249 lines
37 KiB
XML
249 lines
37 KiB
XML
<document id="B33A7E8D6C952AD4EBB85091E93DC918" ID-DOI="10.11646/zootaxa.3637.3.3" ID-GBIF-Dataset="04f506ff-e980-47b6-ac5c-33b5fdbea26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="216877" ID-ZooBank="C86DD907-723F-4EF1-87D7-2F511DCC1CBC" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460591433277" checkinUser="plazi" docAuthor="Maier, Crystal A. & Short, Andrew E. Z." docDate="2013" docId="B43B879AFFEFFFF7D794F982FE7FF906" docLanguage="en" docName="zt03637p307.pdf" docOrigin="Zootaxa 3637 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lutrochus meridaensis Maier & Short, 2013, n. sp." docType="treatment" docVersion="7" lastPageNumber="301" masterDocId="4802FFE2FFE1FFE7D703FF96FFF5FFB9" masterDocTitle="A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species" masterLastPageNumber="307" masterPageNumber="285" pageNumber="299" updateTime="1698326039231" updateUser="plazi">
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<mods:mods id="4A9F35F489D069F9E1815675BFDD14F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="093A6B6DBC6D51CE1749D5A1CA0DCEDC">
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<mods:title id="FAA71A8B782B5620B17BA3AA0108A017">A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species</mods:title>
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</mods:titleInfo>
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<mods:name id="01CB71CCB44EFEE27D1873EAF41DA2D5" type="personal">
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<mods:roleTerm id="38D444474D07B8BFEE81231DA46498A8">Author</mods:roleTerm>
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<mods:namePart id="E65E187680C17D194409886E51A0CD53">Maier, Crystal A.</mods:namePart>
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<mods:name id="EEF8573BD146EB8EBD308190E4144FC1" type="personal">
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<mods:roleTerm id="DE47A4992BE3DC04B4586F4FB02226CD">Author</mods:roleTerm>
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<mods:namePart id="3F506A5545888CE72E34EC25C102D390">Short, Andrew E. Z.</mods:namePart>
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<mods:typeOfResource id="E5CC600E9B1ACE6301F3D9BA610EBD4F">text</mods:typeOfResource>
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<mods:title id="D9522E6402E32B5C97524CCD3C00D46C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7F77E329AF5B64ACF68DBDDBB5261FB9">
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<mods:date id="3EE306A962A04DB7D256D1FB608A380F">2013</mods:date>
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<mods:detail id="53F05FA3E85898177FFD1886DFA9D765" type="volume">
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<mods:number id="4304A29B2087A14EF7D0AE8184E57E25">3637</mods:number>
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<mods:number id="849A5CF0205EF30849AB7781B42DBC80">3</mods:number>
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<mods:classification id="DA05801897BECB8552F7F7F7B9CDFF87">journal article</mods:classification>
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<mods:identifier id="5D1B753E58A6C0F29D8C5531604EFC97" type="DOI">10.11646/zootaxa.3637.3.3</mods:identifier>
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<mods:identifier id="FA360CB8A891A1707A55A4FB379F2A99" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E5DCD870A27EA41A5DDB2BF44BF57F90" type="Zenodo-Dep">216877</mods:identifier>
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<treatment id="B43B879AFFEFFFF7D794F982FE7FF906" ID-DOI="http://doi.org/10.5281/zenodo.6156881" ID-GBIF-Taxon="119565844" ID-Zenodo-Dep="6156881" LSID="urn:lsid:plazi:treatment:B43B879AFFEFFFF7D794F982FE7FF906" httpUri="http://treatment.plazi.org/id/B43B879AFFEFFFF7D794F982FE7FF906" lastPageId="16" lastPageNumber="301" pageId="14" pageNumber="299">
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<subSubSection id="74886507FFEFFFE9D794F982FE93F9F7" pageId="14" pageNumber="299" type="nomenclature">
