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Pocock, 1893
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<taxonomicName id="4C104D59FFBEF118FF45072D7211FB3C" authority=": L. Koch, 1867" authorityName="L. Koch" authorityYear="1867" box="[128,362,1133,1152]" class="Insecta" family="Hesperiidae" genus="Telegonus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FF45072D73A2FB3C" box="[128,217,1133,1152]" italics="true" pageId="4" pageNumber="5">Telegonus</emphasis>
: L.
<bibRefCitation id="EF814B2BFFBEF118FEC5072D7211FB3C" author="Koch, L." box="[256,362,1133,1152]" pageId="4" pageNumber="5" pagination="173 - 250" refId="ref30163" refString="Koch, L. 1867. Beschreibungen neuer Arachniden und Myriapoden. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 17: 173 - 250." type="journal article" year="1867">Koch, 1867</bibRefCitation>
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(part): 235.
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<taxonomicName id="4C104D59FFBEF118FF4507C57226FB24" authority=": Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[128,349,1157,1176]" class="Arachnida" family="Iuridae" genus="Hadrurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FF4507C573AFFB24" box="[128,212,1157,1176]" italics="true" pageId="4" pageNumber="5">Hadrurus</emphasis>
:
<bibRefCitation id="EF814B2BFFBEF118FF2707C57226FB24" author="Thorell, T." box="[226,349,1157,1176]" pageId="4" pageNumber="5" pagination="75 - 272" refId="ref31656" refString="Thorell, T. 1876. Etudes scorpiologiques. Atti della Societa Italiana di Scienze Naturali 19: 75 - 272." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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(part): 186;
<bibRefCitation id="EF814B2BFFBEF118FE1607C57136FB24" author="Karsch, F." box="[467,589,1157,1176]" pageId="4" pageNumber="5" pagination="97 - 136" refId="ref30073" refString="Karsch, F. 1879. Scorpionologische Beitrage. Part II. Mitteilungen des Munchener Entomologischen Vereins 3: 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
(part): 135, 1881: 290;
<treatmentCitation id="0AB110CBFFBEF118FEEE07DD72B0FB0C" author="Boeris, G." box="[299,459,1181,1200]" page="125" pageId="4" pageNumber="5" year="1889">
<bibRefCitation id="EF814B2BFFBEF118FEEE07DD72B0FB0C" author="Boeris, G." box="[299,459,1181,1200]" pageId="4" pageNumber="5" pagination="123 - 135" refId="ref28786" refString="Boeris, G. 1889. Aracnidi raccolti nel Sud-America dal Dott. Vincenzo Ragazzi. Atti della Societa dei Naturalisti di Modena Memorie Ser. 3 (8): 123 - 135." type="journal article" year="1889">Boeris, 1889: 125</bibRefCitation>
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.
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<emphasis id="B964EAC8FFBEF118FF4507F57381FB74" box="[128,250,1205,1224]" italics="true" pageId="4" pageNumber="5">Caraboctonus</emphasis>
:
<bibRefCitation id="EF814B2BFFBEF118FEC307F572F0FB74" author="Pocock, R. I." box="[262,395,1205,1224]" pageId="4" pageNumber="5" pagination="77 - 103" refId="ref30967" refString="Pocock, R. I. 1893 a. A contribution to the study of Neotropical scorpions. Annals and Magazine of Natural History 12 (6): 77 - 103." type="journal article" year="1893">Pocock, 1893a</bibRefCitation>
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(part): 92.
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<treatmentCitationGroup id="AB0011F4FFBEF118FF45078E7399F96B" pageId="4" pageNumber="5">
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<taxonomicName id="4C104D59FFBEF118FF45078E72C7FB5D" ID-CoL="635PM" authority="Pocock, 1893 b: 306" authorityName="Pocock" authorityPageNumber="306" authorityYear="1893" box="[128,444,1230,1249]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FF45078E7395FB5D" box="[128,238,1230,1249]" italics="true" pageId="4" pageNumber="5">Hadruroides</emphasis>
<bibRefCitation id="EF814B2BFFBEF118FF3E078E72C7FB5D" author="Pocock, R. I." box="[251,444,1230,1249]" pageId="4" pageNumber="5" pagination="303 - 330" refId="ref31000" refString="Pocock, R. I. 1893 b. Notes on the classification of scorpions, followed by some observations on synonymy, with descriptions of new genera and species. Annals and Magazine of Natural History 12 (6): 303 - 330." type="journal article" year="1893">Pocock, 1893b: 306</bibRefCitation>
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, 309, 329, 330,
<typeStatus id="54AB8878FFBEF118FDA2078E71F5FB5D" box="[615,654,1230,1249]" pageId="4" pageNumber="5">type</typeStatus>
species by original designation:
<taxonomicName id="4C104D59FFBEF118FE1207A67265FAAC" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" class="Arachnida" family="Iuridae" genus="Hadrurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBEF118FE1207A671F5FB45" box="[471,654,1254,1273]" italics="true" pageId="4" pageNumber="5">Hadrurus charcasus</emphasis>
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</taxonomicName>
(
<emphasis id="B964EAC8FFBEF118FEF307BD7154FAAD" box="[310,559,1277,1297]" italics="true" pageId="4" pageNumber="5">
<emphasis id="B964EAC8FFBEF118FEF307BD7233FAAC" bold="true" box="[310,328,1277,1296]" italics="true" pageId="4" pageNumber="5">=</emphasis>
<taxonomicName id="4C104D59FFBEF118FE9F07BE7154FAAD" ID-CoL="3J7D6" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[346,559,1278,1297]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="charcasus">Hadruroides charcasus</taxonomicName>
</emphasis>
[
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]);
<treatmentCitation id="0AB110CBFFBEF118FF24065572DDFA94" author="Kraepelin, K." box="[225,422,1301,1320]" page="182" pageId="4" pageNumber="5" year="1894">
<bibRefCitation id="EF814B2BFFBEF118FF24065572DDFA94" author="Kraepelin, K." box="[225,422,1301,1320]" pageId="4" pageNumber="5" pagination="1 - 248" refId="ref30211" refString="Kraepelin, K. 1894. Revision der Scorpione. II. Scorpionidae und Bothriuridae. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 11: 1 - 248." type="journal article" year="1894">Kraepelin, 1894: 182</bibRefCitation>
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, 206;
<treatmentCitation id="0AB110CBFFBEF118FE26065571F1FA94" author="Laurie, M." box="[483,650,1301,1320]" page="130" pageId="4" pageNumber="5" year="1896">
<bibRefCitation id="EF814B2BFFBEF118FE26065571F1FA94" author="Laurie, M." box="[483,650,1301,1320]" pageId="4" pageNumber="5" pagination="121 - 133" refId="ref30312" refString="Laurie, M. 1896. Further notes on the anatomy and development of scorpions, and their bearing on the classification of the order. Annals and Magazine of Natural History 18 (6): 121 - 133." type="journal article" year="1896">Laurie, 1896: 130</bibRefCitation>
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;
<treatmentCitation id="0AB110CBFFBEF118FF5F066D722CFAFC" author="Kraepelin, K." box="[154,343,1325,1344]" page="188" pageId="4" pageNumber="5" year="1899">
<bibRefCitation id="EF814B2BFFBEF118FF5F066D722CFAFC" author="Kraepelin, K." box="[154,343,1325,1344]" pageId="4" pageNumber="5" pagination="1 - 265" refId="ref30240" refString="Kraepelin, K. 1899. Scorpiones und Pedipalpi. In: F. Dahl (editor), Das Tierreich. 8 (Arachnoidea): 1 - 265. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: Friedlander." type="book chapter" year="1899">Kraepelin, 1899: 188</bibRefCitation>
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;
<treatmentCitation id="0AB110CBFFBEF118FEA5066D717DFAFC" author="Pocock, R. I." box="[352,518,1325,1344]" page="474" pageId="4" pageNumber="5" year="1900">
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</treatmentCitation>
; Birula, 1917a: 163, 189; Birula, 1917b: 44, 48, 51;
<treatmentCitation id="0AB110CBFFBEF118FE1E060573C7FACC" author="Mello-Leitao, C. de" page="119" pageId="4" pageNumber="5" year="1945">
<bibRefCitation id="EF814B2BFFBEF118FE1E060573C7FACC" author="Mello-Leitao, C. de" pageId="4" pageNumber="5" pagination="7 - 468" refId="ref30738" refString="Mello-Leitao, C. de. 1945. Escorpioes sul-americanos. Arquivos do Museu Nacional Rio de Janeiro 40: 7 - 468." type="journal article" year="1945">Mello-Leitão, 1945: 119</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FF0D061D7210FACC" author="Bucherl, W." box="[200,363,1373,1392]" page="61" pageId="4" pageNumber="5" year="1964">
<bibRefCitation id="EF814B2BFFBEF118FF0D061D7210FACC" author="Bucherl, W." box="[200,363,1373,1392]" pageId="4" pageNumber="5" pagination="55 - 66" refId="ref28855" refString="Bucherl, W. 1964. Distribucao geografica dos Aracnoides peconhentos temiveis. Memorias do Instituto de Butantan 31: 55 - 66." type="journal article" year="1964">Bücherl, 1964: 61</bibRefCitation>
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,
<treatmentCitation id="0AB110CBFFBEF118FEB2061D72AAFACC" author="Bucherl, W." box="[375,465,1373,1392]" page="115" pageId="4" pageNumber="5" year="1967">
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;
<treatmentCitation id="0AB110CBFFBEF118FE19061D71F1FACC" author="Williams, S. C." box="[476,650,1373,1392]" page="31" pageId="4" pageNumber="5" year="1970">
<bibRefCitation id="EF814B2BFFBEF118FE19061D71F1FACC" author="Williams, S. C." box="[476,650,1373,1392]" pageId="4" pageNumber="5" pagination="1 - 62" refId="ref31851" refString="Williams, S. C. 1970. A systematic revision of the giant hairy scorpion genus Hadrurus. Occasional Papers of the California Academy of Sciences 87: 1 - 62." type="journal article" year="1970">Williams, 1970: 31</bibRefCitation>
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;
<treatmentCitation id="0AB110CBFFBEF118FF5F06357233FA34" author="Bucherl, W." box="[154,328,1397,1416]" page="328" pageId="4" pageNumber="5" year="1971">
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<treatmentCitation id="0AB110CBFFBEF118FE910635717DFA34" author="Stahnke, H. L." box="[340,518,1397,1416]" page="122" pageId="4" pageNumber="5" year="1974">
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;
<treatmentCitation id="0AB110CBFFBEF118FDD4063573CAFA1C" author="Maury, E. A." page="10" pageId="4" pageNumber="5" year="1975">
<bibRefCitation id="EF814B2BFFBEF118FDD4063573CAFA1C" author="Maury, E. A." pageId="4" pageNumber="5" pagination="9 - 21" refId="ref30635" refString="Maury, E. A. 1975. Escorpiones y escorpionismo en el Peru IV: Revision del genero Hadruroides Pocock, 1893 (Scorpiones, Vejovidae). Revista Peruana de Entomologia 17 (1): 9 - 21." type="journal article" year="1975">Maury, 1975: 10</bibRefCitation>
</treatmentCitation>
, 11;
<treatmentCitation id="0AB110CBFFBEF118FF1806CD72FAFA1C" author="Francke, O. F." box="[221,385,1421,1440]" page="75" pageId="4" pageNumber="5" year="1977">
<bibRefCitation id="EF814B2BFFBEF118FF1806CD72FAFA1C" author="Francke, O. F." box="[221,385,1421,1440]" pageId="4" pageNumber="5" pagination="73 - 76" refId="ref29752" refString="Francke, O. F. 1977. Escorpiones y escorpionismo en el Peru VI: Lista de especies y claves para identificar las familias y los generos. Revista Peruana de Entomologia 20: 73 - 76." type="journal article" year="1977">Francke, 1977: 75</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FE4E06CD73C7FA04" author="Francke, O. F. &amp; M. E. Soleglad" page="235" pageId="4" pageNumber="5" year="1981">
<bibRefCitation id="EF814B2BFFBEF118FE4E06CD73C7FA04" author="Francke, O. F. &amp; M. E. Soleglad" pageId="4" pageNumber="5" pagination="233 - 258" refId="ref29914" refString="Francke, O. F., and M. E. Soleglad. 1981. The family Iuridae Thorell (Arachnida, Scorpiones). Journal of Arachnology 9: 233 - 258." type="journal article" year="1981">Francke and Soleglad, 1981: 235</bibRefCitation>
</treatmentCitation>
, 256,
<figureCitation id="132B2A5FFFBEF118FF3506E57253FA04" box="[240,296,1445,1464]" captionStart="Fig" captionStartId="13.[112,150,1582,1604]" captionTargetBox="[139,1136,210,1548]" captionTargetId="figure-1@13.[139,1136,210,1548]" captionText="Fig. 9. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides chinchaysuyu, n. sp. A. Paratype ♂ (MHNC), leg III, dextral patella, dorsal aspect. B. Paratype ♀ (MHNC), dextral pedipalp chela, external aspect. C. Paratype ♂ (MHNC), metasomal segments IIIV, ventral aspect showing pigmentation pattern. D. Hadruroides maculatus (Thorell, 1876), ♂ (MHNC), leg III, dextral patella, dorsal aspect. EH. H. chinchaysuyu, holotype ♂ (MHNC), sinistral hemispermatophore. E. Ental aspect. F. Dorsal aspect. G. Ectal aspect. H. Ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682527" httpUri="https://zenodo.org/record/5682527/files/figure.png" pageId="4" pageNumber="5">figs. 9</figureCitation>
,
<figureCitation id="132B2A5FFFBEF118FEF706E57232FA04" box="[306,329,1445,1464]" captionStart="Fig" captionStartId="17.[112,150,896,918]" captionTargetBox="[137,1138,210,862]" captionTargetId="figure-337@17.[137,1138,210,862]" captionText="Fig. 13. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5682539" httpUri="https://zenodo.org/record/5682539/files/figure.png" pageId="4" pageNumber="5">13</figureCitation>
, 2733, 58;
<treatmentCitation id="0AB110CBFFBEF118FE7706E57132FA04" author="Francke, O. F." box="[434,585,1445,1464]" page="8" pageId="4" pageNumber="5" year="1985">
