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<document id="7C7D3E5B19C113CCE9256A1F2028D52F" ID-CLB-Dataset="7175" ID-DOI="10.5252/geodiversitas2021v43a9" ID-GBIF-Dataset="04426a8e-c179-4af0-8e23-d72908e8b4cc" ID-ISSN="1638-9395" ID-Zenodo-Dep="4720776" ID-ZooBank="urn:lsid:zoobank.org:pub:11D0D852-39D7-449C-9EB3-C3D804114556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445770811" checkinUser="felipe" docAuthor="Georgalis, Georgios L., Čerňanský, Andrej &amp; Klembara, Jozef" docDate="2021" docId="03A1633BFF8AFFF53154F8E0F9563D80" docLanguage="en" docName="Geodiversitas.43.9.219-293.pdf" docOrigin="Geodiversitas 43 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Palaeovaranus sp." docType="treatment" docVersion="6" lastPageNumber="269" masterDocId="FF981B43FFA3FFC1303EFFB8FA5E380D" masterDocTitle="Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material" masterLastPageNumber="293" masterPageNumber="219" pageNumber="258" updateTime="1698937615995" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods id="1ECFCDD095F374E72B6B6591AFD7D4B1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CF356BE937654394FE58C0385FEB5504">Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material</mods:title>
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<mods:namePart id="4DDD1F4F90D6046A026CEFA569B9FD19">Georgalis, Georgios L.</mods:namePart>
<mods:affiliation id="B9CC69F98A45B87D277A981656834A7F">Palaeontological Institute and Museum, University of Zurich, Karl Schmid-Strasse 4, 8006 Zurich (Switzerland) and Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia) and Department of Earth Sciences, University of Torino, Via Valperga Caluso 35, 10125 Turin (Italy)</mods:affiliation>
<mods:nameIdentifier id="89A8826B1E15DBFE159348EE3356069E" type="email">georgios.georgalis@pim.uzh.ch</mods:nameIdentifier>
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<mods:namePart id="BBEB724979D7E442D420564666283869">Čerňanský, Andrej</mods:namePart>
<mods:affiliation id="6127E4DE76FECB843BC42B50BAA3A299">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:namePart id="CC36FCC82433F4E47FF092E4A4F012BE">Klembara, Jozef</mods:namePart>
<mods:affiliation id="2628DF140CB9BACA0188909A1D08CD91">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:title id="34366C000C1D0C8B3AA4F2F6259B78F4">Geodiversitas</mods:title>
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<mods:date id="311F23ACA2B15AD0E70EF1DC0825E2C1">2021</mods:date>
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<mods:number id="5221B06C27B1106A836DF6DEF5E83A46">2021-04-22</mods:number>
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<mods:number id="3D9D44F666A42C2037920965080D438A">43</mods:number>
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<treatment id="03A1633BFF8AFFF53154F8E0F9563D80" ID-DOI="http://doi.org/10.5281/zenodo.4721477" ID-GBIF-Taxon="181601112" ID-Zenodo-Dep="4721477" LSID="urn:lsid:plazi:treatment:03A1633BFF8AFFF53154F8E0F9563D80" httpUri="http://treatment.plazi.org/id/03A1633BFF8AFFF53154F8E0F9563D80" lastPageId="52" lastPageNumber="269" pageId="41" pageNumber="258">
<subSubSection id="C31281A6FF8AFFE83154F8E0F87C3F7E" box="[362,546,1880,1907]" pageId="41" pageNumber="258" type="nomenclature">
<paragraph id="8BB7D22DFF8AFFE83154F8E0F87C3F7E" blockId="41.[362,546,1880,1907]" box="[362,546,1880,1907]" pageId="41" pageNumber="258">
<heading id="D0FF6541FF8AFFE83154F8E0F87C3F7E" box="[362,546,1880,1907]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="258" reason="2">
<taxonomicName id="4C08A9AEFF8AFFE83154F8E0F87C3F7E" box="[362,546,1880,1907]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="sp.">
<emphasis id="B97C0E3FFF8AFFE83154F8E0FBA33F7F" box="[362,509,1880,1906]" italics="true" pageId="41" pageNumber="258">Palaeovaranus</emphasis>
sp.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF8AFFE831BDF8C0F8563F9E" box="[387,520,1912,1939]" pageId="41" pageNumber="258" type="description">
<paragraph id="8BB7D22DFF8AFFE831BDF8C0F8563F9E" blockId="41.[387,520,1912,1939]" box="[387,520,1912,1939]" pageId="41" pageNumber="258">
(
<figureCitation id="1333CEA8FF8AFFE831B2F8C0F85E3F9E" box="[396,512,1912,1939]" captionStart-0="FIG" captionStart-1="FIG" captionStart-10="FIG" captionStart-11="FIG" captionStart-12="FIG" captionStart-13="FIG" captionStart-14="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStart-8="FIG" captionStart-9="FIG" captionStartId-0="49.[131,142,1429,1446]" captionStartId-1="50.[132,143,1985,2002]" captionStartId-10="59.[145,156,838,855]" captionStartId-11="60.[132,143,1441,1458]" captionStartId-12="61.[132,143,1598,1615]" captionStartId-13="62.[132,143,1488,1505]" captionStartId-14="63.[132,143,1940,1957]" captionStartId-2="51.[132,143,925,942]" captionStartId-3="52.[132,143,991,1008]" captionStartId-4="53.[132,143,1541,1558]" captionStartId-5="54.[132,143,1446,1463]" captionStartId-6="55.[132,143,1481,1498]" captionStartId-7="56.[132,143,1040,1057]" captionStartId-8="57.[132,143,1262,1279]" captionStartId-9="58.[132,143,1894,1911]" captionTargetBox-0="[139,1448,214,1385]" captionTargetBox-1="[147,1438,219,1839]" captionTargetBox-10="[132,1455,211,801]" captionTargetBox-11="[140,1445,215,1398]" captionTargetBox-12="[132,1455,211,1558]" captionTargetBox-13="[136,1447,217,1443]" captionTargetBox-14="[135,1452,217,1871]" captionTargetBox-2="[132,1455,203,895]" captionTargetBox-3="[132,1456,182,970]" captionTargetBox-4="[138,1449,216,1479]" captionTargetBox-5="[142,1448,215,1407]" captionTargetBox-6="[136,1452,214,1439]" captionTargetBox-7="[139,1449,216,996]" captionTargetBox-8="[132,1455,207,1225]" captionTargetBox-9="[136,1450,216,1841]" captionTargetId-0="figure-335@49.[132,1455,205,1490]" captionTargetId-1="figure-15@50.[138,1449,217,1845]" captionTargetId-10="figure-693@59.[132,1455,211,801]" captionTargetId-11="figure-317@60.[132,1455,205,1403]" captionTargetId-12="figure-210@61.[132,1455,211,1558]" captionTargetId-13="figure-279@62.[132,1455,215,1448]" captionTargetId-14="figure-16@63.[132,1455,215,1878]" captionTargetId-2="figure-582@51.[132,1455,203,895]" captionTargetId-3="figure-475@52.[132,1455,211,892]" captionTargetId-4="figure-270@53.[132,1455,207,1492]" captionTargetId-5="figure-235@54.[132,1455,207,1356]" captionTargetId-6="figure-315@55.[132,1455,209,1447]" captionTargetId-7="figure-573@56.[132,1455,201,1002]" captionTargetId-8="figure-386@57.[132,1455,207,1225]" captionTargetId-9="figure-15@58.[132,1455,211,1845]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionTargetPageId-10="59" captionTargetPageId-11="60" captionTargetPageId-12="61" captionTargetPageId-13="62" captionTargetPageId-14="63" captionTargetPageId-2="51" captionTargetPageId-3="52" captionTargetPageId-4="53" captionTargetPageId-5="54" captionTargetPageId-6="55" captionTargetPageId-7="56" captionTargetPageId-8="57" captionTargetPageId-9="58" captionText-0="FIG. 44. — Palaeovaranus sp. A-C, photographs of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), and dorsal (C) views; D, close up of the posteriormost preserved tooth of the same specimen. Scale bars: A-C, 5 mm; D, 1 mm." captionText-1="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." captionText-10="FIG. 54. — Palaeovaranus sp. Photographs of two articulated sacral vertebrae NHMW 2019/0058/0028 in right (A) and left lateral (B) views. Scale bar: 2 mm." captionText-11="FIG. 55. — Palaeovaranus sp. Virtual 3D models of two articulated sacral vertebrae NHMW 2019/0058/0028 in anterior (A), posterior (B), anterolateral (C), dorsal (D), ventral (E), ventrolateral (F), right lateral (G), left lateral (H), and posteroventral (I) views. Scale bar: 5 mm." captionText-12="FIG. 56. — Palaeovaranus sp. A-F, photographs of sacral vertebra NHMW 2019/0058/0029 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views; G-K, photographs of sacral vertebra NHMW 2019/0058/0047 in anterior (G), posterior (H), dorsal (I), ventral (J), and left lateral (K) views. Scale bars: 2 mm." captionText-13="FIG. 57. — Palaeovaranus sp. Photographs of caudal vertebra NHMW 2019/0058/0035 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F), views. Scale bar: 2 mm." captionText-14="FIG. 58. — Palaeovaranus sp. A-E, photographs of caudal vertebra NHMW 2019/0058/0036 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views;F-J, photographs of caudal vertebra NHMW 2019/0058/0037 in anterior (F), posterior (G), dorsal (H), ventral (I), and right lateral (J) views;K-O, photographs of caudal vertebra NHMW 2019/0058/0048 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of caudal vertebra NHMW 2019/0058/0049 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views. Scale bar: 2 mm." captionText-2="FIG. 46. — Palaeovaranus sp. Photographs of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and posterior (D) views. Scale bar: 5 mm." captionText-3="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." captionText-4="FIG. 48. — Palaeovaranus sp. A, B, photographs of right dentary NHMW 2019/0058/0055 in lateral (A) and medial (B) views; C, D, photographs of right dentary NHMW 2019/0058/0056 in lateral (C) and medial (D) views; E, close up photograph of a tooth of right dentary NHMW 2019/0058/0056, showing the presence of plicidentine. Scale bars: A-D, 5 mm; E, 1 mm." captionText-5="FIG. 49. — Palaeovaranus sp. A-D, virtual 3D models of right dentary NHMW 2019/0058/0055 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views; E, F, virtual 3D models of right dentary NHMW 2019/0058/0056 in medial (E) and dorsal (F) views. Scale bars: 5 mm." captionText-6="FIG. 50. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), dorsal (C), right lateral (D), left lateral (E), and ventral (F) views. Scale bar: 2 mm." captionText-7="FIG. 51. — Palaeovaranus sp. Virtual 3D models of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), right lateral (C), left lateral (D), dorsal (E), ventral (F), ventrolateral (G), and anterolateral (H) views. Scale bar: 5 mm." captionText-8="FIG. 52. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0002 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar: 2 mm." captionText-9="FIG. 53. — Palaeovaranus sp. A-E, photographs of presacral vertebra NHMW 2019/0058/0003 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views; F-J, photographs of presacral vertebra NHMW 2019/0058/0014 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views; K-O, photographs of presacral vertebra NHMW 2019/0058/0045 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of presacral vertebra NHMW 2019/0058/0004 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views; U-Y, photographs of presacral vertebra NHMW 2019/0058/0020 in anterior (U), posterior (V), dorsal (W), ventral (X), and right lateral (Y) views; Z-D’, photographs of presacral vertebra NHMW 2019/0058/0012 in anterior (Z), posterior (A’), dorsal (B’), ventral (C’), and left lateral (D’) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720915" figureDoi-1="http://doi.org/10.5281/zenodo.4720921" figureDoi-10="http://doi.org/10.5281/zenodo.4720949" figureDoi-11="http://doi.org/10.5281/zenodo.4720953" figureDoi-12="http://doi.org/10.5281/zenodo.4720957" figureDoi-13="http://doi.org/10.5281/zenodo.4720961" figureDoi-14="http://doi.org/10.5281/zenodo.4720965" figureDoi-2="http://doi.org/10.5281/zenodo.4720923" figureDoi-3="http://doi.org/10.5281/zenodo.4720925" figureDoi-4="http://doi.org/10.5281/zenodo.4720927" figureDoi-5="http://doi.org/10.5281/zenodo.4720931" figureDoi-6="http://doi.org/10.5281/zenodo.4720935" figureDoi-7="http://doi.org/10.5281/zenodo.4720937" figureDoi-8="http://doi.org/10.5281/zenodo.4720941" figureDoi-9="http://doi.org/10.5281/zenodo.4720945" httpUri-0="https://zenodo.org/record/4720915/files/figure.png" httpUri-1="https://zenodo.org/record/4720921/files/figure.png" httpUri-10="https://zenodo.org/record/4720949/files/figure.png" httpUri-11="https://zenodo.org/record/4720953/files/figure.png" httpUri-12="https://zenodo.org/record/4720957/files/figure.png" httpUri-13="https://zenodo.org/record/4720961/files/figure.png" httpUri-14="https://zenodo.org/record/4720965/files/figure.png" httpUri-2="https://zenodo.org/record/4720923/files/figure.png" httpUri-3="https://zenodo.org/record/4720925/files/figure.png" httpUri-4="https://zenodo.org/record/4720927/files/figure.png" httpUri-5="https://zenodo.org/record/4720931/files/figure.png" httpUri-6="https://zenodo.org/record/4720935/files/figure.png" httpUri-7="https://zenodo.org/record/4720937/files/figure.png" httpUri-8="https://zenodo.org/record/4720941/files/figure.png" httpUri-9="https://zenodo.org/record/4720945/files/figure.png" pageId="41" pageNumber="258">Figs 44-58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF8AFFE830BAF801FF363E94" pageId="41" pageNumber="258" type="materials_examined">
<paragraph id="8BB7D22DFF8AFFE830BAF801FF363E94" blockId="41.[130,776,1976,2027]" lastBlockId="41.[810,1458,1426,1689]" pageId="41" pageNumber="258">
REFERRED SPECIMENS. —
<materialsCitation id="3B60D870FF8AFFE83151F801FF363E94" ID-GBIF-Occurrence="3091185315" pageId="41" pageNumber="258">A left maxilla (NHMW 2019/0048/0002); a left dentary (NHMW 2019/0058/0054); two right dentaries (NHMW 2019/0058/0055 and NHMW 2019/0058/0056); 35 presacral vertebrae (NHMW 2019/0058/0001-NHMW 2019/0058/0009, NHMW 2019/0058/0011-NHMW2019/0058/0027,NHMW2019/0058/0030, NHMW 2019/0058/0041-NHMW 2019/0058/0046, NHMW 2019/0058/0051,and NHMW 2019/0067/0001);two articulated sacral vertebrae (NHMW 2019/0058/0028); two sacral vertebrae (NHMW 2019/0058/0029 and NHMW 2019/0058/0047); 16 caudal vertebrae (NHMW 2019/0058/0031-NHMW 2019/0058/0040, NHMW 2019/0058/0048-NHMW2019/0058/0050,NHMW2019/0058/0052, NHMW 2019/0058/0053, and NHMW 2019/0067/0002).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31281A6FF8AFFF23313F96AF8363DA3" lastPageId="51" lastPageNumber="268" pageId="41" pageNumber="258" type="description">
<paragraph id="8BB7D22DFF8AFFE83313F96AF9E33EE7" blockId="41.[813,1457,1746,2028]" box="[813,957,1746,1772]" pageId="41" pageNumber="258">DESCRIPTION</paragraph>
<paragraph id="8BB7D22DFF8AFFE83313F94AFF733F01" blockId="41.[813,1457,1746,2028]" box="[813,1325,1778,1804]" pageId="41" pageNumber="258">
<emphasis id="B97C0E3FFF8AFFE83313F94AFF733F01" box="[813,1325,1778,1804]" italics="true" pageId="41" pageNumber="258">
Maxilla NHMW 2019/0048/0002 (
