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<document id="FD3865E868F929D3318F13E570893BF5" ID-CLB-Dataset="56108" ID-DOI="10.6620/ZS.2019.58-14" ID-GBIF-Dataset="de0647eb-9b73-4bac-9ff7-f7bff7518dc0" ID-PMC="PMC6759935" ID-PubMed="31966315" ID-Zenodo-Dep="8065869" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="tatiana" checkinTime="1687182873363" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Nguyen, Manh-Ha, Nguyen, Thu-Anh T. &amp; Phung, Hong-Luong T." docDate="2019" docId="03B987C4FFE3AE55FCA2FE38FC89A9B4" docLanguage="en" docName="ZoolStud.58.14.1-31.pdf" docOrigin="Zoological Studies 58 (14)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Hylomus proximus" docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="FF80FFBCFFEAAE5FFFF0FFA5FF8AAB21" masterDocTitle="Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="10" updateTime="1699534267754" updateUser="ExternalLinkService">
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<mods:title id="3647FF915880D90409CFAF6F37EE2520">Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species</mods:title>
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<mods:namePart id="01D64414BE3ED59EA0CF8EDA735458E7">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="770733B33EB9E0CB1C492B885C1A5409">Nguyen, Manh-Ha</mods:namePart>
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<mods:namePart id="D199D1396DD1BA1C3365B0625AB1CFE2">Nguyen, Thu-Anh T.</mods:namePart>
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<mods:namePart id="9B7DA0B14FF4FB8DC41498BA25DE1E4B">Phung, Hong-Luong T.</mods:namePart>
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<mods:title id="96947AFACEACE7B356F3E386AE637062">Zoological Studies</mods:title>
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<mods:date id="67F391355430D9986EBA9326E0FDF1B7">2019</mods:date>
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<treatment id="03B987C4FFE3AE55FCA2FE38FC89A9B4" ID-DOI="http://doi.org/10.5281/zenodo.8065922" ID-GBIF-Taxon="210357018" ID-Zenodo-Dep="8065922" LSID="urn:lsid:plazi:treatment:03B987C4FFE3AE55FCA2FE38FC89A9B4" httpUri="http://treatment.plazi.org/id/03B987C4FFE3AE55FCA2FE38FC89A9B4" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C30A6559FFE3AE56FCA2FE38FB31AAF7" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BAF36D2FFE3AE56FCA2FE38FB31AAF7" blockId="9.[850,1376,412,501]" pageId="9" pageNumber="10">
<taxonomicName id="4C104D51FFE3AE56FCA2FE38FB31AAF7" ID-CoL="3NGDD" authority="(Nguyen, Golovatch &amp; Anichkin, 2005)" baseAuthorityName="Nguyen, Golovatch &amp; Anichkin" baseAuthorityYear="2005" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B964EAC0FFE3AE56FCA2FE38FB31AAF7" bold="true" pageId="9" pageNumber="10">
<heading id="D0E781BEFFE3AE56FCA2FE38FAEAAA97" bold="true" box="[850,1376,412,438]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B964EAC0FFE3AE56FCA2FE38FBC8AA97" bold="true" box="[850,1090,413,438]" italics="true" pageId="9" pageNumber="10">Hylomus proximus</emphasis>
(
<bibRefCitation id="EF814B23FFE3AE56FBA1FE38FAEAAA97" author="Nguyen AD &amp; Golovatch SI &amp; Anichkin AE" box="[1105,1376,412,438]" pageId="9" pageNumber="10" pagination="251 - 257" refId="ref12004" refString="Nguyen AD, Golovatch SI, Anichkin AE. 2005. The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta 14 (3): 251 - 257." type="journal article" year="2005">Nguyen, Golovatch &amp;</bibRefCitation>