<bibRefCitation id="EF814B2BFFBEF118FE7706E57132FA04" author="Francke, O. F." box="[434,585,1445,1464]" pageId="4" pageNumber="5" pagination="1 - 32" refId="ref29791" refString="Francke, O. F. 1985. Conspectus Genericus Scorpionorum 1758 - 1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University 98: 1 - 32." type="journal article" year="1985">Francke, 1985: 8</bibRefCitation>
</treatmentCitation>
, 17, 20;
<treatmentCitation id="0AB110CBFFBEF118FF5F06FD7232FA6C" author="Sissom, W. D." box="[154,329,1469,1488]" page="130" pageId="4" pageNumber="5" year="1990">
<bibRefCitation id="EF814B2BFFBEF118FF5F06FD7232FA6C" author="Sissom, W. D." box="[154,329,1469,1488]" pageId="4" pageNumber="5" pagination="64 - 160" refId="ref31230" refString="Sissom, W. D. 1990. Systematics, biogeography and paleontology. In: G. A. Polis (editor), The Biology of Scorpions. Stanford, CA: Stanford University Press, 64 - 160." type="book chapter" year="1990">Sissom, 1990: 130</bibRefCitation>
</treatmentCitation>
, 131;
<treatmentCitation id="0AB110CBFFBEF118FE4C06FD71F1FA6C" author="Nenilin, A. B. &amp; V. Fet" box="[393,650,1469,1488]" page="14" pageId="4" pageNumber="5" year="1992">
<bibRefCitation id="EF814B2BFFBEF118FE4C06FD71F1FA6C" author="Nenilin, A. B. &amp; V. Fet" box="[393,650,1469,1488]" pageId="4" pageNumber="5" pagination="3 - 31" refId="ref30806" refString="Nenilin, A. B., and V. Fet. 1992. Zoogeographical analysis of the world scorpion fauna (Arachnida: Scorpiones). Arthropoda Selecta 1 (2): 3 - 31. [in Russian, English summary]." type="journal article" year="1992">Nenilin and Fet, 1992: 14</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FF5F0694721BFA5B" author="Lourenco, W. R." box="[154,352,1492,1511]" page="157" pageId="4" pageNumber="5" year="1994">
<bibRefCitation id="EF814B2BFFBEF118FF5F0694721BFA5B" author="Lourenco, W. R." box="[154,352,1492,1511]" pageId="4" pageNumber="5" pagination="155 - 160" refId="ref30353" refString="Lourenco, W. R. 1994. Diversity and endemism in tropical versus temperate scorpion communities. Biogeographica 70 (3): 155 - 160." type="journal article" year="1994">Lourenço, 1994: 157</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AB110CBFFBEF118FEAA06947299FA5B" author="Lourenco, W. R." box="[367,482,1492,1511]" page="74" pageId="4" pageNumber="5" year="1995">
<bibRefCitation id="EF814B2BFFBEF118FEAA06947299FA5B" author="Lourenco, W. R." box="[367,482,1492,1511]" pageId="4" pageNumber="5" pagination="61 - 88" refId="ref30381" refString="Lourenco, W. R. 1995. Les scorpions (Chelicerata, Scorpiones) de l'Equateur avec quelques considerations sur la biogeographie et la diversite des especes. Revue Suisse de Zoologie 102 (1): 61 - 88." type="journal article" year="1995">1995: 7476</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AB110CBFFBEF118FE3406947122FA5B" author="Lourenco, W. R." box="[497,601,1492,1511]" page="601" pageId="4" pageNumber="5" year="1997">
<bibRefCitation id="EF814B2BFFBEF118FE3406947122FA5B" author="Lourenco, W. R." box="[497,601,1492,1511]" pageId="4" pageNumber="5" pagination="587 - 604" refId="ref30423" refString="Lourenco, W. R. 1997 a. Additions a la faune de scorpions neotropicaux. Revue Suisse de Zoologie 104 (3): 587 - 604." type="journal article" year="1997">1997a: 601</bibRefCitation>
</treatmentCitation>
, 602, 1998: 135;
<treatmentCitation id="0AB110CBFFBEF118FEC306AC72BBFA43" author="Kovarik, F." box="[262,448,1516,1536]" page="135" pageId="4" pageNumber="5" year="1998">
<bibRefCitation id="EF814B2BFFBEF118FEC306AC72BBFA43" author="Kovarik, F." box="[262,448,1516,1536]" pageId="4" pageNumber="5" refId="ref30189" refString="Kovarik, F. 1998. Stiri (Scorpions). Madagaskar, Jihlava, 175 pp. [in Czech]" type="book" year="1998">Kovařík, 1998: 135</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FE0A06AC71F1FA43" author="Lourenco, W. R." box="[463,650,1516,1535]" page="25" pageId="4" pageNumber="5" year="2000">
<bibRefCitation id="EF814B2BFFBEF118FE0A06AC71F1FA43" author="Lourenco, W. R." box="[463,650,1516,1535]" pageId="4" pageNumber="5" pagination="21 - 39" refId="ref30529" refString="Lourenco, W. R. 2000. Panbiogeographie, les familles des scorpions et leur repartition geographique. Biogeographica 76 (1): 21 - 39." type="journal article" year="2000">Lourenço, 2000: 25</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FF5F054472CAF9AB" author="Sissom, W. D. &amp; V. Fet" box="[154,433,1540,1559]" page="411" pageId="4" pageNumber="5" year="2000">
<bibRefCitation id="EF814B2BFFBEF118FF5F054472CAF9AB" author="Sissom, W. D. &amp; V. Fet" box="[154,433,1540,1559]" pageId="4" pageNumber="5" pagination="409 - 420" refId="ref31272" refString="Sissom, W. D., and V. Fet. 2000. Family Iuridae Thorell, 1876. In V. Fet, W. D. Sissom, G. Lowe, and M. E. Braunwalder, Catalog of the Scorpions of the World (1758 - 1998). New York: New York Entomological Society, 409 - 420." type="book chapter" year="2000">Sissom and Fet, 2000: 411413</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FE7C05447390F993" author="Lourenco, W. R. &amp; H. Dastych" page="55" pageId="4" pageNumber="5" year="2001">
<bibRefCitation id="EF814B2BFFBEF118FE7C05447390F993" author="Lourenco, W. R. &amp; H. Dastych" pageId="4" pageNumber="5" pagination="51 - 62" refId="ref30558" refString="Lourenco, W. R., and H. Dastych. 2001. A contribution to the scorpion fauna of Peru, with a description of Chactas koepckei sp. nov. (Arachnida: Scorpiones). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 98: 51 - 62." type="journal article" year="2001">Lourenço and Dastych, 2001: 55</bibRefCitation>
</treatmentCitation>
, 56;
<treatmentCitation id="0AB110CBFFBEF118FEDB055C713BF993" author="Soleglad, M. E. &amp; W. D. Sissom" box="[286,576,1564,1583]" page="31" pageId="4" pageNumber="5" year="2001">
<bibRefCitation id="EF814B2BFFBEF118FEDB055C713BF993" author="Soleglad, M. E. &amp; W. D. Sissom" box="[286,576,1564,1583]" pageId="4" pageNumber="5" pagination="25 - 111" refId="ref31460" refString="Soleglad, M. E., and W. D. Sissom. 2001. Phylogeny of the family Euscorpiidae Laurie, 1896: A major revision. In: V. Fet and P. A. Selden (editors), Scorpions 2001. In memoriam Gary A. Polis. Burnham Beeches, Bucks, UK: British Arachnological Society, 25 - 111." type="book chapter" year="2001">Soleglad and Sissom, 2001: 31</bibRefCitation>
</treatmentCitation>
, fig. 34;
<treatmentCitation id="0AB110CBFFBEF118FF5F057472DBF9FB" author="Acosta, L. E. &amp; J. A. Ochoa" box="[154,416,1588,1607]" page="19" pageId="4" pageNumber="5" year="2002">
<bibRefCitation id="EF814B2BFFBEF118FF5F057472DBF9FB" author="Acosta, L. E. &amp; J. A. Ochoa" box="[154,416,1588,1607]" pageId="4" pageNumber="5" pagination="15 - 23" refId="ref28532" refString="Acosta, L. E., and J. A. Ochoa. 2002. Lista de los escorpiones bolivianos (Chelicerata: Scorpiones), con notas sobre su distribucion. Revista de la Sociedad Argentina de Entomologia 61 (3 - 4): 15 - 23." type="journal article" year="2002">Acosta and Ochoa, 2002: 19</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FE6E057473DDF9E3" author="Soleglad, M. E. &amp; V. Fet" page="2" pageId="4" pageNumber="5" year="2003">
<bibRefCitation id="EF814B2BFFBEF118FE6E057473DDF9E3" author="Soleglad, M. E. &amp; V. Fet" pageId="4" pageNumber="5" pagination="1 - 34" refId="ref31339" refString="Soleglad, M. E., and V. Fet. 2003 a. The scorpion sternum: Structure and phylogeny (Scorpiones: Orthosterni). Euscorpius 5: 1 - 34." type="journal article" year="2003">Soleglad and Fet, 2003a: 2</bibRefCitation>
</treatmentCitation>
, 5, 12, 2003b: 8, 111;
<treatmentCitation id="0AB110CBFFBEF118FE40050C7144F9E3" author="Fet, V. &amp; M. E. Soleglad &amp; D. P. A. Neff &amp; I. Stathi" box="[389,575,1612,1631]" page="18" pageId="4" pageNumber="5" year="2004">
<bibRefCitation id="EF814B2BFFBEF118FE40050C7144F9E3" author="Fet, V. &amp; M. E. Soleglad &amp; D. P. A. Neff &amp; I. Stathi" box="[389,575,1612,1631]" pageId="4" pageNumber="5" pagination="17 - 40" refId="ref29676" refString="Fet, V., M. E. Soleglad, D. P. A. Neff, and I. Stathi. 2004. Tarsal armature in the superfamily Iuroidea (Scorpiones: Iurida). Revista Iberica de Aracnologia 10: 17 - 40." type="journal article" year="2004">Fet et al., 2004: 18</bibRefCitation>
</treatmentCitation>
, 23, 24;
<treatmentCitation id="0AB110CBFFBEF118FF5F052472ADF9CB" author="Prendini, L. &amp; W. C. Wheeler" box="[154,470,1636,1655]" page="482" pageId="4" pageNumber="5" year="2005">
<bibRefCitation id="EF814B2BFFBEF118FF5F052472ADF9CB" author="Prendini, L. &amp; W. C. Wheeler" box="[154,470,1636,1655]" pageId="4" pageNumber="5" pagination="446 - 494" refId="ref31150" refString="Prendini, L., and W. C. Wheeler. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics 21: 446 - 494." type="journal article" year="2005">Prendini and Wheeler, 2005: 482</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FE2005247392F933" author="Fet, V. &amp; M. E. Soleglad" page="12" pageId="4" pageNumber="5" year="2005">
<bibRefCitation id="EF814B2BFFBEF118FE2005247392F933" author="Fet, V. &amp; M. E. Soleglad" pageId="4" pageNumber="5" pagination="1 - 13" refId="ref29598" refString="Fet, V., and M. E. Soleglad. 2005. Contributions to scorpion systematics. 1. On recent changes in high-level taxonomy. Euscorpius 31: 1 - 13." type="journal article" year="2005">Fet and Soleglad, 2005: 12</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FF31053C72F0F933" author="Ochoa, J. A." box="[244,395,1660,1679]" page="65" pageId="4" pageNumber="5" year="2005">
<bibRefCitation id="EF814B2BFFBEF118FF31053C72F0F933" author="Ochoa, J. A." box="[244,395,1660,1679]" pageId="4" pageNumber="5" pagination="49 - 68" refId="ref30852" refString="Ochoa, J. A. 2005. Patrones de distribucion de escorpiones de la region andina en el sur peruano. Revista Peruana de Biologia 12 (1): 49 - 68." type="journal article" year="2005">Ochoa, 2005: 65</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="132B2A5FFFBEF118FE53053C72AAF933" box="[406,465,1660,1679]" captionStart="Fig" captionStartId="21.[112,150,971,993]" captionTargetBox="[137,1138,210,932]" captionTargetId="figure-376@21.[137,1138,210,937]" captionText="Fig. 17. Hadruroides graceae, n. sp., paratype ♂ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5713956" httpUri="https://zenodo.org/record/5713956/files/figure.png" pageId="4" pageNumber="5">fig. 17</figureCitation>
; Dupre´, 2007: 5, 14;
<treatmentCitation id="0AB110CBFFBEF118FF5F05D472C9F91B" author="Ochoa, J. A. &amp; J. C. Chaparro" box="[154,434,1684,1703]" page="5" pageId="4" pageNumber="5" year="2008">
<bibRefCitation id="EF814B2BFFBEF118FF5F05D472C9F91B" author="Ochoa, J. A. &amp; J. C. Chaparro" box="[154,434,1684,1703]" pageId="4" pageNumber="5" pagination="5 - 10" refId="ref30887" refString="Ochoa, J. A., and J. C. Chaparro. 2008. Nueva especie de escorpion del genero Hadruroides (Scorpiones: Caraboctoninae) de los valles interandinos de Peru. Revista Peruana de Biologia 15 (1): 5 - 10." type="journal article" year="2008">Ochoa and Chaparro, 2008: 5</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FE7B05D47388F903" author="Francke, O. F. &amp; L. Prendini" page="210" pageId="4" pageNumber="5" year="2008">
<bibRefCitation id="EF814B2BFFBEF118FE7B05D47388F903" author="Francke, O. F. &amp; L. Prendini" pageId="4" pageNumber="5" pagination="205 - 223" refId="ref29826" refString="Francke, O. F., and L. Prendini. 2008. Phylogeny and classification of the giant hairy scorpions, Hadrurus Thorell (Iuridae Thorell): A reappraisal. Systematics and Biodiversity 6 (2): 205 - 223." type="journal article" year="2008">Francke and Prendini, 2008: 210</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FF3805EC7286F903" author="Fet, V. &amp; M. E. Soleglad" box="[253,509,1708,1727]" page="255" pageId="4" pageNumber="5" year="2008">
<bibRefCitation id="EF814B2BFFBEF118FF3805EC7286F903" author="Fet, V. &amp; M. E. Soleglad" box="[253,509,1708,1727]" pageId="4" pageNumber="5" pagination="255 - 281" refId="ref29633" refString="Fet, V., and M. E. Soleglad. 2008. Cladistic analysis of superfamily Iuroidea, with emphasis on subfamily Hadrurinae (Scorpiones: Iurida). Boletin de la Sociedad Entomologica Aragonesa 43: 255 - 281." type="journal article" year="2008">Fet and Soleglad, 2008: 255</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB110CBFFBEF118FDC205EC73A6F96B" author="Soleglad, M. E. &amp; F. Kovarik &amp; V. Fet" page="1" pageId="4" pageNumber="5" year="2009">
<bibRefCitation id="EF814B2BFFBEF118FDC205EC73A6F96B" author="Soleglad, M. E. &amp; F. Kovarik &amp; V. Fet" pageId="4" pageNumber="5" pagination="1 - 21" refId="ref31410" refString="Soleglad, M. E., F. Kovarik, and V. Fet. 2009. Etudes on Iurids. 1. The orthobothriotaxic pattern of Iuridae, with observations on neobothriotaxy in genus Iurus (Scorpiones: Iuroidea). Euscorpius 79: 1 - 21." type="journal article" year="2009">Soleglad et al., 2009: 1</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="C30A6551FFBEF119FD1D03937163FE07" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BAF36DAFFBEF118FD1D0393772CFB40" blockId="4.[701,1228,211,1753]" pageId="4" pageNumber="5">
DIAGNOSIS: The genus
<taxonomicName id="4C104D59FFBEF118FC3C039377F8FF57" authorityName="Pocock" authorityYear="1893" box="[1017,1155,211,235]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FC3C039377F8FF57" box="[1017,1155,211,235]" italics="true" pageId="4" pageNumber="5">Hadruroides</emphasis>
</taxonomicName>
comprises small- to medium-sized scorpions, varying from ca.