<figureCitation id="1333CEA8FF8AFFE83490F94BFEA43F01" box="[1198,1274,1778,1804]" captionStart="FIG" captionStartId="49.[131,142,1429,1446]" captionTargetBox="[139,1448,214,1385]" captionTargetId="figure-335@49.[132,1455,205,1490]" captionTargetPageId="49" captionText="FIG. 44. — Palaeovaranus sp. A-C, photographs of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), and dorsal (C) views; D, close up of the posteriormost preserved tooth of the same specimen. Scale bars: A-C, 5 mm; D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720915" httpUri="https://zenodo.org/record/4720915/files/figure.png" pageId="41" pageNumber="258">Figs 44</figureCitation>
;
<figureCitation id="1333CEA8FF8AFFE83536F94AFF7A3F01" box="[1288,1316,1778,1804]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="41" pageNumber="258">45</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BB7D22DFF8AFFE83313F8AAFFF13FE6" blockId="41.[813,1457,1746,2028]" pageId="41" pageNumber="258">
The left maxilla NHMW 2019/0048/0002 is not fully preserved; its anterior half is well preserved, however, the anteriormost and the posterior portions are missing (
<figureCitation id="1333CEA8FF8AFFE83562F8E9FFF63F61" box="[1372,1448,1873,1900]" captionStart="FIG" captionStartId="49.[131,142,1429,1446]" captionTargetBox="[139,1448,214,1385]" captionTargetId="figure-335@49.[132,1455,205,1490]" captionTargetPageId="49" captionText="FIG. 44. — Palaeovaranus sp. A-C, photographs of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), and dorsal (C) views; D, close up of the posteriormost preserved tooth of the same specimen. Scale bars: A-C, 5 mm; D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720915" httpUri="https://zenodo.org/record/4720915/files/figure.png" pageId="41" pageNumber="258">Figs 44</figureCitation>
;
<figureCitation id="1333CEA8FF8AFFE83313F8C9F9143F81" box="[813,842,1905,1932]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="41" pageNumber="258">45</figureCitation>
). The external surface of the maxilla consists of large and low mounds divided by narrow grooves. Between the dorsal and ventral margins, the wall of the maxilla is distinctly concave. Immediately dorsal to the ventral maxilla
</paragraph>
<caption id="DF7782A5FF89FFEB30BAFBB3FFEF3C11" ID-DOI="http://doi.org/10.5281/zenodo.4720883" ID-Zenodo-Dep="4720883" box="[132,1457,1035,1052]" httpUri="https://zenodo.org/record/4720883/files/figure.png" pageId="42" pageNumber="259" startId="42.[132,143,1035,1052]" targetBox="[142,1444,217,992]" targetPageId="42">
<paragraph id="8BB7D22DFF89FFEB30BAFBB3FFEF3C11" blockId="42.[132,1457,1035,1052]" box="[132,1457,1035,1052]" pageId="42" pageNumber="259">
FIG. 35. —
<taxonomicName id="4C08A9AEFF89FFEB30E2FBB3FBAE3C11" authority="Zittel, 1887" authorityName="Zittel" authorityYear="1887" box="[220,496,1035,1052]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="42" pageNumber="259" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF89FFEB30E2FBB3FBD13C11" box="[220,399,1035,1052]" italics="true" pageId="42" pageNumber="259">Palaeovaranus cayluxi</emphasis>
<bibRefCitation id="EF99AFDCFF89FFEB31ACFBB3FBAE3C11" author="ZITTEL K. A." box="[402,496,1035,1052]" pageId="42" pageNumber="259" refId="ref40223" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen, Leipzig, 900 p. https: // www. biodiversitylibrary. org / page / 40393265" type="book" year="1887">Zittel,1887</bibRefCitation>
</taxonomicName>
-1890.Virtual 3D models of parietal NHMW 2019/0048/0001 in dorsolateral (
<emphasis id="B97C0E3FFF89FFEB3467FBB3FE383C11" bold="true" box="[1113,1126,1035,1052]" pageId="42" pageNumber="259">A</emphasis>
) and anterior (
<emphasis id="B97C0E3FFF89FFEB34E7FBB3FEB83C11" bold="true" box="[1241,1254,1035,1052]" pageId="42" pageNumber="259">B</emphasis>
) views.Scale bar:5 mm.
</paragraph>
</caption>
<caption id="DF7782A5FF89FFEB30BAF879F8D93FE4" ID-DOI="http://doi.org/10.5281/zenodo.4720889" ID-Zenodo-Dep="4720889" httpUri="https://zenodo.org/record/4720889/files/figure.png" pageId="42" pageNumber="259" startId="42.[132,143,1985,2002]" targetBox="[131,1457,1129,1961]" targetPageId="42">
<paragraph id="8BB7D22DFF89FFEB30BAF879F8D93FE4" blockId="42.[132,1455,1985,2025]" pageId="42" pageNumber="259">
FIG. 36. —
<taxonomicName id="4C08A9AEFF89FFEB30E1F879FBA23FDF" authority="Zittel, 1887" authorityName="Zittel" authorityYear="1887" box="[223,508,1985,2002]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="42" pageNumber="259" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF89FFEB30E1F879FBC83FDF" box="[223,406,1985,2002]" italics="true" pageId="42" pageNumber="259">Palaeovaranus cayluxi</emphasis>
<bibRefCitation id="EF99AFDCFF89FFEB31A2F879FBA23FDF" author="ZITTEL K. A." box="[412,508,1985,2002]" pageId="42" pageNumber="259" refId="ref40223" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen, Leipzig, 900 p. https: // www. biodiversitylibrary. org / page / 40393265" type="book" year="1887">Zittel, 1887</bibRefCitation>
</taxonomicName>
-1890. Photographs of parietal MNHN.F.QU17176 in dorsal (
<emphasis id="B97C0E3FFF89FFEB33CAF879FE5F3FDF" bold="true" box="[1012,1025,1985,2002]" pageId="42" pageNumber="259">A</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF89FFEB344DF879FEDE3FDF" bold="true" box="[1139,1152,1985,2002]" pageId="42" pageNumber="259">B</emphasis>
) views. This specimen originally appeared as drawings in
<bibRefCitation id="EF99AFDCFF89FFEB317FF860FBFE3FE4" author="AUGE M." box="[321,416,2008,2025]" pageId="42" pageNumber="259" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé (2005</bibRefCitation>
: fig. 188). Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF7782A5FF88FFEA30BAF8A4F8363F57" ID-DOI="http://doi.org/10.5281/zenodo.4720891" ID-Zenodo-Dep="4720891" httpUri="https://zenodo.org/record/4720891/files/figure.png" pageId="43" pageNumber="260" startId="43.[132,143,1820,1837]" targetBox="[124,1472,185,1797]" targetPageId="43">
<paragraph id="8BB7D22DFF88FFEA30BAF8A4F8363F57" blockId="43.[132,1455,1820,1882]" pageId="43" pageNumber="260">
FIG. 37. —
<taxonomicName id="4C08A9AEFF88FFEA30DEF8A4FBA33F20" authority="Zittel, 1887" authorityName="Zittel" authorityYear="1887" box="[224,509,1820,1837]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="260" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF88FFEA30DEF8A4FBC93F20" box="[224,407,1820,1837]" italics="true" pageId="43" pageNumber="260">Palaeovaranus cayluxi</emphasis>
<bibRefCitation id="EF99AFDCFF88FFEA31A0F8A4FBA33F20" author="ZITTEL K. A." box="[414,509,1820,1837]" pageId="43" pageNumber="260" refId="ref40223" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen, Leipzig, 900 p. https: // www. biodiversitylibrary. org / page / 40393265" type="book" year="1887">Zittel, 1887</bibRefCitation>
</taxonomicName>
-1890 and
<taxonomicName id="4C08A9AEFF88FFEA3269F8A4F9163F20" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[599,840,1820,1837]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="260" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF88FFEA3269F8A4F9163F20" box="[599,840,1820,1837]" italics="true" pageId="43" pageNumber="260">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF88FFEA3371F8A4F9DE3F20" box="[847,896,1820,1837]" pageId="43" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B97C0E3FFF88FFEA33BDF8A4F9CE3F20" bold="true" box="[899,912,1820,1837]" pageId="43" pageNumber="260">A</emphasis>
,
<emphasis id="B97C0E3FFF88FFEA33A5F8A4F9F73F20" bold="true" box="[923,937,1820,1837]" pageId="43" pageNumber="260">C</emphasis>
, virtual 3D models of parietal NHMW 2019/0048/0001 of
<taxonomicName id="4C08A9AEFF88FFEA35B0F8A4FB7F3F4E" authorityName="Zittel" authorityYear="1887" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="260" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF88FFEA35B0F8A4FB7F3F4E" italics="true" pageId="43" pageNumber="260">Palaeovaranus cayluxi</emphasis>
</taxonomicName>
in dorsal (
<emphasis id="B97C0E3FFF88FFEA3147F88AFBD83F4E" bold="true" box="[377,390,1842,1859]" pageId="43" pageNumber="260">A</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF88FFEA31C9F88AF85B3F4E" bold="true" box="[503,517,1842,1859]" pageId="43" pageNumber="260">C</emphasis>
) views;
<emphasis id="B97C0E3FFF88FFEA3277F88AF8083F4E" bold="true" box="[585,598,1842,1859]" pageId="43" pageNumber="260">B</emphasis>
,
<emphasis id="B97C0E3FFF88FFEA325FF88AF8313F4E" bold="true" box="[609,623,1842,1859]" pageId="43" pageNumber="260">D</emphasis>
, virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of
<taxonomicName id="4C08A9AEFF88FFEA3480F88AFFF13F4E" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[1214,1455,1842,1859]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="260" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF88FFEA3480F88AFFF13F4E" box="[1214,1455,1842,1859]" italics="true" pageId="43" pageNumber="260">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF88FFEA30BAF8F1FAEA3F57" box="[132,180,1865,1882]" pageId="43" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
in dorsal (
<emphasis id="B97C0E3FFF88FFEA3137F8F1FB483F57" bold="true" box="[265,278,1865,1882]" pageId="43" pageNumber="260">B</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF88FFEA31B9F8F1FBCB3F57" bold="true" box="[391,405,1865,1882]" pageId="43" pageNumber="260">D</emphasis>
) views. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFF88FFED30BAF829FF513E81" blockId="43.[132,775,1937,2028]" lastBlockId="44.[811,1457,1427,1677]" lastPageId="44" lastPageNumber="261" pageId="43" pageNumber="260">
margin, five labial foramina are preserved. The most distinguished feature of the dorsomedial surface of the nasal process (prefrontal process sensu
<bibRefCitation id="EF99AFDCFF88FFEA3239F869F8EE3FE1" author="FEJERVARY DE G. J." box="[519,688,2001,2028]" pageId="43" pageNumber="260" pagination="1 - 130" refId="ref34838" refString="FEJERVARY DE G. J. 1935. - Further contributions to a monograph of the Megalanidae and fossil Varanidae, with notes on recent varanians. Annales historico-naturales Musei nationalis hungarici, Pars Zoologica 29: 1 - 130." type="journal article" year="1935">Fejérváry 1935</bibRefCitation>
) of the maxilla is a shelf extending medially and running some distance ventral to the dorsal margin of the nasal process of the maxilla (
<figureCitation id="1333CEA8FF88FFEA33EAF869FE273FE1" box="[980,1145,2001,2028]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="43" pageNumber="260">Fig. 45B, D, F</figureCitation>
). This shelf is the nasal crest which articulated with the nasal (a distinguished feature for
<taxonomicName id="4C08A9AEFF8FFFED30BDFA0BFB453DC0" authorityName="Zittel" authorityYear="1887" box="[131,283,1459,1485]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="261" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF8FFFED30BDFA0BFB453DC0" box="[131,283,1459,1485]" italics="true" pageId="44" pageNumber="261">Palaeovaranus</emphasis>
</taxonomicName>
). There is a deep sulcus between the medial margin of the nasal crest and the dorsal margin of the nasal process of the maxilla; the sulcus represents an articulating surface for the maxillary crest of the nasal. Immediately ventral to the mid-length of the nasal crest lies a semilunar impression in the medial surface of the nasal process of the maxilla; this is the articulating surface for the maxillary lamina of the prefrontal. Anteroventrally and anteriorly to the nasal crest, the medial surface of the maxilla is massively constructed;
<bibRefCitation id="EF99AFDCFF8FFFED312CF96AFB963EE1" author="FEJERVARY DE G. J." box="[274,456,1746,1773]" pageId="44" pageNumber="261" pagination="1 - 130" refId="ref34838" refString="FEJERVARY DE G. J. 1935. - Further contributions to a monograph of the Megalanidae and fossil Varanidae, with notes on recent varanians. Annales historico-naturales Musei nationalis hungarici, Pars Zoologica 29: 1 - 130." type="journal article" year="1935">Fejérváry (1935)</bibRefCitation>
called this structure the basal torus (
<figureCitation id="1333CEA8FF8FFFED30F0F94AFB6C3F01" box="[206,306,1778,1805]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="44" pageNumber="261">Fig. 45B</figureCitation>
). Ventral to the semilunar impression and dorsal to the supradental shelf lies a large, elliptical depression in the medial wall of the nasal representing a lacrimal recess. Its anteriormost portion bears a foramen (
<figureCitation id="1333CEA8FF8FFFED32A9F8E9F8A73F61" box="[663,761,1873,1900]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="44" pageNumber="261">Fig. 45C</figureCitation>
). The supradental shelf is straight. Its dorsal surface bears a distinct sulcus for the nasolacrimal duct (
<figureCitation id="1333CEA8FF8FFFED325FF829F8983FA1" box="[609,710,1937,1964]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="44" pageNumber="261">Fig. 45E</figureCitation>
). The anterior portion of the supradental shelf bears an elliptical and slightly dorsomedially inclined surface, an oblique inter- nasal lamella (
<figureCitation id="1333CEA8FF8FFFED33F7FA2BFE723DA3" box="[969,1068,1427,1454]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="44" pageNumber="261">Fig. 45B</figureCitation>
). The maxilla immediately laterally to the supradental shelf is burrowed by the superior alveolar canal; its anterior and posterior openings are well preserved (
<figureCitation id="1333CEA8FF8FFFED3308FA4BF9E13E00" box="[822,959,1522,1549]" captionStart="FIG" captionStartId="50.[132,143,1985,2002]" captionTargetBox="[147,1438,219,1839]" captionTargetId="figure-15@50.[138,1449,217,1845]" captionTargetPageId="50" captionText="FIG. 45. — Palaeovaranus sp. A-G, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), posteromedial (C), dorsomedial (D), dorsal (E) and posterior (F) views; G, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="44" pageNumber="261">Fig. 45D, F</figureCitation>
). The bases of three teeth of the maxilla are well preserved. The bases are mesiodistally broad and bear well preserved basal striae, i.e., plicidentine (
<figureCitation id="1333CEA8FF8FFFED3525F98AFFFF3E40" box="[1307,1441,1586,1613]" captionStart="FIG" captionStartId="49.[131,142,1429,1446]" captionTargetBox="[139,1448,214,1385]" captionTargetId="figure-335@49.[132,1455,205,1490]" captionTargetPageId="49" captionText="FIG. 44. — Palaeovaranus sp. A-C, photographs of left maxilla NHMW 2019/0048/0002 in lateral (A), medial (B), and dorsal (C) views; D, close up of the posteriormost preserved tooth of the same specimen. Scale bars: A-C, 5 mm; D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720915" httpUri="https://zenodo.org/record/4720915/files/figure.png" pageId="44" pageNumber="261">Fig. 44B, D</figureCitation>
). The teeth are strongly curved distally and slightly lingually. The distal cutting edges are well developed.
