth that are again small, reduced), the teeth in the anterior portion of the tooth row are small and slender. The teeth in the posterior region are robust, forming blunt cylinders (
<figureCitation id="1333CEA8FFB2FFD031D0F92AF8713EA1" box="[494,559,1682,1708]" captionStart="FIG" captionStartId="13.[132,143,1410,1427]" captionTargetBox="[137,1450,213,1368]" captionTargetId="figure-360@13.[699,1262,746,1279]" captionTargetPageId="13" captionText="FIG. 8. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the holotype left dentary NHMW 2019/0051/0001 in medial (A) and dorsal (B) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720800" httpUri="https://zenodo.org/record/4720800/files/figure.png" pageId="17" pageNumber="234">Figs 8</figureCitation>
; 
<figureCitation id="1333CEA8FFB2FFD03201F92AF8053EA1" box="[575,603,1682,1708]" captionStart="FIG" captionStartId="17.[132,143,1376,1393]" captionTargetBox="[132,1455,198,1349]" captionTargetId="figure-309@17.[132,1455,198,1349]" captionTargetPageId="17" captionText="FIG. 12. — Pseudeumeces kyrillomethodicus n. sp. Close up photographs of posterior teeth of the paratype right dentary NHMW 2019/0051/0002 in medial (A) and dorsal (B) view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720813" httpUri="https://zenodo.org/record/4720813/files/figure.png" pageId="17" pageNumber="234">12</figureCitation>
; 
<figureCitation id="1333CEA8FFB2FFD03255F92AF8D93EA1" box="[619,647,1682,1708]" captionStart="FIG" captionStartId="18.[813,823,1078,1095]" captionTargetBox="[819,1449,215,1036]" captionTargetId="figure-833@18.[813,1456,211,1050]" captionTargetPageId="18" captionText="FIG. 13. — Pseudeumeces kyrillomethodicus n. sp. Close up photograph of the most robust tooth of the paratype right dentary NHMW 2019/0051/0002 in medial view. Note the prominent striations. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720815" httpUri="https://zenodo.org/record/4720815/files/figure.png" pageId="17" pageNumber="234">13</figureCitation>
). The teeth (especially those in the posterior half of the tooth row) are anteroposteriorly compressed. The tooth crowns bear delicate striations.
</paragraph>
</subSection>
</subSubSection>
<subSubSection id="C31281A6FFB2FFD230BAF88AFE233D80" lastPageId="19" lastPageNumber="236" pageId="17" pageNumber="234" type="discussion">
<paragraph id="8BB7D22DFFB2FFD030BAF88AFAB63F47" blockId="17.[132,775,1842,2027]" box="[132,232,1842,1868]" pageId="17" pageNumber="234">REMARKS</paragraph>
<paragraph id="8BB7D22DFFB2FFD030BAF8E9FE3A3F41" blockId="17.[132,775,1842,2027]" lastBlockId="17.[811,1458,1491,2028]" pageId="17" pageNumber="234">
<taxonomicName id="4C08A9AEFFB2FFD030BAF8E9FBD63F66" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[132,392,1873,1899]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB2FFD030BAF8E9FBD63F66" box="[132,392,1873,1899]" italics="true" pageId="17" pageNumber="234">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
was originally established by 
<bibRefCitation id="EF99AFDCFFB2FFD032F6F8E9FA873F81" author="FILHOL H." pageId="17" pageNumber="234" pagination="1 - 340" refId="ref34973" refString="FILHOL H. 1877 a. - Recherches sur les Phosphorites du Quercy. Etude des fossiles qu'on y rencontre et specialement des mammiferes. Pt. II. Annales des Sciences geologiques 8: 1 - 340. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 432584 w / f 5. item" type="journal article" year="1877">Filhol (1877a)</bibRefCitation>
as a scincid of the extant genus 
<taxonomicName id="4C08A9AEFFB2FFD0321EF8C9FB493FA6" authority="Dumeril &amp; Bibron, 1839" authorityName="Dumeril &amp; Bibron" authorityYear="1839" class="Reptilia" family="Scincidae" genus="Plestiodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB2FFD0321EF8C9F8D63F86" box="[544,648,1905,1931]" italics="true" pageId="17" pageNumber="234">Plestiodon</emphasis>
<bibRefCitation id="EF99AFDCFFB2FFD032B1F8C9FB493FA6" author="DUMERIL A. M. C. &amp; BIBRON G." pageId="17" pageNumber="234" refId="ref34611" refString="DUMERIL A. M. C. &amp; BIBRON G. 1839. - Erpetologie generale ou histoire naturelle complete des reptiles. Tome cinquieme. Contenant l'histoire de quatre-vingt-trois genres et de deux cent sept especes des trois dernieres familles de l'ordre des sauriens, savoir: les lacertiens, les chalcidiens et les scincoIdiens. Librairie encyclopedique de Roret, Paris, 854 p. https: // doi. org / 10.5962 / bhl. title. 45973" type="book" year="1839">Duméril &amp; Bibron, 1839</bibRefCitation>
</taxonomicName>
. This taxonomic opinion was subsequently followed by others (e.g., 
<bibRefCitation id="EF99AFDCFFB2FFD0314AF80AF85A3FC6" author="NOPCSA F." box="[372,516,1969,1996]" pageId="17" pageNumber="234" pagination="33 - 62" refId="ref38326" refString="NOPCSA F. 1908. - Zur Kenntnis der fossilen Eidechsen. Beitrage zur Palaontologie und Geologie Osterreich-ungarns und des Orients 21: 33 - 62. https: // www. biodiversitylibrary. org / page / 15264653" type="journal article" year="1908">Nopcsa 1908</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB2FFD0322CF809F8D13FC6" author="KUHN O." box="[530,655,1969,1995]" pageId="17" pageNumber="234" refId="ref37553" refString="KUHN O. 1939. - Squamata: Lacertilia et Ophidia. Fossilium Catalogus. I: Animalia. Pars 86. Verlag Gustav Feller, Neubrandenburg, 89 p. [Lacertilia] + 3 pp [Ophidia]." type="book" year="1939">Kuhn 1939</bibRefCitation>
), although affinities with the glyptosaurine 
<taxonomicName id="4C08A9AEFFB2FFD031E2F869F80A3FE6" authorityName="Gervais" authorityYear="1848" box="[476,596,2001,2027]" class="Reptilia" family="Anguidae" genus="Placosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB2FFD031E2F869F80A3FE6" box="[476,596,2001,2027]" italics="true" pageId="17" pageNumber="234">Placosaurus</emphasis>
</taxonomicName>
also appeared in the literature (
<bibRefCitation id="EF99AFDCFFB2FFD033FCFA6BFE313DE0" author="LYDEKKER R." box="[962,1135,1491,1517]" pageId="17" pageNumber="234" pagination="110 - 113" refId="ref38098" refString="LYDEKKER R. 1888 b. - Notes on Tertiary Lacertilia and Ophidia. Geological Magazine 5: 110 - 113. https: // doi. org / 10.1017 / S 0016756800173480" type="journal article" year="1888">Lydekker 1888b</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB2FFD03445FA6BFF783DE0" author="LEENHARDT H." box="[1147,1318,1491,1517]" pageId="17" pageNumber="234" pagination="371 - 374" refId="ref37759" refString="LEENHARDT H. 1926. - Sur quelques sauriens de l'Eocene superieur de la France. Bulletin de la Societe geologique de France, serie 4, 26: 371 - 374." type="journal article" year="1926">Leenhardt 1926</bibRefCitation>
). 