<quantity id="4CE89B3FFFBEF118FC56024E7091FE9A" box="[915,1002,270,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="mm" value="30.0">30 mm</quantity>
, e.g.,
<taxonomicName id="4C104D59FFBEF118FBF2024F7027FEF8" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="leopardus">
<emphasis id="B964EAC8FFBEF118FBF2024F77B0FE9B" box="[1079,1227,271,295]" italics="true" pageId="4" pageNumber="5">H. leopardus</emphasis>
<bibRefCitation id="EF814B2BFFBEF118FD78026C7027FEF8" author="Pocock, R. I." box="[701,860,300,324]" pageId="4" pageNumber="5" pagination="469 - 478" refId="ref31046" refString="Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History 5 (7): 469 - 478." type="journal article" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C104D59FFBEF118FC67026C775BFEF8" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[930,1056,300,324]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="graceae" status="sp. nov.">
<emphasis id="B964EAC8FFBEF118FC67026C775BFEF8" box="[930,1056,300,324]" italics="true" pageId="4" pageNumber="5">H. graceae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBEF118FBF6026C770CFEF8" box="[1075,1143,300,324]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
, to ca.
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in total length, e.g.,
<taxonomicName id="4C104D59FFBEF118FBF7020A7740FE3C" authority="(Karsch, 1879) (Maury, 1975)" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBEF118FBF7020A77B0FEDE" box="[1074,1227,330,354]" italics="true" pageId="4" pageNumber="5">H. charcasus</emphasis>
(
<bibRefCitation id="EF814B2BFFBEF118FD0002287011FE3C" author="Karsch, F." box="[709,874,360,384]" pageId="4" pageNumber="5" pagination="97 - 136" refId="ref30073" refString="Karsch, F. 1879. Scorpionologische Beitrage. Part II. Mitteilungen des Munchener Entomologischen Vereins 3: 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
) (
<bibRefCitation id="EF814B2BFFBEF118FC4A02287748FE3C" author="Maury, E. A." box="[911,1075,360,384]" pageId="4" pageNumber="5" pagination="9 - 21" refId="ref30635" refString="Maury, E. A. 1975. Escorpiones y escorpionismo en el Peru IV: Revision del genero Hadruroides Pocock, 1893 (Scorpiones, Vejovidae). Revista Peruana de Entomologia 17 (1): 9 - 21." type="journal article" year="1975">Maury, 1975</bibRefCitation>
)
</taxonomicName>
. The most important diagnostic characters for the genus are as follows: cheliceral movable fingers, internal surface with two subdistal teeth and one prominent basal tooth; carapace anterior margin slightly convex, with three pairs of lateral ocelli; sternum subpentagonal, with Yshaped sulcus; pedipalp chela smooth, acarinate in most species, except
<taxonomicName id="4C104D59FFBEF118FBD7011677DFFDD2" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[1042,1188,598,622]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBEF118FBD7011677DFFDD2" box="[1042,1188,598,622]" italics="true" pageId="4" pageNumber="5">H. charcasus</emphasis>
</taxonomicName>
, in which granular carinae are present on internal surface; chela movable finger, median denticle row with six or seven median subrows of denticles and variable number of accessory denticles; neobothriotaxic major
<typeStatus id="54AB8878FFBEF118FBA201AA77DBFCBE" box="[1127,1184,746,770]" pageId="4" pageNumber="5">Type</typeStatus>
C trichobothrial pattern (Vachon, 1974): femur with three trichobothria (
<emphasis id="B964EAC8FFBEF118FC2300667096FC82" box="[998,1005,806,830]" italics="true" pageId="4" pageNumber="5">i</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC3A00667770FC82" box="[1023,1035,806,830]" italics="true" pageId="4" pageNumber="5">e</emphasis>
,
<emphasis id="B964EAC8FFBEF118FBD900667752FC82" box="[1052,1065,806,830]" italics="true" pageId="4" pageNumber="5">d</emphasis>
); patella with 20: one internal (
<emphasis id="B964EAC8FFBEF118FC48000470EFFCE0" box="[909,916,836,860]" italics="true" pageId="4" pageNumber="5">i</emphasis>
), two dorsal (
<emphasis id="B964EAC8FFBEF118FB860004772BFCE0" box="[1091,1104,836,860]" italics="true" pageId="4" pageNumber="5">d</emphasis>
<subScript id="1794349FFFBEF118FB95000E770DFCE2" attach="left" box="[1104,1142,846,862]" fontSize="7" pageId="4" pageNumber="5">1, 2</subScript>
), three ventral (
<emphasis id="B964EAC8FFBEF118FCE500227057FCC6" box="[800,812,866,890]" italics="true" pageId="4" pageNumber="5">v</emphasis>
<subScript id="1794349FFFBEF118FCEE002C7032FCC0" attach="left" box="[811,841,876,892]" fontSize="7" pageId="4" pageNumber="5">13</subScript>
),
<emphasis id="B964EAC8FFBEF118FCA600227014FCC6" box="[867,879,866,890]" italics="true" pageId="4" pageNumber="5">v</emphasis>
<subScript id="1794349FFFBEF118FCAA002C7002FCC0" attach="left" box="[879,889,876,892]" fontSize="7" pageId="4" pageNumber="5">3</subScript>
situated on external surface, and 14 external trichobothria (
<emphasis id="B964EAC8FFBEF118FBDC00C07756FC24" box="[1049,1069,896,920]" italics="true" pageId="4" pageNumber="5">et</emphasis>
<subScript id="1794349FFFBEF118FBE800CA7730FC26" attach="left" box="[1069,1099,906,922]" fontSize="7" pageId="4" pageNumber="5">13</subScript>
,
<emphasis id="B964EAC8FFBEF118FB9C00C07703FC24" box="[1113,1144,896,920]" italics="true" pageId="4" pageNumber="5">est</emphasis>
,
<emphasis id="B964EAC8FFBEF118FB4300C077DDFC24" box="[1158,1190,896,920]" italics="true" pageId="4" pageNumber="5">em</emphasis>
<subScript id="1794349FFFBEF118FB6300CA77BFFC26" attach="left" box="[1190,1220,906,922]" fontSize="7" pageId="4" pageNumber="5">13</subScript>
,
<emphasis id="B964EAC8FFBEF118FD7800DD719BFC09" box="[701,736,925,949]" italics="true" pageId="4" pageNumber="5">esb</emphasis>
<subScript id="1794349FFFBEF118FD2400E7707FFC0B" attach="right" box="[737,772,935,951]" fontSize="7" pageId="4" pageNumber="5">1, 2</subScript>
,
<emphasis id="B964EAC8FFBEF118FCD100DD7056FC09" box="[788,813,925,949]" italics="true" pageId="4" pageNumber="5">eb</emphasis>
<subScript id="1794349FFFBEF118FCE800E77030FC0B" attach="left" box="[813,843,935,951]" fontSize="7" pageId="4" pageNumber="5">15</subScript>
); chela with 26 trichobothria: 14 on manus (
<emphasis id="B964EAC8FFBEF118FCFB00FB7022FC6F" box="[830,857,955,979]" italics="true" pageId="4" pageNumber="5">Et</emphasis>
<subScript id="1794349FFFBEF118FC9D0085700DFC69" attach="left" box="[856,886,965,981]" fontSize="7" pageId="4" pageNumber="5">15</subScript>
,
<emphasis id="B964EAC8FFBEF118FC4000FB70D1FC6F" box="[901,938,955,979]" italics="true" pageId="4" pageNumber="5">Est</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC7C00FB7099FC6F" box="[953,994,955,979]" italics="true" pageId="4" pageNumber="5">Esb</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC3400FB776BFC6F" box="[1009,1040,955,979]" italics="true" pageId="4" pageNumber="5">Eb</emphasis>
<subScript id="1794349FFFBEF118FBD500857755FC69" attach="left" box="[1040,1070,965,981]" fontSize="7" pageId="4" pageNumber="5">13</subScript>
,
<emphasis id="B964EAC8FFBEF118FBF800FB772BFC6F" box="[1085,1104,955,979]" italics="true" pageId="4" pageNumber="5">V</emphasis>
<subScript id="1794349FFFBEF118FB9500857715FC69" attach="left" box="[1104,1134,965,981]" fontSize="7" pageId="4" pageNumber="5">14</subScript>
) and 12 on fixed finger (
<emphasis id="B964EAC8FFBEF118FCB0009970E9FC4D" box="[885,914,985,1009]" italics="true" pageId="4" pageNumber="5">Dt</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC67009970BFFC4D" box="[930,964,985,1009]" italics="true" pageId="4" pageNumber="5">Db</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC160099709CFC4D" box="[979,999,985,1009]" italics="true" pageId="4" pageNumber="5">et</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC330099776EFC4D" box="[1014,1045,985,1009]" italics="true" pageId="4" pageNumber="5">est</emphasis>
,
<emphasis id="B964EAC8FFBEF118FBE10099773CFC4D" box="[1060,1095,985,1009]" italics="true" pageId="4" pageNumber="5">esb</emphasis>
,
<emphasis id="B964EAC8FFBEF118FB9300997714FC4D" box="[1110,1135,985,1009]" italics="true" pageId="4" pageNumber="5">eb</emphasis>
,
<emphasis id="B964EAC8FFBEF118FBBB009977EFFC4D" box="[1150,1172,985,1009]" italics="true" pageId="4" pageNumber="5">dt</emphasis>
,
<emphasis id="B964EAC8FFBEF118FB61009977BEFC4D" box="[1188,1221,985,1009]" italics="true" pageId="4" pageNumber="5">dst</emphasis>
,
<emphasis id="B964EAC8FFBEF118FD7800B77199FBB3" box="[701,738,1015,1039]" italics="true" pageId="4" pageNumber="5">dsb</emphasis>
,
<emphasis id="B964EAC8FFBEF118FD3000B7706BFBB3" box="[757,784,1015,1039]" italics="true" pageId="4" pageNumber="5">db</emphasis>
,
<emphasis id="B964EAC8FFBEF118FCE700B77049FBB3" box="[802,818,1015,1039]" italics="true" pageId="4" pageNumber="5">it</emphasis>
,
<emphasis id="B964EAC8FFBEF118FC8000B77022FBB3" box="[837,857,1015,1039]" italics="true" pageId="4" pageNumber="5">ib</emphasis>
); leg telotarsus with ventromedian row of spinule clusters (setaceous tufts); metasomal segment V with complete, granular VL and VM carinae; telson slightly concave dorsally, with short aculeus; hemispermatophore lamelliform, with elongated crest, accompanied externally by medial slit and one free lobe, truncal flexure absent, internobasal reflection of the sperm duct absent.
</paragraph>
<paragraph id="8BAF36DAFFBEF119FD1D06437163FE07" blockId="4.[701,1228,211,1753]" lastBlockId="5.[88,614,211,1542]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<taxonomicName id="4C104D59FFBEF118FD1D06437019FAA7" authorityName="Pocock" authorityYear="1893" box="[728,866,1283,1307]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FD1D06437019FAA7" box="[728,866,1283,1307]" italics="true" pageId="4" pageNumber="5">Hadruroides</emphasis>
</taxonomicName>
appears to be most closely related to
<taxonomicName id="4C104D59FFBEF118FC890661709EFA85" authorityName=": Pocock" authorityYear="1893" box="[844,997,1313,1337]" class="Arachnida" family="Caraboctonidae" genus="Caraboctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FC890661709EFA85" box="[844,997,1313,1337]" italics="true" pageId="4" pageNumber="5">Caraboctonus</emphasis>
</taxonomicName>
. The two genera exhibit several similarities, e.g., trichobothrial pattern, ventromedian row of spinule clusters on telotarsus, dentition of chelicerae and pedipalp chela fingers. They may be distinguished by means of the ventral carinae of metasomal segment V: in
<taxonomicName id="4C104D59FFBEF118FC3F069377E8FA57" authorityName=": Pocock" authorityYear="1893" box="[1018,1171,1491,1515]" class="Arachnida" family="Caraboctonidae" genus="Caraboctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FC3F069377E8FA57" box="[1018,1171,1491,1515]" italics="true" pageId="4" pageNumber="5">Caraboctonus</emphasis>
</taxonomicName>
, the VL carinae are restricted to the posterior half of the segment and the VM carina is absent, whereas in
<taxonomicName id="4C104D59FFBEF118FC89056C70ADF9F8" authorityName="Pocock" authorityYear="1893" box="[844,982,1580,1604]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBEF118FC89056C70ADF9F8" box="[844,982,1580,1604]" italics="true" pageId="4" pageNumber="5">Hadruroides</emphasis>
</taxonomicName>
, both VL and VM carinae are present and complete (except in
<taxonomicName id="4C104D59FFBEF118FB6B050A7184F93C" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tishqu" status="sp. nov.">
<emphasis id="B964EAC8FFBEF118FB6B050A7184F93C" italics="true" pageId="4" pageNumber="5">H. tishqu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBEF118FCC805287037F93C" box="[781,844,1640,1664]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
, in which the VL and VM carinae are restricted to the posterior two-thirds of the segment). The two genera may be further distinguished on the basis of pedipalp chela finger dentition: internal and external accessory denticles, flanking the median denticle row in
<taxonomicName id="4C104D59FFBFF119FF26024F7216FE9B" authorityName="Pocock" authorityYear="1893" box="[227,365,271,295]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FF26024F7216FE9B" box="[227,365,271,295]" italics="true" pageId="5" pageNumber="6">Hadruroides</emphasis>
</taxonomicName>
, are absent in
<taxonomicName id="4C104D59FFBFF119FF9D026C738AFEF8" authorityName=": Pocock" authorityYear="1893" box="[88,241,300,324]" class="Arachnida" family="Caraboctonidae" genus="Caraboctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FF9D026C738AFEF8" box="[88,241,300,324]" italics="true" pageId="5" pageNumber="6">Caraboctonus</emphasis>
</taxonomicName>
. Furthermore, the hemispermatophore of
<taxonomicName id="4C104D59FFBFF119FF19020A721DFEDE" authorityName="Pocock" authorityYear="1893" box="[220,358,330,354]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FF19020A721DFEDE" box="[220,358,330,354]" italics="true" pageId="5" pageNumber="6">Hadruroides</emphasis>
</taxonomicName>
exhibits an elongated crest and a medial slit, both absent from the hemispermatophore of
<taxonomicName id="4C104D59FFBFF119FEB702C57170FE21" authorityName=": Pocock" authorityYear="1893" box="[370,523,389,413]" class="Arachnida" family="Caraboctonidae" genus="Caraboctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FEB702C57170FE21" box="[370,523,389,413]" italics="true" pageId="5" pageNumber="6">Caraboctonus</emphasis>
</taxonomicName>
, which instead exhibits a digit-shaped process.