</paragraph>
<caption id="DF7782A5FF8FFFED30BAFA93F95A3D5E" ID-DOI="http://doi.org/10.5281/zenodo.4720893" ID-Zenodo-Dep="4720893" httpUri="https://zenodo.org/record/4720893/files/figure.png" pageId="44" pageNumber="261" startId="44.[132,143,1323,1340]" targetBox="[149,1430,217,1279]" targetPageId="44">
<paragraph id="8BB7D22DFF8FFFED30BAFA93F95A3D5E" blockId="44.[132,1455,1323,1363]" pageId="44" pageNumber="261">
FIG. 38. —
<taxonomicName id="4C08A9AEFF8FFFED30E1FA93FB913D31" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[223,463,1323,1340]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="261" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8FFFED30E1FA93FB913D31" box="[223,463,1323,1340]" italics="true" pageId="44" pageNumber="261">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8FFFED31EAFA93F85A3D31" box="[468,516,1323,1340]" pageId="44" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
Virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of
<taxonomicName id="4C08A9AEFF8FFFED3472FA93FF623D31" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[1100,1340,1323,1340]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="261" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8FFFED3472FA93FF623D31" box="[1100,1340,1323,1340]" italics="true" pageId="44" pageNumber="261">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8FFFED357FFA93FF2F3D31" box="[1345,1393,1323,1340]" pageId="44" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
in posteroventral (
<emphasis id="B97C0E3FFF8FFFED30D9FAFAFAAA3D5E" bold="true" box="[231,244,1346,1363]" pageId="44" pageNumber="261">A</emphasis>
), dorsolateral (
<emphasis id="B97C0E3FFF8FFFED314EFAFAFB233D5E" bold="true" box="[368,381,1346,1363]" pageId="44" pageNumber="261">B</emphasis>
), and anterodorsal (
<emphasis id="B97C0E3FFF8FFFED321DFAFAF86F3D5E" bold="true" box="[547,561,1346,1363]" pageId="44" pageNumber="261">C</emphasis>
) views. Scale bar: 5 mm.
</paragraph>
</caption>
<subSection id="E287CBC6FF8FFFEC3313F90BFE5D3F01" lastPageId="45" lastPageNumber="262" pageId="44" pageNumber="261" type="multiple">
<paragraph id="8BB7D22DFF8FFFED3313F90BFE473EC1" blockId="44.[813,1049,1714,1740]" box="[813,1049,1714,1740]" pageId="44" pageNumber="261">
<heading id="D0FF6541FF8FFFED3313F90BFE473EC1" box="[813,1049,1714,1740]" fontSize="11" level="2" pageId="44" pageNumber="261" reason="8">
<emphasis id="B97C0E3FFF8FFFED3313F90BFE473EC1" box="[813,1049,1714,1740]" italics="true" pageId="44" pageNumber="261">
Dentaries (
<figureCitation id="1333CEA8FF8FFFED339EF90BFE4E3EC1" box="[928,1040,1714,1740]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="51.[132,143,925,942]" captionStartId-1="52.[132,143,991,1008]" captionStartId-2="53.[132,143,1541,1558]" captionStartId-3="54.[132,143,1446,1463]" captionTargetBox-0="[132,1455,203,895]" captionTargetBox-1="[132,1456,182,970]" captionTargetBox-2="[138,1449,216,1479]" captionTargetBox-3="[142,1448,215,1407]" captionTargetId-0="figure-582@51.[132,1455,203,895]" captionTargetId-1="figure-475@52.[132,1455,211,892]" captionTargetId-2="figure-270@53.[132,1455,207,1492]" captionTargetId-3="figure-235@54.[132,1455,207,1356]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionTargetPageId-3="54" captionText-0="FIG. 46. — Palaeovaranus sp. Photographs of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and posterior (D) views. Scale bar: 5 mm." captionText-1="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." captionText-2="FIG. 48. — Palaeovaranus sp. A, B, photographs of right dentary NHMW 2019/0058/0055 in lateral (A) and medial (B) views; C, D, photographs of right dentary NHMW 2019/0058/0056 in lateral (C) and medial (D) views; E, close up photograph of a tooth of right dentary NHMW 2019/0058/0056, showing the presence of plicidentine. Scale bars: A-D, 5 mm; E, 1 mm." captionText-3="FIG. 49. — Palaeovaranus sp. A-D, virtual 3D models of right dentary NHMW 2019/0058/0055 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views; E, F, virtual 3D models of right dentary NHMW 2019/0058/0056 in medial (E) and dorsal (F) views. Scale bars: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720923" figureDoi-1="http://doi.org/10.5281/zenodo.4720925" figureDoi-2="http://doi.org/10.5281/zenodo.4720927" figureDoi-3="http://doi.org/10.5281/zenodo.4720931" httpUri-0="https://zenodo.org/record/4720923/files/figure.png" httpUri-1="https://zenodo.org/record/4720925/files/figure.png" httpUri-2="https://zenodo.org/record/4720927/files/figure.png" httpUri-3="https://zenodo.org/record/4720931/files/figure.png" pageId="44" pageNumber="261">Figs 46-49</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7D22DFF8FFFED3313F96AFFEE3FE6" blockId="44.[810,1457,1746,2028]" pageId="44" pageNumber="261">
All three dentaries are incomplete, missing the anteriormost and posteriormost edges.The specimens NHMW 2019/0058/0054 and NHMW 2019/0058/0055 are relatively large and robust, with preserved maximum lengths 27.6 and
<quantity id="4CF07FC8FF8FFFED352FF88AFF3B3F41" box="[1297,1381,1842,1868]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="44" pageNumber="261" unit="mm" value="21.0">21 mm</quantity>
, while NHMW 2019/0058/0056 is smaller and slenderer, with a maximum preserved length of
<quantity id="4CF07FC8FF8FFFED344BF8C9FE993F81" box="[1141,1223,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="44" pageNumber="261" unit="mm" value="18.0">18 mm</quantity>
. In all three dentaries, the maximum height is achieved at their posterior portions. NHMW 2019/0058/0054 bears four preserved teeth, one more tooth base, and three other empty tooth positions
</paragraph>
<caption id="DF7782A5FF8EFFEC30BAFA9EFB4A3D43" ID-DOI="http://doi.org/10.5281/zenodo.4720899" ID-Zenodo-Dep="4720899" httpUri="https://zenodo.org/record/4720899/files/figure.png" pageId="45" pageNumber="262" startId="45.[132,143,1318,1335]" targetBox="[132,1455,181,1289]" targetPageId="45">
<paragraph id="8BB7D22DFF8EFFEC30BAFA9EFB4A3D43" blockId="45.[132,1457,1318,1358]" pageId="45" pageNumber="262">
FIG. 39. —
<taxonomicName id="4C08A9AEFF8EFFEC30E0FA9EFB903D3A" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[222,462,1318,1335]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="262" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8EFFEC30E0FA9EFB903D3A" box="[222,462,1318,1335]" italics="true" pageId="45" pageNumber="262">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8EFFEC31EAFA9EF85A3D3A" box="[468,516,1318,1335]" pageId="45" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
Photographs of the holotype parietal NHMW 2019/0047/0001 in dorsal (
<emphasis id="B97C0E3FFF8EFFEC3463FA9EFE343D3A" bold="true" box="[1117,1130,1318,1335]" pageId="45" pageNumber="262">A</emphasis>
), ventral (
<emphasis id="B97C0E3FFF8EFFEC3485FA9EFE963D3A" bold="true" box="[1211,1224,1318,1335]" pageId="45" pageNumber="262">B</emphasis>
), and right lateral (
<emphasis id="B97C0E3FFF8EFFEC355DFA9EFF2F3D3A" bold="true" box="[1379,1393,1318,1335]" pageId="45" pageNumber="262">C</emphasis>
) views. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFF8EFFEC30BAFA2BF9BF3E81" blockId="45.[129,776,1427,2028]" lastBlockId="45.[810,1457,1427,1804]" pageId="45" pageNumber="262">
(
<figureCitation id="1333CEA8FF8EFFEC30B3FA2BFA843DA0" box="[141,218,1427,1453]" captionStart="FIG" captionStartId="51.[132,143,925,942]" captionTargetBox="[132,1455,203,895]" captionTargetId="figure-582@51.[132,1455,203,895]" captionTargetPageId="51" captionText="FIG. 46. — Palaeovaranus sp. Photographs of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and posterior (D) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720923" httpUri="https://zenodo.org/record/4720923/files/figure.png" pageId="45" pageNumber="262">Figs 46</figureCitation>
;
<figureCitation id="1333CEA8FF8EFFEC30D9FA2BFB5D3DA0" box="[231,259,1427,1453]" captionStart="FIG" captionStartId="52.[132,143,991,1008]" captionTargetBox="[132,1456,182,970]" captionTargetId="figure-475@52.[132,1455,211,892]" captionTargetPageId="52" captionText="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720925" httpUri="https://zenodo.org/record/4720925/files/figure.png" pageId="45" pageNumber="262">47</figureCitation>
), NHMW 2019/0058/0055 bears six preserved teeth and two other empty tooth positions (
<figureCitation id="1333CEA8FF8EFFEC3272FA0BF8993DC0" box="[588,711,1459,1485]" captionStart="FIG" captionStartId="53.[132,143,1541,1558]" captionTargetBox="[138,1449,216,1479]" captionTargetId="figure-270@53.[132,1455,207,1492]" captionTargetPageId="53" captionText="FIG. 48. — Palaeovaranus sp. A, B, photographs of right dentary NHMW 2019/0058/0055 in lateral (A) and medial (B) views; C, D, photographs of right dentary NHMW 2019/0058/0056 in lateral (C) and medial (D) views; E, close up photograph of a tooth of right dentary NHMW 2019/0058/0056, showing the presence of plicidentine. Scale bars: A-D, 5 mm; E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720927" httpUri="https://zenodo.org/record/4720927/files/figure.png" pageId="45" pageNumber="262">Figs 48A, B</figureCitation>
;
<figureCitation id="1333CEA8FF8EFFEC32F1FA0BF8B53DC0" box="[719,747,1459,1485]" captionStart="FIG" captionStartId="54.[132,143,1446,1463]" captionTargetBox="[142,1448,215,1407]" captionTargetId="figure-235@54.[132,1455,207,1356]" captionTargetPageId="54" captionText="FIG. 49. — Palaeovaranus sp. A-D, virtual 3D models of right dentary NHMW 2019/0058/0055 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views; E, F, virtual 3D models of right dentary NHMW 2019/0058/0056 in medial (E) and dorsal (F) views. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720931" httpUri="https://zenodo.org/record/4720931/files/figure.png" pageId="45" pageNumber="262">49</figureCitation>
A- D), and NHMW 2019/0058/0056 bears two preserved teeth, fragments of two other teeth, and two other empty tooth positions (
<figureCitation id="1333CEA8FF8EFFEC30CBF9AAFB1C3E20" box="[245,322,1554,1581]" captionStart="FIG" captionStartId="53.[132,143,1541,1558]" captionTargetBox="[138,1449,216,1479]" captionTargetId="figure-270@53.[132,1455,207,1492]" captionTargetPageId="53" captionText="FIG. 48. — Palaeovaranus sp. A, B, photographs of right dentary NHMW 2019/0058/0055 in lateral (A) and medial (B) views; C, D, photographs of right dentary NHMW 2019/0058/0056 in lateral (C) and medial (D) views; E, close up photograph of a tooth of right dentary NHMW 2019/0058/0056, showing the presence of plicidentine. Scale bars: A-D, 5 mm; E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720927" httpUri="https://zenodo.org/record/4720927/files/figure.png" pageId="45" pageNumber="262">Figs 48</figureCitation>
C-E; 49E, F). All teeth are subpleurodont, recurved, and rather pointed. Their bases are broad and are marked by the distinct presence of plicidentine. The latter feature is significantly most prominent in the anteriormost tooth of NHMW 2019/0058/0056 (
<figureCitation id="1333CEA8FF8EFFEC3233F92AF8343EA0" box="[525,618,1682,1709]" captionStart="FIG" captionStartId="53.[132,143,1541,1558]" captionTargetBox="[138,1449,216,1479]" captionTargetId="figure-270@53.[132,1455,207,1492]" captionTargetPageId="53" captionText="FIG. 48. — Palaeovaranus sp. A, B, photographs of right dentary NHMW 2019/0058/0055 in lateral (A) and medial (B) views; C, D, photographs of right dentary NHMW 2019/0058/0056 in lateral (C) and medial (D) views; E, close up photograph of a tooth of right dentary NHMW 2019/0058/0056, showing the presence of plicidentine. Scale bars: A-D, 5 mm; E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720927" httpUri="https://zenodo.org/record/4720927/files/figure.png" pageId="45" pageNumber="262">Fig. 48E</figureCitation>
). Teeth possess cutting edges, with this serration being most prominent in NHMW 2019/0058/0056. In the largest specimen (NHMW 2019/0058/0054), remains of the coronoid process are visible (
<figureCitation id="1333CEA8FF8EFFEC3082F8AAFB403F21" box="[188,286,1810,1836]" captionStart="FIG" captionStartId="52.[132,143,991,1008]" captionTargetBox="[132,1456,182,970]" captionTargetId="figure-475@52.[132,1455,211,892]" captionTargetPageId="52" captionText="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720925" httpUri="https://zenodo.org/record/4720925/files/figure.png" pageId="45" pageNumber="262">Fig. 47A</figureCitation>
). The coronoid articulation facet is preserved in NHMW 2019/0058/0054 (
<figureCitation id="1333CEA8FF8EFFEC31E8F889F8653F41" box="[470,571,1841,1868]" captionStart="FIG" captionStartId="52.[132,143,991,1008]" captionTargetBox="[132,1456,182,970]" captionTargetId="figure-475@52.[132,1455,211,892]" captionTargetPageId="52" captionText="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720925" httpUri="https://zenodo.org/record/4720925/files/figure.png" pageId="45" pageNumber="262">Fig. 47D</figureCitation>
). Splenial articulation facets are visible in all specimens. The Meckel’s groove is open throughout the preserved lengths of all specimens. This structure becomes much widened posteriorly in all three dentaries, whereas in the anterior half of these specimens it is partially “blanketed” (but not closed) by the dental crest. In the posterior portion of the two largest dentaries, the characteristic alveolar canal (foramen dentale pro introitu partis alveolaris inferioris nervi mandibularis sensuFejérváry 1935) is prominent, delimiting a large longitudinal groove, which in NHMW 2019/0058/0055 extends and is visible also in its anterior edge. The posterior spine is thin, with its termination being acute. The intramandibular septum is wide (
<figureCitation id="1333CEA8FF8EFFEC334EF9CAF98D3E81" box="[880,979,1650,1676]" captionStart="FIG" captionStartId="52.[132,143,991,1008]" captionTargetBox="[132,1456,182,970]" captionTargetId="figure-475@52.[132,1455,211,892]" captionTargetPageId="52" captionText="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720925" httpUri="https://zenodo.org/record/4720925/files/figure.png" pageId="45" pageNumber="262">Fig. 47C</figureCitation>
).