<bibRefCitation id="EF99AFDCFFB2FFD03504FA6BF9293E00" author="HOFFSTETTER R." pageId="17" pageNumber="234" pagination="547 - 553" refId="ref36751" refString="HOFFSTETTER R. 1944. - Sur les Scincidae fossiles. I. Formes europeennes et nord-americaines. Bulletin du Museum national d'Histoire naturelle, 2 eme serie, 16: 547 - 553. https: // www. biodiversitylibrary. org / page / 54154543" type="journal article" year="1944">Hoffstetter (1944)</bibRefCitation>
was the first to realize its lacertid affinities and placed it into its own genus, 
<taxonomicName id="4C08A9AEFFB2FFD03413F9AAFEE23E21" authorityName="Hoffstetter" authorityYear="1944" box="[1069,1212,1554,1580]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB2FFD03413F9AAFEE23E21" box="[1069,1212,1554,1580]" italics="true" pageId="17" pageNumber="234">Pseudeumeces</emphasis>
</taxonomicName>
. So far, 
<taxonomicName id="4C08A9AEFFB2FFD0351FF9AAF9C13E41" baseAuthorityName="Filhol" baseAuthorityYear="1877" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB2FFD0351FF9AAF9C13E41" italics="true" pageId="17" pageNumber="234">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
represented the only currently recognized species of this genus, as other two species that have in the past been referred to 
<taxonomicName id="4C08A9AEFFB2FFD0339BF9CAFE6E3E81" authorityName="Hoffstetter" authorityYear="1944" box="[933,1072,1650,1676]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB2FFD0339BF9CAFE6E3E81" box="[933,1072,1650,1676]" italics="true" pageId="17" pageNumber="234">Pseudeumeces</emphasis>
</taxonomicName>
are now known to pertain to other genera or represent indeterminate lizards (i.e., 
<taxonomicName id="4C08A9AEFFB2FFD03512F92AFE653EC1" authority="Kuhn, 1940" authorityName="Kuhn" authorityYear="1940" class="Reptilia" family="Anguidae" genus="Glyptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="walbeckensis">
<emphasis id="B97C0E3FFFB2FFD03512F92AF9F03EC1" italics="true" pageId="17" pageNumber="234">Glyptosaurus walbeckensis</emphasis>
Kuhn, 1940
</taxonomicName>
, which was recombined into 
<taxonomicName id="4C08A9AEFFB2FFD03547F90BF9D03EE1" authorityName="Hoffstetter" authorityYear="1944" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB2FFD03547F90BF9D03EE1" italics="true" pageId="17" pageNumber="234">Pseudeumeces</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF99AFDCFFB2FFD03385F96AFE143EE1" author="ESTES R." box="[955,1098,1745,1772]" pageId="17" pageNumber="234" refId="ref34692" refString="ESTES R. 1983. - Sauria Terrestria, Amphisbaenia, in WELL- NHOFER P. (ed.), Encyclopedia of Paleoherpetology. Part 10 a. Gustav Fischer, Stuttgart, New York, 249 p." type="book" year="1983">Estes [1983]</bibRefCitation>
, and 
<taxonomicName id="4C08A9AEFFB2FFD034B1F96AF9343F01" authority="Auge, 1993" authorityName="Auge" authorityYear="1993" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="pouiti">
<emphasis id="B97C0E3FFFB2FFD034B1F96AFF363EE1" box="[1167,1384,1746,1772]" italics="true" pageId="17" pageNumber="234">Pseudeumeces pouiti</emphasis>
<bibRefCitation id="EF99AFDCFFB2FFD03551F96AF9343F01" author="AUGE M." pageId="17" pageNumber="234" pagination="69 - 74" refId="ref31944" refString="AUGE M. 1993. - Une nouvelle espece de Lacertide (Reptilia, Lacertilia) des Faluns miocenes de l'Anjou-Touraine. Bulletin de la Societe de Sciences naturelles de l'Ouest de la France 15: 69 - 74." type="journal article" year="1993">Augé, 1993</bibRefCitation>
</taxonomicName>
, which was subsequently recombined to its own genus 
<taxonomicName id="4C08A9AEFFB2FFD03312F8AAFF5D3F21" authority="Auge, Bailon &amp; Malfay, 2003" authorityName="Auge, Bailon &amp; Malfay" authorityYear="2003" box="[812,1283,1810,1836]" class="Reptilia" family="Lacertidae" genus="Ligerosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB2FFD03312F8AAF9F03F26" box="[812,942,1810,1835]" italics="true" pageId="17" pageNumber="234">Ligerosaurus</emphasis>
<bibRefCitation id="EF99AFDCFFB2FFD03389F8AAFF5D3F21" author="AUGE M. &amp; BAILON S. &amp; MALFAY J. - P." box="[951,1283,1810,1836]" pageId="17" pageNumber="234" pagination="289 - 295" refId="ref32537" refString="AUGE M., BAILON S. &amp; MALFAY J. - P. 2003. - Un nouveau genre de Lacertidae (Reptilia, Lacertilia) dans les faluns miocenes de l'Anjou- Touraine (Maine-et-Loire, France). Geodiversitas 25: 289 - 295." type="journal article" year="2003">Augé, Bailon &amp; Malfay, 2003</bibRefCitation>
</taxonomicName>
, as 
<taxonomicName id="4C08A9AEFFB2FFD03511F8AAF9343F46" class="Reptilia" family="Lacertidae" genus="Ligerosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="pouiti">
<emphasis id="B97C0E3FFFB2FFD03511F8AAF9343F46" italics="true" pageId="17" pageNumber="234">Ligerosaurus pouiti</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF99AFDCFFB2FFD033ACF88AFE0A3F41" author="AUGE M. &amp; BAILON S. &amp; MALFAY J. - P." box="[914,1108,1841,1868]" pageId="17" pageNumber="234" pagination="289 - 295" refId="ref32537" refString="AUGE M., BAILON S. &amp; MALFAY J. - P. 2003. - Un nouveau genre de Lacertidae (Reptilia, Lacertilia) dans les faluns miocenes de l'Anjou- Touraine (Maine-et-Loire, France). Geodiversitas 25: 289 - 295." type="journal article" year="2003">
Augé 
<emphasis id="B97C0E3FFFB2FFD033EEF88AFE5D3F46" box="[976,1027,1841,1867]" italics="true" pageId="17" pageNumber="234">et al.</emphasis>
[2003]
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB7D22DFFB2FFD3337DF8EAF8CE3B82" blockId="17.[811,1458,1491,2028]" lastBlockId="18.[130,777,215,2028]" lastPageId="18" lastPageNumber="235" pageId="17" pageNumber="234">
We consider that the dentition and overall morphology of our new species 
<taxonomicName id="4C08A9AEFFB2FFD033EBF8C9FF443F86" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[981,1306,1905,1931]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB2FFD033EBF8C9FF443F86" box="[981,1306,1905,1931]" italics="true" pageId="17" pageNumber="234">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB2FFD03521F8CAFF063F81" box="[1311,1368,1906,1932]" pageId="17" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
appears to bear a resemblance with 
<taxonomicName id="4C08A9AEFFB2FFD0347AF829FF1C3FA6" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[1092,1346,1937,1963]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB2FFD0347AF829FF1C3FA6" box="[1092,1346,1937,1963]" italics="true" pageId="17" pageNumber="234">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
. Similarly to the case of our new taxon, the type material of 