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FFBFF119FFB702817136F9BA" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BAF36DAFFBFF119FFB702817136F9BA" blockId="5.[88,614,211,1542]" pageId="5" pageNumber="6">
DISTRIBUTION: The genus
<taxonomicName id="4C104D59FFBFF119FE7C02817138FE65" authorityName="Pocock" authorityYear="1893" box="[441,579,449,473]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FE7C02817138FE65" box="[441,579,449,473]" italics="true" pageId="5" pageNumber="6">Hadruroides</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F307764AFFBFF119FF29029F722BFE4B" box="[236,336,479,503]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
,
<collectingCountry id="F307764AFFBFF119FEA1029F72E7FE4B" box="[356,412,479,503]" name="Peru" pageId="5" pageNumber="6">Peru</collectingCountry>
, northern
<collectingCountry id="F307764AFFBFF119FDE6029F7119FE4B" box="[547,610,479,503]" name="Chile" pageId="5" pageNumber="6">Chile</collectingCountry>
, and several offshore islands, including the Galápagos (
<tableCitation id="C6920361FFBFF119FF34015A723BFD8E" box="[241,320,538,562]" captionStart="TABLE 1" captionStartId="1.[592,665,211,230]" captionTargetBox="[88,987,324,742]" captionText="TABLE 1 Currently recognized species of the scorpion genus Hadruroides Pocock, 1893, with countries and departments or provinces from which they have been recorded" httpUri="http://table.plazi.org/id/DF6F6652FFBBF11DFD9503937080FEA1" pageId="5" pageNumber="6" tableUuid="DF6F6652FFBBF11DFD9503937080FEA1">table 1</tableCitation>
). Thirteen of the 16 described species occur in
<collectingCountry id="F307764AFFBFF119FE65017872AEFDEC" box="[416,469,568,592]" name="Peru" pageId="5" pageNumber="6">Peru</collectingCountry>
(figs. 1, 2). Four species inhabit dry forest in northern
<collectingCountry id="F307764AFFBFF119FF9D013373EBFD37" box="[88,144,627,651]" name="Peru" pageId="5" pageNumber="6">Peru</collectingCountry>
:
<taxonomicName id="4C104D59FFBFF119FF5901337255FD37" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[156,302,627,651]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBFF119FF5901337255FD37" box="[156,302,627,651]" italics="true" pageId="5" pageNumber="6">H. charcasus</emphasis>
</taxonomicName>
(
<figureCitation id="132B2A5FFFBFF119FEFA013372F4FD37" box="[319,399,627,651]" captionStart="Fig" captionStartId="7.[112,150,1615,1637]" captionTargetBox="[106,1168,210,1581]" captionTargetId="figure-1@7.[106,1168,210,1581]" captionText="Fig. 4. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides carinatus Pocock, 1900, ♂. B. Hadruroides chinchaysuyu, n. sp., ♀. C. Hadruroides geckoi, n. sp., ♂. D. Hadruroides graceae, n. sp., ♂. E. Hadruroides charcasus (Karsch, 1879), ♂." figureDoi="http://doi.org/10.5281/zenodo.5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="5" pageNumber="6">fig. 4E</figureCitation>
),
<taxonomicName id="4C104D59FFBFF119FE6001337125FD37" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[421,606,627,651]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chinchaysuyu" status="sp. nov.">
<emphasis id="B964EAC8FFBFF119FE6001337125FD37" box="[421,606,627,651]" italics="true" pageId="5" pageNumber="6">H. chinchaysuyu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBFF119FF9D01D173EEFD15" box="[88,149,657,681]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="132B2A5FFFBFF119FF6101D1738EFD15" box="[164,245,657,681]" captionStart="Fig" captionStartId="7.[112,150,1615,1637]" captionTargetBox="[106,1168,210,1581]" captionTargetId="figure-1@7.[106,1168,210,1581]" captionText="Fig. 4. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides carinatus Pocock, 1900, ♂. B. Hadruroides chinchaysuyu, n. sp., ♀. C. Hadruroides geckoi, n. sp., ♂. D. Hadruroides graceae, n. sp., ♂. E. Hadruroides charcasus (Karsch, 1879), ♂." figureDoi="http://doi.org/10.5281/zenodo.5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="5" pageNumber="6">fig. 4B</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FECC01D1720FFD15" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[265,372,657,681]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="geckoi" status="sp. nov.">
<emphasis id="B964EAC8FFBFF119FECC01D1720FFD15" box="[265,372,657,681]" italics="true" pageId="5" pageNumber="6">H. geckoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBFF119FE4601D172BBFD15" box="[387,448,657,681]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(figs. 3B,
<figureCitation id="132B2A5FFFBFF119FDF001D17123FD15" box="[565,600,657,681]" captionStart="Fig" captionStartId="7.[112,150,1615,1637]" captionTargetBox="[106,1168,210,1581]" captionTargetId="figure-1@7.[106,1168,210,1581]" captionText="Fig. 4. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides carinatus Pocock, 1900, ♂. B. Hadruroides chinchaysuyu, n. sp., ♀. C. Hadruroides geckoi, n. sp., ♂. D. Hadruroides graceae, n. sp., ♂. E. Hadruroides charcasus (Karsch, 1879), ♂." figureDoi="http://doi.org/10.5281/zenodo.5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="5" pageNumber="6">4C</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FF9D01EF72DCFD7A" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[88,423,686,711]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leopardus">
<emphasis id="B964EAC8FFBFF119FF9D01EF7389FD7B" box="[88,242,687,711]" italics="true" pageId="5" pageNumber="6">H. leopardus</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FEC101EE72DCFD7A" author="Pocock, R. I." box="[260,423,686,710]" pageId="5" pageNumber="6" pagination="469 - 478" refId="ref31046" refString="Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History 5 (7): 469 - 478." type="journal article" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
(figs. 3A,
<figureCitation id="132B2A5FFFBFF119FDF301EE7122FD7A" box="[566,601,686,710]" captionStart="Fig" captionStartId="8.[151,189,1615,1637]" captionTargetBox="[136,1218,210,1581]" captionTargetId="figure-0@8.[136,1218,210,1581]" captionText="Fig. 5. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides juanchaparroi, n. sp., ♂. B. Hadruroides leopardus Pocock, 1900, ♂. C. Hadruroides lunatus (L. Koch, 1867), ♀. D. Hadruroides vichayitos, n. sp., ♂. E. Hadruroides tishqu, n. sp., ♂." figureDoi="http://doi.org/10.5281/zenodo.5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="5" pageNumber="6">5B</figureCitation>
). Three species occur in inter-Andean valleys along the Cordillera:
<taxonomicName id="4C104D59FFBFF119FE9801AA7230FC9C" authority="Ochoa and Chaparro, 2008" authorityName="Ochoa and Chaparro" authorityYear="2008" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bustamantei">
<emphasis id="B964EAC8FFBFF119FE9801AA7175FCBE" box="[349,526,746,770]" italics="true" pageId="5" pageNumber="6">H. bustamantei</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FDDE01AA7230FC9C" author="Ochoa, J. A. &amp; J. C. Chaparro" pageId="5" pageNumber="6" pagination="5 - 10" refId="ref30887" refString="Ochoa, J. A., and J. C. Chaparro. 2008. Nueva especie de escorpion del genero Hadruroides (Scorpiones: Caraboctoninae) de los valles interandinos de Peru. Revista Peruana de Biologia 15 (1): 5 - 10." type="journal article" year="2008">Ochoa and Chaparro, 2008</bibRefCitation>
</taxonomicName>
(fig. 3D);
<taxonomicName id="4C104D59FFBFF119FE110048738CFC81" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B964EAC8FFBFF119FE110048711EFC9C" box="[468,613,776,800]" italics="true" pageId="5" pageNumber="6">H. carinatus</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FF9D0065738CFC81" author="Pocock, R. I." box="[88,247,805,829]" pageId="5" pageNumber="6" pagination="469 - 478" refId="ref31046" refString="Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History 5 (7): 469 - 478." type="journal article" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
(
<figureCitation id="132B2A5FFFBFF119FECB0065721AFC81" box="[270,353,805,829]" captionStart="Fig" captionStartId="7.[112,150,1615,1637]" captionTargetBox="[106,1168,210,1581]" captionTargetId="figure-1@7.[106,1168,210,1581]" captionText="Fig. 4. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides carinatus Pocock, 1900, ♂. B. Hadruroides chinchaysuyu, n. sp., ♀. C. Hadruroides geckoi, n. sp., ♂. D. Hadruroides graceae, n. sp., ♂. E. Hadruroides charcasus (Karsch, 1879), ♂." figureDoi="http://doi.org/10.5281/zenodo.5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="5" pageNumber="6">fig. 4A</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FEBB0066724CFCE7" authority="Francke and Soleglad, 1980" authorityName=": Francke and Soleglad" authorityYear="1980" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mauryi">
<emphasis id="B964EAC8FFBFF119FEBB00667283FC82" box="[382,504,806,830]" italics="true" pageId="5" pageNumber="6">H. mauryi</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FDCD0065724CFCE7" author="Francke, O. F. &amp; M. E. Soleglad" pageId="5" pageNumber="6" pagination="1 - 13" refId="ref29871" refString="Francke, O. F., and M. E. Soleglad. 1980. Two new Hadruroides Pocock from Peru (Scorpiones, Vaejovidae). Occasional Papers of the Museum, Texas Tech University 69: 1 - 13." type="journal article" year="1980">Francke and Soleglad, 1980</bibRefCitation>
</taxonomicName>
. Six species inhabit desert along the Pacific coast:
<taxonomicName id="4C104D59FFBFF119FEB900217229FC2A" authority="Francke and Soleglad, 1980" authorityName="Francke and Soleglad" authorityYear="1980" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aguilari">
<emphasis id="B964EAC8FFBFF119FEB900217280FCC5" box="[380,507,865,889]" italics="true" pageId="5" pageNumber="6">H. aguilari</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FDCD00217229FC2A" author="Francke, O. F. &amp; M. E. Soleglad" pageId="5" pageNumber="6" pagination="1 - 13" refId="ref29871" refString="Francke, O. F., and M. E. Soleglad. 1980. Two new Hadruroides Pocock from Peru (Scorpiones, Vaejovidae). Occasional Papers of the Museum, Texas Tech University 69: 1 - 13." type="journal article" year="1980">Francke and Soleglad, 1980</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C104D59FFBFF119FEAB003F7283FC2B" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[366,504,895,919]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="graceae" status="sp. nov.">
<emphasis id="B964EAC8FFBFF119FEAB003F7283FC2B" box="[366,504,895,919]" italics="true" pageId="5" pageNumber="6">H. graceae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBFF119FDDD003E711DFC2A" box="[536,614,894,918]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="132B2A5FFFBFF119FFA500DC73C8FC08" box="[96,179,924,948]" captionStart="Fig" captionStartId="7.[112,150,1615,1637]" captionTargetBox="[106,1168,210,1581]" captionTargetId="figure-1@7.[106,1168,210,1581]" captionText="Fig. 4. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides carinatus Pocock, 1900, ♂. B. Hadruroides chinchaysuyu, n. sp., ♀. C. Hadruroides geckoi, n. sp., ♂. D. Hadruroides graceae, n. sp., ♂. E. Hadruroides charcasus (Karsch, 1879), ♂." figureDoi="http://doi.org/10.5281/zenodo.5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="5" pageNumber="6">fig. 4D</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FF1500DC72EEFC08" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[208,405,924,948]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="juanchaparroi" status="sp. nov.">
<emphasis id="B964EAC8FFBFF119FF1500DC72EEFC08" box="[208,405,924,948]" italics="true" pageId="5" pageNumber="6">H. juanchaparroi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBFF119FE6F00DC7294FC08" box="[426,495,924,948]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(figs. 3C,
<figureCitation id="132B2A5FFFBFF119FF9D00FA7307FC6E" box="[88,124,954,978]" captionStart="Fig" captionStartId="8.[151,189,1615,1637]" captionTargetBox="[136,1218,210,1581]" captionTargetId="figure-0@8.[136,1218,210,1581]" captionText="Fig. 5. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides juanchaparroi, n. sp., ♂. B. Hadruroides leopardus Pocock, 1900, ♂. C. Hadruroides lunatus (L. Koch, 1867), ♀. D. Hadruroides vichayitos, n. sp., ♂. E. Hadruroides tishqu, n. sp., ♂." figureDoi="http://doi.org/10.5281/zenodo.5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="5" pageNumber="6">5A</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FF5500FA72C6FC6E" authority="(L. Koch, 1867)" baseAuthorityName="L. Koch" baseAuthorityYear="1867" box="[144,445,954,978]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FFBFF119FF5500FA727FFC6E" box="[144,260,954,978]" italics="true" pageId="5" pageNumber="6">H. lunatus</emphasis>
(L.