</paragraph>
<paragraph id="8BB7D22DFF8EFFEC337DF92BFE5D3F01" blockId="45.[810,1457,1427,1804]" pageId="45" pageNumber="262">
In lateral view, several dorsoventrally elongated labial foramina pierce the dentaries’ surfaces, situated across irregular intervals, but otherwise these surfaces are relatively smooth (
<figureCitation id="1333CEA8FF8EFFEC3577F96AFFF53EE1" box="[1353,1451,1746,1772]" captionStart="FIG" captionStartId="51.[132,143,925,942]" captionTargetBox="[132,1455,203,895]" captionTargetId="figure-582@51.[132,1455,203,895]" captionTargetPageId="51" captionText="FIG. 46. — Palaeovaranus sp. Photographs of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and posterior (D) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720923" httpUri="https://zenodo.org/record/4720923/files/figure.png" pageId="45" pageNumber="262">Figs 46A</figureCitation>
;
<figureCitation id="1333CEA8FF8EFFEC3313F94AF9033F01" box="[813,861,1778,1804]" captionStart="FIG" captionStartId="52.[132,143,991,1008]" captionTargetBox="[132,1456,182,970]" captionTargetId="figure-475@52.[132,1455,211,892]" captionTargetPageId="52" captionText="FIG. 47. — Palaeovaranus sp.Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720925" httpUri="https://zenodo.org/record/4720925/files/figure.png" pageId="45" pageNumber="262">47A</figureCitation>
;
<figureCitation id="1333CEA8FF8EFFEC3356F94AF9E73F01" box="[872,953,1778,1804]" captionStart="FIG" captionStartId="53.[132,143,1541,1558]" captionTargetBox="[138,1449,216,1479]" captionTargetId="figure-270@53.[132,1455,207,1492]" captionTargetPageId="53" captionText="FIG. 48. — Palaeovaranus sp. A, B, photographs of right dentary NHMW 2019/0058/0055 in lateral (A) and medial (B) views; C, D, photographs of right dentary NHMW 2019/0058/0056 in lateral (C) and medial (D) views; E, close up photograph of a tooth of right dentary NHMW 2019/0058/0056, showing the presence of plicidentine. Scale bars: A-D, 5 mm; E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720927" httpUri="https://zenodo.org/record/4720927/files/figure.png" pageId="45" pageNumber="262">48A, C</figureCitation>
;
<figureCitation id="1333CEA8FF8EFFEC33FBF94AF9A93F01" box="[965,1015,1778,1804]" captionStart="FIG" captionStartId="54.[132,143,1446,1463]" captionTargetBox="[142,1448,215,1407]" captionTargetId="figure-235@54.[132,1455,207,1356]" captionTargetPageId="54" captionText="FIG. 49. — Palaeovaranus sp. A-D, virtual 3D models of right dentary NHMW 2019/0058/0055 in lateral (A), dorsal (B), medial (C), and ventromedial (D) views; E, F, virtual 3D models of right dentary NHMW 2019/0058/0056 in medial (E) and dorsal (F) views. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720931" httpUri="https://zenodo.org/record/4720931/files/figure.png" pageId="45" pageNumber="262">49A</figureCitation>
).
</paragraph>
</subSection>
<subSection id="E287CBC6FF8EFFEE3313F889F9583F66" lastPageId="47" lastPageNumber="264" pageId="45" pageNumber="262" type="multiple">
<paragraph id="8BB7D22DFF8EFFEC3313F889FE2A3F41" blockId="45.[813,1140,1841,1868]" box="[813,1140,1841,1868]" pageId="45" pageNumber="262">
<heading id="D0FF6541FF8EFFEC3313F889FE2A3F41" box="[813,1140,1841,1868]" fontSize="11" level="2" pageId="45" pageNumber="262" reason="8">
<emphasis id="B97C0E3FFF8EFFEC3313F889FE2A3F41" box="[813,1140,1841,1868]" italics="true" pageId="45" pageNumber="262">
Presacral vertebrae (
<figureCitation id="1333CEA8FF8EFFEC33C5F88AFE353F41" box="[1019,1131,1842,1868]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="55.[132,143,1481,1498]" captionStartId-1="56.[132,143,1040,1057]" captionStartId-2="57.[132,143,1262,1279]" captionStartId-3="58.[132,143,1894,1911]" captionTargetBox-0="[136,1452,214,1439]" captionTargetBox-1="[139,1449,216,996]" captionTargetBox-2="[132,1455,207,1225]" captionTargetBox-3="[136,1450,216,1841]" captionTargetId-0="figure-315@55.[132,1455,209,1447]" captionTargetId-1="figure-573@56.[132,1455,201,1002]" captionTargetId-2="figure-386@57.[132,1455,207,1225]" captionTargetId-3="figure-15@58.[132,1455,211,1845]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionTargetPageId-3="58" captionText-0="FIG. 50. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), dorsal (C), right lateral (D), left lateral (E), and ventral (F) views. Scale bar: 2 mm." captionText-1="FIG. 51. — Palaeovaranus sp. Virtual 3D models of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), right lateral (C), left lateral (D), dorsal (E), ventral (F), ventrolateral (G), and anterolateral (H) views. Scale bar: 5 mm." captionText-2="FIG. 52. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0002 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar: 2 mm." captionText-3="FIG. 53. — Palaeovaranus sp. A-E, photographs of presacral vertebra NHMW 2019/0058/0003 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views; F-J, photographs of presacral vertebra NHMW 2019/0058/0014 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views; K-O, photographs of presacral vertebra NHMW 2019/0058/0045 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of presacral vertebra NHMW 2019/0058/0004 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views; U-Y, photographs of presacral vertebra NHMW 2019/0058/0020 in anterior (U), posterior (V), dorsal (W), ventral (X), and right lateral (Y) views; Z-D’, photographs of presacral vertebra NHMW 2019/0058/0012 in anterior (Z), posterior (A’), dorsal (B’), ventral (C’), and left lateral (D’) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720935" figureDoi-1="http://doi.org/10.5281/zenodo.4720937" figureDoi-2="http://doi.org/10.5281/zenodo.4720941" figureDoi-3="http://doi.org/10.5281/zenodo.4720945" httpUri-0="https://zenodo.org/record/4720935/files/figure.png" httpUri-1="https://zenodo.org/record/4720937/files/figure.png" httpUri-2="https://zenodo.org/record/4720941/files/figure.png" httpUri-3="https://zenodo.org/record/4720945/files/figure.png" pageId="45" pageNumber="262">Figs 50-53</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7D22DFF8EFFEC3313F8E9FFF13FE1" blockId="45.[811,1456,1873,2028]" pageId="45" pageNumber="262">
The size of the presacral vertebrae varies, with centrum lengths ranging between 7.0 and
<quantity id="4CF07FC8FF8EFFEC3498F8C9FF533F81" box="[1190,1293,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="45" pageNumber="262" unit="mm" value="10.8">10.8 mm</quantity>
(
<figureCitation id="1333CEA8FF8EFFEC3523F8C9FFC13F86" box="[1309,1439,1905,1931]" captionStart="APPENDIX" captionStartId="75.[427,439,219,236]" captionTargetPageId="75" captionText="APPENDIX 1. — Centrum lengths of all lizard vertebrae from the NHMW Quercy collection." pageId="45" pageNumber="262">Appendix 1</figureCitation>
). There are usually two distinct ridges in the ventral surface of the centrum that commence at around the level of the cotyle and terminate slightly prior to the condyle (
<figureCitation id="1333CEA8FF8EFFEC3532F869FFDE3FE1" box="[1292,1408,2001,2028]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="55.[132,143,1481,1498]" captionStartId-1="56.[132,143,1040,1057]" captionStartId-2="57.[132,143,1262,1279]" captionStartId-3="58.[132,143,1894,1911]" captionTargetBox-0="[136,1452,214,1439]" captionTargetBox-1="[139,1449,216,996]" captionTargetBox-2="[132,1455,207,1225]" captionTargetBox-3="[136,1450,216,1841]" captionTargetId-0="figure-315@55.[132,1455,209,1447]" captionTargetId-1="figure-573@56.[132,1455,201,1002]" captionTargetId-2="figure-386@57.[132,1455,207,1225]" captionTargetId-3="figure-15@58.[132,1455,211,1845]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionTargetPageId-3="58" captionText-0="FIG. 50. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), dorsal (C), right lateral (D), left lateral (E), and ventral (F) views. Scale bar: 2 mm." captionText-1="FIG. 51. — Palaeovaranus sp. Virtual 3D models of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), right lateral (C), left lateral (D), dorsal (E), ventral (F), ventrolateral (G), and anterolateral (H) views. Scale bar: 5 mm." captionText-2="FIG. 52. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0002 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar: 2 mm." captionText-3="FIG. 53. — Palaeovaranus sp. A-E, photographs of presacral vertebra NHMW 2019/0058/0003 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views; F-J, photographs of presacral vertebra NHMW 2019/0058/0014 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views; K-O, photographs of presacral vertebra NHMW 2019/0058/0045 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of presacral vertebra NHMW 2019/0058/0004 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views; U-Y, photographs of presacral vertebra NHMW 2019/0058/0020 in anterior (U), posterior (V), dorsal (W), ventral (X), and right lateral (Y) views; Z-D’, photographs of presacral vertebra NHMW 2019/0058/0012 in anterior (Z), posterior (A’), dorsal (B’), ventral (C’), and left lateral (D’) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720935" figureDoi-1="http://doi.org/10.5281/zenodo.4720937" figureDoi-2="http://doi.org/10.5281/zenodo.4720941" figureDoi-3="http://doi.org/10.5281/zenodo.4720945" httpUri-0="https://zenodo.org/record/4720935/files/figure.png" httpUri-1="https://zenodo.org/record/4720937/files/figure.png" httpUri-2="https://zenodo.org/record/4720941/files/figure.png" httpUri-3="https://zenodo.org/record/4720945/files/figure.png" pageId="45" pageNumber="262">Figs 50-53</figureCitation>
). In
</paragraph>
<caption id="DF7782A5FF8DFFEF30BBFCC9F8DC3B94" ID-DOI="http://doi.org/10.5281/zenodo.4720901" ID-Zenodo-Dep="4720901" httpUri="https://zenodo.org/record/4720901/files/figure.png" pageId="46" pageNumber="263" startId="46.[133,143,881,898]" targetBox="[360,1236,153,859]" targetPageId="46">
<paragraph id="8BB7D22DFF8DFFEF30BBFCC9F8DC3B94" blockId="46.[132,1455,881,921]" pageId="46" pageNumber="263">
FIG. 40. —
<taxonomicName id="4C08A9AEFF8DFFEF30E0FCC9FB903B8F" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[222,462,881,898]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="263" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8DFFEF30E0FCC9FB903B8F" box="[222,462,881,898]" italics="true" pageId="46" pageNumber="263">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8DFFEF31EDFCC9F85D3B8F" box="[467,515,881,898]" pageId="46" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
Close up photograph of the sculpturing pattern of the holotype parietal NHMW 2019/0047/0001 and the crenula- tion of its dorsolateral crests in dorsal view. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF7782A5FF8DFFEF30BAF998FF703E45" ID-DOI="http://doi.org/10.5281/zenodo.4720903" ID-Zenodo-Dep="4720903" httpUri="https://zenodo.org/record/4720903/files/figure.png" pageId="46" pageNumber="263" startId="46.[132,143,1568,1585]" targetBox="[264,1324,974,1525]" targetPageId="46">
<paragraph id="8BB7D22DFF8DFFEF30BAF998FF703E45" blockId="46.[132,1455,1568,1608]" pageId="46" pageNumber="263">
FIG. 41. —
<taxonomicName id="4C08A9AEFF8DFFEF30DFF998F85E3E3C" authority="Zittel, 1887" authorityName="Zittel" authorityYear="1887" box="[225,512,1568,1585]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="263" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF8DFFEF30DFF998FBC63E3C" box="[225,408,1568,1585]" italics="true" pageId="46" pageNumber="263">Palaeovaranus cayluxi</emphasis>
<bibRefCitation id="EF99AFDCFF8DFFEF319EF998F85E3E3C" author="ZITTEL K. A." box="[416,512,1568,1585]" pageId="46" pageNumber="263" refId="ref40223" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen, Leipzig, 900 p. https: // www. biodiversitylibrary. org / page / 40393265" type="book" year="1887">Zittel, 1887</bibRefCitation>
</taxonomicName>
-1890 and
<taxonomicName id="4C08A9AEFF8DFFEF3264F998F9153E3C" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[602,843,1568,1585]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="263" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8DFFEF3264F998F9153E3C" box="[602,843,1568,1585]" italics="true" pageId="46" pageNumber="263">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8DFFEF336DF998F9DA3E3C" box="[851,900,1568,1585]" pageId="46" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
Transverse sections of 3D models of parietals.
<emphasis id="B97C0E3FFF8DFFEF3528F998FF7D3E3C" bold="true" box="[1302,1315,1568,1585]" pageId="46" pageNumber="263">A</emphasis>
, parietal NHMW 2019/0048/0001 of
<taxonomicName id="4C08A9AEFF8DFFEF3116F98FFB803E45" authorityName="Zittel" authorityYear="1887" box="[296,478,1591,1608]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="263" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF8DFFEF3116F98FFB803E45" box="[296,478,1591,1608]" italics="true" pageId="46" pageNumber="263">Palaeovaranus cayluxi</emphasis>
</taxonomicName>
;
<emphasis id="B97C0E3FFF8DFFEF31D6F98FFBAB3E45" bold="true" box="[488,501,1591,1608]" pageId="46" pageNumber="263">B</emphasis>
, holotype parietal NHMW 2019/0047/0001 of
<taxonomicName id="4C08A9AEFF8DFFEF334BF98FFE3B3E45" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[885,1125,1591,1608]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="263" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8DFFEF334BF98FFE3B3E45" box="[885,1125,1591,1608]" italics="true" pageId="46" pageNumber="263">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8DFFEF3455F98FFEC53E45" box="[1131,1179,1591,1608]" pageId="46" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
Scale bar: 3 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFF8DFFEF30BAF92AFFEE3FE1" blockId="46.[131,777,1682,2028]" lastBlockId="46.[811,1457,1682,2028]" pageId="46" pageNumber="263">
certain large specimens (e.g., NHMW 2019/0058/0005) a distinct deep groove is clearly delimited between these two ridges. Subcentral foramina are usually present and can be, at certain specimens, rather prominent (e.g., NHMW 2019/0058/0001;
<figureCitation id="1333CEA8FF8DFFEF316FF8AAFB993F21" box="[337,455,1809,1836]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="55.[132,143,1481,1498]" captionStartId-1="56.[132,143,1040,1057]" captionStartId-2="57.[132,143,1262,1279]" captionStartId-3="58.[132,143,1894,1911]" captionTargetBox-0="[136,1452,214,1439]" captionTargetBox-1="[139,1449,216,996]" captionTargetBox-2="[132,1455,207,1225]" captionTargetBox-3="[136,1450,216,1841]" captionTargetId-0="figure-315@55.[132,1455,209,1447]" captionTargetId-1="figure-573@56.[132,1455,201,1002]" captionTargetId-2="figure-386@57.[132,1455,207,1225]" captionTargetId-3="figure-15@58.[132,1455,211,1845]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionTargetPageId-3="58" captionText-0="FIG. 50. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), dorsal (C), right lateral (D), left lateral (E), and ventral (F) views. Scale bar: 2 mm." captionText-1="FIG. 51. — Palaeovaranus sp. Virtual 3D models of presacral vertebra NHMW 2019/0058/0001 in anterior (A), posterior (B), right lateral (C), left lateral (D), dorsal (E), ventral (F), ventrolateral (G), and anterolateral (H) views. Scale bar: 5 mm." captionText-2="FIG. 52. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0002 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar: 2 mm." captionText-3="FIG. 53. — Palaeovaranus sp. A-E, photographs of presacral vertebra NHMW 2019/0058/0003 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views; F-J, photographs of presacral vertebra NHMW 2019/0058/0014 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views; K-O, photographs of presacral vertebra NHMW 2019/0058/0045 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of presacral vertebra NHMW 2019/0058/0004 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views; U-Y, photographs of presacral vertebra NHMW 2019/0058/0020 in anterior (U), posterior (V), dorsal (W), ventral (X), and right lateral (Y) views; Z-D’, photographs of presacral vertebra NHMW 2019/0058/0012 in anterior (Z), posterior (A’), dorsal (B’), ventral (C’), and left lateral (D’) views. Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720935" figureDoi-1="http://doi.org/10.5281/zenodo.4720937" figureDoi-2="http://doi.org/10.5281/zenodo.4720941" figureDoi-3="http://doi.org/10.5281/zenodo.4720945" httpUri-0="https://zenodo.org/record/4720935/files/figure.png" httpUri-1="https://zenodo.org/record/4720937/files/figure.png" httpUri-2="https://zenodo.org/record/4720941/files/figure.png" httpUri-3="https://zenodo.org/record/4720945/files/figure.png" pageId="46" pageNumber="263">Figs 50-53</figureCitation>
). The neural spine is high in lateral view and is posteriorly inclined. The neural spine mostly develops and augments in height in the posterior half of the neural arch, but nevertheless, its base crosses, in all specimens, the entire midline of the neural arch, in the shape of longitudinal ridge. In certain large specimens, a distinct dorsal thickening of the neural spine is obvious (e.g., NHMW 2019/0058/0001; NHMW 2019/0058/0003; NHMW 2019/0067/0001). The neural arch is vaulted (or at least moderately vaulted) in all specimens. Foramina on the neural spine can be present (e.g., NHMW 2019/0058/0001). The cotyle is rather depressed; in fact, it appears to be exceedingly depressed in certain small-sized specimens (e.g., NHMW 2019/0058/0002;
<figureCitation id="1333CEA8FF8DFFEF33C7F8E9FE073F61" box="[1017,1113,1873,1900]" captionStart="FIG" captionStartId="57.[132,143,1262,1279]" captionTargetBox="[132,1455,207,1225]" captionTargetId="figure-386@57.[132,1455,207,1225]" captionTargetPageId="57" captionText="FIG. 52. — Palaeovaranus sp. Photographs of presacral vertebra NHMW 2019/0058/0002 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720941" httpUri="https://zenodo.org/record/4720941/files/figure.png" pageId="46" pageNumber="263">Fig. 52A</figureCitation>
) but also in certain rather large ones (e.g., NHMW 2019/0058/0003;
<figureCitation id="1333CEA8FF8DFFEF34EFF8C9FF6A3F81" box="[1233,1332,1905,1932]" captionStart="FIG" captionStartId="58.[132,143,1894,1911]" captionTargetBox="[136,1450,216,1841]" captionTargetId="figure-15@58.[132,1455,211,1845]" captionTargetPageId="58" captionText="FIG. 53. — Palaeovaranus sp. A-E, photographs of presacral vertebra NHMW 2019/0058/0003 in anterior (A), posterior (B), dorsal (C), ventral (D), and right lateral (E) views; F-J, photographs of presacral vertebra NHMW 2019/0058/0014 in anterior (F), posterior (G), dorsal (H), ventral (I), and left lateral (J) views; K-O, photographs of presacral vertebra NHMW 2019/0058/0045 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of presacral vertebra NHMW 2019/0058/0004 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views; U-Y, photographs of presacral vertebra NHMW 2019/0058/0020 in anterior (U), posterior (V), dorsal (W), ventral (X), and right lateral (Y) views; Z-D’, photographs of presacral vertebra NHMW 2019/0058/0012 in anterior (Z), posterior (A’), dorsal (B’), ventral (C’), and left lateral (D’) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720945" httpUri="https://zenodo.org/record/4720945/files/figure.png" pageId="46" pageNumber="263">Fig. 53A</figureCitation>
). In certain specimens, the cotyle is rather anteroventrally directed, so that its dorsal level can be observed even in ventral view of the vertebra (e.g., NHMW 2019/0058/0005; NHMW
</paragraph>
<caption id="DF7782A5FF8CFFEE30BAF938FB873EB3" ID-DOI="http://doi.org/10.5281/zenodo.4720907" ID-Zenodo-Dep="4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="47" pageNumber="264" startId="47.[132,143,1664,1681]" targetBox="[122,1466,175,1637]" targetPageId="47">
<paragraph id="8BB7D22DFF8CFFEE30BAF938FB873EB3" blockId="47.[132,1457,1664,1726]" pageId="47" pageNumber="264">
FIG. 42. —
<taxonomicName id="4C08A9AEFF8CFFEE30E3F938FB923E9C" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[221,460,1664,1681]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="47" pageNumber="264" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF8CFFEE30E3F938FB923E9C" box="[221,460,1664,1681]" italics="true" pageId="47" pageNumber="264">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF8CFFEE31EEF938FBA13E9C" box="[464,511,1664,1681]" pageId="47" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
Parietals of juvenile individuals.
<emphasis id="B97C0E3FFF8CFFEE3339F938F94A3E9C" bold="true" box="[775,788,1664,1681]" pageId="47" pageNumber="264">A</emphasis>
,
<emphasis id="B97C0E3FFF8CFFEE3323F938F9743E9C" bold="true" box="[797,810,1664,1681]" pageId="47" pageNumber="264">B</emphasis>
, photographs of parietal MNHN.F.QU17177 in dorsal (
<emphasis id="B97C0E3FFF8CFFEE34D4F938FEA93E9C" bold="true" box="[1258,1271,1664,1681]" pageId="47" pageNumber="264">A</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF8CFFEE355BF938FF2C3E9C" bold="true" box="[1381,1394,1664,1681]" pageId="47" pageNumber="264">B</emphasis>
) views;
<emphasis id="B97C0E3FFF8CFFEE30BAF92FFACC3EA5" bold="true" box="[132,146,1687,1704]" pageId="47" pageNumber="264">C</emphasis>
,
<emphasis id="B97C0E3FFF8CFFEE30A3F92FFAF53EA5" bold="true" box="[157,171,1687,1704]" pageId="47" pageNumber="264">D</emphasis>
, photographs of parietal UM BFI 1873 in dorsal (
<emphasis id="B97C0E3FFF8CFFEE3278F92FF80A3EA5" bold="true" box="[582,596,1687,1704]" pageId="47" pageNumber="264">C</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF8CFFEE32F8F92FF88A3EA5" bold="true" box="[710,724,1687,1704]" pageId="47" pageNumber="264">D</emphasis>
) views. Both these specimens originally appeared as drawings in
<bibRefCitation id="EF99AFDCFF8CFFEE34C3F92FFF023EA5" author="AUGE M." box="[1277,1372,1687,1704]" pageId="47" pageNumber="264" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé (2005</bibRefCitation>
: Figs 194 and 195 respectively). Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFF8CFFEE30BAF94AF9583F66" blockId="47.[131,777,1778,1900]" pageId="47" pageNumber="264">2019/0058/0006), a condition reminiscent of varanids. In certain specimens (NHMW 2019/0058/0001; NHMW 2019/0058/0005), distinct anocotylar foramina are present above the cotyle. The synapophyses are massive and elongated.</paragraph>
</subSection>
<subSection id="E287CBC6FF8CFFF030BAF829FBF93F61" lastPageId="49" lastPageNumber="266" pageId="47" pageNumber="264" type="multiple">
<paragraph id="8BB7D22DFF8CFFEE30BAF829FBEE3FA1" blockId="47.[132,432,1937,1964]" box="[132,432,1937,1964]" pageId="47" pageNumber="264">
<heading id="D0FF6541FF8CFFEE30BAF829FBEE3FA1" box="[132,432,1937,1964]" fontSize="11" level="2" pageId="47" pageNumber="264" reason="8">
<emphasis id="B97C0E3FFF8CFFEE30BAF829FBEE3FA1" box="[132,432,1937,1964]" italics="true" pageId="47" pageNumber="264">
Sacral vertebrae (
<figureCitation id="1333CEA8FF8CFFEE3109F82AFBF93FA6" box="[311,423,1937,1963]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="59.[145,156,838,855]" captionStartId-1="60.[132,143,1441,1458]" captionStartId-2="61.[132,143,1598,1615]" captionTargetBox-0="[132,1455,211,801]" captionTargetBox-1="[140,1445,215,1398]" captionTargetBox-2="[132,1455,211,1558]" captionTargetId-0="figure-693@59.[132,1455,211,801]" captionTargetId-1="figure-317@60.[132,1455,205,1403]" captionTargetId-2="figure-210@61.[132,1455,211,1558]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionTargetPageId-2="61" captionText-0="FIG. 54. — Palaeovaranus sp. Photographs of two articulated sacral vertebrae NHMW 2019/0058/0028 in right (A) and left lateral (B) views. Scale bar: 2 mm." captionText-1="FIG. 55. — Palaeovaranus sp. Virtual 3D models of two articulated sacral vertebrae NHMW 2019/0058/0028 in anterior (A), posterior (B), anterolateral (C), dorsal (D), ventral (E), ventrolateral (F), right lateral (G), left lateral (H), and posteroventral (I) views. Scale bar: 5 mm." captionText-2="FIG. 56. — Palaeovaranus sp. A-F, photographs of sacral vertebra NHMW 2019/0058/0029 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views; G-K, photographs of sacral vertebra NHMW 2019/0058/0047 in anterior (G), posterior (H), dorsal (I), ventral (J), and left lateral (K) views. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720949" figureDoi-1="http://doi.org/10.5281/zenodo.4720953" figureDoi-2="http://doi.org/10.5281/zenodo.4720957" httpUri-0="https://zenodo.org/record/4720949/files/figure.png" httpUri-1="https://zenodo.org/record/4720953/files/figure.png" httpUri-2="https://zenodo.org/record/4720957/files/figure.png" pageId="47" pageNumber="264">Figs 54-56</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7D22DFF8CFFEE30BAF80AF9583FE6" blockId="47.[132,774,1969,2028]" pageId="47" pageNumber="264">Two of these specimens (one isolated [NHMW 2019/0058/0029] and two articulated sacral vertebrae</paragraph>
<paragraph id="8BB7D22DFF8CFFF03313F94AFBF93F61" blockId="47.[811,1455,1777,2028]" lastBlockId="49.[132,777,1523,1900]" lastPageId="49" lastPageNumber="266" pageId="47" pageNumber="264">
[NHMW 2019/0058/0028]) are rather large in size (
<figureCitation id="1333CEA8FF8CFFEE3562F94AFFF63F00" box="[1372,1448,1778,1805]" captionStart="FIG" captionStartId="59.[145,156,838,855]" captionTargetBox="[132,1455,211,801]" captionTargetId="figure-693@59.[132,1455,211,801]" captionTargetPageId="59" captionText="FIG. 54. — Palaeovaranus sp. Photographs of two articulated sacral vertebrae NHMW 2019/0058/0028 in right (A) and left lateral (B) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720949" httpUri="https://zenodo.org/record/4720949/files/figure.png" pageId="47" pageNumber="264">Figs 54</figureCitation>
;
<figureCitation id="1333CEA8FF8CFFEE3313F8A9F9143F21" box="[813,842,1809,1836]" captionStart="FIG" captionStartId="60.[132,143,1441,1458]" captionTargetBox="[140,1445,215,1398]" captionTargetId="figure-317@60.[132,1455,205,1403]" captionTargetPageId="60" captionText="FIG. 55. — Palaeovaranus sp. Virtual 3D models of two articulated sacral vertebrae NHMW 2019/0058/0028 in anterior (A), posterior (B), anterolateral (C), dorsal (D), ventral (E), ventrolateral (F), right lateral (G), left lateral (H), and posteroventral (I) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720953" httpUri="https://zenodo.org/record/4720953/files/figure.png" pageId="47" pageNumber="264">55</figureCitation>
;
<figureCitation id="1333CEA8FF8CFFEE3362F8A9F9273F21" box="[860,889,1809,1836]" captionStart="FIG" captionStartId="61.[132,143,1598,1615]" captionTargetBox="[132,1455,211,1558]" captionTargetId="figure-210@61.[132,1455,211,1558]" captionTargetPageId="61" captionText="FIG. 56. — Palaeovaranus sp. A-F, photographs of sacral vertebra NHMW 2019/0058/0029 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views; G-K, photographs of sacral vertebra NHMW 2019/0058/0047 in anterior (G), posterior (H), dorsal (I), ventral (J), and left lateral (K) views. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720957" httpUri="https://zenodo.org/record/4720957/files/figure.png" pageId="47" pageNumber="264">56</figureCitation>
A-F), especially the latter one, which exceeds in centrum length all the above described presacral vertebrae (see
<figureCitation id="1333CEA8FF8CFFEE335FF8E9F9B83F61" box="[865,998,1873,1900]" captionStart="APPENDIX" captionStartId="75.[427,439,219,236]" captionTargetPageId="75" captionText="APPENDIX 1. — Centrum lengths of all lizard vertebrae from the NHMW Quercy collection." pageId="47" pageNumber="264">Appendix 1</figureCitation>
). The much smaller specimen NHMW 2019/0058/0047 is almost identical with the two other specimens, with their only major difference being this size contrast (
<figureCitation id="1333CEA8FF8CFFEE33A2F809F9B13FC1" box="[924,1007,1969,1996]" captionStart="FIG" captionStartId="61.[132,143,1598,1615]" captionTargetBox="[132,1455,211,1558]" captionTargetId="figure-210@61.[132,1455,211,1558]" captionTargetPageId="61" captionText="FIG. 56. — Palaeovaranus sp. A-F, photographs of sacral vertebra NHMW 2019/0058/0029 in anterior (A), posterior (B), right lateral (C), dorsal (D), ventral (E), and left lateral (F) views; G-K, photographs of sacral vertebra NHMW 2019/0058/0047 in anterior (G), posterior (H), dorsal (I), ventral (J), and left lateral (K) views. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720957" httpUri="https://zenodo.org/record/4720957/files/figure.png" pageId="47" pageNumber="264">Fig. 56</figureCitation>
G-J); apparently it pertains to a much smaller (?younger) individual, however, any taxonomic implication for this size deviation cannot be excluded. Similarly to the presacral vertebrae, there are also in these specimens two distinct ridges in the ventral surface of the centrum, that commence at around the level of the cotyle and terminate slightly prior to the condyle. The centrum is convex in lateral view, with the ventral levels of both cotyle and condyle situated below the level of the ventral surface of the centrum. The neural arch is relatively vaulted. The neural spine develops mostly at the posterior half of the neural arch. The cotyle and condyle are depressed. “Pseudozygosphenes” and “pseudozygantra” are visible. The pleurapophyses extend much laterally and their distal termination is rather broad, so that in the two large articulated vertebrae (NHMW 2019/0058/0028), these elements from both vertebrae are united to each other (
<figureCitation id="1333CEA8FF92FFF030B0F96AFA823EE0" box="[142,220,1746,1773]" captionStart="FIG" captionStartId="60.[132,143,1441,1458]" captionTargetBox="[140,1445,215,1398]" captionTargetId="figure-317@60.[132,1455,205,1403]" captionTargetPageId="60" captionText="FIG. 55. — Palaeovaranus sp. Virtual 3D models of two articulated sacral vertebrae NHMW 2019/0058/0028 in anterior (A), posterior (B), anterolateral (C), dorsal (D), ventral (E), ventrolateral (F), right lateral (G), left lateral (H), and posteroventral (I) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720953" httpUri="https://zenodo.org/record/4720953/files/figure.png" pageId="49" pageNumber="266">Fig. 55</figureCitation>
D-F). A large fenestra is being formed by the unification of pleurapophyses of succeeding vertebrae. Also in this larger specimen, two distinct foramina are present below the neural arch in posterior view, inside the “pseudozygantrum” (
<figureCitation id="1333CEA8FF92FFF0310BF8E9FBC43F61" box="[309,410,1873,1900]" captionStart="FIG" captionStartId="60.[132,143,1441,1458]" captionTargetBox="[140,1445,215,1398]" captionTargetId="figure-317@60.[132,1455,205,1403]" captionTargetPageId="60" captionText="FIG. 55. — Palaeovaranus sp. Virtual 3D models of two articulated sacral vertebrae NHMW 2019/0058/0028 in anterior (A), posterior (B), anterolateral (C), dorsal (D), ventral (E), ventrolateral (F), right lateral (G), left lateral (H), and posteroventral (I) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720953" httpUri="https://zenodo.org/record/4720953/files/figure.png" pageId="49" pageNumber="266">Fig. 55B</figureCitation>
).
</paragraph>
</subSection>
<caption id="DF7782A5FF93FFF130BAF8A5FFEF3F48" ID-DOI="http://doi.org/10.5281/zenodo.4720911" ID-Zenodo-Dep="4720911" httpUri="https://zenodo.org/record/4720911/files/figure.png" pageId="48" pageNumber="265" startId="48.[132,142,1821,1838]" targetBox="[263,1345,214,1779]" targetPageId="48">
<paragraph id="8BB7D22DFF93FFF130BAF8A5FFEF3F48" blockId="48.[132,1457,1821,1861]" pageId="48" pageNumber="265">
FIG. 43. —
<taxonomicName id="4C08A9AEFF93FFF130E0F8A5FB923F23" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[222,460,1821,1838]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="265" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF93FFF130E0F8A5FB923F23" box="[222,460,1821,1838]" italics="true" pageId="48" pageNumber="265">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF93FFF131EDF8A5F85D3F23" box="[467,515,1821,1838]" pageId="48" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
frontals:
<emphasis id="B97C0E3FFF93FFF1326EF8A5F8033F23" bold="true" box="[592,605,1821,1838]" pageId="48" pageNumber="265">A</emphasis>
,
<emphasis id="B97C0E3FFF93FFF13256F8A5F82B3F23" bold="true" box="[616,629,1821,1838]" pageId="48" pageNumber="265">B</emphasis>
, photographs of frontal UM PRA 8 in dorsal (
<emphasis id="B97C0E3FFF93FFF133CCF8A5F9A13F23" bold="true" box="[1010,1023,1821,1838]" pageId="48" pageNumber="265">A</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF93FFF1344FF8A5FE203F23" bold="true" box="[1137,1150,1821,1838]" pageId="48" pageNumber="265">B</emphasis>
) views;
<emphasis id="B97C0E3FFF93FFF134FDF8A5FE8F3F23" bold="true" box="[1219,1233,1821,1838]" pageId="48" pageNumber="265">C</emphasis>
,
<emphasis id="B97C0E3FFF93FFF134E2F8A5FEB43F23" bold="true" box="[1244,1258,1821,1838]" pageId="48" pageNumber="265">D</emphasis>
, photographs of frontal MNHN.F.QU17175 in dorsal (
<emphasis id="B97C0E3FFF93FFF13158F88CFB2D3F48" bold="true" box="[358,371,1844,1861]" pageId="48" pageNumber="265">C</emphasis>
) and ventral (
<emphasis id="B97C0E3FFF93FFF131E6F88CFBBB3F48" bold="true" box="[472,485,1844,1861]" pageId="48" pageNumber="265">D</emphasis>
) views.Both these specimens originally appeared as drawings in
<bibRefCitation id="EF99AFDCFF93FFF133E6F88CFE6F3F48" author="AUGE M." box="[984,1073,1844,1861]" pageId="48" pageNumber="265" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé (2005</bibRefCitation>
: figs196 and 189 respectively).Scale bars:5 mm.
</paragraph>
</caption>
<caption id="DF7782A5FF92FFF030BDFA2DF97B3DB1" ID-DOI="http://doi.org/10.5281/zenodo.4720915" ID-Zenodo-Dep="4720915" httpUri="https://zenodo.org/record/4720915/files/figure.png" pageId="49" pageNumber="266" startId="49.[131,142,1429,1446]" targetBox="[139,1448,214,1385]" targetPageId="49">
<paragraph id="8BB7D22DFF92FFF030BDFA2DF97B3DB1" blockId="49.[131,1455,1429,1468]" pageId="49" pageNumber="266">
FIG. 44. —
<taxonomicName id="4C08A9AEFF92FFF030E0FA2DFBD63DAB" box="[222,392,1429,1446]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="266" phylum="Chordata" rank="species" species="undefined-a">
<emphasis id="B97C0E3FFF92FFF030E0FA2DFB093DAB" box="[222,343,1429,1446]" italics="true" pageId="49" pageNumber="266">Palaeovaranus</emphasis>
sp.
<emphasis id="B97C0E3FFF92FFF03145FA2DFBD63DAB" bold="true" box="[379,392,1429,1446]" pageId="49" pageNumber="266">A</emphasis>
</taxonomicName>
<emphasis id="B97C0E3FFF92FFF031B6FA2DFBC33DAB" bold="true" box="[392,413,1429,1446]" pageId="49" pageNumber="266">-C</emphasis>
, photographs of left maxilla NHMW 2019/0048/0002 in lateral (
<emphasis id="B97C0E3FFF92FFF03394FA2DF9E93DAB" bold="true" box="[938,951,1429,1446]" pageId="49" pageNumber="266">A</emphasis>
), medial (
<emphasis id="B97C0E3FFF92FFF03439FA2DFE4A3DAB" bold="true" box="[1031,1044,1429,1446]" pageId="49" pageNumber="266">B</emphasis>
), and dorsal (
<emphasis id="B97C0E3FFF92FFF034BBFA2DFECD3DAB" bold="true" box="[1157,1171,1429,1446]" pageId="49" pageNumber="266">C</emphasis>
) views; D, close up of the posteriormost preserved tooth of the same specimen. Scale bars: A-C, 5 mm; D, 1 mm.