<taxonomicName id="4C08A9AEFFB2FFD0351BF809F9C13FE6" baseAuthorityName="Filhol" baseAuthorityYear="1877" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="234" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB2FFD0351BF809F9C13FE6" italics="true" pageId="17" pageNumber="234">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
also originates from an imprecise locality within the Phosphorites du Quercy (
<bibRefCitation id="EF99AFDCFFB1FFD33181FF6FF81938FC" author="FILHOL H." box="[447,583,215,241]" pageId="18" pageNumber="235" pagination="1 - 340" refId="ref34973" refString="FILHOL H. 1877 a. - Recherches sur les Phosphorites du Quercy. Etude des fossiles qu'on y rencontre et specialement des mammiferes. Pt. II. Annales des Sciences geologiques 8: 1 - 340. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 432584 w / f 5. item" type="journal article" year="1877">Filhol 1877a</bibRefCitation>
; for this taxon see also 
<bibRefCitation id="EF99AFDCFFB1FFD33088FF4FFB73391F" author="AUGE M." box="[182,301,247,274]" pageId="18" pageNumber="235" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé 2005</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD33102FF4FF878391C" author="AUGE M. &amp; HERVET S." box="[316,550,247,274]" pageId="18" pageNumber="235" pagination="191 - 201" refId="ref32254" refString="AUGE M. &amp; HERVET S. 2009. - Fossil lizards from the locality of Gannat (late Oligocene-early Miocene, France) and a revision of the genus Pseudeumeces (Squamata, Lacertidae). Palaeobiodiversity and Palaeoenvironments 89: 191 - 201. https: // doi. org / 10.1007 / s 12549 - 009 - 0009 - 1" type="journal article" year="2009">Augé &amp; Hervet 2009</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD3320BFF41FAE3393C" author="CERNANSKY A. &amp; AUGE M." pageId="18" pageNumber="235" pagination="11 - 19" refId="ref33347" refString="CERNANSKY A. &amp; AUGE M. 2012. - Additions to the lizard fauna (Squamata: Lacertilia) of the Upper Oligocene (MP 28) of Herrlingen 8, Southern Germany. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 264: 11 - 19." type="journal article" year="2012">Čerňanský &amp; Augé 2012</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD330F5FEA0FB9D393C" author="CERNANSKY A. &amp; KLEMBARA J. &amp; MULLER J." box="[203,451,279,308]" pageId="18" pageNumber="235" pagination="559 - 587" refId="ref33649" refString="CERNANSKY A., KLEMBARA J. &amp; MULLER J. 2016 a. - The new rare record of the late Oligocene lizards and amphisbaenians from Germany and its impact on our knowledge of the European terminal Palaeogene. Palaeobiodiversity and Palaeoenvironments 96: 559 - 587. https: // doi. org / 10.1007 / s 12549 - 015 - 0226 - 8" type="journal article" year="2016">
Čerňanský 
<emphasis id="B97C0E3FFFB1FFD3317BFEA0FB27393C" box="[325,377,279,305]" italics="true" pageId="18" pageNumber="235">et al.</emphasis>
2016a
</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD331EFFEAFF8DA393C" author="BOLET A. &amp; RAGE J. - C. &amp; CONRAD J. L." box="[465,644,279,305]" pageId="18" pageNumber="235" pagination="1315669" refId="ref33040" refString="BOLET A., RAGE J. - C. &amp; CONRAD J. L. 2017. - Rediscovery of the long-lost holotype of the lacertid lizard Pseudeumeces cadurcensis (Filhol, 1877). Journal of Vertebrate Paleontology 37: e 1315669. https: // doi. org / 10.1080 / 02724634.2017.1315669" type="journal article" year="2017">
Bolet 
<emphasis id="B97C0E3FFFB1FFD3322FFEA0F81B393C" box="[529,581,279,305]" italics="true" pageId="18" pageNumber="235">et al.</emphasis>
2017
</bibRefCitation>
). Nevertheless, the material described herein clearly exhibits several obvious differences relative to 
<taxonomicName id="4C08A9AEFFB1FFD331ECFEEFF886397C" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[466,728,343,369]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD331ECFEEFF886397C" box="[466,728,343,369]" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
(see Diagnosis above). Besides the features stated in the diagnosis, there is one additional difference – the dentary tooth number in 
<taxonomicName id="4C08A9AEFFB1FFD3309EFE0EFBFA39DD" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[160,420,438,464]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD3309EFE0EFBFA39DD" box="[160,420,438,464]" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
is usually 17 (note that the tooth number in that taxon can range from 15-17; the 
<typeStatus id="54B36C8FFFB1FFD332B2FE6EF8B539FD" box="[652,747,470,496]" pageId="18" pageNumber="235" type="holotype">holotype</typeStatus>
of the species [the left dentary AMNH FARB 241A] is incomplete but the preserved portion bears seven teeth and at least five other empty tooth positions), whereas the dentary tooth number of 
<taxonomicName id="4C08A9AEFFB1FFD3313AFDEEF80E3A7D" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[260,592,598,624]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB1FFD3313AFDEEF80E3A7D" box="[260,592,598,624]" italics="true" pageId="18" pageNumber="235">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB1FFD33266FDEEF8C93A7D" box="[600,663,598,624]" pageId="18" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
, based on material described here, is around 14-15. Although such small differences in tooth counts can be informative in some cases, it should be noted that the tooth number in lacertids (see e.g., 
<bibRefCitation id="EF99AFDCFFB1FFD330BCFD6FF85E3AFD" author="CERNANSKY A. &amp; SYROMYATNIKOVA E. V." box="[130,512,725,755]" pageId="18" pageNumber="235" pagination="0216191" refId="ref33461" refString="CERNANSKY A. &amp; SYROMYATNIKOVA E. V. 2019. - The first Miocene fossils of Lacerta cf. trilineata (Squamata, Lacertidae) with a comparative study of the main cranial osteological differences in green lizards and their relatives. PLoS ONE 14: e 0216191. https: // doi. org / 10.1371 / journal. pone. 0216191" type="journal article" year="2019">Čerňanský &amp; Syromyatnikova 2019</bibRefCitation>
), like virtually all lizards, should not be interpreted as absolute due to its variability. So, whether it seems that the tooth number of 
<taxonomicName id="4C08A9AEFFB1FFD33242FCADFAA83B42" baseAuthorityName="Filhol" baseAuthorityYear="1877" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD33242FCADFAA83B42" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
appears to be slightly higher than the new species, we cannot fully demonstrate it and we refrain from formally considering this feature as a diagnostic character.
</paragraph>
<paragraph id="8BB7D22DFFB1FFD330A5FC2DF85F3D03" blockId="18.[130,777,215,2028]" pageId="18" pageNumber="235">
Besides the resemblance with 
<taxonomicName id="4C08A9AEFFB1FFD331C8FC2DF9563BA2" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[502,776,917,943]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD331C8FC2DF9563BA2" box="[502,776,917,943]" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
discussed above, it should be noted also that the dentaries of 
<taxonomicName id="4C08A9AEFFB1FFD330BAFC6CFB943BE3" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[132,458,980,1006]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB1FFD330BAFC6CFB943BE3" box="[132,458,980,1006]" italics="true" pageId="18" pageNumber="235">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB1FFD331ECFC6DF8533BE2" box="[466,525,981,1007]" pageId="18" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
slightly resemble those of 
<taxonomicName id="4C08A9AEFFB1FFD330A1FC4DFB8F3C03" authority="Pomel, 1846" authorityName="Pomel" authorityYear="1846" box="[159,465,1012,1038]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB1FFD330A1FC4DFB1C3C03" box="[159,322,1013,1038]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus</emphasis>
<bibRefCitation id="EF99AFDCFFB1FFD33179FC4CFB8F3C03" author="POMEL A." box="[327,465,1012,1038]" pageId="18" pageNumber="235" pagination="353 - 373" refId="ref38504" refString="POMEL A. 1846. - Memoire pour servir a la geologie paleontologique des terrains tertiaires du departement de l'Allier. Bulletin de la Societe geologique de France, 2 eme serie, 3: 353 - 373. https: // www. biodiversitylibrary. org / page / 54525673" type="journal article" year="1846">Pomel, 1846</bibRefCitation>
</taxonomicName>
, in the following features (see 
<bibRefCitation id="EF99AFDCFFB1FFD330BCFBACFB513C23" author="MULLER J." box="[130,271,1044,1070]" pageId="18" pageNumber="235" pagination="253 - 266" refId="ref38270" refString="MULLER J. 2004. - Cranial osteology of Dracaenosaurus croizeti, a lacertid lizard from the Oligocene of France (Reptilia, Squamata). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 232: 253 - 266. https: // doi. org / 10.1127 / njgpa / 232 / 2004 / 253" type="journal article" year="2004">Müller 2004</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD3311FFBACFBC53C22" author="AUGE M." box="[289,411,1044,1071]" pageId="18" pageNumber="235" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé 2005</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD33195FBAEF8F33C23" author="CERNANSKY A. &amp; KLEMBARA J. &amp; MULLER J." box="[427,685,1044,1074]" pageId="18" pageNumber="235" pagination="559 - 587" refId="ref33649" refString="CERNANSKY A., KLEMBARA J. &amp; MULLER J. 2016 a. - The new rare record of the late Oligocene lizards and amphisbaenians from Germany and its impact on our knowledge of the European terminal Palaeogene. Palaeobiodiversity and Palaeoenvironments 96: 559 - 587. https: // doi. org / 10.1007 / s 12549 - 015 - 0226 - 8" type="journal article" year="2016">
Čerňanský 
<emphasis id="B97C0E3FFFB1FFD33214FBADF83E3C23" box="[554,608,1044,1070]" italics="true" pageId="18" pageNumber="235">et al.</emphasis>
2016 
<bibRefCitation id="EF99AFDCFFB1FFD3329FFBACF8F33C23" author="CERNANSKY A. &amp; BOLET A. &amp; MULLER J. &amp; RAGE J. - C. &amp; AUGE M. &amp; HERREL A." box="[673,685,1044,1070]" pageId="18" pageNumber="235" pagination="1384738" refId="ref33776" refString="CERNANSKY A., BOLET A., MULLER J., RAGE J. - C., AUGE M. &amp; HERREL A. 2017. - A new exceptionally preserved specimen of Dracaenosaurus (Squamata, Lacertidae) from the Oligocene of France as revealed by micro-computed tomography. Journal of Vertebrate Paleontology 37: e 1384738. https: // doi. org / 10.1 080 / 02724634.2017.1384738" type="journal article" year="2017">a</bibRefCitation>
</bibRefCitation>
, 2017): 1) dentary is a rather short, massive, and deep element; 2) the presence of a dorsally elevated posterior portion of the dentary; 3) the presence of the amblyodont dentition, where the posterior robust teeth are low and form blunt cylinders (this is more pronounced in 
<taxonomicName id="4C08A9AEFFB1FFD3315AFB0DF8593CC3" authorityName="Pomel" authorityYear="1846" box="[356,519,1205,1230]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB1FFD3315AFB0DF8593CC3" box="[356,519,1205,1230]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus</emphasis>
</taxonomicName>
); and 4) the presence of striations on the tooth crown (note that the last two features are not unique to these two forms).
</paragraph>
<caption id="DF7782A5FFB1FFD33313FB8EFF783C79" ID-DOI="http://doi.org/10.5281/zenodo.4720815" ID-Zenodo-Dep="4720815" httpUri="https://zenodo.org/record/4720815/files/figure.png" pageId="18" pageNumber="235" startId="18.[813,823,1078,1095]" targetBox="[819,1449,215,1036]" targetPageId="18">
<paragraph id="8BB7D22DFFB1FFD33313FB8EFF783C79" blockId="18.[813,1456,1078,1140]" pageId="18" pageNumber="235">
FIG. 13. — 
<taxonomicName id="4C08A9AEFFB1FFD333B4FB8EFEFB3C4A" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[906,1189,1078,1095]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB1FFD333B4FB8EFEFB3C4A" box="[906,1189,1078,1095]" italics="true" pageId="18" pageNumber="235">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB1FFD33493FB8EFE813C4A" box="[1197,1247,1078,1095]" pageId="18" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
Close up photograph of the most robust tooth of the paratype right dentary NHMW 2019/0051/0002 in medial view. Note the prominent striations. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFFB1FFD330A5FAACF8FB3FA6" blockId="18.[130,777,215,2028]" pageId="18" pageNumber="235">
However, there are some important differences between these two forms, where 
<taxonomicName id="4C08A9AEFFB1FFD331BFFA8BF8983D40" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[385,710,1331,1357]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB1FFD331BFFA8BF8983D40" box="[385,710,1331,1357]" italics="true" pageId="18" pageNumber="235">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB1FFD332F2FA8CF9593D43" box="[716,775,1332,1358]" pageId="18" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
can be differentiated from 
<taxonomicName id="4C08A9AEFFB1FFD33190FAECFAE03D83" authority="Gervais, 1848" authorityName="Gervais" authorityYear="1848" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="croizeti">
<emphasis id="B97C0E3FFFB1FFD33190FAECF8F93D60" box="[430,679,1364,1389]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus croizeti</emphasis>
<bibRefCitation id="EF99AFDCFFB1FFD3328EFAEBFAE03D83" author="GERVAIS P." pageId="18" pageNumber="235" refId="ref36227" refString="GERVAIS P. 1848 - 1852. - Zoologie et Paleontologie francaises (animaux vertebres): ou nouvelles recherches sur les animaux vivants et fossiles de la France. Arthus Bertrand, Paris, 271 p. https: // doi. org / 10.5962 / bhl. title. 39473" type="book" year="1848">Gervais, 1848</bibRefCitation>
</taxonomicName>
-1852, by the following combination of features (for 
<taxonomicName id="4C08A9AEFFB1FFD330B8FA2CFB7A3DA0" authorityName="Pomel" authorityYear="1846" box="[134,292,1428,1453]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB1FFD330B8FA2CFB7A3DA0" box="[134,292,1428,1453]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus</emphasis>
</taxonomicName>
, see 
<bibRefCitation id="EF99AFDCFFB1FFD3316CFA2BFB8B3DA0" author="MULLER J." box="[338,469,1427,1453]" pageId="18" pageNumber="235" pagination="253 - 266" refId="ref38270" refString="MULLER J. 2004. - Cranial osteology of Dracaenosaurus croizeti, a lacertid lizard from the Oligocene of France (Reptilia, Squamata). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 232: 253 - 266. https: // doi. org / 10.1127 / njgpa / 232 / 2004 / 253" type="journal article" year="2004">Müller 2004</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD331DDFA2BF80B3DA3" author="AUGE M." box="[483,597,1427,1454]" pageId="18" pageNumber="235" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé 2005</bibRefCitation>
; 
<bibRefCitation id="EF99AFDCFFB1FFD3325FFA2CFA963DC0" author="CERNANSKY A. &amp; KLEMBARA J. &amp; MULLER J." pageId="18" pageNumber="235" pagination="559 - 587" refId="ref33649" refString="CERNANSKY A., KLEMBARA J. &amp; MULLER J. 2016 a. - The new rare record of the late Oligocene lizards and amphisbaenians from Germany and its impact on our knowledge of the European terminal Palaeogene. Palaeobiodiversity and Palaeoenvironments 96: 559 - 587. https: // doi. org / 10.1007 / s 12549 - 015 - 0226 - 8" type="journal article" year="2016">
Čerňanský 
<emphasis id="B97C0E3FFFB1FFD332E9FA2CF9563DA0" box="[727,776,1427,1453]" italics="true" pageId="18" pageNumber="235">et al.</emphasis>
2016 
<bibRefCitation id="EF99AFDCFFB1FFD33082FA0BFA963DC0" author="CERNANSKY A. &amp; BOLET A. &amp; MULLER J. &amp; RAGE J. - C. &amp; AUGE M. &amp; HERREL A." box="[188,200,1459,1485]" pageId="18" pageNumber="235" pagination="1384738" refId="ref33776" refString="CERNANSKY A., BOLET A., MULLER J., RAGE J. - C., AUGE M. &amp; HERREL A. 2017. - A new exceptionally preserved specimen of Dracaenosaurus (Squamata, Lacertidae) from the Oligocene of France as revealed by micro-computed tomography. Journal of Vertebrate Paleontology 37: e 1384738. https: // doi. org / 10.1 080 / 02724634.2017.1384738" type="journal article" year="2017">a</bibRefCitation>
</bibRefCitation>
, 2017): 1) the dentary tooth number is around 14-15 rather than seven or eight; 2) the tooth size increases posteriorly, however, the largest tooth is the 4th or 5th one (counted from posterior), whereas the further posterior teeth decrease in size (to the contrary, the largest tooth in 
<taxonomicName id="4C08A9AEFFB1FFD33237F98BF8283E41" authorityName="Gervais" authorityYear="1848" box="[521,630,1587,1612]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="croizeti">
<emphasis id="B97C0E3FFFB1FFD33237F98BF8283E41" box="[521,630,1587,1612]" italics="true" pageId="18" pageNumber="235">D. croizeti</emphasis>
</taxonomicName>
is usually the posteriormost one or sometimes the penultimate one); 3) the alveolar foramen, although its position can vary, is located further anteriorly (at the level between the 4th and 5th tooth positions in the 
<typeStatus id="54B36C8FFFB1FFD33114F90AFBD43EC1" box="[298,394,1714,1740]" pageId="18" pageNumber="235" type="holotype">holotype</typeStatus>
; counted from posterior) rather than at the level of the posteriormost tooth (or between last and penultimate tooth positions); 4) the facet for the anterolateral process of coronoid reaches around the level of the 4 
<superScript id="7C7D7F65FFB1FFD3328BF8ABF89B3F29" attach="left" box="[693,709,1811,1828]" fontSize="7" pageId="18" pageNumber="235">th</superScript>
tooth position (counted from posterior) rather than terminating posterior to the tooth row; and 5) teeth (especially those in the posterior half of the tooth row) are anteroposteriorly compressed rather than mediolateraly compressed.
</paragraph>
<paragraph id="8BB7D22DFFB1FFD330A5F80AFF3A3D23" blockId="18.[130,777,215,2028]" lastBlockId="18.[810,1457,1204,2028]" pageId="18" pageNumber="235">
Note that for 
<taxonomicName id="4C08A9AEFFB1FFD33103F80AFBB83FC6" authorityName="Pomel" authorityYear="1846" box="[317,486,1970,1995]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB1FFD33103F80AFBB83FC6" box="[317,486,1970,1995]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus</emphasis>
</taxonomicName>
we follow recent workers and treat 
<taxonomicName id="4C08A9AEFFB1FFD330D3F86AF8D13FE6" authority="(Filhol, 1882)" baseAuthorityName="Filhol" baseAuthorityYear="1882" box="[237,655,2001,2027]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="sauvagei">
<emphasis id="B97C0E3FFFB1FFD330D3F86AFBB13FE6" box="[237,495,2002,2027]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus sauvagei</emphasis>
(Filhol, 1882)
</taxonomicName>
as a junior synonym of the type species 
<taxonomicName id="4C08A9AEFFB1FFD33467FB0DFF103CC3" authorityName="Gervais" authorityYear="1848" box="[1113,1358,1205,1230]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="croizeti">
<emphasis id="B97C0E3FFFB1FFD33467FB0DFF103CC3" box="[1113,1358,1205,1230]" italics="true" pageId="18" pageNumber="235">Dracaenosaurus croizeti</emphasis>
</taxonomicName>
(see e.g., 
<bibRefCitation id="EF99AFDCFFB1FFD33315FB6CF9FE3CE3" author="AUGE M." box="[811,928,1235,1262]" pageId="18" pageNumber="235" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé 2005</bibRefCitation>
). In any case, the 
<typeStatus id="54B36C8FFFB1FFD3345EFB6CFEE13CE3" box="[1120,1215,1236,1262]" pageId="18" pageNumber="235" type="holotype">holotype</typeStatus>
dentary of 
<taxonomicName id="4C08A9AEFFB1FFD33508FB6CFFF03CE0" baseAuthorityName="Filhol" baseAuthorityYear="1882" box="[1334,1454,1236,1261]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="sauvagei">
<emphasis id="B97C0E3FFFB1FFD33508FB6CFFF03CE0" box="[1334,1454,1236,1261]" italics="true" pageId="18" pageNumber="235">D. sauvagei</emphasis>
</taxonomicName>
is different from that of 
<taxonomicName id="4C08A9AEFFB1FFD33414FB4BFF313D00" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[1066,1391,1267,1293]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB1FFD33414FB4BFF313D00" box="[1066,1391,1267,1293]" italics="true" pageId="18" pageNumber="235">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB1FFD3354BFB4CFFF03D03" box="[1397,1454,1268,1294]" pageId="18" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
and its tooth count is within the range of 
<taxonomicName id="4C08A9AEFFB1FFD334CEFAACFF033D20" authorityName="Gervais" authorityYear="1848" box="[1264,1373,1300,1325]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="croizeti">
<emphasis id="B97C0E3FFFB1FFD334CEFAACFF033D20" box="[1264,1373,1300,1325]" italics="true" pageId="18" pageNumber="235">D. croizeti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB7D22DFFB1FFD3337DFA8CFFCC3E80" blockId="18.[810,1457,1204,2028]" pageId="18" pageNumber="235">
In regards to our material, the specimen MNHN.F.QU17169 (see 
<bibRefCitation id="EF99AFDCFFB1FFD33362FAEBFE1D3D60" author="AUGE M. &amp; HERVET S." box="[860,1091,1363,1390]" pageId="18" pageNumber="235" pagination="191 - 201" refId="ref32254" refString="AUGE M. &amp; HERVET S. 2009. - Fossil lizards from the locality of Gannat (late Oligocene-early Miocene, France) and a revision of the genus Pseudeumeces (Squamata, Lacertidae). Palaeobiodiversity and Palaeoenvironments 89: 191 - 201. https: // doi. org / 10.1007 / s 12549 - 009 - 0009 - 1" type="journal article" year="2009">Augé &amp; Hervet 2009</bibRefCitation>
: fig. 1) deserves a comment. This specimen, which has been allocated to 
<taxonomicName id="4C08A9AEFFB1FFD33482FACBF91D3DA0" baseAuthorityName="Filhol" baseAuthorityYear="1877" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD33482FACBF91D3DA0" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF99AFDCFFB1FFD33352FA2BF9AA3DA3" author="AUGE M." box="[876,1012,1427,1454]" pageId="18" pageNumber="235" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé (2005)</bibRefCitation>
and 
<bibRefCitation id="EF99AFDCFFB1FFD33412FA2BFF7B3DA3" author="AUGE M. &amp; HERVET S." box="[1068,1317,1427,1454]" pageId="18" pageNumber="235" pagination="191 - 201" refId="ref32254" refString="AUGE M. &amp; HERVET S. 2009. - Fossil lizards from the locality of Gannat (late Oligocene-early Miocene, France) and a revision of the genus Pseudeumeces (Squamata, Lacertidae). Palaeobiodiversity and Palaeoenvironments 89: 191 - 201. https: // doi. org / 10.1007 / s 12549 - 009 - 0009 - 1" type="journal article" year="2009">Augé &amp; Hervet (2009)</bibRefCitation>
, has only 12 tooth positions instead of usual 16-17. Moreover, the dentary of this specimen appears to be robust rather than narrow. This would point to a huge level of variability. However, in MNHN.F.QU17169, only the last posterior tooth is reduced and the coronoid reaches the level of this last posterior tooth position on both sides as it is typical, indeed, for 
<taxonomicName id="4C08A9AEFFB1FFD3351AF9EAF9C13E81" baseAuthorityName="Filhol" baseAuthorityYear="1877" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD3351AF9EAF9C13E81" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
(in contrast to our material described herein).