<bibRefCitation id="EF814B2BFFBFF119FEF700FA72CEFC6E" author="Koch, L." box="[306,437,954,978]" pageId="5" pageNumber="6" pagination="173 - 250" refId="ref30163" refString="Koch, L. 1867. Beschreibungen neuer Arachniden und Myriapoden. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 17: 173 - 250." type="journal article" year="1867">Koch, 1867</bibRefCitation>
)
</taxonomicName>
(figs. 3G,
<figureCitation id="132B2A5FFFBFF119FDF000FA7123FC6E" box="[565,600,954,978]" captionStart="Fig" captionStartId="8.[151,189,1615,1637]" captionTargetBox="[136,1218,210,1581]" captionTargetId="figure-0@8.[136,1218,210,1581]" captionText="Fig. 5. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides juanchaparroi, n. sp., ♂. B. Hadruroides leopardus Pocock, 1900, ♂. C. Hadruroides lunatus (L. Koch, 1867), ♀. D. Hadruroides vichayitos, n. sp., ♂. E. Hadruroides tishqu, n. sp., ♂." figureDoi="http://doi.org/10.5281/zenodo.5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="5" pageNumber="6">5C</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FF9D009873C6FC4C" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[88,189,984,1008]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tishqu" status="sp. nov.">
<emphasis id="B964EAC8FFBFF119FF9D009873C6FC4C" box="[88,189,984,1008]" italics="true" pageId="5" pageNumber="6">H. tishqu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBFF119FF0E0097727CFC53" box="[203,263,983,1007]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(figs. 3E,
<figureCitation id="132B2A5FFFBFF119FEBE009772E6FC53" box="[379,413,983,1007]" captionStart="Fig" captionStartId="8.[151,189,1615,1637]" captionTargetBox="[136,1218,210,1581]" captionTargetId="figure-0@8.[136,1218,210,1581]" captionText="Fig. 5. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides juanchaparroi, n. sp., ♂. B. Hadruroides leopardus Pocock, 1900, ♂. C. Hadruroides lunatus (L. Koch, 1867), ♀. D. Hadruroides vichayitos, n. sp., ♂. E. Hadruroides tishqu, n. sp., ♂." figureDoi="http://doi.org/10.5281/zenodo.5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="5" pageNumber="6">5E</figureCitation>
);
<taxonomicName id="4C104D59FFBFF119FE7400987139FC4C" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[433,578,984,1008]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vichayitos" status="sp. nov.">
<emphasis id="B964EAC8FFBFF119FE7400987139FC4C" box="[433,578,984,1008]" italics="true" pageId="5" pageNumber="6">H. vichayitos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBFF119FD9500977303FBB1" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(figs. 3F,
<figureCitation id="132B2A5FFFBFF119FF3D00B57265FBB1" box="[248,286,1013,1037]" captionStart="Fig" captionStartId="8.[151,189,1615,1637]" captionTargetBox="[136,1218,210,1581]" captionTargetId="figure-0@8.[136,1218,210,1581]" captionText="Fig. 5. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides juanchaparroi, n. sp., ♂. B. Hadruroides leopardus Pocock, 1900, ♂. C. Hadruroides lunatus (L. Koch, 1867), ♀. D. Hadruroides vichayitos, n. sp., ♂. E. Hadruroides tishqu, n. sp., ♂." figureDoi="http://doi.org/10.5281/zenodo.5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="5" pageNumber="6">5D</figureCitation>
). We exclude
<taxonomicName id="4C104D59FFBFF119FE0900B57275FB97" authority="(Thorell, 1876)" baseAuthorityName="Thorell" baseAuthorityYear="1876" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B964EAC8FFBFF119FE0900B5711DFBB1" box="[460,614,1013,1037]" italics="true" pageId="5" pageNumber="6">H. maculatus</emphasis>
(
<bibRefCitation id="EF814B2BFFBFF119FF9A0753727DFB97" author="Thorell, T." box="[95,262,1043,1067]" pageId="5" pageNumber="6" pagination="75 - 272" refId="ref31656" refString="Thorell, T. 1876. Etudes scorpiologiques. Atti della Societa Italiana di Scienze Naturali 19: 75 - 272." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
)
</taxonomicName>
, which is restricted to the central coastline of
<collectingCountry id="F307764AFFBFF119FE85077172DFFBF5" box="[320,420,1073,1097]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
, from the list of Peruvian species. Two other
<taxonomicName id="4C104D59FFBFF119FE19070E73DEFB38" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B964EAC8FFBFF119FE19070E711DFBDA" box="[476,614,1102,1126]" italics="true" pageId="5" pageNumber="6">Hadruroides</emphasis>
species
</taxonomicName>
are endemic to
<collectingCountry id="F307764AFFBFF119FEB6072C72A3FB38" box="[371,472,1132,1156]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
:
<taxonomicName id="4C104D59FFBFF119FE2C072C7237FB1E" authority="Maury, 1975" authorityName="Maury" authorityYear="1975" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="galapagoensis">
<emphasis id="B964EAC8FFBFF119FE2C072C73DCFB1E" italics="true" pageId="5" pageNumber="6">H. galapagoensis</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FF7607CA7237FB1E" author="Maury, E. A." box="[179,332,1162,1186]" pageId="5" pageNumber="6" pagination="9 - 21" refId="ref30635" refString="Maury, E. A. 1975. Escorpiones y escorpionismo en el Peru IV: Revision del genero Hadruroides Pocock, 1893 (Scorpiones, Vejovidae). Revista Peruana de Entomologia 17 (1): 9 - 21." type="journal article" year="1975">Maury, 1975</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C104D59FFBFF119FE9907CA73F6FB03" authority="Lourenco, 1995" authorityName="Lourenco" authorityYear="1995" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="udvardyi">
<emphasis id="B964EAC8FFBFF119FE9907CA729EFB1E" box="[348,485,1162,1186]" italics="true" pageId="5" pageNumber="6">H. udvardyi</emphasis>
<bibRefCitation id="EF814B2BFFBFF119FE3507CA73F6FB03" author="Lourenco, W. R." pageId="5" pageNumber="6" pagination="61 - 88" refId="ref30381" refString="Lourenco, W. R. 1995. Les scorpions (Chelicerata, Scorpiones) de l'Equateur avec quelques considerations sur la biogeographie et la diversite des especes. Revue Suisse de Zoologie 102 (1): 61 - 88." type="journal article" year="1995">Lourenco, 1995</bibRefCitation>
</taxonomicName>
(
<tableCitation id="C6920361FFBFF119FF6607E77389FB03" box="[163,242,1191,1215]" captionStart="TABLE 1" captionStartId="1.[592,665,211,230]" captionTargetBox="[88,987,324,742]" captionText="TABLE 1 Currently recognized species of the scorpion genus Hadruroides Pocock, 1893, with countries and departments or provinces from which they have been recorded" httpUri="http://table.plazi.org/id/DF6F6652FFBBF11DFD9503937080FEA1" pageId="5" pageNumber="6" tableUuid="DF6F6652FFBBF11DFD9503937080FEA1">table 1</tableCitation>
). Most species of
<taxonomicName id="4C104D59FFBFF119FE1907E8711DFB7C" authorityName="Pocock" authorityYear="1893" box="[476,614,1192,1216]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FE1907E8711DFB7C" box="[476,614,1192,1216]" italics="true" pageId="5" pageNumber="6">Hadruroides</emphasis>
</taxonomicName>
have restricted distributions, except
<taxonomicName id="4C104D59FFBFF119FDC6078573E8FB47" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBFF119FDC6078573E8FB47" italics="true" pageId="5" pageNumber="6">H. charcasus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D59FFBFF119FF1C07A37228FB47" baseAuthorityName="L. Koch" baseAuthorityYear="1867" box="[217,339,1251,1275]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FFBFF119FF1C07A37228FB47" box="[217,339,1251,1275]" italics="true" pageId="5" pageNumber="6">H. lunatus</emphasis>
</taxonomicName>
, which are apparently more widely distributed. We consider it necessary to reassess all previous records of the latter two species, because we suspect several are based on misidentifications. Other records of
<taxonomicName id="4C104D59FFBFF119FF1406377220FA33" authorityName="Pocock" authorityYear="1893" box="[209,347,1399,1423]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FF1406377220FA33" box="[209,347,1399,1423]" italics="true" pageId="5" pageNumber="6">Hadruroides</emphasis>
</taxonomicName>
from the coastal desert of southern
<collectingCountry id="F307764AFFBFF119FF2306D57260FA11" box="[230,283,1429,1453]" name="Peru" pageId="5" pageNumber="6">Peru</collectingCountry>
and northern
<collectingCountry id="F307764AFFBFF119FE0006D5717AFA11" box="[453,513,1429,1453]" name="Chile" pageId="5" pageNumber="6">Chile</collectingCountry>
(
<bibRefCitation id="EF814B2BFFBFF119FDD606D573EBFA76" author="Ochoa, J. A." pageId="5" pageNumber="6" pagination="49 - 68" refId="ref30852" refString="Ochoa, J. A. 2005. Patrones de distribucion de escorpiones de la region andina en el sur peruano. Revista Peruana de Biologia 12 (1): 49 - 68." type="journal article" year="2005">Ochoa, 2005</bibRefCitation>
; unpubl. data) correspond to new species related to
<taxonomicName id="4C104D59FFBFF119FF1406917233FA55" baseAuthorityName="L. Koch" baseAuthorityYear="1867" box="[209,328,1489,1513]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FFBFF119FF1406917233FA55" box="[209,328,1489,1513]" italics="true" pageId="5" pageNumber="6">H. lunatus</emphasis>
</taxonomicName>
(fig. 2), the descriptions of which are in preparation by the authors.
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FFBFF119FF9D05797141F93F" pageId="5" pageNumber="6" type="description">
<paragraph id="8BAF36DAFFBFF119FF9D0579711EF9EE" blockId="5.[88,613,1593,1618]" box="[88,613,1593,1618]" pageId="5" pageNumber="6">
<heading id="D0E781B6FFBFF119FF9D0579711EF9EE" allCaps="true" box="[88,613,1593,1618]" centered="true" fontSize="10" level="1" pageId="5" pageNumber="6" reason="1">
PERUVIAN SPECIES OF
<taxonomicName id="4C104D59FFBFF119FE5D057A711EF9EE" authorityName="Pocock" authorityYear="1893" box="[408,613,1594,1618]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBFF119FE5D057A711EF9EE" box="[408,613,1594,1618]" italics="true" pageId="5" pageNumber="6">HADRUROIDES</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FF41052D7141F93F" blockId="5.[132,570,1645,1669]" box="[132,570,1645,1669]" pageId="5" pageNumber="6">
<heading id="D0E781B6FFBFF119FF41052D7141F93F" box="[132,570,1645,1669]" pageId="5" pageNumber="6">KEY TO IDENTIFICATION OF THE SPECIES</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FFBFF115FF9F05E977D8FEBE" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6" type="key">
<key id="FF8161EBFFBFF115FF9F05E977D8FEBE" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
<keyStep id="30E42E7FFFBFF119FF9F05E977DFFE84" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FF9F05E977D8FEBE" blockId="5.[90,614,1705,1754]" lastBlockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FF9F05E977D8FEBE" pageId="5" pageNumber="6">
1. Adult pedipalp chela robust, internomedian, dorsointernal, and dorsal marginal carinae well developed and comprising strong granules (
<figureCitation id="132B2A5FFFBFF119FCC903AC7029FEBE" box="[780,850,236,258]" captionStart="Fig" captionStartId="7.[112,150,1615,1637]" captionTargetBox="[106,1168,210,1581]" captionTargetId="figure-1@7.[106,1168,210,1581]" captionText="Fig. 4. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides carinatus Pocock, 1900, ♂. B. Hadruroides chinchaysuyu, n. sp., ♀. C. Hadruroides geckoi, n. sp., ♂. D. Hadruroides graceae, n. sp., ♂. E. Hadruroides charcasus (Karsch, 1879), ♂." figureDoi="http://doi.org/10.5281/zenodo.5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="5" pageNumber="6">fig. 4E</figureCitation>
).............
<taxonomicName id="4C104D59FFBFF119FBE403AD77D8FEBE" ID-CoL="3J7D6" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[1057,1187,237,258]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBFF119FBE403AD77D8FEBE" box="[1057,1187,237,258]" italics="true" pageId="5" pageNumber="6">H. charcasus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D024777DFFE84" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D024777DFFE84" pageId="5" pageNumber="6">
Pedipalp chela smooth, acarinate (
<figureCitation id="132B2A5FFFBFF119FB8B024777E4FEA1" box="[1102,1183,263,285]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="5" pageNumber="6">figs. 8E</figureCitation>
,
<figureCitation id="132B2A5FFFBFF119FD100262707FFE84" box="[725,772,290,312]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="5" pageNumber="6">11G</figureCitation>
).. .. .. .. ... .. .. .. ... .. .. ... 2
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<keyStep id="30E42E7FFFBFF119FD5D027D77DFFE35" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D027D77DFFEEF" blockId="5.[664,1188,210,1754]" box="[664,1188,317,339]" pageId="5" pageNumber="6">
<key id="FF8161EBFFBFF119FD5D027D77DFFEEF" box="[664,1188,317,339]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D027D77DFFEEF" box="[664,1188,317,339]" pageId="5" pageNumber="6">2. Sternite VII acarinate................ 3</keyLead>
</key>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D021877DFFE35" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D021877DFFE35" pageId="5" pageNumber="6">
Sternite VII with two or four carinae (
<figureCitation id="132B2A5FFFBFF119FB8802187190FE35" captionStart="Fig" captionStartId="11.[112,150,1356,1378]" captionTargetBox="[110,1165,214,1322]" captionTargetId="figure-181@11.[110,1165,214,1322]" captionText="Fig. 7. Hadruroides Pocock, 1893, diagnostic characters. A, C. Hadruroides chinchaysuyu, n. sp., paratype ♂ (MHNC). B, D. Hadruroides maculatus (Thorell, 1876), ♂ (MHNC). A, B. Carapace. C, D. Sternite VII and metasomal segment I, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682518" httpUri="https://zenodo.org/record/5682518/files/figure.png" pageId="5" pageNumber="6">figs. 7C, D</figureCitation>
, 27C)........................ 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFBFF119FD5D02CE77DFFDC5" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D02CE77D8FE4F" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D02CE77D8FE4F" pageId="5" pageNumber="6">
3. Metasomal segments IIV elongated; segment II longer than wide; segment V (♂) approximately three times longer than wide, length:width ratio: 3.18......
<taxonomicName id="4C104D59FFBFF119FBF0029E77D8FE4F" ID-CoL="3J7D3" authorityName="Francke and Soleglad" authorityYear="1980" box="[1077,1187,478,499]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aguilari">
<emphasis id="B964EAC8FFBFF119FBF0029E77D8FE4F" box="[1077,1187,478,499]" italics="true" pageId="5" pageNumber="6">H. aguilari</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D02B977DFFDC5" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D02B977DFFDC5" pageId="5" pageNumber="6">
Metasomal segments IIV relatively short (
<figureCitation id="132B2A5FFFBFF119FD190153704AFD95" box="[732,817,531,553]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="5" pageNumber="6">fig. 20E</figureCitation>
); segment II as wide as long; segment V (♂) approximately twice longer than wide, length:width ratio: 2.042.59 (
<figureCitation id="132B2A5FFFBFF119FD1901237046FDC5" box="[732,829,611,633]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="5" pageNumber="6">figs. 19G</figureCitation>
,
<figureCitation id="132B2A5FFFBFF119FC8C01237009FDC5" box="[841,882,611,633]" captionStart="Fig" captionStartId="44.[151,189,1617,1639]" captionTargetBox="[167,1187,210,1583]" captionTargetId="figure-0@44.[167,1187,210,1583]" captionText="Fig. 23. Hadruroides tishqu, n. sp., diagnostic characters. A. Paratype ♀ (AMNH), telson, lateral aspect. BH. Holotype ³ (MHNC). B. Metasomal segment V, ventral aspect. C. Metasomal segment V and telson, lateral aspect. D. Dextral pedipalp femur, dorsal aspect. E. Dextral pedipalp patella, dorsal aspect. F. Dextral pedipalp chela, ventrointernal aspect. G. Dextral pedipalp chela, external aspect. H. Metasomal segment IV, lateral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713968" httpUri="https://zenodo.org/record/5713968/files/figure.png" pageId="5" pageNumber="6">23B</figureCitation>
)................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFBFF119FD5D013E77DFFC05" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D013E77D8FCA5" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D013E77D8FCA5" pageId="5" pageNumber="6">
4. Pigmentation reduced to faint spots on carapace and tergites (
<figureCitation id="132B2A5FFFBFF119FC0501D9776EFD13" box="[960,1045,665,687]" captionStart="Fig" captionStartId="45.[112,150,1039,1061]" captionTargetBox="[139,1136,210,1005]" captionTargetId="figure-214@45.[139,1137,210,1005]" captionText="Fig. 24. Hadruroides tishqu, n. sp., diagnostic characters. A. Subadult ³ paratype (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. B. Paratype ³ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. C. Paratype ♀ (AMNH), dextral pedipalp chela, external aspect. D. Holotype ³ (MHNC). Dextral pedipalp chela, ventral aspect. E. Paratype ♀ (AMNH), dextral pedipalp chela, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713970" httpUri="https://zenodo.org/record/5713970/files/figure.png" pageId="5" pageNumber="6">fig. 24B</figureCitation>
), absent on metasomal segments; segment V, VM and VL carinae reduced to posterior two-thirds of segment (
<figureCitation id="132B2A5FFFBFF119FCF801A870E9FD42" box="[829,914,744,766]" captionStart="Fig" captionStartId="44.[151,189,1617,1639]" captionTargetBox="[167,1187,210,1583]" captionTargetId="figure-0@44.[167,1187,210,1583]" captionText="Fig. 23. Hadruroides tishqu, n. sp., diagnostic characters. A. Paratype ♀ (AMNH), telson, lateral aspect. BH. Holotype ³ (MHNC). B. Metasomal segment V, ventral aspect. C. Metasomal segment V and telson, lateral aspect. D. Dextral pedipalp femur, dorsal aspect. E. Dextral pedipalp patella, dorsal aspect. F. Dextral pedipalp chela, ventrointernal aspect. G. Dextral pedipalp chela, external aspect. H. Metasomal segment IV, lateral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713968" httpUri="https://zenodo.org/record/5713968/files/figure.png" pageId="5" pageNumber="6">fig. 23B</figureCitation>
); telson, ventral surfaces smooth (♂,
<specimenCount id="9D16FD53FFBFF119FC9300407012FCA6" box="[854,873,768,794]" pageId="5" pageNumber="6" type="female"></specimenCount>
) (
<figureCitation id="132B2A5FFFBFF119FCBA0043708FFCA5" box="[895,1012,771,793]" captionStart="Fig" captionStartId="44.[151,189,1617,1639]" captionTargetBox="[167,1187,210,1583]" captionTargetId="figure-0@44.[167,1187,210,1583]" captionText="Fig. 23. Hadruroides tishqu, n. sp., diagnostic characters. A. Paratype ♀ (AMNH), telson, lateral aspect. BH. Holotype ³ (MHNC). B. Metasomal segment V, ventral aspect. C. Metasomal segment V and telson, lateral aspect. D. Dextral pedipalp femur, dorsal aspect. E. Dextral pedipalp patella, dorsal aspect. F. Dextral pedipalp chela, ventrointernal aspect. G. Dextral pedipalp chela, external aspect. H. Metasomal segment IV, lateral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713968" httpUri="https://zenodo.org/record/5713968/files/figure.png" pageId="5" pageNumber="6">fig. 23A, C</figureCitation>
)....
<taxonomicName id="4C104D59FFBFF119FB83004377D8FCA5" ID-CoL="3J7DL" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[1094,1187,771,793]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tishqu">
<emphasis id="B964EAC8FFBFF119FB83004377D8FCA5" bold="true" box="[1094,1187,771,793]" italics="true" pageId="5" pageNumber="6">H. tishqu</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D005E77DFFC05" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D005E77DFFC05" pageId="5" pageNumber="6">
Pigmentation pronounced on carapace, tergites, and metasomal segments (
<figureCitation id="132B2A5FFFBFF119FB8F00797191FCD5" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="5" pageNumber="6">fig. 20C, D</figureCitation>
); segment V, VM and VL carinae complete (
<figureCitation id="132B2A5FFFBFF119FD19002E704FFC38" box="[732,820,878,900]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="5" pageNumber="6">fig. 19G</figureCitation>
); telson, ventral surfaces smooth or sparsely granular anteriorly (♂), granular (
<specimenCount id="9D16FD53FFBFF119FD1800E0718BFC06" box="[733,752,928,954]" pageId="5" pageNumber="6" type="female"></specimenCount>
) ... .. .. .. ... .. .. .. ... .. .. ... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFBFF119FD5D00FE77D8FB45" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D00FE77D8FB82" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D00FE77D8FB82" pageId="5" pageNumber="6">
5. Tergites IVI with rectangular dorsosubmedian spots of pigmentation (
<figureCitation id="132B2A5FFFBFF119FC0300997761FC53" box="[966,1050,985,1007]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="5" pageNumber="6">fig. 20C</figureCitation>
); sternite VII and metasomal segment I, ventral surfaces unpigmented; hemispermatophore broader basally with relatively long crest...
<taxonomicName id="4C104D59FFBFF119FBF8076977D8FB82" ID-CoL="3J7DG" baseAuthorityName="L. Koch" baseAuthorityYear="1867" box="[1085,1187,1065,1086]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FFBFF119FBF8076977D8FB82" box="[1085,1187,1065,1086]" italics="true" pageId="5" pageNumber="6">H. lunatus</emphasis>
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</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D070377D8FB45" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D070377D8FB45" pageId="5" pageNumber="6">
Tergites IVI with irregular dorsosubmedian spots of pigmentation (
<figureCitation id="132B2A5FFFBFF119FC01071E7760FBC8" box="[964,1051,1118,1140]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="5" pageNumber="6">fig. 20D</figureCitation>
); sternite VII and metasomal segment I, ventral surfaces with several spots of pigmentation surrounding insertion of setae (
<figureCitation id="132B2A5FFFBFF119FC7F07EE7775FB78" box="[954,1038,1198,1220]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="5" pageNumber="6">fig. 20E</figureCitation>
); hemispermatophore slender with short crest (
<figureCitation id="132B2A5FFFBFF119FBE307887713FB62" box="[1062,1128,1224,1246]" captionStart="Fig" captionStartId="36.[151,189,1044,1066]" captionTargetBox="[184,1170,210,1010]" captionTargetId="figure-297@36.[184,1171,210,1010]" captionText="Fig. 21. Hadruroides juanchaparroi, n. sp., paratype ³ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5713964" httpUri="https://zenodo.org/record/5713964/files/figure.png" pageId="5" pageNumber="6">fig. 21</figureCitation>
)........................
<taxonomicName id="4C104D59FFBFF119FC3F07A377D8FB45" ID-CoL="3J7DD" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[1018,1187,1251,1273]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FFBFF119FC3F07A377D8FB45" bold="true" box="[1018,1187,1251,1273]" italics="true" pageId="5" pageNumber="6">H. juanchaparroi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFBFF119FD5D07BE77DFFA25" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D07BE77DFFAF5" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D07BE77DFFAF5" pageId="5" pageNumber="6">
6. Chela fixed finger (adult ♂) curved, creating distinct proximal gap with movable finger when fingers closed (
<figureCitation id="132B2A5FFFBFF119FCAC067370C3FAF5" box="[873,952,1331,1353]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="5" pageNumber="6">figs. 8C</figureCitation>
,
<figureCitation id="132B2A5FFFBFF119FC060673708BFAF5" box="[963,1008,1331,1353]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="5" pageNumber="6">11H</figureCitation>
, 27H)....... 8
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D060E77DFFA25" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D060E77DFFA25" pageId="5" pageNumber="6">
Chela fixed and movable fingers (adult ♂) straight, no proximal gap evident when fingers closed (
<figureCitation id="132B2A5FFFBFF119FCB506C370C0FA25" box="[880,955,1411,1433]" captionStart="Fig" captionStartId="19.[112,150,1613,1635]" captionTargetBox="[147,1128,210,1579]" captionTargetId="figure-0@19.[147,1128,210,1579]" captionText="Fig. 15. Hadruroides graceae, n. sp., holotype ♂ (MHNC). A. Metasomal segment V and telson, lateral aspect. B. Metasomal segment IV, lateral aspect. C. Metasomal segment IV, ventral aspect. D. Metasomal segment V, ventral aspect. E. Dextral pedipalp patella, dorsal aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, ventral aspect. I. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713952" httpUri="https://zenodo.org/record/5713952/files/figure.png" pageId="5" pageNumber="6">fig. 15I</figureCitation>
).............. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFBFF119FD5D06DE77D8F91F" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D06DE77DFF9A3" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D06DE77DFF9A3" pageId="5" pageNumber="6">
7. Metasomal segment IV, ventral surface densely granular (
<figureCitation id="132B2A5FFFBFF119FC8006F970E2FA73" box="[837,921,1465,1487]" captionStart="Fig" captionStartId="19.[112,150,1613,1635]" captionTargetBox="[147,1128,210,1579]" captionTargetId="figure-0@19.[147,1128,210,1579]" captionText="Fig. 15. Hadruroides graceae, n. sp., holotype ♂ (MHNC). A. Metasomal segment V and telson, lateral aspect. B. Metasomal segment IV, lateral aspect. C. Metasomal segment IV, ventral aspect. D. Metasomal segment V, ventral aspect. E. Dextral pedipalp patella, dorsal aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, ventral aspect. I. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713952" httpUri="https://zenodo.org/record/5713952/files/figure.png" pageId="5" pageNumber="6">fig. 15C</figureCitation>
); segment V with 78 ventral setae and 68 lateral setae; pedipalp chela length:width ratio: 5.005.41 (♂), 4.46 5.12 (
<specimenCount id="9D16FD53FFBFF119FCCB0546705BF99C" box="[782,800,1542,1568]" pageId="5" pageNumber="6" type="female"></specimenCount>
)...................
<taxonomicName id="4C104D59FFBFF119FBF2054977DFF9A3" ID-CoL="3J7DC" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[1079,1188,1545,1567]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="graceae">
<emphasis id="B964EAC8FFBFF119FBF2054977DFF9A3" bold="true" box="[1079,1188,1545,1567]" italics="true" pageId="5" pageNumber="6">H. graceae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFBFF119FD5D056477D8F91F" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D056477D8F91F" pageId="5" pageNumber="6">
Metasomal segment IV, ventral surface smooth; segment V with 1218 ventral setae and 35 lateral setae; pedipalp chela length:width ratio: 3.934.54 (♂), 4.234.64 (
<specimenCount id="9D16FD53FFBFF119FD1805CB718BF919" box="[733,752,1675,1701]" pageId="5" pageNumber="6" type="female"></specimenCount>
)....................
<taxonomicName id="4C104D59FFBFF119FBE705CE77D8F91F" ID-CoL="6L926" authorityName="Pocock" authorityYear="1900" box="[1058,1187,1678,1699]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leopardus">
<emphasis id="B964EAC8FFBFF119FBE705CE77D8F91F" box="[1058,1187,1678,1699]" italics="true" pageId="5" pageNumber="6">H. leopardus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFBFF115FD5D05E9711DFEA2" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
<paragraph id="8BAF36DAFFBFF119FD5D05E977DFF966" blockId="5.[664,1188,210,1754]" pageId="5" pageNumber="6">
<keyLead id="30E195EFFFBFF119FD5D05E977DFF966" pageId="5" pageNumber="6">8. Metasomal segments, VSM carinae absent on segments IIII, present on IV........ 12</keyLead>
</paragraph>
<caption id="DF6F6652FFBCF11AFF520551727DF97B" ID-DOI="http://doi.org/10.5281/zenodo.4735537" ID-Zenodo-Dep="4735537" httpUri="https://zenodo.org/record/4735537/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,189,1553,1575]" targetBox="[215,1140,214,1519]" targetPageId="6">
<paragraph id="8BAF36DAFFBCF11AFF520551727DF97B" blockId="6.[128,1228,1553,1735]" pageId="6" pageNumber="7">
Fig. 3.
<taxonomicName id="4C104D59FFBCF11AFF3505527287F99B" ID-CoL="635PM" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[240,508,1553,1575]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBCF11AFF3505527217F99B" box="[240,364,1554,1575]" italics="true" pageId="6" pageNumber="7">Hadruroides</emphasis>
Pocock, 1893
</taxonomicName>
, habitats in Peru.
<emphasis id="B964EAC8FFBCF11AFD76055171B0F99B" bold="true" box="[691,715,1553,1575]" pageId="6" pageNumber="7">A.</emphasis>
Puchaca Alto (Lambayeque Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFF59056C7168F9FE" ID-CoL="6L926" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[156,531,1580,1602]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="leopardus">
<emphasis id="B964EAC8FFBCF11AFF59056C7204F9FD" box="[156,383,1580,1601]" italics="true" pageId="6" pageNumber="7">Hadruroides leopardus</emphasis>
<bibRefCitation id="EF814B2BFFBCF11AFE42056C7168F9FE" author="Pocock, R. I." box="[391,531,1580,1602]" pageId="6" pageNumber="7" pagination="469 - 478" refId="ref31046" refString="Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History 5 (7): 469 - 478." type="journal article" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
.
<emphasis id="B964EAC8FFBCF11AFDDB056C714FF9FE" bold="true" box="[542,564,1580,1602]" pageId="6" pageNumber="7">B.</emphasis>
Balsas (Cajamarca Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFBC4056C77BEF9FD" ID-CoL="3J7DB" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[1025,1221,1580,1601]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="geckoi" status="sp. nov.">
<emphasis id="B964EAC8FFBCF11AFBC4056C77BEF9FD" box="[1025,1221,1580,1601]" italics="true" pageId="6" pageNumber="7">Hadruroides geckoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBCF11AFF45050773C2F9E1" box="[128,185,1607,1629]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B964EAC8FFBCF11AFF05050773A3F9E1" bold="true" box="[192,216,1607,1629]" pageId="6" pageNumber="7">C.</emphasis>
Cerro Campana (La Libertad Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFCDE05077753F9E0" ID-CoL="3J7DD" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[795,1064,1607,1628]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="juanchaparroi" status="sp. nov.">
<emphasis id="B964EAC8FFBCF11AFCDE05077753F9E0" box="[795,1064,1607,1628]" italics="true" pageId="6" pageNumber="7">Hadruroides juanchaparroi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBCF11AFBF30507770BF9E1" box="[1078,1136,1607,1629]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B964EAC8FFBCF11AFBB2050777EBF9E1" bold="true" box="[1143,1168,1607,1629]" pageId="6" pageNumber="7">D.</emphasis>
Wari (Ayacucho Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFDC705217749F9CB" ID-CoL="3J7D4" authority="Ochoa and Chaparro, 2008" authorityName="Ochoa and Chaparro" authorityYear="2008" box="[514,1074,1633,1655]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bustamantei">
<emphasis id="B964EAC8FFBCF11AFDC70521707BF9CA" box="[514,768,1633,1654]" italics="true" pageId="6" pageNumber="7">Hadruroides bustamantei</emphasis>
<bibRefCitation id="EF814B2BFFBCF11AFCCE05217749F9CB" author="Ochoa, J. A. &amp; J. C. Chaparro" box="[779,1074,1633,1655]" pageId="6" pageNumber="7" pagination="5 - 10" refId="ref30887" refString="Ochoa, J. A., and J. C. Chaparro. 2008. Nueva especie de escorpion del genero Hadruroides (Scorpiones: Caraboctoninae) de los valles interandinos de Peru. Revista Peruana de Biologia 15 (1): 5 - 10." type="journal article" year="2008">Ochoa and Chaparro, 2008</bibRefCitation>
</taxonomicName>
.
<emphasis id="B964EAC8FFBCF11AFB850521772CF9CB" bold="true" box="[1088,1111,1633,1655]" pageId="6" pageNumber="7">E.</emphasis>
Isla Santa (Ancash Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFE13053C71EFF92D" ID-CoL="3J7DL" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[470,660,1660,1681]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tishqu" status="sp. nov.">
<emphasis id="B964EAC8FFBCF11AFE13053C71EFF92D" box="[470,660,1660,1681]" italics="true" pageId="6" pageNumber="7">Hadruroides tishqu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBCF11AFD64053C71A2F92E" box="[673,729,1660,1682]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B964EAC8FFBCF11AFD1A053C718FF92E" bold="true" box="[735,756,1660,1682]" pageId="6" pageNumber="7">F.</emphasis>
Playa Vichayitos (Piura Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFF5F05D77204F910" ID-CoL="3J7DP" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[154,383,1687,1708]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vichayitos" status="sp. nov.">
<emphasis id="B964EAC8FFBCF11AFF5F05D77204F910" box="[154,383,1687,1708]" italics="true" pageId="6" pageNumber="7">Hadruroides vichayitos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBCF11AFE4E05D672B9F910" box="[395,450,1686,1708]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B964EAC8FFBCF11AFE0205D6729AF910" bold="true" box="[455,481,1686,1708]" pageId="6" pageNumber="7">G.</emphasis>
Lomas de Lachay (Lima Department), habitat of
<taxonomicName id="4C104D59FFBCF11AFC1B05D7727AF97B" ID-CoL="3J7DG" authority="(L. Koch, 1867)" baseAuthorityName="L. Koch" baseAuthorityYear="1867" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FFBCF11AFC1B05D777DCF910" box="[990,1191,1687,1708]" italics="true" pageId="6" pageNumber="7">Hadruroides lunatus</emphasis>
(L.
<bibRefCitation id="EF814B2BFFBCF11AFF4505F17381F97B" author="Koch, L." box="[128,250,1713,1735]" pageId="6" pageNumber="7" pagination="173 - 250" refId="ref30163" refString="Koch, L. 1867. Beschreibungen neuer Arachniden und Myriapoden. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 17: 173 - 250." type="journal article" year="1867">Koch, 1867</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF6F6652FFBDF11BFFB5050F717DF926" ID-DOI="http://doi.org/10.5281/zenodo.5794285" ID-Zenodo-Dep="5794285" httpUri="https://zenodo.org/record/5794285/files/figure.png" pageId="7" pageNumber="8" startId="7.[112,150,1615,1637]" targetBox="[106,1168,210,1581]" targetPageId="7">
<paragraph id="8BAF36DAFFBDF11BFFB5050F717DF926" blockId="7.[88,1188,1610,1691]" pageId="7" pageNumber="8">
Fig. 4.
<taxonomicName id="4C104D59FFBDF11BFF15050F7290F9D9" ID-CoL="635PM" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[208,491,1615,1637]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFBDF11BFF15050F7237F9D8" box="[208,332,1615,1636]" italics="true" pageId="7" pageNumber="8">Hadruroides</emphasis>
Pocock, 1893
</taxonomicName>
, habitus in life.
<emphasis id="B964EAC8FFBDF11BFD6F050F71B9F9D9" bold="true" box="[682,706,1615,1637]" pageId="7" pageNumber="8">A.</emphasis>
<taxonomicName id="4C104D59FFBDF11BFD0A050F7728F9D9" ID-CoL="3J7D5" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[719,1107,1615,1637]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B964EAC8FFBDF11BFD0A050F70C8F9D8" box="[719,947,1615,1636]" italics="true" pageId="7" pageNumber="8">Hadruroides carinatus</emphasis>
<bibRefCitation id="EF814B2BFFBDF11BFC04050F7728F9D9" author="Pocock, R. I." box="[961,1107,1615,1637]" pageId="7" pageNumber="8" pagination="469 - 478" refId="ref31046" refString="Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History 5 (7): 469 - 478." type="journal article" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
, ♂.
<emphasis id="B964EAC8FFBDF11BFB48050F77DFF9D9" bold="true" box="[1165,1188,1615,1637]" pageId="7" pageNumber="8">B.</emphasis>
<taxonomicName id="4C104D59FFBDF11BFF9D052A7224F9C3" ID-CoL="3J7D7" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[88,351,1642,1663]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chinchaysuyu" status="sp. nov.">
<emphasis id="B964EAC8FFBDF11BFF9D052A7224F9C3" box="[88,351,1642,1663]" italics="true" pageId="7" pageNumber="8">Hadruroides chinchaysuyu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBDF11BFEA9052972DDF9C3" box="[364,422,1641,1663]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
,
<specimenCount id="9D16FD53FFBDF11BFE74052672BFF93C" box="[433,452,1638,1664]" pageId="7" pageNumber="8" type="female"></specimenCount>
.
<emphasis id="B964EAC8FFBDF11BFE1405297292F9C3" bold="true" box="[465,489,1641,1663]" pageId="7" pageNumber="8">C.</emphasis>
<taxonomicName id="4C104D59FFBDF11BFE35052A71C8F9C3" ID-CoL="3J7DB" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[496,691,1642,1663]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="geckoi" status="sp. nov.">
<emphasis id="B964EAC8FFBDF11BFE35052A71C8F9C3" box="[496,691,1642,1663]" italics="true" pageId="7" pageNumber="8">Hadruroides geckoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBDF11BFD0505297181F9C3" box="[704,762,1641,1663]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
, ♂.
<emphasis id="B964EAC8FFBDF11BFCED0529703AF9C3" bold="true" box="[808,833,1641,1663]" pageId="7" pageNumber="8">D.</emphasis>
<taxonomicName id="4C104D59FFBDF11BFC8D052A776CF9C3" ID-CoL="3J7DC" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[840,1047,1642,1663]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="graceae" status="sp. nov.">
<emphasis id="B964EAC8FFBDF11BFC8D052A776CF9C3" box="[840,1047,1642,1663]" italics="true" pageId="7" pageNumber="8">Hadruroides graceae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFBDF11BFBE005297724F9C3" box="[1061,1119,1641,1663]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
, ♂.
<emphasis id="B964EAC8FFBDF11BFB48052977DFF9C3" bold="true" box="[1165,1188,1641,1663]" pageId="7" pageNumber="8">E.</emphasis>
<taxonomicName id="4C104D59FFBDF11BFF9D05C472A5F926" ID-CoL="3J7D6" authority="(Karsch, 1879)" authorityName="Maury" authorityYear="1975" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[88,478,1668,1690]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="charcasus">
<emphasis id="B964EAC8FFBDF11BFF9D05C47247F925" box="[88,316,1668,1689]" italics="true" pageId="7" pageNumber="8">Hadruroides charcasus</emphasis>
(
<bibRefCitation id="EF814B2BFFBDF11BFE8E05C472ACF926" author="Karsch, F." box="[331,471,1668,1690]" pageId="7" pageNumber="8" pagination="97 - 136" refId="ref30073" refString="Karsch, F. 1879. Scorpionologische Beitrage. Part II. Mitteilungen des Munchener Entomologischen Vereins 3: 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
)
</taxonomicName>
, ♂.
</paragraph>
</caption>
<caption id="DF6F6652FFB2F114FF52050F72B5F927" ID-DOI="http://doi.org/10.5281/zenodo.5794287" ID-Zenodo-Dep="5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,189,1615,1637]" targetBox="[136,1218,210,1581]" targetPageId="8">
<paragraph id="8BAF36DAFFB2F114FF52050F72B5F927" blockId="8.[128,1227,1610,1692]" pageId="8" pageNumber="9">
Fig. 5.
<taxonomicName id="4C104D59FFB2F114FF3505107280F9D9" ID-CoL="635PM" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[240,507,1615,1637]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC8FFB2F114FF3505107217F9D9" box="[240,364,1616,1637]" italics="true" pageId="8" pageNumber="9">Hadruroides</emphasis>
Pocock, 1893
</taxonomicName>
, habitus in life.
<emphasis id="B964EAC8FFB2F114FD5E050F71C8F9D9" bold="true" box="[667,691,1615,1637]" pageId="8" pageNumber="9">A.</emphasis>
<taxonomicName id="4C104D59FFB2F114FD7D051070B8F9D9" ID-CoL="3J7DD" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[696,963,1616,1637]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="juanchaparroi" status="sp. nov.">
<emphasis id="B964EAC8FFB2F114FD7D051070B8F9D9" box="[696,963,1616,1637]" italics="true" pageId="8" pageNumber="9">Hadruroides juanchaparroi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFB2F114FC0A050F7773F9D9" box="[975,1032,1615,1637]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
, ♂.
<emphasis id="B964EAC8FFB2F114FBF6050F7731F9D9" bold="true" box="[1075,1098,1615,1637]" pageId="8" pageNumber="9">B.</emphasis>
<taxonomicName id="4C104D59FFB2F114FB8A0510720BF93C" ID-CoL="6L926" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="leopardus">
<emphasis id="B964EAC8FFB2F114FB8A051073A4F9C3" italics="true" pageId="8" pageNumber="9">Hadruroides leopardus</emphasis>
<bibRefCitation id="EF814B2BFFB2F114FF20052A720BF93C" author="Pocock, R. I." box="[229,368,1642,1664]" pageId="8" pageNumber="9" pagination="469 - 478" refId="ref31046" refString="Pocock, R. I. 1900. Some new or little-known Neotropical scorpions in the British Museum. Annals and Magazine of Natural History 5 (7): 469 - 478." type="journal article" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
, ♂.
<emphasis id="B964EAC8FFB2F114FE59052A72CFF93C" bold="true" box="[412,436,1642,1664]" pageId="8" pageNumber="9">C.</emphasis>
<taxonomicName id="4C104D59FFB2F114FE7F052A7054F93C" ID-CoL="3J7DG" authority="(L. Koch, 1867)" baseAuthorityName="L. Koch" baseAuthorityYear="1867" box="[442,815,1642,1664]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FFB2F114FE7F052A71FFF9C3" box="[442,644,1642,1663]" italics="true" pageId="8" pageNumber="9">Hadruroides lunatus</emphasis>
(L.
<bibRefCitation id="EF814B2BFFB2F114FD75052A7053F93C" author="Koch, L." box="[688,808,1642,1664]" pageId="8" pageNumber="9" pagination="173 - 250" refId="ref30163" refString="Koch, L. 1867. Beschreibungen neuer Arachniden und Myriapoden. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 17: 173 - 250." type="journal article" year="1867">Koch, 1867</bibRefCitation>
)
</taxonomicName>
,
<specimenCount id="9D16FD53FFB2F114FCFF05277036F93D" box="[826,845,1639,1665]" pageId="8" pageNumber="9" type="female"></specimenCount>
.
<emphasis id="B964EAC8FFB2F114FC9F052A7008F93C" bold="true" box="[858,883,1642,1664]" pageId="8" pageNumber="9">D.</emphasis>
<taxonomicName id="4C104D59FFB2F114FCBC052A7724F9C3" ID-CoL="3J7DP" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[889,1119,1642,1663]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vichayitos" status="sp. nov.">
<emphasis id="B964EAC8FFB2F114FCBC052A7724F9C3" box="[889,1119,1642,1663]" italics="true" pageId="8" pageNumber="9">Hadruroides vichayitos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFB2F114FBAE052A77DEF93C" box="[1131,1189,1642,1664]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
, ♂.
<emphasis id="B964EAC8FFB2F114FF4505C573EDF927" bold="true" box="[128,150,1669,1691]" pageId="8" pageNumber="9">E.</emphasis>
<taxonomicName id="4C104D59FFB2F114FF5B05C57226F926" ID-CoL="3J7DL" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[158,349,1669,1690]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tishqu" status="sp. nov.">
<emphasis id="B964EAC8FFB2F114FF5B05C57226F926" box="[158,349,1669,1690]" italics="true" pageId="8" pageNumber="9">Hadruroides tishqu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FFB2F114FEA905C572DDF927" box="[364,422,1669,1691]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
, ♂.
</paragraph>
</caption>
<paragraph id="8BAF36DAFFB3F115FF9F0392711DFEA2" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F0392711DFEA2" pageId="9" pageNumber="10"> Metasomal segments, VSM carinae present on segment I and, usually, II and III, absent on IV............................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFB3F115FF9F0264711DFD27" pageId="9" pageNumber="10">
<paragraph id="8BAF36DAFFB3F115FF9F0264711DFE60" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<key id="FF8161EBFFB3F115FF9F0264711DFE60" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F0264711DFE60" pageId="9" pageNumber="10">
9. Pigmentation pronounced on carapace, tergites, and metasomal segments (
<figureCitation id="132B2A5FFFB3F115FE1E027F715FFEE9" box="[475,548,319,341]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="9" pageNumber="10">figs. 8I</figureCitation>
,
<figureCitation id="132B2A5FFFB3F115FD87027F711AFEE9" box="[578,609,319,341]" captionStart="Fig" captionStartId="13.[112,150,1582,1604]" captionTargetBox="[139,1136,210,1548]" captionTargetId="figure-1@13.[139,1136,210,1548]" captionText="Fig. 9. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides chinchaysuyu, n. sp. A. Paratype ♂ (MHNC), leg III, dextral patella, dorsal aspect. B. Paratype ♀ (MHNC), dextral pedipalp chela, external aspect. C. Paratype ♂ (MHNC), metasomal segments IIIV, ventral aspect showing pigmentation pattern. D. Hadruroides maculatus (Thorell, 1876), ♂ (MHNC), leg III, dextral patella, dorsal aspect. EH. H. chinchaysuyu, holotype ♂ (MHNC), sinistral hemispermatophore. E. Ental aspect. F. Dorsal aspect. G. Ectal aspect. H. Ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682527" httpUri="https://zenodo.org/record/5682527/files/figure.png" pageId="9" pageNumber="10">9C</figureCitation>
,
<figureCitation id="132B2A5FFFB3F115FF52021A7399FECC" box="[151,226,346,368]" captionStart="Fig" captionStartId="16.[151,189,1537,1559]" captionTargetBox="[225,1129,210,1504]" captionTargetId="figure-0@16.[225,1129,210,1504]" captionText="Fig. 12. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides geckoi, n. sp., paratype ♀ (MHNC). A. Metasomal segment V and telson, lateral aspect. B. Dextral pedipalp chela, ventrointernal aspect. C. Dextral pedipalp chela, external aspect. D. H. geckoi, paratype ♂ (MHNC), sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. E. Hadruroides carinatus Pocock, 1900, ♂ (MHNC), dextral pedipalp chela, ventral aspect. F. H. geckoi, paratype ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. G. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. H. H. carinatus, ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682535" httpUri="https://zenodo.org/record/5682535/files/figure.png" pageId="9" pageNumber="10">12F, D</figureCitation>
); VSM carinae present on metasomal segments IIII; pedipalp chela fixed finger (adult ♂) strongly curved, creating well developed proximal gap with movable finger when fingers closed (
<figureCitation id="132B2A5FFFB3F115FEA9028672BBFE60" box="[364,448,454,476]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="9" pageNumber="10">fig. 11H</figureCitation>
)......... 10
</keyLead>
</key>
</paragraph>
<paragraph id="8BAF36DAFFB3F115FF9F02A2711DFD27" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F02A2711DFD27" pageId="9" pageNumber="10">
Pigmentation absent, except for small, faint spots on carapace and tergites (
<figureCitation id="132B2A5FFFB3F115FDC102BD7121FDAF" box="[516,602,509,531]" captionStart="Fig" captionStartId="16.[151,189,1537,1559]" captionTargetBox="[225,1129,210,1504]" captionTargetId="figure-0@16.[225,1129,210,1504]" captionText="Fig. 12. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides geckoi, n. sp., paratype ♀ (MHNC). A. Metasomal segment V and telson, lateral aspect. B. Dextral pedipalp chela, ventrointernal aspect. C. Dextral pedipalp chela, external aspect. D. H. geckoi, paratype ♂ (MHNC), sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. E. Hadruroides carinatus Pocock, 1900, ♂ (MHNC), dextral pedipalp chela, ventral aspect. F. H. geckoi, paratype ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. G. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. H. H. carinatus, ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682535" httpUri="https://zenodo.org/record/5682535/files/figure.png" pageId="9" pageNumber="10">fig. 12G</figureCitation>
); VSM carinae obsolete on metasomal segment I, absent on segments II and III; pedipalp chela fixed finger (adult ♂) slightly curved, creating weak proximal gap with movable finger when fingers closed (fig. 27H)......
<taxonomicName id="4C104D59FFB3F115FE2101C5711DFD27" ID-CoL="3J7DP" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[484,614,645,667]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vichayitos">
<emphasis id="B964EAC8FFB3F115FE2101C5711DFD27" bold="true" box="[484,614,645,667]" italics="true" pageId="9" pageNumber="10">H. vichayitos</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFB3F115FF9F01E0711EFC7A" pageId="9" pageNumber="10">
<paragraph id="8BAF36DAFFB3F115FF9F01E0711EFC9E" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F01E0711EFC9E" pageId="9" pageNumber="10">
10. Sternite VII, VL and VSM carinae well developed; metasomal segment V, DL carinae smooth; telson, ventral surface (
<specimenCount id="9D16FD53FFB3F115FD8E01937125FD51" box="[587,606,723,749]" pageId="9" pageNumber="10" type="female"></specimenCount>
) smooth; hemispermatophore lamina apically rounded................
<taxonomicName id="4C104D59FFB3F115FE2D004D711EFC9E" ID-CoL="3J7D5" authorityName="Pocock" authorityYear="1900" box="[488,613,781,802]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B964EAC8FFB3F115FE2D004D711EFC9E" box="[488,613,781,802]" italics="true" pageId="9" pageNumber="10">H. carinatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFB3F115FF9F0068711EFC7A" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F0068711EFC7A" pageId="9" pageNumber="10">
Sternite VII, VL carinae distinct, VSM carinae obsolete (
<figureCitation id="132B2A5FFFB3F115FF3C00037244FCE5" box="[249,319,835,857]" captionStart="Fig" captionStartId="11.[112,150,1356,1378]" captionTargetBox="[110,1165,214,1322]" captionTargetId="figure-181@11.[110,1165,214,1322]" captionText="Fig. 7. Hadruroides Pocock, 1893, diagnostic characters. A, C. Hadruroides chinchaysuyu, n. sp., paratype ♂ (MHNC). B, D. Hadruroides maculatus (Thorell, 1876), ♂ (MHNC). A, B. Carapace. C, D. Sternite VII and metasomal segment I, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682518" httpUri="https://zenodo.org/record/5682518/files/figure.png" pageId="9" pageNumber="10">fig. 7C</figureCitation>
); metasomal segment V, DL carinae sparsely to densely granular (
<figureCitation id="132B2A5FFFB3F115FDCC001E73D7FC33" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="9" pageNumber="10">figs. 8 A, D</figureCitation>
,
<figureCitation id="132B2A5FFFB3F115FF7E0039739DFC33" box="[187,230,889,911]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="9" pageNumber="10">11A</figureCitation>
); telson, ventral surface (
<specimenCount id="9D16FD53FFB3F115FE2800377284FC2D" box="[493,511,887,913]" pageId="9" pageNumber="10" type="female"></specimenCount>
) granular (
<figureCitation id="132B2A5FFFB3F115FF5B00D57398FC17" box="[158,227,917,939]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="9" pageNumber="10">fig. 8B</figureCitation>
); hemispermatophore lamina apically acuminate (
<figureCitation id="132B2A5FFFB3F115FECE00F07222FC7A" box="[267,345,944,966]" captionStart="Fig" captionStartId="13.[112,150,1582,1604]" captionTargetBox="[139,1136,210,1548]" captionTargetId="figure-1@13.[139,1136,210,1548]" captionText="Fig. 9. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides chinchaysuyu, n. sp. A. Paratype ♂ (MHNC), leg III, dextral patella, dorsal aspect. B. Paratype ♀ (MHNC), dextral pedipalp chela, external aspect. C. Paratype ♂ (MHNC), metasomal segments IIIV, ventral aspect showing pigmentation pattern. D. Hadruroides maculatus (Thorell, 1876), ♂ (MHNC), leg III, dextral patella, dorsal aspect. EH. H. chinchaysuyu, holotype ♂ (MHNC), sinistral hemispermatophore. E. Ental aspect. F. Dorsal aspect. G. Ectal aspect. H. Ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682527" httpUri="https://zenodo.org/record/5682527/files/figure.png" pageId="9" pageNumber="10">figs. 9E</figureCitation>
,
<figureCitation id="132B2A5FFFB3F115FEA600F07200FC7A" box="[355,379,944,966]" captionStart="Fig" captionStartId="17.[112,150,896,918]" captionTargetBox="[137,1138,210,862]" captionTargetId="figure-337@17.[137,1138,210,862]" captionText="Fig. 13. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5682539" httpUri="https://zenodo.org/record/5682539/files/figure.png" pageId="9" pageNumber="10">13</figureCitation>
).............. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFB3F115FF9F008B711EFA76" pageId="9" pageNumber="10">
<paragraph id="8BAF36DAFFB3F115FF9F008B711DFB69" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F008B711DFB69" pageId="9" pageNumber="10">
11. Metasomal segment V elongated (♂): length:width ratio: 2.402.89 (
<figureCitation id="132B2A5FFFB3F115FE1000A6715DFC40" box="[469,550,998,1020]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="9" pageNumber="10">fig. 11B</figureCitation>
); segment V, ventral surface sparsely granular, comprising scattered granules in posterior third (
<figureCitation id="132B2A5FFFB3F115FF5B07787394FBF2" box="[158,239,1080,1102]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="9" pageNumber="10">fig. 11B</figureCitation>
); segment V with 1014 DL setae and 913 VL setae; pedipalp chela slender, length:width ratio: 4.414.40 (♂), 4.534.74 (
<specimenCount id="9D16FD53FFB3F115FF5A07C673CAFB1C" box="[159,177,1158,1184]" pageId="9" pageNumber="10" type="female"></specimenCount>
) (
<figureCitation id="132B2A5FFFB3F115FF0207C97259FB23" box="[199,290,1161,1183]" captionStart="Fig" captionStartId="15.[112,150,1615,1637]" captionTargetBox="[121,1154,210,1581]" captionTargetId="figure-0@15.[121,1154,210,1581]" captionText="Fig. 11. Hadruroides geckoi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Metasomal segment V, ventral aspect. C. Dextral pedipalp patella, dorsal aspect. D. Dextral pedipalp patella, ventral aspect. E. Dextral pedipalp chela, ventral aspect. F. Dextral pedipalp femur, dorsal aspect. G. Dextral pedipalp chela, ventrointernal aspect. H. Dextral pedipalp chela, external aspect. Scale bars 5 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5682533" httpUri="https://zenodo.org/record/5682533/files/figure.png" pageId="9" pageNumber="10">figs. 11E</figureCitation>
,
<figureCitation id="132B2A5FFFB3F115FEE807C9722CFB23" box="[301,343,1161,1183]" captionStart="Fig" captionStartId="16.[151,189,1537,1559]" captionTargetBox="[225,1129,210,1504]" captionTargetId="figure-0@16.[225,1129,210,1504]" captionText="Fig. 12. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides geckoi, n. sp., paratype ♀ (MHNC). A. Metasomal segment V and telson, lateral aspect. B. Dextral pedipalp chela, ventrointernal aspect. C. Dextral pedipalp chela, external aspect. D. H. geckoi, paratype ♂ (MHNC), sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. E. Hadruroides carinatus Pocock, 1900, ♂ (MHNC), dextral pedipalp chela, ventral aspect. F. H. geckoi, paratype ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. G. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. H. H. carinatus, ♂ (MHNC), tergite IV, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682535" httpUri="https://zenodo.org/record/5682535/files/figure.png" pageId="9" pageNumber="10">12B</figureCitation>
); pectinal tooth count: 21 23 (♂), 1821 (
<specimenCount id="9D16FD53FFB3F115FEEB07E1723BFB07" box="[302,320,1185,1211]" pageId="9" pageNumber="10" type="female"></specimenCount>
); hemispermatophore lamina curved in distal half (
<figureCitation id="132B2A5FFFB3F115FEA907FF72D6FB69" box="[364,429,1215,1237]" captionStart="Fig" captionStartId="17.[112,150,896,918]" captionTargetBox="[137,1138,210,862]" captionTargetId="figure-337@17.[137,1138,210,862]" captionText="Fig. 13. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5682539" httpUri="https://zenodo.org/record/5682539/files/figure.png" pageId="9" pageNumber="10">fig. 13</figureCitation>
).....
<taxonomicName id="4C104D59FFB3F115FDC307FF711DFB69" ID-CoL="3J7DB" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[518,614,1215,1237]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="geckoi">
<emphasis id="B964EAC8FFB3F115FDC307FF711DFB69" bold="true" box="[518,614,1215,1237]" italics="true" pageId="9" pageNumber="10">H. geckoi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFB3F115FF9F079B711EFA76" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F079B711EFA76" pageId="9" pageNumber="10">
Metasomal segment V short (♂): length:width ratio: 1.972.14 (
<figureCitation id="132B2A5FFFB3F115FE8207B672F5FAB0" box="[327,398,1270,1292]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="9" pageNumber="10">fig. 8F</figureCitation>
); segment V, ventral surface densely granular (
<figureCitation id="132B2A5FFFB3F115FE5A0651729DFA9B" box="[415,486,1297,1319]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="9" pageNumber="10">fig. 8F</figureCitation>
); segment V with 58 DL setae and 610 VL setae; pedipalp chela relatively broad, length:width ratio: 3.664.02 (♂), 4.04.18 (
<specimenCount id="9D16FD53FFB3F115FE20061F7283FAC5" box="[485,504,1375,1401]" pageId="9" pageNumber="10" type="female"></specimenCount>
) (
<figureCitation id="132B2A5FFFB3F115FDD706227122FAC4" box="[530,601,1378,1400]" captionStart="Fig" captionStartId="12.[151,189,1597,1619]" captionTargetBox="[181,1175,191,1563]" captionTargetId="figure-0@12.[180,1175,191,1563]" captionText="Fig. 8. Hadruroides chinchaysuyu, n. sp., diagnostic characters. A, B. Paratype ♀ (MHNC). A. Metasomal segment V, lateral aspect. B. Telson, lateral aspect. CI. Paratype ♂ (MHNC). C. Dextral pedipalp chela, external aspect. D. Metasomal segments IV, V and telson, lateral aspect. E. Dextral pedipalp chela, ventral aspect. F. Metasomal segment IV, ventral aspect. G. Dextral pedipalp femur, dorsal aspect. H. Dextral pedipalp patella, dorsal aspect. I. Tergite V, dorsal aspect showing pigmentation pattern. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682521" httpUri="https://zenodo.org/record/5682521/files/figure.png" pageId="9" pageNumber="10">fig. 8E</figureCitation>
); pectinal tooth count: 1720 (♂), 16 (
<specimenCount id="9D16FD53FFB3F115FD81063A712CFA28" box="[580,599,1402,1428]" pageId="9" pageNumber="10" type="female"></specimenCount>
); hemispermatophore lamina straight in distal half (
<figureCitation id="132B2A5FFFB3F115FF0B06F4724FFA76" box="[206,308,1460,1482]" captionStart="Fig" captionStartId="13.[112,150,1582,1604]" captionTargetBox="[139,1136,210,1548]" captionTargetId="figure-1@13.[139,1136,210,1548]" captionText="Fig. 9. Hadruroides Pocock, 1893, diagnostic characters. AC. Hadruroides chinchaysuyu, n. sp. A. Paratype ♂ (MHNC), leg III, dextral patella, dorsal aspect. B. Paratype ♀ (MHNC), dextral pedipalp chela, external aspect. C. Paratype ♂ (MHNC), metasomal segments IIIV, ventral aspect showing pigmentation pattern. D. Hadruroides maculatus (Thorell, 1876), ♂ (MHNC), leg III, dextral patella, dorsal aspect. EH. H. chinchaysuyu, holotype ♂ (MHNC), sinistral hemispermatophore. E. Ental aspect. F. Dorsal aspect. G. Ectal aspect. H. Ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5682527" httpUri="https://zenodo.org/record/5682527/files/figure.png" pageId="9" pageNumber="10">fig. 9EH</figureCitation>
)........
<taxonomicName id="4C104D59FFB3F115FE7A06F4711EFA76" ID-CoL="3J7D7" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[447,613,1460,1482]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chinchaysuyu">
<emphasis id="B964EAC8FFB3F115FE7A06F4711EFA76" bold="true" box="[447,613,1460,1482]" italics="true" pageId="9" pageNumber="10">H. chinchaysuyu</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E42E7FFFB3F115FF9F068F77D8FEBE" pageId="9" pageNumber="10">
<paragraph id="8BAF36DAFFB3F115FF9F068F711EF93B" blockId="9.[90,614,210,1755]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F068F711EF93B" pageId="9" pageNumber="10">
12. Tergites IIV each with paired dorsosubmedian and dorsolateral spots of pigmentation, forming four longitudinal stripes along mesosoma; legs IIV each with several prolateral spots of pigmentation; pedipalp chela (♂) relatively broad, length:width ratio: 3.093.36.............
<taxonomicName id="4C104D59FFB3F115FE0E0532711EF93B" ID-CoL="3J7D4" authorityName="Ochoa and Chaparro" authorityYear="2008" box="[459,613,1650,1671]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bustamantei">
<emphasis id="B964EAC8FFB3F115FE0E0532711EF93B" box="[459,613,1650,1671]" italics="true" pageId="9" pageNumber="10">H. bustamantei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF36DAFFB3F115FF9F05CD77D8FEBE" blockId="9.[90,614,210,1755]" lastBlockId="9.[725,1188,210,259]" pageId="9" pageNumber="10">
<keyLead id="30E195EFFFB3F115FF9F05CD77D8FEBE" pageId="9" pageNumber="10">
Tergites IIV unpigmented, mesosoma without longitudinal stripes; legs without prolateral spots of pigmentation; pedipalp chela (♂) relatively slender, length:width ratio: 2.7 2.8......................
<taxonomicName id="4C104D59FFB3F115FBFE03AD77D8FEBE" ID-CoL="3J7DJ" authorityName=": Francke and Soleglad" authorityYear="1980" box="[1083,1187,237,258]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mauryi">
<emphasis id="B964EAC8FFB3F115FBFE03AD77D8FEBE" box="[1083,1187,237,258]" italics="true" pageId="9" pageNumber="10">H. mauryi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>