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<document id="95F5694D00803C02F47C1A75DD338149" ID-CLB-Dataset="24150" ID-DOI="10.5252/zoosystema2020v42a3" ID-GBIF-Dataset="7bdbb02c-ce5d-4df5-846a-b45c54fedc5a" ID-Zenodo-Dep="3652791" ID-ZooBank="urn:lsid:zoobank.org:pub:27C94F0F-BB9B-40A3-B615-4DB19C94F042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581020160253" checkinUser="plazi" docAuthor="Kantor, Yuri, Kosyan, Alisa &amp; Sorokin, Pavel" docDate="2020" docId="03D4B94FFFC22024FEF3F95EFD49C5EB" docLanguage="en" docName="zoosystema2020v42a3.pdf" docOrigin="Zoosystema 42 (3)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Phaenomenella samadiae Kantor &amp; Kosyan &amp; Sorokin 2020, n. sp." docType="treatment" docUuid="E2A6F480-9882-4221-BA7D-84760D903882" docUuidSource="ZooBank" docVersion="8" lastPageNumber="43" masterDocId="FFEDC137FFCB2028FFEFFFCDFFA9C206" masterDocTitle="On the taxonomic position of Phaenomenella Fraussen &amp; Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species" masterLastPageNumber="55" masterPageNumber="33" pageNumber="40" updateTime="1698780820290" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="6A061759BC7099CF53A78094C3A91525" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3517141CCAC6154C989EC7EA96A74ED4">On the taxonomic position of Phaenomenella Fraussen &amp; Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species</mods:title>
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<mods:name id="B1D5857B551BCABC012A251D433263C0" type="personal">
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<mods:namePart id="6FEA4A5A4E6852C7BADBD72990066804">Kantor, Yuri</mods:namePart>
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<mods:namePart id="681F34A4082E548DEE6225AEDC423B77">Kosyan, Alisa</mods:namePart>
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<mods:role id="A083EE7008C5EE7789385EFC2C717755">
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<mods:namePart id="C654526454466A424CEE0957E42B4205">Sorokin, Pavel</mods:namePart>
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<mods:typeOfResource id="E1FF55BD819F602D8829B4D2950040F9">text</mods:typeOfResource>
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<mods:title id="352AE5992026C124C40F567AB38ACD42">Zoosystema</mods:title>
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<mods:part id="AFB2231C5F0AE5A86D5B818F4E6C9C96">
<mods:date id="8F6A60657DBF8AB4C7CB09C9142C7C3B">2020</mods:date>
<mods:detail id="DBAD1A64176C8F3A3A228F3E063FBA9F" type="pubDate">
<mods:number id="61E5905C2DB5B8F44560048E04E2887D">2020-02-04</mods:number>
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<mods:detail id="18AADD44B813EEE3F05FA4217718E6AC" type="volume">
<mods:number id="EBD7E7496CDEEB663DBF318550BC7BFC">42</mods:number>
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<mods:number id="5200237C337251CF9BC613E1FA43A2C5">3</mods:number>
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<mods:classification id="8DCE1E0D59A2331565A5344B031121E9">journal article</mods:classification>
<mods:identifier id="EF6C657D75A82E7A8BE92FFEB31BB55E" type="CLB-Dataset">24150</mods:identifier>
<mods:identifier id="040F089FE4AE6AF1FDF38FBB08775813" type="DOI">10.5252/zoosystema2020v42a3</mods:identifier>
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<mods:identifier id="A00C5DF89477CB4CB41DACDE6EC9DE78" type="Zenodo-Dep">3652791</mods:identifier>
<mods:identifier id="BFFB89347668CBA0E20867BE60F3BE0F" type="ZooBank">urn:lsid:zoobank.org:pub:27C94F0F-BB9B-40A3-B615-4DB19C94F042</mods:identifier>
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<treatment id="03D4B94FFFC22024FEF3F95EFD49C5EB" ID-DOI="http://doi.org/10.5281/zenodo.3665205" ID-GBIF-Taxon="162054099" ID-Zenodo-Dep="3665205" LSID="urn:lsid:zoobank.org:act:E2A6F480-9882-4221-BA7D-84760D903882" httpUri="http://treatment.plazi.org/id/03D4B94FFFC22024FEF3F95EFD49C5EB" lastPageId="12" lastPageNumber="43" pageId="9" pageNumber="40">
<subSubSection id="C3675BD2FFC22021FEF3F95EFDC6C4A9" box="[284,623,1683,1711]" pageId="9" pageNumber="40" type="nomenclature">
<paragraph id="8BC20859FFC22021FEF3F95EFDC6C4A9" blockId="9.[284,623,1683,1743]" box="[284,623,1683,1711]" pageId="9" pageNumber="40">
<heading id="D08ABF35FFC22021FEF3F95EFDC6C4A9" box="[284,623,1683,1711]" centered="true" fontSize="11" level="4" pageId="9" pageNumber="40" reason="1">
<taxonomicName id="4C7D73DAFFC22021FEF3F95EFD84C4A8" ID-CoL="7SNZS" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[284,557,1683,1710]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="40" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
<emphasis id="B909D44BFFC22021FEF3F95EFD84C4A8" bold="true" box="[284,557,1683,1710]" italics="true" pageId="9" pageNumber="40">Phaenomenella samadiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFC22021FDDBF958FDC6C4A9" box="[564,623,1685,1711]" pageId="9" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC22021FE87F979FD8AC4C9" box="[360,547,1716,1743]" pageId="9" pageNumber="40" type="description">