</paragraph>
</caption>
<subSection id="E287CBC6FF92FFF230BAF829F8363DA3" lastPageId="51" lastPageNumber="268" pageId="49" pageNumber="266" type="multiple">
<paragraph id="8BB7D22DFF92FFF030BAF829FB8C3FA1" blockId="49.[132,466,1937,1964]" box="[132,466,1937,1964]" pageId="49" pageNumber="266">
<heading id="D0FF6541FF92FFF030BAF829FB8C3FA1" box="[132,466,1937,1964]" fontSize="11" level="2" pageId="49" pageNumber="266" reason="8">
<emphasis id="B97C0E3FFF92FFF030BAF829FB8C3FA1" box="[132,466,1937,1964]" italics="true" pageId="49" pageNumber="266">
Caudal vertebrae (
<figureCitation id="1333CEA8FF92FFF03170F82AFBC33FA1" box="[334,413,1938,1964]" captionStart="FIG" captionStartId="62.[132,143,1488,1505]" captionTargetBox="[136,1447,217,1443]" captionTargetId="figure-279@62.[132,1455,215,1448]" captionTargetPageId="62" captionText="FIG. 57. — Palaeovaranus sp. Photographs of caudal vertebra NHMW 2019/0058/0035 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F), views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720961" httpUri="https://zenodo.org/record/4720961/files/figure.png" pageId="49" pageNumber="266">Figs 57</figureCitation>
;
<figureCitation id="1333CEA8FF92FFF03192F829FB973FA6" box="[428,457,1937,1963]" captionStart="FIG" captionStartId="63.[132,143,1940,1957]" captionTargetBox="[135,1452,217,1871]" captionTargetId="figure-16@63.[132,1455,215,1878]" captionTargetPageId="63" captionText="FIG. 58. — Palaeovaranus sp. A-E, photographs of caudal vertebra NHMW 2019/0058/0036 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views;F-J, photographs of caudal vertebra NHMW 2019/0058/0037 in anterior (F), posterior (G), dorsal (H), ventral (I), and right lateral (J) views;K-O, photographs of caudal vertebra NHMW 2019/0058/0048 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of caudal vertebra NHMW 2019/0058/0049 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720965" httpUri="https://zenodo.org/record/4720965/files/figure.png" pageId="49" pageNumber="266">58</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7D22DFF92FFF030BAF809F95B3FE6" blockId="49.[132,775,1969,2028]" pageId="49" pageNumber="266">
These are extremely elongated and rather slender (
<figureCitation id="1333CEA8FF92FFF032B8F809F88E3FC1" box="[646,720,1969,1996]" captionStart="FIG" captionStartId="62.[132,143,1488,1505]" captionTargetBox="[136,1447,217,1443]" captionTargetId="figure-279@62.[132,1455,215,1448]" captionTargetPageId="62" captionText="FIG. 57. — Palaeovaranus sp. Photographs of caudal vertebra NHMW 2019/0058/0035 in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F), views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720961" httpUri="https://zenodo.org/record/4720961/files/figure.png" pageId="49" pageNumber="266">Figs 57</figureCitation>
;
<figureCitation id="1333CEA8FF92FFF032E2F809F8A93FC1" box="[732,759,1969,1996]" captionStart="FIG" captionStartId="63.[132,143,1940,1957]" captionTargetBox="[135,1452,217,1871]" captionTargetId="figure-16@63.[132,1455,215,1878]" captionTargetPageId="63" captionText="FIG. 58. — Palaeovaranus sp. A-E, photographs of caudal vertebra NHMW 2019/0058/0036 in anterior (A), posterior (B), dorsal (C), ventral (D), and left lateral (E) views;F-J, photographs of caudal vertebra NHMW 2019/0058/0037 in anterior (F), posterior (G), dorsal (H), ventral (I), and right lateral (J) views;K-O, photographs of caudal vertebra NHMW 2019/0058/0048 in anterior (K), posterior (L), dorsal (M), ventral (N), and left lateral (O) views; P-T, photographs of caudal vertebra NHMW 2019/0058/0049 in anterior (P), posterior (Q), dorsal (R), ventral (S), and left lateral (T) views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4720965" httpUri="https://zenodo.org/record/4720965/files/figure.png" pageId="49" pageNumber="266">58</figureCitation>
), with the elongation becoming more prominent in the
</paragraph>
<paragraph id="8BB7D22DFF92FFF03313FA4BFFEE3FE6" blockId="49.[813,1457,1522,2028]" pageId="49" pageNumber="266">
slenderer specimens (e.g., NHMW 2019/0058/0031; NHMW 2019/0058/0033), whereas the more robust ones tend to have a smaller centrum length but nevertheless wider vertebral centra (NHMW 2019/0058/0035; NHMW 2019/0058/0036; NHMW 2019/0058/0048). Their centrum lengths vary between 8.9 and
<quantity id="4CF07FC8FF92FFF0357AF92AFFF13EA0" box="[1348,1455,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="49" pageNumber="266" unit="mm" value="15.9">15.9 mm</quantity>
(see
<figureCitation id="1333CEA8FF92FFF0335CF90AF9B73EC1" box="[866,1001,1714,1740]" captionStart="APPENDIX" captionStartId="75.[427,439,219,236]" captionTargetPageId="75" captionText="APPENDIX 1. — Centrum lengths of all lizard vertebrae from the NHMW Quercy collection." pageId="49" pageNumber="266">Appendix 1</figureCitation>
). The neural spine, when preserved, is rather high in lateral view, it augments in height rapidly but only at the posterior portion of the neural arch, and is inclined posteriorly (e.g., NHMW 2019/0058/0035; NHMW 2019/0058/0036). Nevertheless, the base of the neural spine extends much anteriorly in dorsal view, in the shape of a thin longitudinal ridge. The cotyle is large and elliptical. The prezygapophyses are dorsally inclined. The postzygapophyses are rather small. The pedicles for articulation with the chevrons are rather thick in the robust
</paragraph>
<caption id="DF7782A5FF91FFF330BAF879FE343FE4" ID-DOI="http://doi.org/10.5281/zenodo.4720921" ID-Zenodo-Dep="4720921" httpUri="https://zenodo.org/record/4720921/files/figure.png" pageId="50" pageNumber="267" startId="50.[132,143,1985,2002]" targetBox="[147,1438,219,1839]" targetPageId="50">
<paragraph id="8BB7D22DFF91FFF330BAF879FE343FE4" blockId="50.[132,1456,1985,2025]" pageId="50" pageNumber="267">
FIG. 45. —
<taxonomicName id="4C08A9AEFF91FFF330E1F879FBD73FDF" box="[223,393,1985,2002]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="50" pageNumber="267" phylum="Chordata" rank="species" species="undefined-a">
<emphasis id="B97C0E3FFF91FFF330E1F879FB063FDF" box="[223,344,1985,2002]" italics="true" pageId="50" pageNumber="267">Palaeovaranus</emphasis>
sp.
<emphasis id="B97C0E3FFF91FFF33142F879FBD73FDF" bold="true" box="[380,393,1985,2002]" pageId="50" pageNumber="267">A</emphasis>
</taxonomicName>
<emphasis id="B97C0E3FFF91FFF331B7F879FBC03FDF" bold="true" box="[393,414,1985,2002]" pageId="50" pageNumber="267">-G</emphasis>
, virtual 3D models of left maxilla NHMW 2019/0048/0002 in lateral (
<emphasis id="B97C0E3FFF91FFF333EAF879F9BF3FDF" bold="true" box="[980,993,1985,2002]" pageId="50" pageNumber="267">A</emphasis>
), medial (
<emphasis id="B97C0E3FFF91FFF3340CF879FE613FDF" bold="true" box="[1074,1087,1985,2002]" pageId="50" pageNumber="267">B</emphasis>
), posteromedial (
<emphasis id="B97C0E3FFF91FFF334EEF879FE803FDF" bold="true" box="[1232,1246,1985,2002]" pageId="50" pageNumber="267">C</emphasis>
), dorsomedial (
<emphasis id="B97C0E3FFF91FFF33560F879FF323FDF" bold="true" box="[1374,1388,1985,2002]" pageId="50" pageNumber="267">D</emphasis>
), dorsal (
<emphasis id="B97C0E3FFF91FFF330B7F860FACB3FE4" bold="true" box="[137,149,2008,2025]" pageId="50" pageNumber="267">E</emphasis>
) and posterior (
<emphasis id="B97C0E3FFF91FFF33126F860FB7D3FE4" bold="true" box="[280,291,2008,2025]" pageId="50" pageNumber="267">F</emphasis>
) views;
<emphasis id="B97C0E3FFF91FFF33159F860FB2B3FE4" bold="true" box="[359,373,2008,2025]" pageId="50" pageNumber="267">G</emphasis>
, close up view of posteriormost preserved tooth of the same specimen. Scale bars: 5 mm.
</paragraph>
</caption>
<caption id="DF7782A5FF90FFF230BAFC25FFEF3BA3" ID-DOI="http://doi.org/10.5281/zenodo.4720923" ID-Zenodo-Dep="4720923" box="[132,1457,925,942]" httpUri="https://zenodo.org/record/4720923/files/figure.png" pageId="51" pageNumber="268" startId="51.[132,143,925,942]" targetBox="[132,1455,203,895]" targetPageId="51">
<paragraph id="8BB7D22DFF90FFF230BAFC25FFEF3BA3" blockId="51.[132,1457,925,942]" box="[132,1457,925,942]" pageId="51" pageNumber="268">
FIG. 46. —
<taxonomicName id="4C08A9AEFF90FFF230E0FC25FB293BA3" box="[222,375,925,942]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B97C0E3FFF90FFF230E0FC25FB093BA3" box="[222,343,925,942]" italics="true" pageId="51" pageNumber="268">Palaeovaranus</emphasis>
sp.
</taxonomicName>
Photographs of left dentary NHMW 2019/0058/0054 in lateral (
<emphasis id="B97C0E3FFF90FFF233BAFC25F9CF3BA3" bold="true" box="[900,913,925,942]" pageId="51" pageNumber="268">A</emphasis>
), dorsal (
<emphasis id="B97C0E3FFF90FFF233E3FC25F9B43BA3" bold="true" box="[989,1002,925,942]" pageId="51" pageNumber="268">B</emphasis>
), medial (
<emphasis id="B97C0E3FFF90FFF23405FC25FE173BA3" bold="true" box="[1083,1097,925,942]" pageId="51" pageNumber="268">C</emphasis>
), and posterior (
<emphasis id="B97C0E3FFF90FFF234EEFC25FE803BA3" bold="true" box="[1232,1246,925,942]" pageId="51" pageNumber="268">D</emphasis>
) views. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFF90FFF230BAFBACF8363DA3" blockId="51.[131,777,1044,1454]" pageId="51" pageNumber="268">
vertebrae and are located rather close to the condyle. A longitudinal groove runs throughout the centrum, originating posteriorly between the two pedicles and reaching anteriorly prior to the level of the cotyle. In the most robust specimens, this groove becomes rather thin in the anterior half of the centrum, while it is almost constantly wide in the slender, elongated specimens. There are no signs of autotomic septa. The transverse processes differ in size. In the most robust specimens, they occupy a larger surface at the lateral sides of the centrum, whereas in the more slender ones, they are rather thin. They extend laterally. In several specimens, there are “pseudozygosphenes” and “pseudozygantra” (sensu
<bibRefCitation id="EF99AFDCFF90FFF231A2FA2BF8063DA0" author="HOFFSTETTER R." box="[412,600,1427,1453]" pageId="51" pageNumber="268" pagination="1051 - 1064" refId="ref37022" refString="HOFFSTETTER R. 1969. - Presence de Varanidae (Reptilia, Sauria) dans le Miocene de Catalogne. Considerations sur l'histoire de la famille. Bulletin du Museum national d'Histoire naturelle, 2 eme serie, 40: 1051 - 1064. https: // www. biodiversitylibrary. org / page / 55344240" type="journal article" year="1969">Hoffstetter 1969</bibRefCitation>
).
</paragraph>
</subSection>
</subSubSection>
<subSubSection id="C31281A6FF90FFF530BAFA6BF9563D80" lastPageId="52" lastPageNumber="269" pageId="51" pageNumber="268" type="discussion">
<paragraph id="8BB7D22DFF90FFF230BAFA6BFAB63DE6" blockId="51.[130,776,1491,2028]" box="[132,232,1491,1517]" pageId="51" pageNumber="268">REMARKS</paragraph>
<paragraph id="8BB7D22DFF90FFF230BAFA4BFBB73FE1" blockId="51.[130,776,1491,2028]" pageId="51" pageNumber="268">
The distinctly developed nasal crest on the dorsomedial surface of the nasal process of maxilla represents the most distinctive maxillary structure of
<taxonomicName id="4C08A9AEFF90FFF23152F98AFBA13E41" authorityName="Zittel" authorityYear="1887" box="[364,511,1586,1612]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF90FFF23152F98AFBA13E41" box="[364,511,1586,1612]" italics="true" pageId="51" pageNumber="268">Palaeovaranus</emphasis>
</taxonomicName>
. This crest was not mentioned by
<bibRefCitation id="EF99AFDCFF90FFF230D5F9EAFB353E61" author="ZITTEL K. A." box="[235,363,1618,1644]" pageId="51" pageNumber="268" refId="ref40223" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen, Leipzig, 900 p. https: // www. biodiversitylibrary. org / page / 40393265" type="book" year="1887">Zittel (1887</bibRefCitation>
-1890) in the original, brief description of the
<typeStatus id="54B36C8FFF90FFF230F5F9CAFB743E81" box="[203,298,1650,1676]" pageId="51" pageNumber="268" type="holotype">holotype</typeStatus>
maxilla of
<taxonomicName id="4C08A9AEFF90FFF23197F9CAF8D23E81" authorityName="Zittel" authorityYear="1887" box="[425,652,1650,1676]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF90FFF23197F9CAF8D23E81" box="[425,652,1650,1676]" italics="true" pageId="51" pageNumber="268">Palaeovaranus cayluxi</emphasis>
</taxonomicName>
, but it was extensively described and discussed for the same specimen by
<bibRefCitation id="EF99AFDCFF90FFF2309AF90AFB083EC0" author="FEJERVARY DE G. J." box="[164,342,1714,1741]" pageId="51" pageNumber="268" pagination="1 - 130" refId="ref34838" refString="FEJERVARY DE G. J. 1935. - Further contributions to a monograph of the Megalanidae and fossil Varanidae, with notes on recent varanians. Annales historico-naturales Musei nationalis hungarici, Pars Zoologica 29: 1 - 130." type="journal article" year="1935">Fejérváry (1935)</bibRefCitation>
, and we here use mostly his terminology.