</paragraph>
<paragraph id="8BB7D22DFFB1FFD2337DF92BFB223DE0" blockId="18.[810,1457,1204,2028]" lastBlockId="19.[131,777,1204,2028]" lastPageId="19" lastPageNumber="236" pageId="18" pageNumber="235">
In general, it can be expected stratigraphically that not identical but slightly similar forms of a lineage (the exact age of our material is unfortunately unknown, as is also that of the 
<typeStatus id="54B36C8FFFB1FFD3339BF94AF98A3F01" box="[933,980,1778,1804]" pageId="18" pageNumber="235">type</typeStatus>
material of 
<taxonomicName id="4C08A9AEFFB1FFD3346FF94AFF043F01" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[1105,1370,1778,1804]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="235" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB1FFD3346FF94AFF043F01" box="[1105,1370,1778,1804]" italics="true" pageId="18" pageNumber="235">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
) would exhibit a higher degree of morphological disparity reflecting the evolution through time than specimens collected from a single stratigraphic level. In those cases, it is of course difficult to add an exact border between such forms to distinguish taxa as units for science. However, we are convinced that all the above mentioned differences allow to erect a new taxon based on our 
<typeStatus id="54B36C8FFFB1FFD33488F86AFEB83FE1" box="[1206,1254,2002,2028]" pageId="18" pageNumber="235">type</typeStatus>
material. Even in extant herpetofaunas, lacertids include several morphologically cryptic species for which determination based on morphology can be even more difficult than the situation discussed here. Therefore, we consider the obvious differences in our material relative to the previously described forms as sufficient. Due to a high level of similarities of the currently known limited material with 
<taxonomicName id="4C08A9AEFFB0FFD23246FACBFAA33DA0" baseAuthorityName="Filhol" baseAuthorityYear="1877" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="species" species="cadurcensis">
<emphasis id="B97C0E3FFFB0FFD23246FACBFAA33DA0" italics="true" pageId="19" pageNumber="236">Pseudeumeces cadurcensis</emphasis>
</taxonomicName>
, we decided to allocate this new taxon 
<taxonomicName id="4C08A9AEFFB0FFD232F0FA2CFBF83DC0" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB0FFD232F0FA2CFBF83DC0" italics="true" pageId="19" pageNumber="236">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB0FFD23192FA0BFBB93DC0" box="[428,487,1459,1485]" pageId="19" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
to the same genus instead of erecting a new one.
</paragraph>
<caption id="DF7782A5FFB0FFD230BAFBE2F8EB3C8F" ID-DOI="http://doi.org/10.5281/zenodo.4720819" ID-Zenodo-Dep="4720819" httpUri="https://zenodo.org/record/4720819/files/figure.png" pageId="19" pageNumber="236" startId="19.[132,143,1114,1131]" targetBox="[141,1447,215,1070]" targetPageId="19">
<paragraph id="8BB7D22DFFB0FFD230BAFBE2F8EB3C8F" blockId="19.[132,1455,1114,1154]" pageId="19" pageNumber="236">
FIG. 14. — 
<taxonomicName id="4C08A9AEFFB0FFD230E1FBE2FBA83C66" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[223,502,1114,1131]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB0FFD230E1FBE2FBA83C66" box="[223,502,1114,1131]" italics="true" pageId="19" pageNumber="236">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB0FFD231C5FBE2F8753C66" box="[507,555,1114,1131]" pageId="19" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
Virtual slices of the paratype right dentary NHMW 2019/0051/0002: horizontal (
<emphasis id="B97C0E3FFFB0FFD234FCFBE2FE913C66" bold="true" box="[1218,1231,1114,1131]" pageId="19" pageNumber="236">A</emphasis>
) and sagittal (
<emphasis id="B97C0E3FFFB0FFD2357BFBE2FF0C3C66" bold="true" box="[1349,1362,1114,1131]" pageId="19" pageNumber="236">B</emphasis>
). Note that the posterior section of the tooth row is missing. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFFB0FFD230A5FA4BFB6D3F86" blockId="19.[131,777,1204,2028]" pageId="19" pageNumber="236">
It should be further noted that the extinct genera 
<taxonomicName id="4C08A9AEFFB0FFD23297FA4BFAE53E21" authorityName="Hoffstetter" authorityYear="1944" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB0FFD23297FA4BFAE53E21" italics="true" pageId="19" pageNumber="236">Pseudeumeces</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C08A9AEFFB0FFD230F6F9ABFB363E21" authorityName="Pomel" authorityYear="1846" box="[200,360,1555,1580]" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB0FFD230F6F9ABFB363E21" box="[200,360,1555,1580]" italics="true" pageId="19" pageNumber="236">Dracaenosaurus</emphasis>
</taxonomicName>
, and 
<emphasis id="B97C0E3FFFB0FFD231A1F9AAFBA53E21" box="[415,507,1554,1580]" italics="true" pageId="19" pageNumber="236">Janosikia</emphasis>
Čerňanský, Klembara, &amp; Smith, 2016, have been recovered as sister taxa to the extant 
<taxonomicName id="4C08A9AEFFB0FFD230EEF9EAFBB43E61" authority="Boulenger, 1916" authorityName="Boulenger" authorityYear="1916" box="[208,490,1618,1644]" class="Reptilia" family="Lacertidae" genus="Gallotia" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB0FFD230EEF9EAFB763E61" box="[208,296,1618,1644]" italics="true" pageId="19" pageNumber="236">Gallotia</emphasis>
<bibRefCitation id="EF99AFDCFFB0FFD23110F9EAFBB43E61" author="BOULENGER G. A." box="[302,490,1618,1644]" pageId="19" pageNumber="236" pagination="1 - 104" refId="ref33171" refString="BOULENGER G. A. 1916. - On the lizards allied to Lacerta muralis with an account of Lacerta agilis and L. parva. Transactions of the Zoological Society of London 21: 1 - 104. https: // doi. org / 10.1111 / j. 1096 - 3642.1916. tb 00480. x" type="journal article" year="1916">Boulenger, 1916</bibRefCitation>
</taxonomicName>
, from the 
<collectingRegion id="49CC1CCFFFB0FFD23264F9EAF95C3E61" box="[602,770,1618,1644]" country="Spain" name="Canarias" pageId="19" pageNumber="236">Canary Islands</collectingRegion>
, and all these taxa together with 
<taxonomicName id="4C08A9AEFFB0FFD231D3F9CAFA9C3EA1" authority="Fitzinger, 1826" authorityName="Fitzinger" authorityYear="1826" class="Reptilia" family="Lacertidae" genus="Psammodromus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB0FFD231D3F9CAF8CB3E81" box="[493,661,1650,1676]" italics="true" pageId="19" pageNumber="236">Psammodromus</emphasis>
<bibRefCitation id="EF99AFDCFFB0FFD232A3F9CAFA9C3EA1" author="FITZINGER L. J. F. J." pageId="19" pageNumber="236" refId="ref35259" refString="FITZINGER L. J. F. J. 1826. - Neue Classification der Reptilien nach ihren Naturlichen Verwandtschaften. Nebst einer Verwandtschafts- Tafel und einem Verzeichnisse der Reptilien-Sammlung des k. k. zoologischen Museums zu Wien. J. G. Huebner, Wien, 66 p. https: // doi. org / 10.5962 / bhl. title. 4683" type="book" year="1826">Fitzinger, 1826</bibRefCitation>
</taxonomicName>
, form the clade 
<taxonomicName id="4C08A9AEFFB0FFD2314FF92AFBAC3EA1" authorityName="Cano, Baez, Lopez-Jurado &amp; Ortega" authorityYear="1984" box="[369,498,1682,1708]" class="Reptilia" family="Lacertidae" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="subFamily" subFamily="Gallotiinae">Gallotiinae</taxonomicName>
. This was firstly observed by 
<bibRefCitation id="EF99AFDCFFB0FFD23099F90BFBE93EC1" author="CERNANSKY A. &amp; KLEMBARA J. &amp; SMITH K. T." box="[167,439,1714,1743]" pageId="19" pageNumber="236" pagination="861 - 877" refId="ref33715" refString="CERNANSKY A., KLEMBARA J. &amp; SMITH K. T. 2016 b. - Fossil lizard from central Europe resolves the origin of large body size and herbivory in giant Canary Island lacertids. Zoological Journal of the Linnean Society 176: 861 - 877. https: // doi. org / 10.1111 / zoj. 12340" type="journal article" year="2016">
Čerňanský 
<emphasis id="B97C0E3FFFB0FFD23119F90BFB033EC1" box="[295,349,1714,1740]" italics="true" pageId="19" pageNumber="236">et al.</emphasis>
(2016b
</bibRefCitation>
, 
<bibRefCitation id="EF99AFDCFFB0FFD231F8F90AF85E3EC1" author="CERNANSKY A. &amp; BOLET A. &amp; MULLER J. &amp; RAGE J. - C. &amp; AUGE M. &amp; HERREL A." box="[454,512,1714,1740]" pageId="19" pageNumber="236" pagination="1384738" refId="ref33776" refString="CERNANSKY A., BOLET A., MULLER J., RAGE J. - C., AUGE M. &amp; HERREL A. 2017. - A new exceptionally preserved specimen of Dracaenosaurus (Squamata, Lacertidae) from the Oligocene of France as revealed by micro-computed tomography. Journal of Vertebrate Paleontology 37: e 1384738. https: // doi. org / 10.1 080 / 02724634.2017.1384738" type="journal article" year="2017">2017</bibRefCitation>
), who also applied this revelation on the principles of the island rules, and later supported by Garcia-Porta 
<emphasis id="B97C0E3FFFB0FFD23187F94BFBB13F01" box="[441,495,1778,1804]" italics="true" pageId="19" pageNumber="236">et al.</emphasis>
(2019) by their analyses based on a supermatrix relying on novel phylogenomic datasets. Therefore, we allocate 
<taxonomicName id="4C08A9AEFFB0FFD231C6F889FA823F66" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB0FFD231C6F889FA823F66" italics="true" pageId="19" pageNumber="236">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB0FFD230DBF8EAFB7B3F61" box="[229,293,1874,1900]" pageId="19" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
, as a member of 
<taxonomicName id="4C08A9AEFFB0FFD231D4F8E9F8253F66" authorityName="Hoffstetter" authorityYear="1944" box="[490,635,1873,1899]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFFB0FFD231D4F8E9F8253F66" box="[490,635,1873,1899]" italics="true" pageId="19" pageNumber="236">Pseudeumeces</emphasis>
</taxonomicName>
, to the 
<taxonomicName id="4C08A9AEFFB0FFD232EBF8E9FA873F86" authorityName="Cano, Baez, Lopez-Jurado &amp; Ortega" authorityYear="1984" class="Reptilia" family="Lacertidae" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="subFamily" subFamily="Gallotiinae">Gallotiinae</taxonomicName>
as well.
</paragraph>
<paragraph id="8BB7D22DFFB0FFD230A5F829FE233D80" blockId="19.[131,777,1204,2028]" lastBlockId="19.[811,1455,1204,1422]" pageId="19" pageNumber="236">
Recently described fossil material from the early Eocene (MP 8-9) French locality of Mutigny (Paris Basin) indicates that not only stem but also morphologically mod- ern-like (potentially crown or close to crown) lacertids were present on the European continent already in the early Eocene (
<bibRefCitation id="EF99AFDCFFB0FFD233F2FB4DFE9C3D03" author="CERNANSKY A. &amp; AUGE M. &amp; PHELIZON A." box="[972,1218,1267,1297]" pageId="19" pageNumber="236" pagination="1768539" refId="ref33980" refString="CERNANSKY A., AUGE M. &amp; PHELIZON A. 2020. - Dawn of lacertids (Squamata, Lacertidae): new finds from the Upper Paleocene and the Lower Eocene. Journal of Vertebrate Paleontology 40: e 1768539. https: // doi. org / 10.1080 / 02724634. 2020.1768539" type="journal article" year="2020">
Čerňanský 
<emphasis id="B97C0E3FFFB0FFD23475FB4CFEDF3D00" box="[1099,1153,1267,1293]" italics="true" pageId="19" pageNumber="236">et al.</emphasis>
2020
</bibRefCitation>
). And later, lacertids were a rather diverse group during the Paleogene. The new taxon described herein, 
<taxonomicName id="4C08A9AEFFB0FFD23475FA8BFFF13D40" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[1099,1455,1331,1357]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="236" phylum="Chordata" rank="species" species="kyrillomethodicus" status="sp. nov.">
<emphasis id="B97C0E3FFFB0FFD23475FA8BFFF13D40" box="[1099,1455,1331,1357]" italics="true" pageId="19" pageNumber="236">Pseudeumeces kyrillomethodicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FFB0FFD23313FAECF9353D63" box="[813,875,1364,1390]" pageId="19" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
, fully supports this high diversity and abundance of European Paleogene lacertids.
</paragraph>
</subSubSection>
</treatment>
</document>