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<paragraph id="3C2D368CFFEFFFE9D794F982FE03F994" blockId="14.[151,502,1555,1614]" box="[151,502,1555,1582]" pageId="14" pageNumber="299">
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<heading id="676581E0FFEFFFE9D794F982FE03F994" bold="true" box="[151,502,1555,1582]" fontSize="11" level="1" pageId="14" pageNumber="299" reason="1">
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<emphasis id="0EE6EA9EFFEFFFE9D794F982FE03F994" bold="true" box="[151,502,1555,1582]" pageId="14" pageNumber="299">
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<taxonomicName id="FB924D0FFFEFFFE9D794F982FE59F997" box="[151,428,1556,1582]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="299" phylum="Arthropoda" rank="species" species="meridaensis" status="sp. nov.">
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<emphasis id="0EE6EA9EFFEFFFE9D794F982FE59F997" bold="true" box="[151,428,1556,1582]" italics="true" pageId="14" pageNumber="299">Lutrochus meridaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="15D557E5FFEFFFE9D6B7F985FE03F994" box="[436,502,1555,1581]" pageId="14" pageNumber="299" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="3C2D368CFFEFFFE9D794F9A0FE93F9F7" blockId="14.[151,502,1555,1614]" box="[151,358,1590,1614]" pageId="14" pageNumber="299">
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||
(
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<figureCitation id="A4A92A09FFEFFFE9D79CF9A0FF1BF9F7" box="[159,238,1590,1614]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[151,1436,733,1379]" captionTargetId="figure@3.[151,1436,733,1380]" captionTargetPageId="3" captionText="FIGURE 3. Habitat of Lutrochus species. A) L. meridaensis: Venezuela, Mérida State, Cascada Velo de la Novia; specimens were collected on the vertical wet rocks adjacent to the waterfall (collecting event VZ 09 - 0719 - 01 A). B) L. vestitus: Venezuela, Falcón State, Cataratas del Hueque; specimens were found in the exposed tree root mats along the margin (collecting event VZ 09 - 0709 - 01 B)." httpUri="https://zenodo.org/record/216880/files/figure.png" pageId="14" pageNumber="299">Figs. 3</figureCitation>
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,
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<figureCitation id="A4A92A09FFEFFFE9D7F9F9A0FEFFF9F7" box="[250,266,1590,1614]" captionStart="FIGURES 4 – 7. 4" captionStartId="5.[151,264,1862,1885]" captionTargetBox="[154,1432,781,1837]" captionTargetId="figure@5.[153,1434,772,1841]" captionTargetPageId="5" captionText="FIGURES 4 – 7. 4. Map of Venezuela showing collection localities of L. acuminatus (red circles) and L. cauraensis (yellow square). 5. Map of Venezuela showing collection localities of L. vestitus (red circles). 6. Map of Venezuela showing collection localities of L. meridaensis (red circles), L. minutus (orange circles), and L. maldonadoi (yellow circles). 7. Map of Venezuela showing collection localities of L. violaceus (red circles) and L. gustafsoni (yellow circles)." httpUri="https://zenodo.org/record/216881/files/figure.png" pageId="14" pageNumber="299">6</figureCitation>
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,
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<figureCitation id="A4A92A09FFEFFFE9D615F9A0FEABF9F7" box="[278,350,1590,1614]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="14" pageNumber="299">40–47</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="74886507FFEFFFE9D794F9E9FE0FF89F" pageId="14" pageNumber="299" type="materials_examined">
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<paragraph id="3C2D368CFFEFFFE9D794F9E9FE0FF89F" blockId="14.[151,1437,1661,2010]" pageId="14" pageNumber="299">
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||
<emphasis id="0EE6EA9EFFEFFFE9D794F9E9FDF1F92E" bold="true" box="[151,516,1662,1687]" pageId="14" pageNumber="299">
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||
<typeStatus id="E329882EFFEFFFE9D794F9E9FF25F92E" box="[151,208,1663,1687]" pageId="14" pageNumber="299">Type</typeStatus>
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Material.