<bibRefCitation id="EF99AFDCFF90FFF230BDF96AFB663EE1" author="FEJERVARY DE G. J." box="[131,312,1746,1773]" pageId="51" pageNumber="268" pagination="1 - 130" refId="ref34838" refString="FEJERVARY DE G. J. 1935. - Further contributions to a monograph of the Megalanidae and fossil Varanidae, with notes on recent varanians. Annales historico-naturales Musei nationalis hungarici, Pars Zoologica 29: 1 - 130." type="journal article" year="1935">Fejérváry (1935)</bibRefCitation>
was able to confirm the morphology of its medial wall and the presence of this nasal crest and continuously highlighted this feature as principally differentiating this genus from
<taxonomicName id="4C08A9AEFF90FFF2313AF88AFB063F46" authorityName="Merrem" authorityYear="1820" box="[260,344,1842,1867]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="51" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF90FFF2313AF88AFB063F46" box="[260,344,1842,1867]" italics="true" pageId="51" pageNumber="268">Varanus</emphasis>
</taxonomicName>
. This crest is also present in the referred maxilla of
<taxonomicName id="4C08A9AEFF90FFF230D3F8E9FB973F66" authorityName="Zittel" authorityYear="1887" box="[237,457,1873,1899]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF90FFF230D3F8E9FB973F66" box="[237,457,1873,1899]" italics="true" pageId="51" pageNumber="268">Palaeovaranus cayluxi</emphasis>
</taxonomicName>
from Sainte Néboule, Quercy, described and figured by
<bibRefCitation id="EF99AFDCFF90FFF231AEF8CAF8463F81" author="RAGE J. - C." box="[400,536,1905,1932]" pageId="51" pageNumber="268" pagination="201 - 215" refId="ref38578" refString="RAGE J. - C. 1978. - Squamates, in GEZE B., RAGE J. - C., VERGNAUD- GRAZZINI F., DE BROIN F., BUFFETAUT E., MOURI- ER- CHAUVIRE C., CROCHET J. - Y., SIGE B., SUDRE J., REMY A., LANGEBADRE L., BONIS L. DE, HARTENBERGER J. L. &amp; VIANEY- LI- AUD M. (eds), La poche a Phosphate de Ste-Neboule (Lot) et sa faune de vertebres du Ludien superior. Palaeovertebrata 8: 201 - 215." type="journal article" year="1978">Rage (1978)</bibRefCitation>
. With the new photographs and the 3D models we provided above, this maxillary feature is evident and this enhances our understanding of this peculiar feature in
<taxonomicName id="4C08A9AEFF90FFF23171F869FBBC3FE6" authorityName="Zittel" authorityYear="1887" box="[335,482,2001,2027]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF90FFF23171F869FBBC3FE6" box="[335,482,2001,2027]" italics="true" pageId="51" pageNumber="268">Palaeovaranus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB7D22DFF90FFF2337DFBACFF103DE0" blockId="51.[811,1457,1044,2028]" pageId="51" pageNumber="268">
The fact that the external side of the maxilla is not smooth but rather appears to bear some kind of ornamentation could hint for a referral of this maxilla to
<taxonomicName id="4C08A9AEFF90FFF234AEFBECFFF33C63" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" box="[1168,1453,1108,1134]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF90FFF234AEFBECFFF33C63" box="[1168,1453,1108,1134]" italics="true" pageId="51" pageNumber="268">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF90FFF23315FBCDF93B3C82" box="[811,869,1141,1167]" pageId="51" pageNumber="268" rank="species">n. sp.</taxonomicNameLabel>
, the
<typeStatus id="54B36C8FFF90FFF233A9FBCCF9AB3C83" box="[919,1013,1140,1166]" pageId="51" pageNumber="268" type="holotype">holotype</typeStatus>
parietal of which is characterized by more prominent sculpturing in comparison with that of
<taxonomicName id="4C08A9AEFF90FFF2351CFB2CF92B3CC3" authorityName="Zittel" authorityYear="1887" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF90FFF2351CFB2CF92B3CC3" italics="true" pageId="51" pageNumber="268">Palaeovaranus cayluxi</emphasis>
</taxonomicName>
described above. Furthermore, the
<typeStatus id="54B36C8FFF90FFF234DBFB0CFF1D3CC3" box="[1253,1347,1204,1230]" pageId="51" pageNumber="268" type="holotype">holotype</typeStatus>
maxilla of
<taxonomicName id="4C08A9AEFF90FFF23315FB6CFE523CE3" authorityName="Zittel" authorityYear="1887" box="[811,1036,1236,1262]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="cayluxi">
<emphasis id="B97C0E3FFF90FFF23315FB6CFE523CE3" box="[811,1036,1236,1262]" italics="true" pageId="51" pageNumber="268">Palaeovaranus cayluxi</emphasis>
</taxonomicName>
does not bear any kind of ornamentation as it can be judged by
<bibRefCitation id="EF99AFDCFF90FFF23402FB4BFEA73D03" author="FEJERVARY DE G. J." box="[1084,1273,1267,1294]" pageId="51" pageNumber="268" pagination="1 - 130" refId="ref34838" refString="FEJERVARY DE G. J. 1935. - Further contributions to a monograph of the Megalanidae and fossil Varanidae, with notes on recent varanians. Annales historico-naturales Musei nationalis hungarici, Pars Zoologica 29: 1 - 130." type="journal article" year="1935">Fejérváry’s (1935)</bibRefCitation>
photographs and especially from his extremely detailed and careful descriptions of this specimen. Neither also exists any kind of ornamentation in the referred maxillary material from Sainte Néboule described and figured byRage (1978). Nevertheless, we avoid of formally assigning this maxilla to
<taxonomicName id="4C08A9AEFF90FFF234FEFA2BF9353DC0" authority="Georgalis &amp; Čerňanský &amp; Klembara, 2021" authorityName="Georgalis &amp; Čerňanský &amp; Klembara" authorityYear="2021" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
<emphasis id="B97C0E3FFF90FFF234FEFA2BF9353DC0" italics="true" pageId="51" pageNumber="268">Palaeovaranus lismonimenos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24FB344FF90FFF2334DFA0BF9EE3DC0" box="[883,944,1459,1485]" pageId="51" pageNumber="268" rank="species">n. sp.</taxonomicNameLabel>
, preferring the most conservative approach and determining the specimen solely to the genus level.
</paragraph>
<paragraph id="8BB7D22DFF90FFF2337DFA4BFF373F01" blockId="51.[811,1457,1044,2028]" pageId="51" pageNumber="268">
The dentaries of
<taxonomicName id="4C08A9AEFF90FFF233C4FA4BFECC3E00" authorityName="Zittel" authorityYear="1887" box="[1018,1170,1523,1549]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF90FFF233C4FA4BFECC3E00" box="[1018,1170,1523,1549]" italics="true" pageId="51" pageNumber="268">Palaeovaranus</emphasis>
</taxonomicName>
described herein show an array of sizes, general morphologies, and tooth shapes. They also seem different and more robust that other
<taxonomicName id="4C08A9AEFF90FFF23524F98AFFF33E41" authorityName="Zittel" authorityYear="1887" box="[1306,1453,1586,1612]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF90FFF23524F98AFFF33E41" box="[1306,1453,1586,1612]" italics="true" pageId="51" pageNumber="268">Palaeovaranus</emphasis>
</taxonomicName>
dentaries described from Quercy (
<bibRefCitation id="EF99AFDCFF90FFF23495F9EBFF393E61" author="DE STEFANO G." box="[1195,1383,1618,1645]" pageId="51" pageNumber="268" pagination="382 - 418" refId="ref34253" refString="DE STEFANO G. 1903. - I sauri del Quercy appartenenti alla collezione Rossignol. Atti della Societa Italiana di Scienze Naturali e del Museo Civili di Storia Naturale, Milan 42: 382 - 418." type="journal article" year="1903">De Stefano 1903</bibRefCitation>
;
<bibRefCitation id="EF99AFDCFF90FFF23549F9EAF9393E80" author="AUGE M." pageId="51" pageNumber="268" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé 2005</bibRefCitation>
) and Dielsdorf,
<collectingCountry id="F31F92BDFF90FFF23414F9CAFEF03E81" box="[1066,1198,1650,1676]" name="Switzerland" pageId="51" pageNumber="268">Switzerland</collectingCountry>
(
<bibRefCitation id="EF99AFDCFF90FFF234FCF9CAF9383EA1" author="GEORGALIS G. L. &amp; SCHEYER T. M." pageId="51" pageNumber="268" pagination="383 - 417" refId="ref36018" refString="GEORGALIS G. L. &amp; SCHEYER T. M. 2019. - A new species of Palaeopython (Serpentes) and other extinct squamates from the Eocene of Dielsdorf (Zurich, Switzerland). Swiss Journal of Geosciences 112: 383 - 417. https: // doi. org / 10.1007 / s 00015 - 019 - 00341 - 6" type="journal article" year="2019">Georgalis &amp; Scheyer 2019</bibRefCitation>
). Whether such differences in robustness reflect some kind of taxonomic or simply intraspecific variation, cannot be evaluated on the basis of this material and our current knowledge of
<taxonomicName id="4C08A9AEFF90FFF233F4F94AFE3C3F01" authorityName="Zittel" authorityYear="1887" box="[970,1122,1778,1804]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="51" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF90FFF233F4F94AFE3C3F01" box="[970,1122,1778,1804]" italics="true" pageId="51" pageNumber="268">Palaeovaranus</emphasis>
</taxonomicName>
mandibular variability.
</paragraph>
<paragraph id="8BB7D22DFF90FFF2337DF8AAFFEF3FE1" blockId="51.[811,1457,1044,2028]" pageId="51" pageNumber="268">
While the cranial material of palaeovaranids resembles in many respects that of varanids, the vertebral morphology of the two groups is relatively distinct, with the former lacking a precondylar constriction and having a relatively more straight centrum, not so widened anteriorly as in the latter group (
<bibRefCitation id="EF99AFDCFF90FFF23545F82AF9383FC6" author="RAGE J. - C." pageId="51" pageNumber="268" pagination="201 - 215" refId="ref38578" refString="RAGE J. - C. 1978. - Squamates, in GEZE B., RAGE J. - C., VERGNAUD- GRAZZINI F., DE BROIN F., BUFFETAUT E., MOURI- ER- CHAUVIRE C., CROCHET J. - Y., SIGE B., SUDRE J., REMY A., LANGEBADRE L., BONIS L. DE, HARTENBERGER J. L. &amp; VIANEY- LI- AUD M. (eds), La poche a Phosphate de Ste-Neboule (Lot) et sa faune de vertebres du Ludien superior. Palaeovertebrata 8: 201 - 215." type="journal article" year="1978">Rage 1978</bibRefCitation>
). Nevertheless, cervical and caudal vertebrae of both palaeovaranids and varanids bear strong resemblance. The two
</paragraph>
<caption id="DF7782A5FF97FFF530BAFC67FFEF3BFD" ID-DOI="http://doi.org/10.5281/zenodo.4720925" ID-Zenodo-Dep="4720925" box="[132,1457,991,1008]" httpUri="https://zenodo.org/record/4720925/files/figure.png" pageId="52" pageNumber="269" startId="52.[132,143,991,1008]" targetBox="[132,1456,182,970]" targetPageId="52">
<paragraph id="8BB7D22DFF97FFF530BAFC67FFEF3BFD" blockId="52.[132,1457,991,1008]" box="[132,1457,991,1008]" pageId="52" pageNumber="269">
FIG. 47. —
<taxonomicName id="4C08A9AEFF97FFF530E5FC67FB303BFD" box="[219,366,991,1008]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="269" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B97C0E3FFF97FFF530E5FC67FB0E3BFD" box="[219,336,991,1008]" italics="true" pageId="52" pageNumber="269">Palaeovaranus</emphasis>
sp.
</taxonomicName>
Virtual 3D models of left dentary NHMW 2019/0058/0054 in lateral (
<emphasis id="B97C0E3FFF97FFF533BDFC67F9D13BFD" bold="true" box="[899,911,991,1008]" pageId="52" pageNumber="269">A</emphasis>
), dorsal (
<emphasis id="B97C0E3FFF97FFF533E9FC67F9BA3BFD" bold="true" box="[983,996,991,1008]" pageId="52" pageNumber="269">B</emphasis>
), medial (
<emphasis id="B97C0E3FFF97FFF53411FC67FE633BFD" bold="true" box="[1071,1085,991,1008]" pageId="52" pageNumber="269">C</emphasis>
), and ventromedial (
<emphasis id="B97C0E3FFF97FFF534E3FC67FEB53BFD" bold="true" box="[1245,1259,991,1008]" pageId="52" pageNumber="269">D</emphasis>
) views.Scale bar:5 mm.
</paragraph>
</caption>
<paragraph id="8BB7D22DFF97FFF530BAFB8CF9563D80" blockId="52.[130,777,1076,1422]" pageId="52" pageNumber="269">
articulated sacral vertebrae resemble much the specimen figured by
<bibRefCitation id="EF99AFDCFF97FFF5309BFBEDFB3C3C63" author="DE STEFANO G." box="[165,354,1108,1135]" pageId="52" pageNumber="269" pagination="382 - 418" refId="ref34253" refString="DE STEFANO G. 1903. - I sauri del Quercy appartenenti alla collezione Rossignol. Atti della Societa Italiana di Scienze Naturali e del Museo Civili di Storia Naturale, Milan 42: 382 - 418." type="journal article" year="1903">De Stefano (1903</bibRefCitation>
: pl. X, fig. 3). Note that due to the fact that vertebrae of certain varanoids, especially varanids (but also observable in our material), possess an anteroventrally directed cotyle, two different centrum lengths can be obtained; we here consider as centrum length the so-called “minimum centrum length” of
<bibRefCitation id="EF99AFDCFF97FFF530C2FB4BFBAA3D03" author="BAILON S. &amp; RAGE J. - C." box="[252,500,1267,1294]" pageId="52" pageNumber="269" pagination="129 - 135" refId="ref32584" refString="BAILON S. &amp; RAGE J. - C. 1994. - Squamates Neogenes et Pleistocenes du Rift occidental, Ouganda, in SENUT B. &amp; PICKFORD M. (eds), Geology and Palaeobiology of the Albertine Rift Valley, Uganda-Zaire. Vol. 2. Palaeobiology. CIFEG, Orleans: 29: 129 - 135 (CIFEG Occasional Publications; 29)." type="journal article" year="1994">Bailon &amp; Rage (1994)</bibRefCitation>
, i.e., the length between the tip of the condyle and the ventral margin of the cotyle (
<bibRefCitation id="EF99AFDCFF97FFF530B3FA8BFB303D40" author="BAILON S. &amp; RAGE J. - C." box="[141,366,1331,1358]" pageId="52" pageNumber="269" pagination="129 - 135" refId="ref32584" refString="BAILON S. &amp; RAGE J. - C. 1994. - Squamates Neogenes et Pleistocenes du Rift occidental, Ouganda, in SENUT B. &amp; PICKFORD M. (eds), Geology and Palaeobiology of the Albertine Rift Valley, Uganda-Zaire. Vol. 2. Palaeobiology. CIFEG, Orleans: 29: 129 - 135 (CIFEG Occasional Publications; 29)." type="journal article" year="1994">Bailon &amp; Rage 1994</bibRefCitation>
;
<bibRefCitation id="EF99AFDCFF97FFF53140FA8BF8003D40" author="GEORGALIS G. L. &amp; RAGE J. - C. &amp; DE BONIS L. &amp; KOUFOS G." box="[382,606,1331,1357]" pageId="52" pageNumber="269" pagination="169 - 181" refId="ref35946" refString="GEORGALIS G. L., RAGE J. - C., DE BONIS L. &amp; KOUFOS G. 2018. - Lizards and snakes from the late Miocene hominoid locality of Ravin de la Pluie (Axios Valley, Greece). Swiss Journal of Geosciences 111: 169 - 181. https: // doi. org / 10.1007 / s 00015 - 017 - 0291 - 2" type="journal article" year="2018">
Georgalis
<emphasis id="B97C0E3FFF97FFF531D5FA8CF8413D40" box="[491,543,1331,1357]" italics="true" pageId="52" pageNumber="269">et al.</emphasis>
2018
</bibRefCitation>
). For a further, detailed discussion about the resemblance and identification of
<taxonomicName id="4C08A9AEFF97FFF530BCFACBFB4F3D80" authorityName="Zittel" authorityYear="1887" box="[130,273,1395,1421]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="52" pageNumber="269" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF97FFF530BCFACBFB4F3D80" box="[130,273,1395,1421]" italics="true" pageId="52" pageNumber="269">Palaeovaranus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C08A9AEFF97FFF5317BFACBFBE43D80" authorityName="Gervais" authorityYear="1848" box="[325,442,1395,1421]" class="Reptilia" family="Anguidae" genus="Placosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="269" phylum="Chordata" rank="genus">
<emphasis id="B97C0E3FFF97FFF5317BFACBFBE43D80" box="[325,442,1395,1421]" italics="true" pageId="52" pageNumber="269">Placosaurus</emphasis>
</taxonomicName>
vertebrae, see Discussion below.
</paragraph>
</subSubSection>
</treatment>
</document>
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