</heading>
Anichkin, 2005)
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE3AE56FBE8FE78FB10AAD4" box="[1048,1178,477,501]" pageId="9" pageNumber="10" type="description">
<paragraph id="8BAF36D2FFE3AE56FBE8FE78FB10AAD4" blockId="9.[850,1376,412,501]" box="[1048,1178,477,501]" pageId="9" pageNumber="10">
(
<figureCitation id="132B2A57FFE3AE56FBD0FE78FBE3AAD4" box="[1056,1129,477,501]" captionStart="Fig" captionStartId="2.[175,206,1813,1832]" captionTargetBox="[440,1185,515,1783]" captionTargetId="figure-96@2.[440,1185,486,1804]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Hylomus species in mainland Vietnam. 1: Hoang Lien National Park (Sa Pa, Lao Cai Province); 2: Nam Xay Commune (Van Ban District, Lao Cai Province); 3: Duc Xuan Commune (Bac Quang District, Ha Giang Province); 4: Xuan Son National Park (Phu Tho Province); 5: Tam Dao (Vinh Phuc Province); 6: Cuc Phuong National Park (Ninh Binh Province); 7: Phong Nha – Ke Bang National Park (Quang Binh Province); 8: Lien Chieu District (Da Nang Province); 9: Ba Na National Park (Da Nang Province); 10: Lo Xo Pass (Kon Tum Province); 11: Ngoc Linh Mt. (Kon Tum Province); 12: Kon Chu Rang Natural Reserve (Gia Lai Province); 13: Buon Ho District (Dak Lak Province); 14: Hon Ba Mt. (Khanh Hoa Province); 15: Phan Rang (Ninh Thuan Province); 16: Cat Tien National Park (Dong Nai Province)." figureDoi="http://doi.org/10.5281/zenodo.8065873" httpUri="https://zenodo.org/record/8065873/files/figure.png" pageId="9" pageNumber="10">Figs. 1</figureCitation>
,
<figureCitation id="132B2A57FFE3AE56FB87FE78FB18AAD4" box="[1143,1170,477,501]" captionStart="Fig" captionStartId="13.[136,167,1926,1945]" captionTargetBox="[231,1319,811,1899]" captionTargetId="figure-260@13.[231,1319,811,1899]" captionTargetPageId="13" captionText="Fig. 13. Hylomus proximus (Nguyen, Golovatch &amp; Anichkin, 2005) from Hoang Lien National Park. Right gonopod, lateral view (A), mesal view (B). Scale bars: A = 0.3 mm; B = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065910" httpUri="https://zenodo.org/record/8065910/files/figure.png" pageId="9" pageNumber="10">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE3AE56FCDDFDBCFB4AA940" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8BAF36D2FFE3AE56FCDDFDBCFC0BA967" blockId="9.[813,1415,536,609]" pageId="9" pageNumber="10">
<taxonomicName id="4C104D51FFE3AE56FCDDFDBCFAD2A90A" ID-CoL="354BW" authority="Nguyen, Golovatch &amp; Anichkin, 2005: 252" authorityName="Nguyen, Golovatch &amp; Anichkin" authorityPageNumber="252" authorityYear="2005" box="[813,1368,536,556]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="proxima">
<emphasis id="B964EAC0FFE3AE56FCDDFDBCFC54A90A" box="[813,990,537,555]" italics="true" pageId="9" pageNumber="10">Desmoxytes proxima</emphasis>
<bibRefCitation id="EF814B23FFE3AE56FC14FDBCFAD2A90A" author="Nguyen AD &amp; Golovatch SI &amp; Anichkin AE" box="[996,1368,536,556]" pageId="9" pageNumber="10" pagination="251 - 257" refId="ref12004" refString="Nguyen AD, Golovatch SI, Anichkin AE. 2005. The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta 14 (3): 251 - 257." type="journal article" year="2005">Nguyen, Golovatch &amp; Anichkin, 2005: 252</bibRefCitation>
</taxonomicName>
, figs. 1−5.
</paragraph>
<paragraph id="8BAF36D2FFE3AE56FCDDFDE8FB4AA940" blockId="9.[813,1415,536,609]" box="[813,1216,589,609]" pageId="9" pageNumber="10">
<taxonomicName id="4C104D51FFE3AE56FCDDFDE8FB36A940" ID-CoL="3NGDD" authority=": Srisonchai et al. 2018 a: 11" authorityName="Srisonchai" authorityPageNumber="11" authorityYear="2018" box="[813,1212,589,609]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B964EAC0FFE3AE56FCDDFDE8FC47A941" box="[813,973,589,608]" italics="true" pageId="9" pageNumber="10">Hylomus proximus</emphasis>
:
<bibRefCitation id="EF814B23FFE3AE56FC28FDE8FB36A940" author="Srisonchai R &amp; Enghoff H &amp; Likhitrakarn N &amp; Panha S." box="[984,1212,589,609]" pageId="9" pageNumber="10" pagination="1 - 177" refId="ref12093" refString="Srisonchai R, Enghoff H, Likhitrakarn N, Panha S. 2018 a. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zookeys 761: 1 - 177. doi: 10.3897 / zookeys. 761.24214." type="journal article" year="2018">Srisonchai et al. 2018a: 11</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE3AE55FC99FD2CFE54AA54" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BAF36D2FFE3AE55FC99FD2CFE54AA54" blockId="9.[813,1414,649,929]" lastBlockId="10.[174,775,253,661]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B964EAC0FFE3AE56FC99FD2CFBCAA980" box="[873,1088,649,673]" italics="true" pageId="9" pageNumber="10">Material examined</emphasis>
:
<specimenCount id="9D16FD5BFFE3AE56FBA2FD2CFB15A980" box="[1106,1183,649,673]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
(IEBR-28)
<collectingRegion id="49D4F830FFE3AE56FADAFD2CFC11A9E0" country="Vietnam" name="Lao Cai" pageId="9" pageNumber="10">Lao Cai Province</collectingRegion>
, Van Ban District, Nam Xay commune, bamboo forest,
<quantity id="4CE89B37FFE3AE56FC12FD6CFBCEA9C1" box="[994,1092,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="1000.0">1,000 m</quantity>
,
<date id="FFAE1012FFE3AE56FBA2FD6CFB63A9C0" box="[1106,1257,713,737]" pageId="9" pageNumber="10" value="2005-04-06">6 April 2005</date>
, coll. Anh D. Nguyen;
<specimenCount id="9D16FD5BFFE3AE56FC7FFD4CFC7CA820" box="[911,1014,745,769]" pageId="9" pageNumber="10" type="female">2 females</specimenCount>
(IEBR-33) same locality and habitat,
<date id="FFAE1012FFE3AE56FCDDFCACFC3DA800" box="[813,951,777,801]" pageId="9" pageNumber="10" value="2005-04-08">8 April 2005</date>
, coll. Anh D. Nguyen;
<specimenCount id="9D16FD5BFFE3AE56FB5FFCACFA87A800" box="[1199,1293,777,801]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
(IEBR-34) same locality, secondary forest,
<quantity id="4CE89B37FFE3AE56FB7AFC8CFB5BA861" box="[1162,1233,809,833]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="10" unit="m" value="850.0">850 m</quantity>
,
<date id="FFAE1012FFE3AE56FB2EFC8CFAE6A860" box="[1246,1388,809,833]" pageId="9" pageNumber="10" value="2005-04-03">3 April 2005</date>
;
<specimenCount id="9D16FD5BFFE3AE56FA88FC8CFCEBA840" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
(IEBR-44)
<collectingRegion id="49D4F830FFE3AE56FC2FFCECFB11A840" box="[991,1179,841,865]" country="Vietnam" name="Lao Cai" pageId="9" pageNumber="10">Lao Cai Province</collectingRegion>
, Hoang Lien National Park, Sa Pa, forest,
<quantity id="4CE89B37FFE3AE56FBE0FCCCFBE5A8A1" box="[1040,1135,873,897]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="m" value="2000.0">2,000 m</quantity>
,
<date id="FFAE1012FFE3AE56FB8DFCCCFAC2A8A0" box="[1149,1352,873,897]" pageId="9" pageNumber="10" value="2005-12-02">2 December 2005</date>
, coll. Anh D. Nguyen;
<specimenCount id="9D16FD5BFFE3AE56FC00FC2CFBC2A880" box="[1008,1096,905,929]" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
,
<specimenCount id="9D16FD5BFFE3AE56FBA6FC2CFB48A880" box="[1110,1218,905,929]" pageId="9" pageNumber="10" type="female">2 females</specimenCount>
(IEBR-Myr 243) same locality, but bamboo forest, near a stream,
<quantity id="4CE89B37FFE0AE55FD5AFF5BFC89AA34" box="[682,771,254,278]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="11" unit="m" value="1800.0">1,800 m</quantity>
,
<date id="FFAE1012FFE0AE55FF5EFEB8FEDEAA17" box="[174,340,285,310]" pageId="10" pageNumber="11" value="2007-03-26">26 March 2007</date>
, coll. Anh D. Nguyen;
<specimenCount id="9D16FD5BFFE0AE55FDBAFEBBFD15AA14" box="[586,671,285,310]" pageId="10" pageNumber="11" type="male">3 males</specimenCount>
,
<specimenCount id="9D16FD5BFFE0AE55FD5AFEBBFC8DAA14" box="[682,775,285,310]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(IEBR-42) same locality, but forest,
<quantity id="4CE89B37FFE0AE55FDACFE9BFD34AA74" box="[604,702,318,342]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2000.0">2,000 m</quantity>
,
<date id="FFAE1012FFE0AE55FD3BFE98FF6CAA57" pageId="10" pageNumber="11" value="2003-04">April 2003</date>
, coll. Nguyen Tri Tien.
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE0AE55FF1AFEDBFD7FA917" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BAF36D2FFE0AE55FF1AFEDBFD7FA917" blockId="10.[174,775,253,661]" pageId="10" pageNumber="11">
<emphasis id="B964EAC0FFE0AE55FF1AFEDBFEDCAAB4" box="[234,342,382,405]" italics="true" pageId="10" pageNumber="11">Diagnosis</emphasis>
: The species differs from its congeners in the transverse sulcus starting from metatergum 2; paraterga antler-shaped; male leg femora 5 and 6 humped ventrally; gonopod femorite subequal to postfemoral region in length; both solenophore and solenomere long; tip of solenophore serrated (
<figureCitation id="132B2A57FFE0AE55FD69FDB8FD62A917" box="[665,744,541,566]" captionStart="Fig" captionStartId="13.[136,167,1926,1945]" captionTargetBox="[231,1319,811,1899]" captionTargetId="figure-260@13.[231,1319,811,1899]" captionTargetPageId="13" captionText="Fig. 13. Hylomus proximus (Nguyen, Golovatch &amp; Anichkin, 2005) from Hoang Lien National Park. Right gonopod, lateral view (A), mesal view (B). Scale bars: A = 0.3 mm; B = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065910" httpUri="https://zenodo.org/record/8065910/files/figure.png" pageId="10" pageNumber="11">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE0AE55FF1AFD9BFC89A9B4" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BAF36D2FFE0AE55FF1AFD9BFE7BA954" blockId="10.[174,775,253,661]" pageId="10" pageNumber="11">
<emphasis id="B964EAC0FFE0AE55FF1AFD9BFE25A974" box="[234,431,573,597]" italics="true" pageId="10" pageNumber="11">Previous records</emphasis>
:
<collectingRegion id="49D4F830FFE0AE55FE32FD9BFD05A974" box="[450,655,573,597]" country="Vietnam" name="Lao Cai" pageId="10" pageNumber="11">Lao Cai Province</collectingRegion>
(Van Ban District) (
<bibRefCitation id="EF814B23FFE0AE55FEE5FDFBFE6CA954" author="Nguyen AD &amp; Golovatch SI &amp; Anichkin AE" box="[277,486,605,630]" pageId="10" pageNumber="11" pagination="251 - 257" refId="ref12004" refString="Nguyen AD, Golovatch SI, Anichkin AE. 2005. The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta 14 (3): 251 - 257." type="journal article" year="2005">Nguyen et al. 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF36D2FFE0AE55FF1AFDD8FC89A9B4" blockId="10.[174,775,253,661]" box="[234,771,637,661]" pageId="10" pageNumber="11">
<emphasis id="B964EAC0FFE0AE55FF1AFDD8FEE6A9B4" box="[234,364,637,661]" italics="true" pageId="10" pageNumber="11">Distribution</emphasis>
: Only known from northern
<collectingCountry id="F3077642FFE0AE55FD51FDD8FD75A9B4" box="[673,767,637,661]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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