<paragraph id="8BC20859FFC22021FE87F979FD8AC4C9" blockId="9.[284,623,1683,1743]" box="[360,547,1716,1743]" pageId="9" pageNumber="40">
<heading id="D08ABF35FFC22021FE87F979FD8AC4C9" box="[360,547,1716,1743]" centered="true" fontSize="11" level="4" pageId="9" pageNumber="40" reason="1">
(
<figureCitation id="134614DCFFC22021FE9FF979FE18C4C8" box="[368,433,1716,1742]" captionStart="FIG" captionStartId="11.[132,143,1759,1777]" captionTargetBox="[132,1455,215,1713]" captionTargetId="figure@11.[132,1455,214,1713]" captionTargetPageId="11" captionText="FIG. 4. — Shells of Phaenomenella samadiae n. sp.: A-C, holotype, MNHN-IM-2013-61617, SL 58.2 mm; D, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm; E, F, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm; G, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="9" pageNumber="40">Figs 4</figureCitation>
;
<figureCitation id="134614DCFFC22021FE50F979FE57C4C9" box="[447,510,1716,1743]" captionStart="FIG" captionStartId="13.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1835]" captionTargetId="figure@13.[132,1455,215,1836]" captionTargetPageId="13" captionText="FIG. 5. — Radulae of Phaenomenella spp.A, B, Phaenomenella samadiae n. sp.; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (A); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (B); C-F. Phaenomenella nicoi n. sp.; C, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C); D, MNHN-IM-2013-59398 (shell on Fig. 7D); E, MNHN-IM-2013-61639 (shell on Fig. 7E); F, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="9" pageNumber="40">5A, B</figureCitation>
;
<figureCitation id="134614DCFFC22021FDE3F979FDB3C4C8" box="[524,538,1716,1742]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="9" pageNumber="40">6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
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<paragraph id="8BC20859FFC22021FF4CF93CFD4EC505" blockId="9.[163,743,1777,1795]" box="[163,743,1777,1795]" pageId="9" pageNumber="40">
<uri id="FFEC045BFFC22021FF4CF93CFD4EC505" box="[163,743,1777,1795]" pageId="9" pageNumber="40">
urn:lsid:zoobank.org:act:
<uuid id="FFDB328CFFC22021FE85F93CFD4EC505" box="[362,743,1777,1795]" pageId="9" pageNumber="40">E2A6F480-9882-4221-BA7D-84760D903882</uuid>
</uri>
</paragraph>
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<subSubSection id="C3675BD2FFC22021FF6BF8EEFAA9C50F" pageId="9" pageNumber="40" type="materials_examined">
<paragraph id="8BC20859FFC22021FF6BF8EEFEAFC58C" blockId="9.[132,775,1827,1931]" pageId="9" pageNumber="40">
<materialsCitation id="3B150204FFC22021FF6BF8EEFEAFC58C" ID-GBIF-Occurrence="2564217915" collectionCode="MNHN" collectorName="S. W. &amp; Dong Sha" country="China" elevation="1035" latitude="19.983334" location="South China Sea" longLatPrecision="1269" longitude="116.4" municipality="Sea" pageId="9" pageNumber="40" specimenCount="1" typeStatus="holotype">
MATERIAL EXAMINED. —
<emphasis id="B909D44BFFC22021FE90F8EEFE77C53D" bold="true" box="[383,478,1827,1851]" pageId="9" pageNumber="40">
<typeStatus id="54C6B6FBFFC22021FE90F8EEFE77C53D" box="[383,478,1827,1851]" pageId="9" pageNumber="40" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B909D44BFFC22021FE04F8EEFD0DC53D" bold="true" box="[491,676,1827,1851]" pageId="9" pageNumber="40">
<location id="8EA25E82FFC22021FE04F8EEFD3AC53D" LSID="urn:lsid:plazi:treatment:03D4B94FFFC22024FEF3F95EFD49C5EB:8EA25E82FFC22021FE04F8EEFD3AC53D" box="[491,659,1827,1851]" country="China" latitude="19.983334" longLatPrecision="1269" longitude="116.4" municipality="Sea" name="South China Sea" pageId="9" pageNumber="40">South China Sea</location>
•
</emphasis>
<collectionCode id="ED6C909CFFC22021FD43F8E9FCAEC53D" box="[684,775,1828,1851]" pageId="9" pageNumber="40">MNHN-</collectionCode>
IM-2013-61617 (sequenced,
<figureCitation id="134614DCFFC22021FE4CF8F3FE72C550" box="[419,475,1854,1878]" captionStart="FIG" captionStartId="11.[132,143,1759,1777]" captionTargetBox="[132,1455,215,1713]" captionTargetId="figure@11.[132,1455,214,1713]" captionTargetPageId="11" captionText="FIG. 4. — Shells of Phaenomenella samadiae n. sp.: A-C, holotype, MNHN-IM-2013-61617, SL 58.2 mm; D, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm; E, F, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm; G, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="9" pageNumber="40">Figs 4</figureCitation>
A-C; 5A; 6A, B, E-G);
<collectorName id="26886D8FFFC22021FD56F8F3FD41C550" box="[697,744,1854,1878]" name="T. SCHWARTZ" pageId="9" pageNumber="40">S.W.</collectorName>
off
<collectorName id="26886D8FFFC22021FF6BF894FF40C576" box="[132,233,1880,1905]" pageId="9" pageNumber="40">Dong Sha</collectorName>
, ZHONGSHA 2015; st. CP4133;
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,
<geoCoordinate id="EE496E9EFFC22021FD4AF895FCAAC576" box="[677,771,1880,1904]" degrees="116" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="40" precision="925" value="116.4">116°24’E</geoCoordinate>
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.
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</paragraph>
<paragraph id="8BC20859FFC22021FF6BF86FFAA9C50F" blockId="9.[132,775,1954,2031]" lastBlockId="9.[813,1456,1671,1801]" pageId="9" pageNumber="40">
<materialsCitation id="3B150204FFC22021FF6BF86FFDC5C5BC" ID-GBIF-Occurrence="2564217916" box="[132,620,1954,1978]" collectionCode="OTHER" country="China" location="South China Sea" municipality="Sea" pageId="9" pageNumber="40" specimenCount="1">
OTHER SEQUENCED MATERIAL. —
<location id="8EA25E82FFC22021FE25F86FFDC5C5BC" LSID="urn:lsid:plazi:treatment:03D4B94FFFC22024FEF3F95EFD49C5EB:8EA25E82FFC22021FE25F86FFDC5C5BC" box="[458,620,1954,1978]" country="China" municipality="Sea" name="South China Sea" pageId="9" pageNumber="40">
<emphasis id="B909D44BFFC22021FE25F86FFDC5C5BC" bold="true" box="[458,620,1954,1978]" pageId="9" pageNumber="40">South China Sea</emphasis>
</location>
</materialsCitation>
<specimenCode id="DBDBA022FFC22021FD9EF86FFD25C5BC" box="[625,652,1954,1978]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="40">
<emphasis id="B909D44BFFC22021FD9EF86FFDD2C5BC" bold="true" box="[625,635,1954,1978]" pageId="9" pageNumber="40">•</emphasis>
<materialsCitation id="3B150204FFC22021FD90F86FFD25C5BC" ID-GBIF-Occurrence="2573220332" box="[639,652,1954,1978]" collectionCode="MNHN" collectorName="S. W. &amp; Dong Sha" country="China" elevation="1203" latitude="19.833334" longLatPrecision="1270" longitude="116.45" pageId="9" pageNumber="40" specimenCode="T 1" specimenCount="1">
<specimenCount id="9D7BC3D0FFC22021FD90F86FFD25C5BC" box="[639,652,1954,1978]" pageId="9" pageNumber="40">1</specimenCount>
</materialsCitation>
</specimenCode>
lv;
<collectionCode id="ED6C909CFFC22021FD43F86EFCAEC5BC" box="[684,775,1955,1978]" pageId="9" pageNumber="40">MNHN-</collectionCode>
IM-2013-61674 (
<figureCitation id="134614DCFFC22021FED4F870FE0EC5D2" box="[315,423,1981,2005]" captionStart="FIG" captionStartId="11.[132,143,1759,1777]" captionTargetBox="[132,1455,215,1713]" captionTargetId="figure@11.[132,1455,214,1713]" captionTargetPageId="11" captionText="FIG. 4. — Shells of Phaenomenella samadiae n. sp.: A-C, holotype, MNHN-IM-2013-61617, SL 58.2 mm; D, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm; E, F, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm; G, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="9" pageNumber="40">Figs 4E, F</figureCitation>
;
<figureCitation id="134614DCFFC22021FE58F871FE7DC5D2" box="[439,468,1980,2004]" captionStart="FIG" captionStartId="13.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1835]" captionTargetId="figure@13.[132,1455,215,1836]" captionTargetPageId="13" captionText="FIG. 5. — Radulae of Phaenomenella spp.A, B, Phaenomenella samadiae n. sp.; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (A); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (B); C-F. Phaenomenella nicoi n. sp.; C, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C); D, MNHN-IM-2013-59398 (shell on Fig. 7D); E, MNHN-IM-2013-61639 (shell on Fig. 7E); F, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="9" pageNumber="40">5B</figureCitation>
;
<figureCitation id="134614DCFFC22021FE0CF870FD8CC5D2" box="[483,549,1981,2005]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="9" pageNumber="40">6C, D</figureCitation>
);
<collectorName id="26886D8FFFC22021FDD2F870FDC7C5D3" box="[573,622,1981,2005]" name="T. SCHWARTZ" pageId="9" pageNumber="40">S.W.</collectorName>
off
<collectorName id="26886D8FFFC22021FD76F870FCAAC5D3" box="[665,771,1981,2005]" pageId="9" pageNumber="40">Dong Sha</collectorName>
, ZHONGSHA 2015; st. CP4134;
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,
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;
<quantity id="4C85A5BCFFC22021FD6FF81AFCAAC5E9" box="[640,771,2007,2031]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" metricValueMax="1.278" metricValueMin="1.128" pageId="9" pageNumber="40" unit="m" value="1203.0" valueMax="1278.0" valueMin="1128.0">
<elevation id="0050EF6AFFC22021FD6FF81AFCAAC5E9" box="[640,771,2007,2031]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" metricValueMax="1.278" metricValueMin="1.128" pageId="9" pageNumber="40" unit="m" value="1203.0" valueMax="1278.0" valueMin="1128.0">1128-1278 m</elevation>
</quantity>
.
<materialsCitation id="3B150204FFC22021FCC2F94AFAA1C499" ID-GBIF-Occurrence="2564217904" box="[813,1288,1671,1695]" collectionCode="OTHER" country="China" location="South China Sea" municipality="Sea" pageId="9" pageNumber="40" specimenCount="1">
OTHER STUDIED MATERIAL. —
<location id="8EA25E82FFC22021FB8FF94AFAA1C499" LSID="urn:lsid:plazi:treatment:03D4B94FFFC22024FEF3F95EFD49C5EB:8EA25E82FFC22021FB8FF94AFAA1C499" box="[1120,1288,1671,1695]" country="China" municipality="Sea" name="South China Sea" pageId="9" pageNumber="40">
<emphasis id="B909D44BFFC22021FB8FF94AFAA1C499" bold="true" box="[1120,1288,1671,1695]" pageId="9" pageNumber="40">South China Sea</emphasis>
</location>
</materialsCitation>
<specimenCode id="DBDBA022FFC22021FAFFF94AFA87C499" box="[1296,1326,1671,1695]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="40">
<emphasis id="B909D44BFFC22021FAFFF94AFAB3C499" bold="true" box="[1296,1306,1671,1695]" pageId="9" pageNumber="40">•</emphasis>
<materialsCitation id="3B150204FFC22021FACEF94AFA87C499" ID-GBIF-Occurrence="2573220325" box="[1313,1326,1671,1695]" collectionCode="MNHN" collectorName="S. W. &amp; Dong Sha" country="China" elevation="1203" latitude="19.833334" longLatPrecision="1270" longitude="116.45" pageId="9" pageNumber="40" specimenCode="T 1" specimenCount="1">
<specimenCount id="9D7BC3D0FFC22021FACEF94AFA87C499" box="[1313,1326,1671,1695]" pageId="9" pageNumber="40">1</specimenCount>
</materialsCitation>
</specimenCode>
lv;
<collectionCode id="ED6C909CFFC22021FABBF945FA06C499" box="[1364,1455,1672,1695]" pageId="9" pageNumber="40">MNHN-</collectionCode>
IM-2013-61670;
<collectorName id="26886D8FFFC22021FC21F96FFC53C4BC" box="[974,1018,1698,1722]" name="T. SCHWARTZ" pageId="9" pageNumber="40">S.W.</collectorName>
off
<collectorName id="26886D8FFFC22021FBF5F96FFBDFC4BC" box="[1050,1142,1698,1722]" pageId="9" pageNumber="40">Dong Sha</collectorName>
, ZHONGSHA 2015; st. CP4134;
<geoCoordinate id="EE496E9EFFC22021FCC2F971FC2BC4D2" box="[813,898,1724,1748]" degrees="19" direction="north" minutes="50" orientation="latitude" pageId="9" pageNumber="40" precision="925" value="19.833334">19°50’N</geoCoordinate>
,
<geoCoordinate id="EE496E9EFFC22021FC64F971FC41C4D2" box="[907,1000,1724,1748]" degrees="116" direction="east" minutes="27" orientation="longitude" pageId="9" pageNumber="40" precision="925" value="116.45">116°27’E</geoCoordinate>
;
<quantity id="4C85A5BCFFC22021FC1FF971FBDCC4D2" box="[1008,1141,1724,1748]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" metricValueMax="1.278" metricValueMin="1.128" pageId="9" pageNumber="40" unit="m" value="1203.0" valueMax="1278.0" valueMin="1128.0">
<elevation id="0050EF6AFFC22021FC1FF971FBDCC4D2" box="[1008,1141,1724,1748]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" metricValueMax="1.278" metricValueMin="1.128" pageId="9" pageNumber="40" unit="m" value="1203.0" valueMax="1278.0" valueMin="1128.0">1128-1278 m</elevation>
</quantity>
<specimenCode id="DBDBA022FFC22021FB96F970FB3DC4D2" box="[1145,1172,1724,1748]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="40">
•
<materialsCitation id="3B150204FFC22021FB68F971FB3DC4D2" ID-GBIF-Occurrence="2573220327" box="[1159,1172,1724,1748]" collectionCode="MNHN" country="China" elevation="1297" latitude="19.8" longLatPrecision="1270" longitude="116.48333" pageId="9" pageNumber="40" specimenCode="2013-59393, 2013-59665" specimenCount="2">
<specimenCount id="9D7BC3D0FFC22021FB68F971FB3DC4D2" box="[1159,1172,1724,1748]" pageId="9" pageNumber="40">2</specimenCount>
</materialsCitation>
</specimenCode>
lv;
<collectionCode id="ED6C909CFFC22021FB5AF970FA9EC4D2" box="[1205,1335,1725,1748]" pageId="9" pageNumber="40">MNHN-IM-</collectionCode>
<specimenCode id="DBDBA022FFC22021FAD9F971FA06C4D2" box="[1334,1455,1724,1748]" pageId="9" pageNumber="40">2013-59393</specimenCode>
(
<figureCitation id="134614DCFFC22021FCDAF91AFC2DC4E8" box="[821,900,1751,1775]" captionStart="FIG" captionStartId="11.[132,143,1759,1777]" captionTargetBox="[132,1455,215,1713]" captionTargetId="figure@11.[132,1455,214,1713]" captionTargetPageId="11" captionText="FIG. 4. — Shells of Phaenomenella samadiae n. sp.: A-C, holotype, MNHN-IM-2013-61617, SL 58.2 mm; D, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm; E, F, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm; G, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="9" pageNumber="40">Fig. 4D</figureCitation>
),
<collectionCode id="ED6C909CFFC22021FC73F91AFB8BC4E8" box="[924,1058,1751,1774]" pageId="9" pageNumber="40">MNHN-IM-</collectionCode>
<specimenCode id="DBDBA022FFC22021FBCDF91AFB36C4E8" box="[1058,1183,1750,1775]" pageId="9" pageNumber="40">2013-59665</specimenCode>
(
<figureCitation id="134614DCFFC22021FB5FF91AFB57C4E9" box="[1200,1278,1751,1775]" captionStart="FIG" captionStartId="11.[132,143,1759,1777]" captionTargetBox="[132,1455,215,1713]" captionTargetId="figure@11.[132,1455,214,1713]" captionTargetPageId="11" captionText="FIG. 4. — Shells of Phaenomenella samadiae n. sp.: A-C, holotype, MNHN-IM-2013-61617, SL 58.2 mm; D, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm; E, F, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm; G, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="9" pageNumber="40">Fig. 4G</figureCitation>
); ZHONGSHA; st. CP4157;
<geoCoordinate id="EE496E9EFFC22021FC47F93CFC56C50F" box="[936,1023,1777,1801]" degrees="19" direction="north" minutes="48" orientation="latitude" pageId="9" pageNumber="40" precision="925" value="19.8">19°48’N</geoCoordinate>
,
<geoCoordinate id="EE496E9EFFC22021FBE6F93CFBCEC50F" box="[1033,1127,1777,1801]" degrees="116" direction="east" minutes="29" orientation="longitude" pageId="9" pageNumber="40" precision="925" value="116.48333">116°29’E</geoCoordinate>
;
<quantity id="4C85A5BCFFC22021FB9EF93CFB55C50F" box="[1137,1276,1777,1801]" metricMagnitude="3" metricUnit="m" metricValue="1.297" metricValueMax="1.389" metricValueMin="1.205" pageId="9" pageNumber="40" unit="m" value="1297.0" valueMax="1389.0" valueMin="1205.0">
<elevation id="0050EF6AFFC22021FB9EF93CFB55C50F" box="[1137,1276,1777,1801]" metricMagnitude="3" metricUnit="m" metricValue="1.297" metricValueMax="1.389" metricValueMin="1.205" pageId="9" pageNumber="40" unit="m" value="1297.0" valueMax="1389.0" valueMin="1205.0">1205-1389 m</elevation>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC22021FCC2F8EDFB2BC58E" pageId="9" pageNumber="40" type="etymology">
<paragraph id="8BC20859FFC22021FCC2F8EDFB2BC58E" blockId="9.[813,1455,1824,1928]" pageId="9" pageNumber="40">
ETYMOLOGY. — Named in honour of Sarah Samadi, professor at MNHN, for her leadership in the France-Taiwan research programme in the context of which research cruises in the South
<collectingCountry id="F36A48C9FFC22021FAA6F89BFA2FC568" box="[1353,1414,1878,1902]" name="China" pageId="9" pageNumber="40">China</collectingCountry>
Sea discovered the present new species.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC22022FCC2F852FDD4C4CD" lastPageId="10" lastPageNumber="41" pageId="9" pageNumber="40" type="diagnosis">
<paragraph id="8BC20859FFC22021FCC2F852FA06C5EB" blockId="9.[813,1456,1951,2029]" pageId="9" pageNumber="40">
DIAGNOSIS. — Shell large for genus, up to
<quantity id="4C85A5BCFFC22021FB51F86DFAB1C5B1" box="[1214,1304,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="5.82" pageId="9" pageNumber="40" unit="mm" value="58.2">58.2 mm</quantity>
, broad fusiform with truncated base, short and strongly left reclined siphon canal. Spiral sculpture of distinct cords, more than 20 on last whorl. Radula
</paragraph>
<caption id="DF0258D1FFC12022FF6AF9E7FC6CC454" ID-DOI="http://doi.org/10.5281/zenodo.3652803" ID-Zenodo-Dep="3652803" httpUri="https://zenodo.org/record/3652803/files/figure.png" pageId="10" pageNumber="41" startId="10.[133,144,1578,1596]" targetBox="[165,1423,235,1540]" targetPageId="10">
<paragraph id="8BC20859FFC12022FF6AF9E7FC6CC454" blockId="10.[133,1455,1577,1618]" pageId="10" pageNumber="41">FIG. 3. — Phylogenetic tree of a dataset of Buccinoidea Rafinesque, 1815 obtained with Bayesian analysis of concatenated sequences of COI and 28S. Support values shown for the supported nodes with posterior probability values between 0.70 and 1.00 only.</paragraph>
</caption>
<paragraph id="8BC20859FFC12022FF6BF954FDD4C4CD" blockId="10.[132,774,1689,1740]" pageId="10" pageNumber="41">with tricuspate central teeth with short rectangular basal part and tricuspate lateral teeth with longest outermost cusp.</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC12022FF6BF92EFE7FC513" pageId="10" pageNumber="41" type="distribution">
<paragraph id="8BC20859FFC12022FF6BF92EFE7FC513" blockId="10.[132,775,1763,1814]" pageId="10" pageNumber="41">
DISTRIBUTION. — Presently the species was recorded only in the South
<collectingCountry id="F36A48C9FFC12022FF2AF933FEABC510" box="[197,258,1790,1814]" name="China" pageId="10" pageNumber="41">China</collectingCountry>
Sea at
<quantity id="4C85A5BCFFC12022FEA8F933FE7BC513" box="[327,466,1790,1814]" metricMagnitude="3" metricUnit="m" metricValue="1.297" metricValueMax="1.389" metricValueMin="1.205" pageId="10" pageNumber="41" unit="m" value="1297.0" valueMax="1389.0" valueMin="1205.0">1205-1389 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC12024FF6BF8FEFEFFC4A8" lastPageId="12" lastPageNumber="43" pageId="10" pageNumber="41" type="description">
<paragraph id="8BC20859FFC12022FF6BF8FEFE34C54B" blockId="10.[132,775,1843,2030]" box="[132,413,1843,1869]" pageId="10" pageNumber="41">
DESCRIPTION (
<typeStatus id="54C6B6FBFFC12022FECFF8FEFE3DC54A" box="[288,404,1843,1869]" pageId="10" pageNumber="41" type="holotype">HOLOTYPE</typeStatus>
)
</paragraph>
<paragraph id="8BC20859FFC12022FF6BF89EFF1FC56B" blockId="10.[132,775,1843,2030]" box="[132,182,1875,1901]" pageId="10" pageNumber="41">
<emphasis id="B909D44BFFC12022FF6BF89EFF1FC56B" box="[132,182,1875,1901]" italics="true" pageId="10" pageNumber="41">Shell</emphasis>
</paragraph>
<paragraph id="8BC20859FFC12023FF6BF8BEFD92C58B" blockId="10.[132,775,1843,2030]" lastBlockId="11.[132,775,1875,2030]" lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41">
Shell broad fusiform with truncated base (
<figureCitation id="134614DCFFC12022FDD1F8BEFDD0C58B" box="[574,633,1907,1933]" captionStart="FIG" captionStartId="11.[132,143,1759,1777]" captionTargetBox="[132,1455,215,1713]" captionTargetId="figure@11.[132,1455,214,1713]" captionTargetPageId="11" captionText="FIG. 4. — Shells of Phaenomenella samadiae n. sp.: A-C, holotype, MNHN-IM-2013-61617, SL 58.2 mm; D, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm; E, F, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm; G, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="10" pageNumber="41">Fig. 4</figureCitation>
A-C), strong, white under periostracum. Spire high, siphonal canal very short, strongly reclined to left. Protoconch and upper teleoconch whorls eroded, remaining teleoconch whorls 6½ in number. Teleoconch whorls convex, last and penultimate whorls less convex than upper ones. Suture distinct, adpressed. Spiral sculpture of distinct rounded on top spiral cords, on first not eroded teleoconch whorl (3
<superScript id="7C08A511FFC12022FB46F938FB10C500" attach="left" box="[1193,1209,1781,1798]" fontSize="7" pageId="10" pageNumber="41">rd</superScript>
remaining) 9 cords, on penultimate 11, on last whorl 24 cords, of which 4 on canal. Cords differing twice in width, the broader ones indistinctly subdivided by shallow spiral groove and with indistinct spiral striation, not visible on most narrow cords. Interspaces between cords from ¼ to more than cord’s width. On shell base and canal cords more broadly spaced. Upper teleoconch whorls with axial ribs, disappearing on last and penultimate whorls. Ribs nearly orthocline, broadly spaced, 12 on first preserved whorl, 14 on antepenultimate.
</paragraph>
<caption id="DF0258D1FFC02023FF6BF912FCE4C518" ID-DOI="http://doi.org/10.5281/zenodo.3652811" ID-Zenodo-Dep="3652811" httpUri="https://zenodo.org/record/3652811/files/figure.png" pageId="11" pageNumber="42" startId="11.[132,143,1759,1777]" targetBox="[132,1455,215,1713]" targetPageId="11">
<paragraph id="8BC20859FFC02023FF6BF912FCE4C518" blockId="11.[132,1457,1758,1822]" pageId="11" pageNumber="42">
FIG. 4. — Shells of
<taxonomicName id="4C7D73DAFFC02023FEC8F912FE57C4F7" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[295,510,1759,1777]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="42" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
<emphasis id="B909D44BFFC02023FEC8F912FE57C4F7" box="[295,510,1759,1777]" italics="true" pageId="11" pageNumber="42">Phaenomenella samadiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFC02023FDEBF912FD9EC4F7" box="[516,567,1759,1777]" pageId="11" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B909D44BFFC02023FDD0F912FDC8C4F7" bold="true" box="[575,609,1759,1777]" pageId="11" pageNumber="42">A -C</emphasis>
, holotype, MNHN-IM-2013-61617, SL 58.2 mm;
<emphasis id="B909D44BFFC02023FBECF912FBB8C4F7" bold="true" box="[1027,1041,1759,1777]" pageId="11" pageNumber="42">D</emphasis>
, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59393, SL 47.2 mm;
<emphasis id="B909D44BFFC02023FE04F93BFE5EC50E" bold="true" box="[491,503,1782,1800]" pageId="11" pageNumber="42">E</emphasis>
,
<emphasis id="B909D44BFFC02023FDEEF93BFDA5C50E" bold="true" box="[513,524,1782,1800]" pageId="11" pageNumber="42">F</emphasis>
, ZHONGSHA 2015, st. CP4134,sequenced spm, MNHN-IM-2013-61674, SL 51.4 mm;
<emphasis id="B909D44BFFC02023FB0CF938FB58C501" bold="true" box="[1251,1265,1781,1799]" pageId="11" pageNumber="42">G</emphasis>
, ZHONGSHA 2015, st. CP4157, non-sequenced spm, MNHN-IM-2013-59665, SL 37.1 mm. Scale bar: 5 cm.
</paragraph>
</caption>
<paragraph id="8BC20859FFC02023FF74F85EFCAEC5EB" blockId="11.[132,775,1875,2030]" pageId="11" pageNumber="42">Aperture broad ovate, white inside, angulated posteriorly, outer lip thick, slightly reflected.Parietal wall and columella with narrow but thick glossy callus with yellowish band along edge.</paragraph>
<paragraph id="8BC20859FFC02023FCACF89EFBB8C5AB" blockId="11.[813,1455,1875,2030]" pageId="11" pageNumber="42">Shell covered with light olive adhering periostracum, forming densely spaced low axial lamellae, obsolete on cords, but visible in interspaces.</paragraph>
<paragraph id="8BC20859FFC02023FCACF87EFA06C5E8" blockId="11.[813,1455,1875,2030]" pageId="11" pageNumber="42">Operculum spanning most of aperture, with distinctly turned leftwards terminal nucleus and weakly angulate in upper part.</paragraph>
<paragraph id="8BC20859FFC72024FF6BFF14FEFDC2F5" blockId="12.[132,775,217,531]" box="[132,340,217,244]" pageId="12" pageNumber="43">
<heading id="D08ABF35FFC72024FF6BFF14FEFDC2F5" box="[132,340,217,244]" fontSize="11" level="5" pageId="12" pageNumber="43" reason="1">
<emphasis id="B909D44BFFC72024FF6BFF14FEFDC2F5" box="[132,340,217,244]" italics="true" pageId="12" pageNumber="43">
Radula (
<figureCitation id="134614DCFFC72024FF30FF17FEE2C2F5" box="[223,331,218,244]" captionStart="FIG" captionStartId="13.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1835]" captionTargetId="figure@13.[132,1455,215,1836]" captionTargetPageId="13" captionText="FIG. 5. — Radulae of Phaenomenella spp.A, B, Phaenomenella samadiae n. sp.; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (A); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (B); C-F. Phaenomenella nicoi n. sp.; C, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C); D, MNHN-IM-2013-59398 (shell on Fig. 7D); E, MNHN-IM-2013-61639 (shell on Fig. 7E); F, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="12" pageNumber="43">Fig. 5A, B</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC20859FFC72024FF6BFF34FEB2C015" blockId="12.[132,775,217,531]" pageId="12" pageNumber="43">
Examined in
<typeStatus id="54C6B6FBFFC72024FEF4FF34FED5C315" box="[283,380,249,275]" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
and sequenced specimen MNHN- IM-2013-61674. Very similar in both specimens; central tooth with rather short rectangular basal part with weakly arcuate anterior margin and three short triangular broad cusps. Lateral teeth tricuspate with weakly curved basal side, attached to membrane. Outermost cusp recurved, medium long, inner cup weakly recurved, about 2/3 of outer cusp length. Intermediate cusp shortest, situated slightly closer to inner cusp.
</paragraph>
<paragraph id="8BC20859FFC72024FF6BFDF4FEBCC054" blockId="12.[132,775,569,690]" box="[132,277,569,594]" pageId="12" pageNumber="43">
<heading id="D08ABF35FFC72024FF6BFDF4FEBCC054" box="[132,277,569,594]" fontSize="11" level="5" pageId="12" pageNumber="43" reason="1">
<emphasis id="B909D44BFFC72024FF6BFDF4FEBCC054" box="[132,277,569,594]" italics="true" pageId="12" pageNumber="43">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="8BC20859FFC72024FF6BFD95FE61C0B4" blockId="12.[132,775,569,690]" pageId="12" pageNumber="43">
<typeStatus id="54C6B6FBFFC72024FF6BFD95FF43C074" box="[132,234,600,626]" pageId="12" pageNumber="43" type="holotype">Holotype</typeStatus>
(largest studied specimen), shell length
<quantity id="4C85A5BCFFC72024FD73FD95FCAAC074" box="[668,771,600,627]" metricMagnitude="-2" metricUnit="m" metricValue="5.82" pageId="12" pageNumber="43" unit="mm" value="58.2">58.2 mm</quantity>
, last whorl length
<quantity id="4C85A5BCFFC72024FEAEFDB5FE00C094" box="[321,425,632,659]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="12" pageNumber="43" unit="mm" value="35.8">35.8 mm</quantity>
, aperture length (without canal)
<quantity id="4C85A5BCFFC72024FF6BFD55FF42C0B4" box="[132,235,664,690]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="12" pageNumber="43" unit="mm" value="23.2">23.2 mm</quantity>
, diameter
<quantity id="4C85A5BCFFC72024FEB3FD55FE6DC0B4" box="[348,452,664,690]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="12" pageNumber="43" unit="mm" value="26.7">26.7 mm</quantity>
.
</paragraph>
<paragraph id="8BC20859FFC72024FF6BFD15FF48C0F4" blockId="12.[132,775,728,1232]" box="[132,225,728,754]" pageId="12" pageNumber="43">
<heading id="D08ABF35FFC72024FF6BFD15FF48C0F4" box="[132,225,728,754]" fontSize="11" level="5" pageId="12" pageNumber="43" reason="1">
<emphasis id="B909D44BFFC72024FF6BFD15FF48C0F4" box="[132,225,728,754]" italics="true" pageId="12" pageNumber="43">Anatomy</emphasis>
</heading>
</paragraph>
<paragraph id="8BC20859FFC72024FF6BFD35FEBCC6D6" blockId="12.[132,775,728,1232]" pageId="12" pageNumber="43">
Two specimens studied — MNHN-IM-2013-61674, male, sequenced
<typeStatus id="54C6B6FBFFC72024FF19FCD5FEFDC134" box="[246,340,792,818]" pageId="12" pageNumber="43" type="paratype">paratype</typeStatus>
; MNHN-IM-2013-61617, female,
<typeStatus id="54C6B6FBFFC72024FD21FCDAFF1FC154" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
. Soft body partly extracted from the shell. Head rather large, with two thick long tentacles. Eye lobes poorly defined, not pigmented in both examined specimens, eyes obviously absent. Mantle of female (
<figureCitation id="134614DCFFC72024FE4DFC5AFE5BC1B7" box="[418,498,919,945]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6B</figureCitation>
) approximately square in shape, with long siphon. Ctenidium comprises ¾ of mantle length and in average 1⁄5 of mantle width; bipectinate symmetric osphradium slightly narrower than ctenidium and ¾ of its length. Capsule gland medium large, covered by thick rectum and terminated by large vagina. In male’s mantle, prostate gland well-developed, situated parallel and equal in size to rectum. Penis (
<figureCitation id="134614DCFFC72024FE7DFBBBFE45C696" box="[402,492,1142,1168]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6 C</figureCitation>
) flattened, terminating in seminal papilla shifted to left side and not surrounded by a circlular fold.
</paragraph>
<paragraph id="8BC20859FFC72024FF6BFB38FEFFC4A8" blockId="12.[132,775,1269,1710]" pageId="12" pageNumber="43">
<emphasis id="B909D44BFFC72024FF6BFB38FEF8C709" bold="true" box="[132,337,1269,1295]" pageId="12" pageNumber="43">Digestive system.</emphasis>
Proboscis almost completely inverted into rhynchodaeum (
<figureCitation id="134614DCFFC72024FE94FAD8FE67C729" box="[379,462,1301,1327]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6E</figureCitation>
, pr). Several bands of proboscis retractors attached at middle part of both sides of rhynchodaeum (
<figureCitation id="134614DCFFC72024FED6FA98FE2CC769" box="[313,389,1365,1391]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6E</figureCitation>
, prr). Anterior oesophagus straight, along ventral side of rhynchodaeum (
<figureCitation id="134614DCFFC72024FDC3FAB8FDD4C789" box="[556,637,1397,1423]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6E</figureCitation>
, aoe). Valve of Leiblein medium large, situated immediately before nerve ring (
<figureCitation id="134614DCFFC72024FEE6FA78FEF7C7C9" box="[265,350,1461,1487]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6G</figureCitation>
, vl). Salivary glands large (about half proboscis length), fused ventrally (
<figureCitation id="134614DCFFC72024FE12FA18FDCEC7E9" box="[509,615,1493,1519]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6E, F</figureCitation>
, sg), with thin salivary ducts following along anterior oesophagus. Gland of Leiblein large, S-twisted, situated beneath salivary glands (
<figureCitation id="134614DCFFC72024FF62F9F9FF72C448" box="[141,219,1588,1614]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6E</figureCitation>
, gl) and following along posterior oesophagus and anterior aorta. Stomach small, occupying about 0.25 whorl (
<figureCitation id="134614DCFFC72024FF62F9B9FF76C489" box="[141,223,1652,1679]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[144,1435,220,1825]" captionTargetId="figure@14.[132,1455,215,1836]" captionTargetPageId="14" captionText="FIG. 6. — Anatomy of Phaenomenella samadiae n. sp.: A, B, E-G, holotype, MNHN-IM-2013-61617, female; C, D, MNHN-IM-2013-61674, male; A, cephalopodium, front view, mantle removed; B, mantle; C, uppermost part of penis; D, stomach, external view; E, foregut, ventral view; F, foregut, lateral view; G, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="12" pageNumber="43">Fig. 6D</figureCitation>
). Posterior mixing area small, posterior oesophagus and intestine wide.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC72024FF6BF91EFD49C5EB" pageId="12" pageNumber="43" type="discussion">
<paragraph id="8BC20859FFC72024FF6BF91EFF4CC4EA" blockId="12.[132,775,1747,2030]" box="[132,229,1747,1773]" pageId="12" pageNumber="43">REMARKS</paragraph>
<paragraph id="8BC20859FFC72024FF6BF939FD6DC528" blockId="12.[132,775,1747,2030]" pageId="12" pageNumber="43">The new species demonstrates some variability in shell shape with more inflated shell outline in smaller specimens.</paragraph>
<paragraph id="8BC20859FFC72024FF74F8FEFD49C5EB" blockId="12.[132,775,1747,2030]" pageId="12" pageNumber="43">
The new species is most similar in the shell shape to
<taxonomicName id="4C7D73DAFFC72024FD29F8F9FE2EC568" authority="Fraussen, 2008" authorityName="Fraussen" authorityYear="2008" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="43" phylum="Mollusca" rank="species" species="mokenorum">
<emphasis id="B909D44BFFC72024FD29F8F9FF49C56B" italics="true" pageId="12" pageNumber="43">P. mokenorum</emphasis>
<bibRefCitation id="EFEC75A8FFC72024FF09F899FE2EC568" author="FRAUSSEN K." box="[230,391,1876,1902]" journalOrPublisher="Veliger" pageId="12" pageNumber="43" pagination="48 - 50" part="50" refId="ref12618" refString="FRAUSSEN K. 2008 - A new Phaenomenella Fraussen &amp; Hadorn, 2006 (Gastropoda: Buccinidae), from the Andaman Sea. Veliger 50: 48 - 50." title="A new Phaenomenella Fraussen &amp; Hadorn, 2006 (Gastropoda: Buccinidae), from the Andaman Sea" type="journal article" year="2008">Fraussen, 2008</bibRefCitation>
</taxonomicName>
from the Andaman Sea, differing in better pronounced spiral cords. Another somewhat similar species is
<taxonomicName id="4C7D73DAFFC72024FF04F859FD10C5AB" authority="(Okutani &amp; Lan, 1994)" baseAuthorityName="Okutani &amp; Lan" baseAuthorityYear="1994" box="[235,697,1939,1966]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="43" phylum="Mollusca" rank="species" species="insulapratasensis">
<emphasis id="B909D44BFFC72024FF04F859FE07C5AB" box="[235,430,1939,1965]" italics="true" pageId="12" pageNumber="43">P. insulapratasensis</emphasis>
(Okutani &amp; Lan, 1994)
</taxonomicName>
, which possesses smaller, more ovoid shell with faster incrementing teleoconch whorls and a longer and more distinct canal.
</paragraph>
</subSubSection>
</treatment>
</document>
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