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<typeStatus id="E329882EFFEFFFE9D64DF9E9FE4EF92E" box="[334,443,1663,1687]" pageId="14" pageNumber="299" type="holotype">Holotype</typeStatus>
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male:
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</emphasis>
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“
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<collectingCountry id="4485761CFFEFFFE9D516F9E8FD35F92F" box="[533,704,1662,1686]" name="Venezuela" pageId="14" pageNumber="299">VENEZUELA</collectingCountry>
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:Merida State/8° 52.423ʹ N, 71° 37.611ʹ W,
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<quantity id="FB6A9B69FFEFFFE9D3A6F9E8FB0DF92F" box="[1189,1272,1662,1686]" metricMagnitude="3" metricUnit="m" metricValue="1.616" pageId="14" pageNumber="299" unit="m" value="1616.0">1616 m</quantity>
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/ Cascada velo de la Novia/
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<date id="482C104CFFEFFFE9D628F934FE5BF903" box="[299,430,1698,1722]" pageId="14" pageNumber="299" value="2009-07-19">19.vii.2009</date>
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; leg. Short, Gustafson/ Camacho, Garcia, & Inciarte/ VZ09-0719-01A; wet rocks/seeps” “SEMC0881687/KUNHM-ENT”.
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<typeStatus id="E329882EFFEFFFE9D51DF950FD73F967" box="[542,646,1734,1758]" pageId="14" pageNumber="299" type="holotype">Holotype</typeStatus>
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deposited in MIZA.
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<emphasis id="0EE6EA9EFFEFFFE9D474F951FA69F966" bold="true" box="[887,1436,1734,1759]" pageId="14" pageNumber="299">
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||
<typeStatus id="E329882EFFEFFFE9D474F951FC1AF966" box="[887,1007,1735,1759]" pageId="14" pageNumber="299" type="paratype">Paratypes</typeStatus>
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(13):
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<collectingCountry id="4485761CFFEFFFE9D335F950FB1CF966" box="[1078,1257,1734,1759]" name="Venezuela" pageId="14" pageNumber="299">VENEZUELA</collectingCountry>
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: Mérida State:
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</emphasis>
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Same locality data as
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<typeStatus id="E329882EFFEFFFE9D68AF97CFE1AF8BB" box="[393,495,1770,1794]" pageId="14" pageNumber="299" type="holotype">holotype</typeStatus>
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.
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<typeStatus id="E329882EFFEFFFE9D6FAF97DFD93F8BA" box="[505,614,1771,1795]" pageId="14" pageNumber="299" type="paratype">Paratypes</typeStatus>
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will be deposited in:
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<quantity id="FB6A9B69FFEFFFE9D456F97CFC75F8BB" box="[853,896,1770,1794]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="299" unit="in" value="2.0">2 in</quantity>
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||
MIZA,
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||
<quantity id="FB6A9B69FFEFFFE9D4D8F97CFBF3F8BB" box="[987,1030,1770,1794]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="299" unit="in" value="1.0">1 in</quantity>
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||
MALUZ,
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<quantity id="FB6A9B69FFEFFFE9D37EF97CFB5DF8BB" box="[1149,1192,1770,1794]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="299" unit="in" value="1.0">1 in</quantity>
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NMW,
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<quantity id="FB6A9B69FFEFFFE9D202F97DFADEF8BB" box="[1281,1323,1770,1794]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="299" unit="in" value="7.0">7 in</quantity>
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||
SEMC (1 on SEM stub), and
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||
<quantity id="FB6A9B69FFEFFFE9D673F898FE6EF89F" box="[368,411,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="299" unit="in" value="2.0">2 in</quantity>
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USNM.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="74886507FFEFFFE9D7C4F8A4FE39F82B" pageId="14" pageNumber="299" type="diagnosis">
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||
<paragraph id="3C2D368CFFEFFFE9D7C4F8A4FE39F82B" blockId="14.[151,1437,1661,2010]" pageId="14" pageNumber="299">
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||
<emphasis id="0EE6EA9EFFEFFFE9D7C4F8A4FEB4F8F2" bold="true" box="[199,321,1842,1867]" pageId="14" pageNumber="299">Diagnosis.</emphasis>
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This species can be distinguished from all other described
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<taxonomicName id="FB924D0FFFEFFFE9D4F4F8A4FB8BF8F3" box="[1015,1150,1842,1866]" class="Insecta" family="Lutrochidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="299" phylum="Arthropoda" rank="family">Lutrochidae</taxonomicName>
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by the unique diamondshaped scutellum, the medial circular glabrous patch on the metasternum, and the presence of seven shallow rows of punctures on the elytron.
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</paragraph>
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</subSubSection>
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<subSubSection id="74886507FFEFFFF7D7C4F808FBD8FA51" lastPageId="16" lastPageNumber="301" pageId="14" pageNumber="299" type="description">
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<paragraph id="3C2D368CFFEFFFE9D7C4F808FB48F863" blockId="14.[151,1437,1661,2010]" pageId="14" pageNumber="299">
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<emphasis id="0EE6EA9EFFEFFFE9D7C4F808FEACF80E" bold="true" box="[199,345,1950,1975]" pageId="14" pageNumber="299">Description.</emphasis>
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MALE.—Length
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<quantity id="FB6A9B69FFEFFFE9D52AF808FD76F80F" box="[553,643,1950,1974]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="299" unit="mm" value="2.6">2.6 mm</quantity>
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; width
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<quantity id="FB6A9B69FFEFFFE9D5D5F808FCC5F80F" box="[726,816,1950,1974]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="299" unit="mm" value="1.4">1.4 mm</quantity>
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. Body wedge-shaped and strongly convex, dark brown to black, densely pubescent; clothed with short, fine, slightly recumbent golden hairs (
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<figureCitation id="A4A92A09FFEFFFE9D35FF854FB45F863" box="[1116,1200,1986,2010]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="14" pageNumber="299">Fig. 40</figureCitation>
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).
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</paragraph>
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<caption id="68ED6604FFEEFFE8D794F964FBC8F8FF" httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="15" pageNumber="300" targetBox="[151,1435,193,1757]" targetPageId="15">
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||
<paragraph id="3C2D368CFFEEFFE8D794F964FBC8F8FF" blockId="15.[151,1436,1778,1862]" pageId="15" pageNumber="300">
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||
<emphasis id="0EE6EA9EFFEEFFE8D794F964FEA1F8B1" bold="true" box="[151,340,1778,1801]" pageId="15" pageNumber="300">FIGURES 40–47.</emphasis>
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<taxonomicName id="FB924D0FFFEEFFE8D65AF965FDCAF8B0" box="[345,575,1779,1801]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="300" phylum="Arthropoda" rank="species" species="meridaensis" status="sp. nov.">
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<emphasis id="0EE6EA9EFFEEFFE8D65AF965FDCAF8B0" box="[345,575,1779,1801]" italics="true" pageId="15" pageNumber="300">Lutrochus meridaensis</emphasis>
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</taxonomicName>
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||
<emphasis id="0EE6EA9EFFEEFFE8D545F965FD8AF8B1" bold="true" box="[582,639,1779,1800]" pageId="15" pageNumber="300">
|
||
<taxonomicNameLabel id="15D557E5FFEEFFE8D545F965FD8AF8B1" box="[582,639,1779,1800]" pageId="15" pageNumber="300" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
|
||
:
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<emphasis id="0EE6EA9EFFEEFFE8D58FF965FD59F8B1" bold="true" box="[652,684,1779,1800]" pageId="15" pageNumber="300">40.</emphasis>
|
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Habitus, dorsal view;
|
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<emphasis id="0EE6EA9EFFEEFFE8D493F965FC45F8B1" bold="true" box="[912,944,1779,1800]" pageId="15" pageNumber="300">41.</emphasis>
|
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Habitus, ventral view;
|
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<emphasis id="0EE6EA9EFFEEFFE8D39EF965FB48F8B1" bold="true" box="[1181,1213,1779,1800]" pageId="15" pageNumber="300">42.</emphasis>
|
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Habitus, lateral view;
|
||
<emphasis id="0EE6EA9EFFEEFFE8D794F884FF42F89E" bold="true" box="[151,183,1810,1831]" pageId="15" pageNumber="300">43.</emphasis>
|
||
Head, three-quarters view, showing shape of apical maxillary palpomere;
|
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<emphasis id="0EE6EA9EFFEEFFE8D4A7F884FC31F89E" bold="true" box="[932,964,1810,1831]" pageId="15" pageNumber="300">44.</emphasis>
|
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Mesotibia;
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<emphasis id="0EE6EA9EFFEEFFE8D33FF884FBA9F89E" bold="true" box="[1084,1116,1810,1831]" pageId="15" pageNumber="300">45.</emphasis>
|
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SEM of metasternum, showing medial bare patch;
|
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<emphasis id="0EE6EA9EFFEEFFE8D65BF8A6FE8DF8FC" bold="true" box="[344,376,1840,1861]" pageId="15" pageNumber="300">46.</emphasis>
|
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SEM of scutellum;
|
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<emphasis id="0EE6EA9EFFEEFFE8D547F8A6FD91F8FC" bold="true" box="[580,612,1840,1861]" pageId="15" pageNumber="300">47.</emphasis>
|
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Aedeagus, lateral view (scale bar = 0.62 mm).
|
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</paragraph>
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</caption>
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<paragraph id="3C2D368CFFEEFFF7D7C4F8E6FBB3FE4D" blockId="15.[151,1436,1904,2036]" lastBlockId="16.[151,1436,151,1727]" lastPageId="16" lastPageNumber="301" pageId="15" pageNumber="300">
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Head broad, broadly rounded, weakly grooved beneath eye for reception of antenna. Eye rounded, small, with short, golden interfacetal setae; eyes smooth within outline of head (
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<figureCitation id="A4A92A09FFEEFFE8D494F802FC19F815" box="[919,1004,1940,1964]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="15" pageNumber="300">Fig. 43</figureCitation>
|
||
). Antenna short, reaching slightly past anterior border of pronotum, apical 9 antennomeres short, dark brown and clavate, clothed with short golden setae (
|
||
<figureCitation id="A4A92A09FFEEFFE8D79CF84AFF01F84D" box="[159,244,2012,2036]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="15" pageNumber="300">Fig. 43</figureCitation>
|
||
). Frontoclypeal suture indistinct. Clypeal margin straight, with brush of long setae apically, setae nearly as long as labrum. Apical margin of labrum sinuate, apical setal brush, lateral setae three times as long as medial setae. Labrum clothed with sparse, blunt setae. Mandibles large, dark brown, sickle-shaped; apex of mandible with one or two blunt teeth, if with two teeth, weakly so; prostheca with apical field of papilliform sensillae. Maxilla with 4-segmented palpus, apical palpomere flattened and triangular, truncate at apex (
|
||
<figureCitation id="A4A92A09FFF1FFF7D36CFE92FB33FEA5" box="[1135,1222,260,284]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 43</figureCitation>
|
||
); galea with dense apical brush of setae, lateral setae two times as long as medial setae; lacinia shorter and smaller than galea or palpus, with dense brush of curved setae on inner margin. Labial palpus 3-segmented; apical palpomere fusiform. Pronotum pubescent, dark brown, two times as wide as long at base, bisinuate at base (
|
||
<figureCitation id="A4A92A09FFF1FFF7D380FEF9FB2CFE3E" box="[1155,1241,367,391]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 40</figureCitation>
|
||
). Lateral edge of pronotum with distinct bead, slightly explanate apically (
|
||
<figureCitation id="A4A92A09FFF1FFF7D42BFE02FC75FE15" box="[808,896,404,428]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 42</figureCitation>
|
||
). Posterior margin notched medially to receive pointed anterior margin of scutellum. Disc of pronotum broadly convex. Scutellum diamond-shaped, about as wide as long (
|
||
<figureCitation id="A4A92A09FFF1FFF7D7F5FE4AFEBEFE4D" box="[246,331,476,500]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 46</figureCitation>
|
||
). Hypomeron excavate posteriorly to accommodate femur (
|
||
<figureCitation id="A4A92A09FFF1FFF7D4E6FE4AFBCCFE4D" box="[997,1081,476,500]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FE69FD99FD3D" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
Elytron pubescent, dark brown, with slight metallic sheen, widest at base (
|
||
<figureCitation id="A4A92A09FFF1FFF7D31EFE69FB86FDAE" box="[1053,1139,511,535]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 40</figureCitation>
|
||
). Elytron broadly convex, with seven rows of widely spaced coarse punctures (3 times diameter of punctures); intervals with dense setae; humerus not protuberant (
|
||
<figureCitation id="A4A92A09FFF1FFF7D6BEFDD1FDE7FDE6" box="[445,530,583,607]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 40</figureCitation>
|
||
). Lateral edge with distinct bead; epipleuron appearing narrow in ventral aspect (
|
||
<figureCitation id="A4A92A09FFF1FFF7D79CFDFAFF07FD3D" box="[159,242,620,644]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 41</figureCitation>
|
||
). Elytral apex slightly acuminate.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FD06FDF5FC81" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
Prosternum transverse, approximately two times as wide as long; anterior edge with strong bead, slightly deflexed to accommodate withdrawn head; prosternal process about as wide as long, with bead laterally, acuminate posteriorly (
|
||
<figureCitation id="A4A92A09FFF1FFF7D62FFD41FE78FD56" box="[300,397,727,751]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 41</figureCitation>
|
||
). Mesoventrite pubescent, with distinct, round glabrous patch medially (
|
||
<figureCitation id="A4A92A09FFF1FFF7D22DFD41FA65FD56" box="[1326,1424,727,751]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 45</figureCitation>
|
||
). Mesepisternum and mesepimeron excavated to accommodate folded front and middle legs. Mesepimeron extended posteriorly nearly to metacoxae.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FCDEFEB6FB1B" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
Legs long and slender. Profemur densely pubescent. Protibia pubescent to apical 1/9th. Protarsus with all tarsomeres glabrous, except for tufts of golden setae ventrally on all tarsomeres; apical tarsomere 1.5 times as long as preceding four tarsomeres combined. Mesocoxa with tuft of long golden setae basally. Mesofemur densely pubescent posteriorly, becoming less so anteriorly. Mesotibia completely glabrous, with small patch of golden setae at apex; apex without excavation laterally for reception of tarsus (
|
||
<figureCitation id="A4A92A09FFF1FFF7D4BBFC41FBE6FC56" box="[952,1043,983,1007]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 44</figureCitation>
|
||
). Mesotarsus with all tarsomeres glabrous, except for tufts of golden setae ventrally on all tarsomeres, with single long seta (twice as long as preceding tarsomere) arising from fourth tarsomere; apical tarsomere as long as preceding four tarsomeres combined. Metatrochanter globose, without posterior extensions. Metafemur densely pubescent. Metatibia entirely glabrous, with small patch of golden setae at apex. Metatarsus glabrous, with apical tarsomere as long as previous four combined.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FB39FD32FB53" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
Abdomen densely pubescent, with five ventrites (
|
||
<figureCitation id="A4A92A09FFF1FFF7D5F9FB39FCBAFB7E" box="[762,847,1199,1223]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 41</figureCitation>
|
||
). First ventrite shallowly excavated for reception of folded hind legs. Apical ventrite broadly rounded.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FB61FB10FA8B" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
Genitalia as illustrated, basal piece + parameres
|
||
<quantity id="FB6A9B69FFF1FFF7D5EDFB61FCB3FAB6" box="[750,838,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="301" unit="mm" value="1.6">1.6 mm</quantity>
|
||
long. Aedeagus slender and curved, becoming more curved at tapered tip (
|
||
<figureCitation id="A4A92A09FFF1FFF7D68FFA8CFE15FA8B" box="[396,480,1306,1330]" captionStart="FIGURES 40 – 47" captionStartId="15.[151,264,1778,1801]" captionTargetBox="[151,1435,193,1757]" captionTargetId="figure@15.[151,1435,193,1757]" captionTargetPageId="15" captionText="FIGURES 40 – 47. Lutrochus meridaensis n. sp.: 40. Habitus, dorsal view; 41. Habitus, ventral view; 42. Habitus, lateral view; 43. Head, three-quarters view, showing shape of apical maxillary palpomere; 44. Mesotibia; 45. SEM of metasternum, showing medial bare patch; 46. SEM of scutellum; 47. Aedeagus, lateral view (scale bar = 0.62 mm)." httpUri="https://zenodo.org/record/216887/files/figure.png" pageId="16" pageNumber="301">Fig. 47</figureCitation>
|
||
). Parameres fused, pointed, with distinct lobes interiorly subapically.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FAA9FAA2FAEE" blockId="16.[151,1436,151,1727]" box="[199,1367,1343,1367]" pageId="16" pageNumber="301">
|
||
<emphasis id="0EE6EA9EFFF1FFF7D7C4FAA9FED0FAEE" bold="true" box="[199,293,1343,1367]" pageId="16" pageNumber="301">Female.</emphasis>
|
||
Length
|
||
<quantity id="FB6A9B69FFF1FFF7D681FAA9FE2EFAEE" box="[386,475,1343,1367]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="301" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; width
|
||
<quantity id="FB6A9B69FFF1FFF7D52EFAA9FD73FAEE" box="[557,646,1343,1367]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="301" unit="mm" value="1.4">1.4 mm</quantity>
|
||
. Externally similar to male, females generally larger than males.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FAF4FBD8FA51" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
<emphasis id="0EE6EA9EFFF1FFF7D7C4FAF4FE28FAC2" bold="true" box="[199,477,1378,1403]" pageId="16" pageNumber="301">Intraspecific Variation.</emphasis>
|
||
This species varies considerably in size, ranging from
|
||
<quantity id="FB6A9B69FFF1FFF7D34DFAF4FB23FAC3" box="[1102,1238,1378,1402]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="16" pageNumber="301" unit="mm" value="2.85" valueMax="3.2" valueMin="2.5">2.5–3.2 mm</quantity>
|
||
long and
|
||
<quantity id="FB6A9B69FFF1FFF7D24BFAF4FF36FA26" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="16" pageNumber="301" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 mm</quantity>
|
||
wide. Additionally, specimens vary slightly in color and luster from dark brown to black, and from slightly metallic to not metallic. Some specimens examined have differing levels of setation, with patches where setae have rubbed off, as well as patches where setae have become matted down with grease.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74886507FFF1FFF7D7C4FA65FE7FF906" pageId="16" pageNumber="301" type="etymology">
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4FA65FAD7F9B3" blockId="16.[151,1436,151,1727]" box="[199,1314,1522,1547]" pageId="16" pageNumber="301">
|
||
<emphasis id="0EE6EA9EFFF1FFF7D7C4FA65FEBBF9B2" bold="true" box="[199,334,1523,1547]" pageId="16" pageNumber="301">Etymology.</emphasis>
|
||
This species,
|
||
<taxonomicName id="FB924D0FFFF1FFF7D6EFFA62FD64F9B2" box="[492,657,1523,1547]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="301" phylum="Arthropoda" rank="species" species="meridaensis">
|
||
<emphasis id="0EE6EA9EFFF1FFF7D6EFFA62FD64F9B2" box="[492,657,1523,1547]" italics="true" pageId="16" pageNumber="301">L. meridaensis</emphasis>
|
||
</taxonomicName>
|
||
, is named for the
|
||
<typeStatus id="E329882EFFF1FFF7D458FA65FC7EF9B2" box="[859,907,1523,1547]" pageId="16" pageNumber="301">type</typeStatus>
|
||
locality in Mérida State,
|
||
<collectingCountry id="4485761CFFF1FFF7D3A4FA64FAEBF9B3" box="[1191,1310,1522,1546]" name="Venezuela" pageId="16" pageNumber="301">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3C2D368CFFF1FFF7D7C4F980FE7FF906" blockId="16.[151,1436,151,1727]" pageId="16" pageNumber="301">
|
||
<emphasis id="0EE6EA9EFFF1FFF7D7C4F980FE0DF996" bold="true" box="[199,504,1558,1583]" pageId="16" pageNumber="301">Habitat and Distribution.</emphasis>
|
||
This species is known only from the
|
||
<typeStatus id="E329882EFFF1FFF7D49EF98EFC38F989" box="[925,973,1560,1584]" pageId="16" pageNumber="301">type</typeStatus>
|
||
locality, the Cascada Velo de la Novia in Mérida State,
|
||
<collectingCountry id="4485761CFFF1FFF7D635F9ACFE5CF9EB" box="[310,425,1594,1618]" name="Venezuela" pageId="16" pageNumber="301">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="A4A92A09FFF1FFF7D6BAF9ACFDF5F9EB" box="[441,512,1594,1618]" captionStart="FIGURES 4 – 7. 4" captionStartId="5.[151,264,1862,1885]" captionTargetBox="[154,1432,781,1837]" captionTargetId="figure@5.[153,1434,772,1841]" captionTargetPageId="5" captionText="FIGURES 4 – 7. 4. Map of Venezuela showing collection localities of L. acuminatus (red circles) and L. cauraensis (yellow square). 5. Map of Venezuela showing collection localities of L. vestitus (red circles). 6. Map of Venezuela showing collection localities of L. meridaensis (red circles), L. minutus (orange circles), and L. maldonadoi (yellow circles). 7. Map of Venezuela showing collection localities of L. violaceus (red circles) and L. gustafsoni (yellow circles)." httpUri="https://zenodo.org/record/216881/files/figure.png" pageId="16" pageNumber="301">Fig. 6</figureCitation>
|
||
). The
|
||
<typeStatus id="E329882EFFF1FFF7D54AF9ADFD8CF9EA" box="[585,633,1595,1619]" pageId="16" pageNumber="301">type</typeStatus>
|
||
series was collected among liverworts on the seepages and spray zone adjacent to a waterfall (
|
||
<figureCitation id="A4A92A09FFF1FFF7D6A4F9C9FE18F9CE" box="[423,493,1631,1655]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[151,1436,733,1379]" captionTargetId="figure@3.[151,1436,733,1380]" captionTargetPageId="3" captionText="FIGURE 3. Habitat of Lutrochus species. A) L. meridaensis: Venezuela, Mérida State, Cascada Velo de la Novia; specimens were collected on the vertical wet rocks adjacent to the waterfall (collecting event VZ 09 - 0719 - 01 A). B) L. vestitus: Venezuela, Falcón State, Cataratas del Hueque; specimens were found in the exposed tree root mats along the margin (collecting event VZ 09 - 0709 - 01 B)." httpUri="https://zenodo.org/record/216880/files/figure.png" pageId="16" pageNumber="301">Fig. 3</figureCitation>
|
||
). This is the first known record of the family from hygropetric habitats. A second visit to this locality and other similar nearby habitats during the dry season (January) in 2012 did not yield additional specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |