198 lines
31 KiB
XML
198 lines
31 KiB
XML
<document id="5318AF52882D79291E8E2EADBA128D08" ID-CLB-Dataset="29359" ID-DOI="10.11646/zootaxa.4486.4.8" ID-GBIF-Dataset="8987221d-2607-4d89-b01d-df350d3d8d61" ID-ISSN="1175-5326" ID-Zenodo-Dep="1442340" ID-ZooBank="DE414CAD-F68C-4967-999C-2EE7688ACEDB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538407593628" checkinUser="plazi" docAuthor="Ulitzka, Manfred R." docDate="2018" docId="DB1E8076826D3E2CDF914D2BFDEEFA94" docLanguage="en" docName="zootaxa.4486.4.8.pdf" docOrigin="Zootaxa 4486 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rohrthrips burmiticus Ulitzka 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="555" masterDocId="2727F80E826B3E2BDF064B63FFEDFF8A" masterDocTitle="A first survey of Cretaceous thrips from Burmese amber including the establishment of a new family of Tubulifera (Insecta: Thysanoptera)" masterLastPageNumber="558" masterPageNumber="548" pageNumber="554" updateTime="1698562237327" updateUser="ExternalLinkService">
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<mods:title id="61CCE8276C1639301CF136BE3C6A9564">A first survey of Cretaceous thrips from Burmese amber including the establishment of a new family of Tubulifera (Insecta: Thysanoptera)</mods:title>
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<mods:namePart id="E17D8D18247E827EC4D552BCA54AC8DA">Ulitzka, Manfred R.</mods:namePart>
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<mods:date id="8C4C5A5A561F60780B6FA32E5D2A3A9C">2018</mods:date>
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<mods:number id="7DDE617A443C2E7B224616F07EA0EA18">2018-10-01</mods:number>
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<treatment id="DB1E8076826D3E2CDF914D2BFDEEFA94" ID-DOI="http://doi.org/10.5281/zenodo.5974670" ID-GBIF-Taxon="148425588" ID-Zenodo-Dep="5974670" LSID="urn:lsid:plazi:treatment:DB1E8076826D3E2CDF914D2BFDEEFA94" httpUri="http://treatment.plazi.org/id/DB1E8076826D3E2CDF914D2BFDEEFA94" lastPageId="7" lastPageNumber="555" pageId="6" pageNumber="554">
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<subSubSection id="1BAD62EB826D3E2DDF914D2BFEF5F90E" pageId="6" pageNumber="554" type="nomenclature">
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<paragraph id="53083160826D3E2DDF914D2BFE03F9E9" blockId="6.[151,494,1608,1669]" box="[151,494,1608,1635]" pageId="6" pageNumber="554">
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<heading id="0840860C826D3E2DDF914D2BFE03F9E9" bold="true" box="[151,494,1608,1635]" fontSize="11" level="1" pageId="6" pageNumber="554" reason="1">
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<emphasis id="61C3ED72826D3E2DDF914D2BFE03F9E9" bold="true" box="[151,494,1608,1635]" pageId="6" pageNumber="554">
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<taxonomicName id="94B74AE3826D3E2DDF914D2BFE49F9E8" authority="Ulitzka, 2018" authorityName="Ulitzka" authorityYear="2018" box="[151,420,1608,1634]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="554" phylum="Arthropoda" rank="species" species="burmiticus" status="sp. nov.">
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<emphasis id="61C3ED72826D3E2DDF914D2BFE49F9E8" bold="true" box="[151,420,1608,1634]" italics="true" pageId="6" pageNumber="554">Rohrthrips burmiticus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7AF05009826D3E2DDEAA4D2AFE03F9E9" box="[428,494,1609,1635]" pageId="6" pageNumber="554" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="53083160826D3E2DDF914D0FFEF5F90E" blockId="6.[151,494,1608,1669]" box="[151,280,1644,1669]" pageId="6" pageNumber="554">
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(
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<figureCitation id="CB8C2DE5826D3E2DDF994D0FFEFCF90E" box="[159,273,1644,1669]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="6" pageNumber="554">Figs 9–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1BAD62EB826D3E2CDF914DCDFC69FCC1" lastPageId="7" lastPageNumber="555" pageId="6" pageNumber="554" type="description">
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<paragraph id="53083160826D3E2DDF914DCDFD81F962" blockId="6.[151,1437,1710,2035]" pageId="6" pageNumber="554">
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Male: Colour (
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<figureCitation id="CB8C2DE5826D3E2DDE3D4DCDFE93F94D" box="[315,382,1710,1735]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="6" pageNumber="554">Fig. 9</figureCitation>
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) uniformly dark brown including antennae and legs; all
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<taxonomicName id="94B74AE3826D3E2DDCF74DCDFBDFF94D" box="[1009,1074,1710,1735]" class="Insecta" family="Melanthripidae" genus="Gymnopollisthrips" higherTaxonomySource="CoL" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="554" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
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setae dark as well as wing veins and fringes; wings shaded greyish-brown.
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</paragraph>
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<paragraph id="53083160826D3E2CDFC14D92FCD2FDEB" blockId="6.[151,1437,1710,2035]" lastBlockId="7.[151,1437,151,1310]" lastPageId="7" lastPageNumber="555" pageId="6" pageNumber="554">
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Head (
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<figureCitation id="CB8C2DE5826D3E2DDE104D92FE81F880" box="[278,364,1777,1802]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="6" pageNumber="554">Fig. 12</figureCitation>
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) with reticulate sculpture at base; longer than wide, with cheeks rounded behind eyes and converging slightly to base; cheeks lacking setae; ocellar setae difficult to assess, one strong pair clearly visible close to base of antennae. Eyes large, their front margin protruding over base of antennae, not prolonged ventrally. Hind ocelli close to compound eyes; far apart from each other. Vertex with two postocular setae far behind eyes. Antennae (
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<figureCitation id="CB8C2DE5826D3E2DDE134C15FE8AF805" box="[277,359,1910,1935]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="6" pageNumber="554">Fig. 11</figureCitation>
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) 9-segmented; segments III–VII inverse conically shaped and tapering distally from level of sense cones; segment II apically with a circular sensorium; number of sense cones of the following segments difficult to assess but at least two on III, IV and V; segment IX broadly joined to VIII. Mouth cone ophistognathus; short but pointed; clypeal suture clearly asymmetric. Maxillary palps with two apical setae; short and very likely 2- segmented; labial palps not visible. Maxillary stylets (
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<figureCitation id="CB8C2DE5826C3E2CDC214BF4FC6DFF3A" box="[807,896,151,176]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 12</figureCitation>
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) slender, deeply retracted into head capsule, extending to base of compound eyes, close together and parallel medially; maxillary bridge not developed. Left mandible strong; right mandible degenerate (a shaded structure possibly its remnant). Pronotum (
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<figureCitation id="CB8C2DE5826C3E2CDBDC4BB9FAA0FF79" box="[1242,1357,218,243]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Figs 9, 12</figureCitation>
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) wider than long, trapezoidal; pronotal chaetotaxy not assessable; epimeral setae long and pointed. Pterothorax as well as basal abdominal segments dented and partly concealed. Fore wings (
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<figureCitation id="CB8C2DE5826C3E2CDC904A7EFC34FEBC" box="[918,985,285,310]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 9</figureCitation>
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) becoming progressively larger in distal third; anterior vein complete, reaching costa near apex and bearing setae; second vein not developed, reduced to a faint basal shading; membrane without microtrichia. Fringe cilia straight; arising from sockets (
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<figureCitation id="CB8C2DE5826C3E2CDBC64A3CFAFEFEF2" box="[1216,1299,351,376]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 10</figureCitation>
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); duplicated cilia present around margin of wing apex and on distal part of posterior margin; sub-basal wing setae at anterior margin lacking; clavus with paired setiform processes at tip (
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<figureCitation id="CB8C2DE5826C3E2CDC454AC1FC7AFE31" box="[835,919,418,443]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 10</figureCitation>
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). Hind wing without microtrichia and without any veins; base with recurved setae. Fore legs (
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<figureCitation id="CB8C2DE5826C3E2CDDAF4AA7FCF6FE56" box="[681,795,452,477]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Figs 9, 12</figureCitation>
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) with femora stout; fore tarsi with a large tooth. Mid and hind legs slender; all tibiae with one long dorsal seta (
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<figureCitation id="CB8C2DE5826C3E2CDDFF4A86FCD3FE74" box="[761,830,485,510]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 9</figureCitation>
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); mid and hind tibiae terminally each with one strong spine; mid and hind tarsi two-segmented. Abdominal tergites without wing-retaining setae; lateral setae long and pointed. Abdominal segment IX conical, sternite with clearly visible subgenital plate. Abdominal segment X elongated tubular, with terminal crown of anal setae (
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<figureCitation id="CB8C2DE5826C3E2CDDEA492AFCDDFDEB" box="[748,816,585,610]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 9</figureCitation>
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).
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</paragraph>
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<paragraph id="53083160826C3E2CDFC14909FC69FCC1" blockId="7.[151,1437,151,1310]" pageId="7" pageNumber="555">
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<emphasis id="61C3ED72826C3E2CDFC14909FE9BFD09" bold="true" box="[199,374,618,643]" pageId="7" pageNumber="555">Measurements</emphasis>
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. Male (in microns): Body, length 1480 (slightly contracted). Head, length 227; width 201. Eyes, length 107, width 50. Hind ocelli, diameter 19; distance between the hind ocelli 57. Ocellar setae 44. Postocular setae 38. Prothorax, length 258; width 441; epimeral setae 176. Abdomen, length 888 (slightly contracted); largest width 391 (segment IV); segment X (tube), length 189; basal width 50. Antennae, length 422; length (largest width) of segment
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<date id="270917A0826C3E2CDD084993FDD7FC82" box="[526,570,752,776]" pageId="7" pageNumber="555" value="1937-01">I 37</date>
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(31) [deformed],
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<date id="270917A0826C3E2CDC064993FCD8FC82" box="[768,821,751,776]" pageId="7" pageNumber="555" value="1965-02">II 65</date>
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(34),
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<date id="270917A0826C3E2CDC714993FC5BFC82" box="[887,950,752,776]" pageId="7" pageNumber="555" value="1971-03">III 71</date>
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(37),
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<date id="270917A0826C3E2CDCFE4993FBD5FC82" box="[1016,1080,751,776]" pageId="7" pageNumber="555" value="1956-04">IV 56</date>
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(37),
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<date id="270917A0826C3E2CDB7C4993FB5CFC82" box="[1146,1201,751,776]" pageId="7" pageNumber="555" value="1953-05">V 53</date>
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(34),
|
||
<date id="270917A0826C3E2CDBF54993FADEFC82" box="[1267,1331,752,776]" pageId="7" pageNumber="555" value="1943-06">VI 43</date>
|
||
(28),
|
||
<date id="270917A0826C3E2CDA734993FF5EFCA0" pageId="7" pageNumber="555" value="1950-07">VII 50</date>
|
||
(22),
|
||
<date id="270917A0826C3E2CDFF04871FEA7FCA0" box="[246,330,786,810]" pageId="7" pageNumber="555" value="1931-08">VIII 31</date>
|
||
(16),
|
||
<date id="270917A0826C3E2CDE8B4871FE23FCA0" box="[397,462,785,810]" pageId="7" pageNumber="555" value="2016-09">IX 16</date>
|
||
(6). Sense cones, length of inner (outer) on segment
|
||
<date id="270917A0826C3E2CDB264871FBB2FCA0" box="[1056,1119,786,810]" pageId="7" pageNumber="555" value="1919-03">III 19</date>
|
||
(16),
|
||
<date id="270917A0826C3E2CDBA54871FB0EFCA0" box="[1187,1251,786,810]" pageId="7" pageNumber="555" value="1919-04">IV 19</date>
|
||
(22),
|
||
<date id="270917A0826C3E2CDA214871FAB2FCA0" box="[1319,1375,785,810]" pageId="7" pageNumber="555" value="2016-05">V 16</date>
|
||
(22); basal width 5–6. Fore wings, length about 1200; largest width 245.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BAD62EB826C3E2CDFC14837FE50FC25" pageId="7" pageNumber="555" type="materials_examined">
|
||
<paragraph id="53083160826C3E2CDFC14837FE50FC25" blockId="7.[151,1437,151,1310]" pageId="7" pageNumber="555">
|
||
<materialsCitation id="E3DF3B3D826C3E2CDFC14837FE54FC25" ID-GBIF-Occurrence="1913894088" collectionCode="SMF" country="Germany" county="Frankfurt am Main" location="Material" municipality="Senckenbergische Naturforschende Gesellschaft" pageId="7" pageNumber="555" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="61C3ED72826C3E2CDFC14837FE79FCE7" bold="true" box="[199,404,852,877]" pageId="7" pageNumber="555">
|
||
<location id="566867BB826C3E2CDFC14837FEC2FCE7" LSID="urn:lsid:plazi:treatment:DB1E8076826D3E2CDF914D2BFDEEFA94:566867BB826C3E2CDFC14837FEC2FCE7" box="[199,303,852,877]" country="Germany" county="Frankfurt am Main" municipality="Senckenbergische Naturforschende Gesellschaft" name="Material" pageId="7" pageNumber="555">Material</location>
|
||
studied.
|
||
</emphasis>
|
||
<typeStatus id="8C0C8FC2826C3E2CDE9C4837FDEFFCE7" box="[410,514,852,877]" pageId="7" pageNumber="555" type="holotype">Holotype</typeStatus>
|
||
male MU-Fos-53/1 (
|
||
<figureCitation id="CB8C2DE5826C3E2CDDE84837FCDFFCE6" box="[750,818,852,877]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 9</figureCitation>
|
||
).
|
||
<location id="566867BB826C3E2CDC414837FBDBFCE6" LSID="urn:lsid:plazi:treatment:DB1E8076826D3E2CDF914D2BFDEEFA94:566867BB826C3E2CDC414837FBDBFCE6" box="[839,1078,852,877]" country="Germany" county="Frankfurt am Main" municipality="Senckenbergische Naturforschende Gesellschaft" name="Inclusion in Burmese" pageId="7" pageNumber="555">Inclusion in Burmese</location>
|
||
amber, donated to the author by Patrick Müller.
|
||
<typeStatus id="8C0C8FC2826C3E2CDE4B4816FE58FC04" box="[333,437,885,910]" pageId="7" pageNumber="555" type="holotype">Holotype</typeStatus>
|
||
deposited in the
|
||
<collectingMunicipality id="B36CAB1A826C3E2CDD854816FB46FC04" box="[643,1195,885,910]" pageId="7" pageNumber="555">Senckenbergische Naturforschende Gesellschaft</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="BA6949EC826C3E2CDBBF4816FA75FC04" box="[1209,1432,885,910]" pageId="7" pageNumber="555">Frankfurt am Main</collectingCounty>
|
||
,
|
||
<collectingCountry id="2BA071F0826C3E2CDF9148F4FF12FC25" box="[151,255,919,943]" name="Germany" pageId="7" pageNumber="555">Germany</collectingCountry>
|
||
(
|
||
<collectionCode id="35A6A9A5826C3E2CDE0848F4FEA5FC25" box="[270,328,919,943]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="555">SMF</collectionCode>
|
||
T 19282)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BAD62EB826C3E2CDFC148D4FB09FC5A" box="[199,1252,951,976]" pageId="7" pageNumber="555" type="etymology">
|
||
<paragraph id="53083160826C3E2CDFC148D4FB09FC5A" blockId="7.[151,1437,151,1310]" box="[199,1252,951,976]" pageId="7" pageNumber="555">
|
||
<emphasis id="61C3ED72826C3E2CDFC148D4FEA3FC45" bold="true" box="[199,334,951,976]" pageId="7" pageNumber="555">Etymology.</emphasis>
|
||
The specific epithet ‘
|
||
<taxonomicName id="94B74AE3826C3E2CDD4448D4FD57FC5A" authorityName="Ulitzka" authorityYear="2018" box="[578,698,951,976]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="555" phylum="Arthropoda" rank="species" species="burmiticus">
|
||
<emphasis id="61C3ED72826C3E2CDD4448D4FD57FC5A" box="[578,698,951,976]" italics="true" pageId="7" pageNumber="555">burmiticus</emphasis>
|
||
</taxonomicName>
|
||
’ refers to the occurrence of the fossil in Burmite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BAD62EB826C3E2CDFC148BAFDEEFA94" pageId="7" pageNumber="555" type="discussion">
|
||
<paragraph id="53083160826C3E2CDFC148BAFE60FB12" blockId="7.[151,1437,151,1310]" pageId="7" pageNumber="555">
|
||
<emphasis id="61C3ED72826C3E2CDFC148BAFEA1FC78" bold="true" box="[199,332,985,1010]" pageId="7" pageNumber="555">Discussion.</emphasis>
|
||
Many features of
|
||
<taxonomicName id="94B74AE3826C3E2CDD1E48B9FD42FC78" authority="Ulitzka, 2018" authorityName="Ulitzka" authorityYear="2018" box="[536,687,985,1010]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="555" phylum="Arthropoda" rank="species" species="burmiticus" status="sp. nov.">
|
||
<emphasis id="61C3ED72826C3E2CDD1E48B9FD42FC78" box="[536,687,985,1010]" italics="true" pageId="7" pageNumber="555">R. burmiticus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7AF05009826C3E2CDDB048BAFD18FC78" box="[694,757,985,1010]" pageId="7" pageNumber="555" rank="species">
|
||
<emphasis id="61C3ED72826C3E2CDDB048BAFD18FC78" bold="true" box="[694,757,985,1010]" pageId="7" pageNumber="555">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
correspond clearly to those of
|
||
<taxonomicName id="94B74AE3826C3E2CDB4948B9FB39FC78" authorityName="Nel, Penalver, Azar, Hodebert & Nel" authorityYear="2010" box="[1103,1236,985,1010]" class="Insecta" family="Thripidae" genus="Tethysthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="555" phylum="Arthropoda" rank="species" species="libanicus">
|
||
<emphasis id="61C3ED72826C3E2CDB4948B9FB39FC78" box="[1103,1236,985,1010]" italics="true" pageId="7" pageNumber="555">R. libanicus</emphasis>
|
||
</taxonomicName>
|
||
, particularly with regard to the plesiomorphic characters of the wings mentioned above. The new species, however, has only one vein developed on the fore wings (
|
||
<figureCitation id="CB8C2DE5826C3E2CDEF74F7FFDD5FBBE" box="[497,568,1052,1077]" captionStart="FIGURES 9–13" captionStartId="8.[151,264,1899,1921]" captionTargetBox="[161,1426,206,1885]" captionTargetId="figure@8.[161,1426,193,1885]" captionTargetPageId="8" captionText="FIGURES 9–13. FIGURES 9–12. Rohrthrips burmiticus sp. n., holotype male (MU-Fos-53/1). (9) dorsal view; (10) coupling mechanism of the wings, fore wing clavus with paired setiform processus (black arrow), hindwing with basal recurved seta (red arrow); (11) right antenna (segments I and II reconstructed); (12) head and prothorax (md: left mandible, mx: maxillary stylets). FIGURE 13. Rohrthrips libanicus, holotype, right fore wing with two veins (indicated)." httpUri="https://zenodo.org/record/1442342/files/figure.png" pageId="7" pageNumber="555">Fig. 9</figureCitation>
|
||
). Furthermore, it has only one ventral spine on the apex of the hind tibiae, whereas
|
||
<taxonomicName id="94B74AE3826C3E2CDFF84F5DFE64FBDC" authorityName="Nel, Penalver, Azar, Hodebert & Nel" authorityYear="2010" box="[254,393,1085,1110]" class="Insecta" family="Thripidae" genus="Tethysthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="555" phylum="Arthropoda" rank="species" species="libanicus">
|
||
<emphasis id="61C3ED72826C3E2CDFF84F5DFE64FBDC" box="[254,393,1085,1110]" italics="true" pageId="7" pageNumber="555">R. libanicus</emphasis>
|
||
</taxonomicName>
|
||
has three (cf.
|
||
<bibRefCitation id="37264C91826C3E2CDD3C4F5EFD1FFBDC" box="[570,754,1085,1110]" pageId="7" pageNumber="555" refId="ref7147" refString="Nel, P., Penalver, E., Azar, D., Hodebert, G., & Nel, A. (2010) Modern thrips families Thripidae and Phlaeothripidae in Early Cretaceous amber (Insecta: Thysanoptera). Annales de la Societe entomologique de France, New Series, 46 (1 - 2), 154 - 163. https: // doi. org / 10.1080 / 00379271.2010.10697651" type="journal article">
|
||
Nel
|
||
<emphasis id="61C3ED72826C3E2CDD684F5DFD46FBDC" box="[622,683,1085,1110]" italics="true" pageId="7" pageNumber="555">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
, p. 155, fig. 3). Establishing a new genus based on these differences was rejected. Further findings that allow comparing the head structures of both species may lead to another classification.
|
||
</paragraph>
|
||
<paragraph id="53083160826C3E2CDFC14FC2FDEEFA94" blockId="7.[151,1437,151,1310]" pageId="7" pageNumber="555">
|
||
Assuming the loss of the wing veins as an apomorphy in the evolution of Tubulifera (Mound
|
||
<emphasis id="61C3ED72826C3E2CDBF44FC1FAC6FB30" box="[1266,1323,1185,1210]" italics="true" pageId="7" pageNumber="555">et al.</emphasis>
|
||
2010),
|
||
<taxonomicName id="94B74AE3826C3E2CDA824FC1FEE2FB51" authorityName="Ulitzka" authorityYear="2018" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="555" phylum="Arthropoda" rank="species" species="burmiticus">
|
||
<emphasis id="61C3ED72826C3E2CDA824FC1FEE2FB51" italics="true" pageId="7" pageNumber="555">R. burmiticus</emphasis>
|
||
</taxonomicName>
|
||
from the late Cretaceous might be interpreted as more modern than its Mid-Cretaceous sister species (cf. figs 9 and 13). Its phylogenetic position as a direct ancestor of recent
|
||
<taxonomicName id="94B74AE3826C3E2CDCF64F87FB4DFB77" box="[1008,1184,1252,1277]" class="Insecta" family="Phlaeothripidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="555" phylum="Arthropoda" rank="family">Phlaeothripidae</taxonomicName>
|
||
or as an evolutionary dead end remains undetermined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |