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<document id="FA87D8387FD0503B9603DC0A0DAD8D05" ID-CLB-Dataset="2793" ID-DOI="10.11646/zootaxa.5084.1.1" ID-GBIF-Dataset="bd42fe03-1a35-4f17-b0fe-55b6ec7fdf80" ID-ISSN="1175-5326" ID-Zenodo-Dep="5800716" ID-ZooBank="53B21C11-CA12-480F-8048-1A0601784172" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1640238335522" checkinUser="plazi" docAuthor="Melika, George, Nicholls, James A., Abrahamson, Warren G., Buss, Eileen A. &amp; Stone, Graham N." docDate="2021" docId="03868785FFCCFFB5FF76F908FB237A48" docLanguage="en" docName="zootaxa.5084.1.1.pdf" docOrigin="Zootaxa 5084 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Andricus williami Melika, Nicholls &amp; Stone 2021, sp. nov." docType="treatment" docUuid="122D3049-1D0A-4527-807C-B89DBFA47209" docUuidSource="ZooBank" docVersion="7" lastPageNumber="74" masterDocId="FFBFFFFDFF89FFFCFFE1FFE3FFE67B09" masterDocTitle="New species of Nearctic oak gall wasps (Hymenoptera: Cynipidae, Cynipini)" masterLastPageNumber="131" masterPageNumber="1" pageNumber="70" updateTime="1699473059019" updateUser="ExternalLinkService">
<mods:mods id="35DE3C9301B631B5AE51879861327C70" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FD8EBEDC88ED04ADB05083FDE299609A">
<mods:title id="3056A2F7A52437DB857CEC142AD26AC3">New species of Nearctic oak gall wasps (Hymenoptera: Cynipidae, Cynipini)</mods:title>
</mods:titleInfo>
<mods:name id="316654B6B311535C2735ACD226CA897F" type="personal">
<mods:role id="FB6E86523BFAE883EDF13178753A3ABD">
<mods:roleTerm id="6EFF7EBB4BDDF864EAF8B0C6F36EFBCD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="95532982B3F5277753DDB0C9B9FE6A69">Melika, George</mods:namePart>
</mods:name>
<mods:name id="EEB21B7F57C4B8010912CBD73695C11A" type="personal">
<mods:role id="C9FA489A92E4102E4B389ED8C4529874">
<mods:roleTerm id="110A98347E3C8404C5060DFE2064BDF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC16DF2C5B61369D29F5BA6C5CD780A7">Nicholls, James A.</mods:namePart>
<mods:nameIdentifier id="1C23077FB53F2944CC10988A7FCA0A9B" type="ORCID">0000-0002-9325-563X</mods:nameIdentifier>
<mods:nameIdentifier id="F32E56E9D553FA7BB58213FCD1763E0E" type="email">james.nicholls@csiro.au</mods:nameIdentifier>
</mods:name>
<mods:name id="5EBBD723050D69F204C9137BF7BD13C6" type="personal">
<mods:role id="8FDBDB91B32D71F584BA7C006239A7CF">
<mods:roleTerm id="69DA817DFD2AA9EEBB9F880435F2F8D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0727D13C4CF6B0DEF73464D1DC2FDDC2">Abrahamson, Warren G.</mods:namePart>
<mods:nameIdentifier id="5228D7D2B3AE920F2D3C0D139422DC1D" type="ORCID">0000-0002-3557-3613</mods:nameIdentifier>
<mods:nameIdentifier id="E610A9F36A13714AE2FA452AEB4B59B0" type="email">abrahmsn@bucknell.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="29E1C27867CD4F58AFCBF2F8CCD21A46" type="personal">
<mods:role id="DE3397A352D862A2599FC827EC4D1A4E">
<mods:roleTerm id="247093CE20A85328FC245B16AC959E20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C94D75AA2D6A42F25C257F247AEBCE6">Buss, Eileen A.</mods:namePart>
<mods:nameIdentifier id="14AC4B04E165B95B57DC68B1E8E2C57A" type="email">eabuss@ufl.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="AB9392E294F63011E1E5CFEB9E909ABD" type="personal">
<mods:role id="36BC63C24B81B95DA13677C8855E8C3C">
<mods:roleTerm id="B0FC95A9C2513BAFC9BD637A52B23F20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="769D0F9BC874C7367DFB070A70BAF2B9">Stone, Graham N.</mods:namePart>
<mods:nameIdentifier id="994052038CD0D29EA6D506623264CB33" type="ORCID">0000-0002-2737-696X</mods:nameIdentifier>
<mods:nameIdentifier id="A449F653938C49872E3741DA4203D431" type="email">gstone@staffmail.ed.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="00D4F3BAAE8C30F3799AE3B851871C6B">text</mods:typeOfResource>
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<mods:titleInfo id="6CAB7CF7A6F8A213BFD1B062588ACABB">
<mods:title id="FD5EBBF97FC0C1AAC0200BE51836BC20">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="22E02A79FECAE8EE7968E631D26E537D">
<mods:date id="A52C5F98814E70F6583430BAF880511C">2021</mods:date>
<mods:detail id="B47B9C2B9BA976B54CB0348C42CF0609" type="pubDate">
<mods:number id="AB5A47D6D7845657E13DF0E6E3C80C8E">2021-12-23</mods:number>
</mods:detail>
<mods:detail id="C95F7EFE9CA0F5036AFBB3B00CED86FF" type="volume">
<mods:number id="45C38CBBEA5EBA9C25AE1E667B4CF710">5084</mods:number>
</mods:detail>
<mods:detail id="FF58B5267855E874169EA78674BDFF4C" type="issue">
<mods:number id="7DAD25968C87ED1EA452EDB39B9B4776">1</mods:number>
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<mods:start id="3F778519E315186EFA40CC51798516C5">1</mods:start>
<mods:end id="9BA43C20DA3A6B114A00FD38D54CC2F2">131</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="DFDEA442F58DD3EFF8BA3F7927972C4C">journal article</mods:classification>
<mods:identifier id="02AA3224E05E7C5488EF38BFA3426470" type="CLB-Dataset">2793</mods:identifier>
<mods:identifier id="C7EDD9107F22976827EDF7CFAD1A5702" type="DOI">10.11646/zootaxa.5084.1.1</mods:identifier>
<mods:identifier id="6044DA9DD9F2C749C9545F0081958976" type="GBIF-Dataset">bd42fe03-1a35-4f17-b0fe-55b6ec7fdf80</mods:identifier>
<mods:identifier id="388D1620DA43758BB78D87341795090B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="75AC70B6CC9E5494934627DA2C386905" type="Zenodo-Dep">5800716</mods:identifier>
<mods:identifier id="56C070041F65C26C1E6665A42FCE2A79" type="ZooBank">53B21C11-CA12-480F-8048-1A0601784172</mods:identifier>
</mods:mods>
<treatment id="03868785FFCCFFB5FF76F908FB237A48" ID-DOI="http://doi.org/10.5281/zenodo.5800982" ID-GBIF-Taxon="191881063" ID-Zenodo-Dep="5800982" LSID="urn:lsid:zoobank.org:act:122D3049-1D0A-4527-807C-B89DBFA47209" httpUri="http://treatment.plazi.org/id/03868785FFCCFFB5FF76F908FB237A48" lastPageId="73" lastPageNumber="74" pageId="69" pageNumber="70">
<subSubSection id="C3356518FFCCFFB9FF76F908FCFA7C0F" box="[151,796,1771,1798]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="8B903693FFCCFFB9FF76F908FCFA7C0F" blockId="69.[151,961,1771,1871]" box="[151,796,1771,1798]" pageId="69" pageNumber="70">
<heading id="D0D881FFFFCCFFB9FF76F908FCFA7C0F" bold="true" box="[151,796,1771,1798]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="4">
<emphasis id="B95BEA81FFCCFFB9FF76F908FCFA7C0F" bold="true" box="[151,796,1771,1798]" pageId="69" pageNumber="70">
<taxonomicName id="4C2F4D10FFCCFFB9FF76F908FD577C0F" authority="Melika, Nicholls &amp; Stone" authorityName="Melika, Nicholls &amp; Stone" authorityYear="2021" box="[151,689,1771,1798]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
<emphasis id="B95BEA81FFCCFFB9FF76F908FE897C0F" bold="true" box="[151,367,1771,1798]" italics="true" pageId="69" pageNumber="70">Andricus williami</emphasis>
Melika, Nicholls &amp; Stone
</taxonomicName>
,
<taxonomicNameLabel id="A26857FAFFCCFFB9FD5FF90FFCFA7C0F" box="[702,796,1772,1798]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFCCFFB9FF76F8F3FC277C46" pageId="69" pageNumber="70" type="description">
<paragraph id="8B903693FFCCFFB9FF76F8F3FEDE7C22" blockId="69.[151,961,1771,1871]" box="[151,312,1808,1835]" pageId="69" pageNumber="70">
<figureCitation id="13142A16FFCCFFB9FF76F8F3FEDE7C22" box="[151,312,1808,1835]" captionStart-0="FIGURES 256–264" captionStart-1="FIGURES 265–269" captionStart-2="FIGURES 270–274" captionStartId-0="71.[151,264,1922,1946]" captionStartId-1="72.[151,264,1918,1942]" captionStartId-2="73.[151,264,1490,1514]" captionTargetBox-0="[160,1434,402,1880]" captionTargetBox-1="[160,1417,470,1881]" captionTargetBox-2="[158,1430,365,1457]" captionTargetId-0="figure-121@71.[150,1437,381,1899]" captionTargetId-1="figure-151@72.[153,1435,453,1895]" captionTargetId-2="figure-92@73.[149,1439,345,1467]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionTargetPageId-2="73" captionText-0="FIGURES 256–264. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 256–258, head, female: 256, frontal view, 257, dorsal view, 258, posterior view. 259–260, head, male: 259, frontal view, 260, dorsal view. 261–262, antenna: 261, female, 262, male. 263, tarsal claw, female. 264, forewing, part, female." captionText-1="FIGURES 265–269. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation, female. 265, mesosoma and propleura, frontal view, 266, mesosoma, lateral view, 267, mesoscutum, dorsal view, 268, mesoscutellum, dorsal view, 269, metascutellum and propodeum, posterodorsal view." captionText-2="FIGURES 270–274. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 270, mesosoma, male, lateral view. 271–272, metasoma, lateral view: 271, female, 272, male. 273–274, galls: 273, general view, infested acorns, 274, dissected acorns with larval chambers inside." figureDoi-0="http://doi.org/10.5281/zenodo.5800854" figureDoi-1="http://doi.org/10.5281/zenodo.5800856" figureDoi-2="http://doi.org/10.5281/zenodo.5800860" httpUri-0="https://zenodo.org/record/5800854/files/figure.png" httpUri-1="https://zenodo.org/record/5800856/files/figure.png" httpUri-2="https://zenodo.org/record/5800860/files/figure.png" pageId="69" pageNumber="70">Figs. 256–274</figureCitation>
</paragraph>
<paragraph id="8B903693FFCCFFB9FF76F8D7FC277C46" blockId="69.[151,961,1771,1871]" box="[151,961,1844,1871]" pageId="69" pageNumber="70">
<uri id="FFBE3A91FFCCFFB9FF76F8D7FC277C46" box="[151,961,1844,1871]" pageId="69" pageNumber="70">
urn:lsid:zoobank.org:act:
<uuid id="FF890C46FFCCFFB9FE49F8D7FC277C46" box="[424,961,1844,1871]" pageId="69" pageNumber="70">122D3049-1D0A-4527-807C-B89DBFA47209</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFCCFFB9FF76F89FFBF87CD7" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="8B903693FFCCFFB9FF76F89FFBF87CD7" blockId="69.[151,1437,1916,2014]" pageId="69" pageNumber="70">
<emphasis id="B95BEA81FFCCFFB9FF76F89FFEA17C9F" bold="true" box="[151,327,1916,1942]" pageId="69" pageNumber="70">Type material:</emphasis>
<materialsCitation id="3B473CCEFFCCFFB9FEAFF89FFD877CB3" ID-GBIF-Occurrence="3426514314" collectingDate="2008-04-07" collectorName="J. Nicholls" country="United States of America" location="Arizona" pageId="69" pageNumber="70" specimenCount="1" stateProvince="Arizona" typeStatus="holotype">
<typeStatus id="54948831FFCCFFB9FEAFF89FFE047C9F" box="[334,482,1916,1942]" pageId="69" pageNumber="70" type="holotype">HOLOTYPE</typeStatus>
female “
<collectingCountry id="F3387603FFCCFFB9FDA7F89FFD657C9F" box="[582,643,1916,1942]" name="United States of America" pageId="69" pageNumber="70">USA</collectingCountry>
,
<collectingRegion id="49EBF871FFCCFFB9FD6DF89FFD0F7C9F" box="[652,745,1916,1942]" country="United States of America" name="Arizona" pageId="69" pageNumber="70">Arizona</collectingRegion>
, Chiricahua Mtns. summit, leg.
<collectorName id="26DA5345FFCCFFB9FBB1F89FFB2C7C9F" box="[1104,1226,1916,1942]" pageId="69" pageNumber="70">J. Nicholls</collectorName>
,
<date id="FF911053FFCCFFB9FB34F89FFABD7C9F" box="[1237,1371,1916,1942]" pageId="69" pageNumber="70" value="2008-04-07">
<collectingDate id="EFD5E9BBFFCCFFB9FB34F89FFABD7C9F" box="[1237,1371,1916,1942]" pageId="69" pageNumber="70" value="2008-04-07">2008.04.07.</collectingDate>
</date>
Code AZ1976, spAZa3; ex
<taxonomicName id="4C2F4D10FFCCFFB9FE67F843FDB57CB3" box="[390,595,1952,1978]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="species" species="arizonica">
<emphasis id="B95BEA81FFCCFFB9FE67F843FDB57CB3" box="[390,595,1952,1978]" italics="true" pageId="69" pageNumber="70">Quercus arizonica</emphasis>
</taxonomicName>
”
</materialsCitation>
.
<materialsCitation id="3B473CCEFFCCFFB9FD8DF843FDB17CD7" ID-GBIF-Occurrence="3426514357" collectionCode="USNM" country="United States of America" pageId="69" pageNumber="70" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Arizona" typeStatus="paratype">
<typeStatus id="54948831FFCCFFB9FD8DF843FCED7CB3" box="[620,779,1952,1978]" pageId="69" pageNumber="70" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D29FD1AFFCCFFB9FCF5F843FC987CB3" box="[788,894,1952,1978]" pageId="69" pageNumber="70" type="female">8 females</specimenCount>
and
<specimenCount id="9D29FD1AFFCCFFB9FC50F843FBE37CB3" box="[945,1029,1952,1978]" pageId="69" pageNumber="70" type="male">3 males</specimenCount>
with the same labels as the
<typeStatus id="54948831FFCCFFB9FAD2F843FA7F7CB3" box="[1331,1433,1952,1978]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
. The
<typeStatus id="54948831FFCCFFB9FF2BF827FECA7CD7" box="[202,300,1988,2014]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
is deposited at the
<collectionCode id="ED3EAE56FFCCFFB9FDE2F827FDB17CD7" box="[515,599,1988,2014]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="69" pageNumber="70" type="Museum">USNM</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B473CCEFFCCFFB9FD83F827FBFC7CD7" ID-GBIF-Occurrence="3426514308" box="[610,1050,1988,2014]" collectionCode="PHDNRL" country="United States of America" pageId="69" pageNumber="70" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Arizona" typeStatus="paratype">
<specimenCount id="9D29FD1AFFCCFFB9FD83F827FD287CD7" box="[610,718,1988,2014]" pageId="69" pageNumber="70" type="female">8 females</specimenCount>
and
<specimenCount id="9D29FD1AFFCCFFB9FCE4F827FCBD7CD7" box="[773,859,1988,2014]" pageId="69" pageNumber="70" type="male">3 males</specimenCount>
at the
<collectionCode id="ED3EAE56FFCCFFB9FC47F827FBFC7CD7" box="[934,1050,1988,2014]" pageId="69" pageNumber="70">PHDNRL</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFCFFFBAFF26FF74FBF87BDF" pageId="70" pageNumber="71" type="etymology">
<paragraph id="8B903693FFCFFFBAFF26FF74FBF87BDF" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">
<emphasis id="B95BEA81FFCFFFBAFF26FF74FEAE7BB8" bold="true" box="[199,328,151,177]" pageId="70" pageNumber="71">Etymology</emphasis>
. Named after William L. Nicholls, son of J.A. Nicholls (Institute of Evolutionary Biology, University of
<collectingRegion id="49EBF871FFCFFFBAFF57FF58FEC97BDC" box="[182,303,187,213]" country="United Kingdom" name="City of Edinburgh" pageId="70" pageNumber="71">Edinburgh</collectingRegion>
,
<collectingCountry id="F3387603FFCFFFBAFED8FF58FE877BDC" box="[313,353,187,213]" name="United Kingdom" pageId="70" pageNumber="71">UK</collectingCountry>
and Australian National Insect Collection, CSIRO, Canberra).
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFCFFFBAFF26FF3DFBBA7AA4" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="8B903693FFCFFFBAFF26FF3DFBBA7AA4" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">
<emphasis id="B95BEA81FFCFFFBAFF26FF3DFEA47BF1" bold="true" box="[199,322,222,248]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
<taxonomicName id="4C2F4D10FFCFFFBAFEABFF3CFDA77BF0" authority="" box="[330,577,223,249]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Callirhytis">
<emphasis id="B95BEA81FFCFFFBAFEABFF3CFE497BF0" box="[330,431,223,249]" italics="true" pageId="70" pageNumber="71">Andricus</emphasis>
(
<emphasis id="B95BEA81FFCFFFBAFE20FF3CFDDE7BF0" box="[449,568,223,249]" italics="true" pageId="70" pageNumber="71">Callirhytis</emphasis>
)
</taxonomicName>
species, known to induce acorn galls in the
<collectingCountry id="F3387603FFCFFFBAFBDFFF3CFB927BF0" box="[1086,1140,223,249]" name="United States of America" pageId="70" pageNumber="71">USA</collectingCountry>
southwest are
<taxonomicName id="4C2F4D10FFCFFFBAFAC1FF3CFEFD7A14" authority="Weld, 1944" authorityName="Weld" authorityYear="1944" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B95BEA81FFCFFFBAFAC1FF3CFA7A7BF1" box="[1312,1436,223,249]" italics="true" pageId="70" pageNumber="71">A. costatus</emphasis>
<bibRefCitation id="EFBE4B62FFCFFFBAFF76FEE0FEFD7A14" author="Weld, L. H." box="[151,283,259,285]" pageId="70" pageNumber="71" pagination="1 - 24" refId="ref70508" refString="Weld, L. H. (1944) New American cynipid wasps from galls. Proceedings of the United States National Museum, 95, 1 - 24." type="journal article" year="1944">Weld, 1944</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2F4D10FFCFFFBAFEC6FEE0FDD57A14" authority="Weld, 1952" authorityName="Weld" authorityYear="1952" box="[295,563,258,285]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="prescotti">
<emphasis id="B95BEA81FFCFFFBAFEC6FEE0FE4E7A15" box="[295,424,259,285]" italics="true" pageId="70" pageNumber="71">A. prescotti</emphasis>
Weld, 1952
</taxonomicName>
,
<taxonomicName id="4C2F4D10FFCFFFBAFDDEFEE0FCB37A14" authority="Weld, 1957" authorityName="Weld" authorityYear="1957" box="[575,853,258,285]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="balanella">
<emphasis id="B95BEA81FFCFFFBAFDDEFEE0FD2C7A14" box="[575,714,259,285]" italics="true" pageId="70" pageNumber="71">C. balanella</emphasis>
Weld, 1957
</taxonomicName>
,
<taxonomicName id="4C2F4D10FFCFFFBAFC80FEE0FBB97A14" authority="Weld, 1952" authorityName="Weld" authorityYear="1952" box="[865,1119,258,285]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cistella">
<emphasis id="B95BEA81FFCFFFBAFC80FEE0FC327A14" box="[865,980,259,285]" italics="true" pageId="70" pageNumber="71">C. cistella</emphasis>
Weld, 1952
</taxonomicName>
, and
<taxonomicName id="4C2F4D10FFCFFFBAFB7AFEE0FA7E7A14" authority="Weld, 1922" authorityName="Weld" authorityYear="1922" box="[1179,1432,259,285]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="petrina">
<emphasis id="B95BEA81FFCFFFBAFB7AFEE0FAEB7A15" box="[1179,1293,259,285]" italics="true" pageId="70" pageNumber="71">C. petrina</emphasis>
Weld, 1922
</taxonomicName>
. However, in
<taxonomicName id="4C2F4D10FFCFFFBAFECBFEC4FE407A48" authorityName="Melika, Nicholls &amp; Stone" authorityYear="2021" box="[298,422,295,321]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="williami">
<emphasis id="B95BEA81FFCFFFBAFECBFEC4FE407A48" box="[298,422,295,321]" italics="true" pageId="70" pageNumber="71">A. williami</emphasis>
</taxonomicName>
the tarsal claws have a basal lobe, while in
<taxonomicName id="4C2F4D10FFCFFFBAFC7CFEC4FBCF7A48" authorityName="Weld" authorityYear="1957" box="[925,1065,295,321]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="balanella">
<emphasis id="B95BEA81FFCFFFBAFC7CFEC4FBCF7A48" box="[925,1065,295,321]" italics="true" pageId="70" pageNumber="71">C. balanella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D10FFCFFFBAFBD9FEC4FB4D7A48" authorityName="Weld" authorityYear="1952" box="[1080,1195,295,321]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cistella">
<emphasis id="B95BEA81FFCFFFBAFBD9FEC4FB4D7A48" box="[1080,1195,295,321]" italics="true" pageId="70" pageNumber="71">C. cistella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D10FFCFFFBAFB5BFEC4FACB7A49" authorityName="Weld" authorityYear="1922" box="[1210,1325,295,321]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="petrina">
<emphasis id="B95BEA81FFCFFFBAFB5BFEC4FACB7A49" box="[1210,1325,295,321]" italics="true" pageId="70" pageNumber="71">C. petrina</emphasis>
</taxonomicName>
the tarsal claws are simple, without a basal lobe. In
<taxonomicName id="4C2F4D10FFCFFFBAFD99FEA8FD137A6C" authorityName="Melika, Nicholls &amp; Stone" authorityYear="2021" box="[632,757,331,357]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="williami">
<emphasis id="B95BEA81FFCFFFBAFD99FEA8FD137A6C" box="[632,757,331,357]" italics="true" pageId="70" pageNumber="71">A. williami</emphasis>
</taxonomicName>
the anterior end of notaulus with rounded deep impression, the mesoscutum uniformly reticulate, while in
<taxonomicName id="4C2F4D10FFCFFFBAFD44FE8CFCC77A81" authorityName="Weld" authorityYear="1944" box="[677,801,367,393]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B95BEA81FFCFFFBAFD44FE8CFCC77A81" box="[677,801,367,393]" italics="true" pageId="70" pageNumber="71">A. costatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F4D10FFCFFFBAFCB9FE8CFC3E7A81" authorityName="Weld" authorityYear="1952" box="[856,984,367,393]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="prescotti">
<emphasis id="B95BEA81FFCFFFBAFCB9FE8CFC3E7A81" box="[856,984,367,393]" italics="true" pageId="70" pageNumber="71">A. prescotti</emphasis>
</taxonomicName>
the anterior end of the notaulus without rounded impression, the mesoscutum entirely glabrous, without any surface sculpture.
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFCFFFB4FF26FE55FE7A7A48" lastPageId="72" lastPageNumber="73" pageId="70" pageNumber="71" type="description">
<paragraph id="8B903693FFCFFFBAFF26FE55FB677AFC" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">
<emphasis id="B95BEA81FFCFFFBAFF26FE55FEBF7AD9" bold="true" box="[199,345,438,464]" pageId="70" pageNumber="71">Description.</emphasis>
Sexual female (
<figureCitation id="13142A16FFCFFFBAFDF7FE54FC8F7AD8" box="[534,873,438,465]" captionStart-0="FIGURES 256–264" captionStart-1="FIGURES 265–269" captionStartId-0="71.[151,264,1922,1946]" captionStartId-1="72.[151,264,1918,1942]" captionTargetBox-0="[160,1434,402,1880]" captionTargetBox-1="[160,1417,470,1881]" captionTargetId-0="figure-121@71.[150,1437,381,1899]" captionTargetId-1="figure-151@72.[153,1435,453,1895]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="FIGURES 256–264. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 256–258, head, female: 256, frontal view, 257, dorsal view, 258, posterior view. 259–260, head, male: 259, frontal view, 260, dorsal view. 261–262, antenna: 261, female, 262, male. 263, tarsal claw, female. 264, forewing, part, female." captionText-1="FIGURES 265–269. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation, female. 265, mesosoma and propleura, frontal view, 266, mesosoma, lateral view, 267, mesoscutum, dorsal view, 268, mesoscutellum, dorsal view, 269, metascutellum and propodeum, posterodorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.5800854" figureDoi-1="http://doi.org/10.5281/zenodo.5800856" httpUri-0="https://zenodo.org/record/5800854/files/figure.png" httpUri-1="https://zenodo.org/record/5800856/files/figure.png" pageId="70" pageNumber="71">Figs. 256–258, 261, 263–269</figureCitation>
,
<figureCitation id="13142A16FFCFFFBAFC96FE54FC437AD8" box="[887,933,439,465]" captionStart="FIGURES 270–274" captionStartId="73.[151,264,1490,1514]" captionTargetBox="[158,1430,365,1457]" captionTargetId="figure-92@73.[149,1439,345,1467]" captionTargetPageId="73" captionText="FIGURES 270–274. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 270, mesosoma, male, lateral view. 271–272, metasoma, lateral view: 271, female, 272, male. 273–274, galls: 273, general view, infested acorns, 274, dissected acorns with larval chambers inside." figureDoi="http://doi.org/10.5281/zenodo.5800860" httpUri="https://zenodo.org/record/5800860/files/figure.png" pageId="70" pageNumber="71">271</figureCitation>
). Head and mesosoma black, antennae dark brown, metasoma reddish brown; mandibles, maxillary and labial palpi, legs light brown.
</paragraph>
<paragraph id="8B903693FFCFFFBAFF26FE1CFAC97F74" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">Head coriaceous to reticulate, with dense white setae, denser on lower face and postgena; 1.3× as broad as high and slightly broader than mesosoma in frontal view, 2.0× as broad as long in dorsal view. Gena reticulate, broadened behind eye in frontal view, broader than transverse diameter of eye in lateral view. Malar space reticulate, with striae radiating from clypeus and reaching eye; eye 2.6× as high as length of malar space. Inner margins of eyes parallel. POL 1.8× as long as OOL, OOL 1.2× as long as diameter of lateral ocellus and shorter than LOL, all ocelli ovate, of same size. Transfacial distance nearly equal to height of eye; diameter of antennal torulus 2.0× as long as distance between them, distance between torulus and eye equal to diameter of torulus; lower face delicately coriaceous, with dense short white setae, without striae; slightly elevated median area coriaceous, without striae, with setae. Clypeus quadrangular as high as broad, smooth, glabrous; ventrally rounded, emarginate, without median incision, with a few long setae; anterior tentorial pit large, rounded; epistomal sulcus and clypeo-pleurostomal line indistinct. Frons and slightly elevated interocellar area reticulate, with short white setae. Vertex reticulate; occiput with delicately transverse parallel striae, with dense white setae; postocciput smooth, glabrous; postgena smooth, glabrous, with dense setae; posterior tentorial pit large, ovate, area below impressed; occipital foramen 3.0× as high as height of postgenal bridge; hypostomal carina emarginate, continuing into postgenal sulci which are not united, running parallel and slightly diverging in anterior 1/3, postgenal bridge anteriorly as broad as occipital foramen. Antenna longer than head+mesosoma, with 12 flagellomeres, scape, pedicel and F1 lighter than all subsequent flagellomeres, pedicel slightly longer than broad; F1 2.0× as long as pedicel; F1=F2; F2 1.3× as long as F3, F3=F4; subsequent flagellomeres shorter, nearly equal in length; F12 slightly longer than F11; placodeal sensilla on F3–F12.</paragraph>
<paragraph id="8B903693FFCFFFBAFF26FB64FB9A7DB4" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">Mesosoma longer than high, with dense white setae. Pronotum glabrous, with delicate striae laterally and dense setae; propleuron coriaceous, with dense setae. Mesoscutum reticulate, with dense setae; slightly longer than broad (greatest width measured across mesoscutum at level of base of tegulae). Notaulus complete, with smooth, glabrous bottom, broadened in posterior 1/3, posteriorly converging; anterior parallel and parapsidal lines marked by smooth glabrous areas; median mesoscutal line absent; circumscutellar carina narrow, reaching notaulus. Anterior end of notaulus with rounded deep impression. Mesoscutellum slightly longer than broad, trapezoid, broadened in posterior 1/3; uniformly rugose, with long white setae, posteriorly rounded, overhanging metanotum. Mesoscutellar foveae ovate, broader than high, with smooth, glabrous bottom, separated by narrow triangular elevated coriaceous central carina. Mesopleuron and speculum smooth, glabrous, with setae in posteroventral quarter, along anterior and ventral margins; mesopleural triangle smooth, glabrous, with dense white setae; dorsal and lateral axillar areas smooth, glabrous, with white setae; subaxillular bar smooth, glabrous, triangular, posterior end higher than height of metanotal trough; metapleural sulcus reaching mesopleuron in upper 1/3 of height, upper part of sulcus indistinct. Metascutellum coriaceous, 2.0× as high as height of smooth, glabrous ventral impressed area; metanotal trough smooth, glabrous, with few setae; central propodeal area smooth, glabrous, with few rugae and dense setae along subparallel lateral propodeal carinae; lateral propodeal area smooth, glabrous, with long white dense setae. Nucha with numerous delicate sulci dorsally and laterally. Tarsal claws toothed, with basal lobe.</paragraph>
<paragraph id="8B903693FFCFFFBAFF26F924FC6A7C20" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">Forewing longer than body, hyaline, margin with long dense cilia, veins dark brown, radial cell open, 3.4× as long as broad; R1 and Rs nearly reaching wing margin; areolet triangular, well-delimited, Rs+M distinct for 1/2 of the distance to basalis, its projection reaching basalis in mid height.</paragraph>
<paragraph id="8B903693FFCFFFBAFF26F8D0FC647CD4" blockId="70.[151,1437,151,2013]" pageId="70" pageNumber="71">
Metasoma slightly longer than head+mesosoma, longer than high in lateral view; second metasomal tergite extending to half the length of metasoma in dorsal view, with a few white setae anterolaterally, without micropunctures posteriorly, subsequent tergites smooth, glabrous, without micropunctures. Hypopygium without micropunctures, prominent part of ventral spine of hypopygium 2.5× as long as broad in ventral view, with long setae ventrally which not extend beyond apex of spine. Body length
<quantity id="4CD79B76FFCFFFBAFD40F820FCCE7CD4" box="[673,808,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="70" pageNumber="71" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
(n = 8).
</paragraph>
<paragraph id="8B903693FFCEFFBBFF26FF74FD677A6C" blockId="71.[151,1436,150,357]" pageId="71" pageNumber="72">
Male (
<figureCitation id="13142A16FFCEFFBBFEF3FF74FE147BB8" box="[274,498,150,177]" captionStart="FIGURES 256–264" captionStartId="71.[151,264,1922,1946]" captionTargetBox="[160,1434,402,1880]" captionTargetId="figure-121@71.[150,1437,381,1899]" captionTargetPageId="71" captionText="FIGURES 256–264. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 256–258, head, female: 256, frontal view, 257, dorsal view, 258, posterior view. 259–260, head, male: 259, frontal view, 260, dorsal view. 261–262, antenna: 261, female, 262, male. 263, tarsal claw, female. 264, forewing, part, female." figureDoi="http://doi.org/10.5281/zenodo.5800854" httpUri="https://zenodo.org/record/5800854/files/figure.png" pageId="71" pageNumber="72">Figs. 259–260, 262</figureCitation>
,
<figureCitation id="13142A16FFCEFFBBFE1FFF74FD817BB8" box="[510,615,151,177]" captionStart="FIGURES 270–274" captionStartId="73.[151,264,1490,1514]" captionTargetBox="[158,1430,365,1457]" captionTargetId="figure-92@73.[149,1439,345,1467]" captionTargetPageId="73" captionText="FIGURES 270–274. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 270, mesosoma, male, lateral view. 271–272, metasoma, lateral view: 271, female, 272, male. 273–274, galls: 273, general view, infested acorns, 274, dissected acorns with larval chambers inside." figureDoi="http://doi.org/10.5281/zenodo.5800860" httpUri="https://zenodo.org/record/5800860/files/figure.png" pageId="71" pageNumber="72">270, 272</figureCitation>
). Similar to female but hind coxae black; transfacial distance as long as height of eye; malar space with numerous striae radiating from clypeus and reaching eye; eye 3.0× as high as length of malar space; POL 1.8× as long as OOL, OOL 1.3× as long as diameter of lateral ocellus and 1.6× as long as LOL; ocelli larger; interocellar area elevated; antenna with 13 flagellomeres, F1 slightly curved, apical 2/3 broader than basal 1/3; F13 slightly longer than F12; placodeal sensilla on all flagellomeres; mesopleuron without white setae anteriorly. Body length
<quantity id="4CD79B76FFCEFFBBFE40FEA8FDCE7A6C" box="[417,552,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="71" pageNumber="72" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
(n = 3).
</paragraph>
<caption id="DF50661BFFCEFFBBFF76F861FC457CEB" ID-DOI="http://doi.org/10.5281/zenodo.5800854" ID-Zenodo-Dep="5800854" httpUri="https://zenodo.org/record/5800854/files/figure.png" pageId="71" pageNumber="72" startId="71.[151,264,1922,1946]" targetBox="[160,1434,402,1880]" targetPageId="71">
<paragraph id="8B903693FFCEFFBBFF76F861FC457CEB" blockId="71.[151,1437,1922,2018]" pageId="71" pageNumber="72">
<emphasis id="B95BEA81FFCEFFBBFF76F861FE897C93" bold="true" box="[151,367,1922,1946]" pageId="71" pageNumber="72">FIGURES 256–264.</emphasis>
<taxonomicName id="4C2F4D10FFCEFFBBFE99F861FCA27C93" authority="Melika, Nicholls &amp; Stone" authorityName="Melika, Nicholls &amp; Stone" authorityYear="2021" box="[376,836,1922,1946]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
<emphasis id="B95BEA81FFCEFFBBFE99F861FDC97C93" box="[376,559,1922,1946]" italics="true" pageId="71" pageNumber="72">Andricus williami</emphasis>
Melika, Nicholls &amp; Stone
</taxonomicName>
,
<emphasis id="B95BEA81FFCEFFBBFCB0F861FC457C93" bold="true" box="[849,931,1922,1946]" pageId="71" pageNumber="72">
<taxonomicNameLabel id="A26857FAFFCEFFBBFCB0F861FC457C93" box="[849,931,1922,1946]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, sexual generation. 256–258, head, female: 256, frontal view, 257, dorsal view, 258, posterior view. 259–260, head, male: 259, frontal view, 260, dorsal view. 261–262, antenna: 261, female, 262, male. 263, tarsal claw, female. 264, forewing, part, female.
</paragraph>
</caption>
<paragraph id="8B903693FFC1FFB4FF26FF75FE7A7A48" blockId="72.[151,1437,150,429]" pageId="72" pageNumber="73">
<emphasis id="B95BEA81FFC1FFB4FF26FF75FEE77BB9" bold="true" box="[199,257,150,176]" pageId="72" pageNumber="73">Gall.</emphasis>
(
<figureCitation id="13142A16FFC1FFB4FEF1FF74FE537BB8" box="[272,437,151,177]" captionStart="FIGURES 270–274" captionStartId="73.[151,264,1490,1514]" captionTargetBox="[158,1430,365,1457]" captionTargetId="figure-92@73.[149,1439,345,1467]" captionTargetPageId="73" captionText="FIGURES 270–274. Andricus williami Melika, Nicholls &amp; Stone, sp. nov., sexual generation. 270, mesosoma, male, lateral view. 271–272, metasoma, lateral view: 271, female, 272, male. 273–274, galls: 273, general view, infested acorns, 274, dissected acorns with larval chambers inside." figureDoi="http://doi.org/10.5281/zenodo.5800860" httpUri="https://zenodo.org/record/5800860/files/figure.png" pageId="72" pageNumber="73">Figs. 273–274</figureCitation>
). An acorn gall, developing as a swelling external to the cotyledons around the base of the acorn inside the acorn cup (see also
<bibRefCitation id="EFBE4B62FFC1FFB4FDCFFF58FD4D7BDC" author="Weld, L. H." box="[558,683,187,213]" pageId="72" pageNumber="73" refId="ref70806" refString="Weld, L. H. (1960) Cynipid galls of the Southwest. Privately printed, Ann Arbor, Michigan, 50 pp." type="book" year="1960">Weld 1960</bibRefCitation>
,
<figureCitation id="13142A16FFC1FFB4FD57FF58FCEF7BDC" box="[694,777,187,213]" captionStart="FIGURES 18–23" captionStartId="11.[151,264,1682,1706]" captionTargetBox="[164,1431,198,1656]" captionTargetId="figure-18@11.[151,1436,181,1658]" captionTargetPageId="11" captionText="FIGURES 18–23. Andricus archboldi Melika &amp; Abrahamson, sp. nov., asexual generation, female. 18–21, head: 18, frontal view, 19, dorsal view, 20, posterior view, 21, lateral view. 22, antenna. 23, forewing, part." figureDoi="http://doi.org/10.5281/zenodo.5800724" httpUri="https://zenodo.org/record/5800724/files/figure.png" pageId="72" pageNumber="73">Fig. 18</figureCitation>
). The larval cell intrudes into the body of the acorn; adults emerge through holes they create along the rim of the acorn cup. The gall is unilocular and consists only of the larval chamber ca
<quantity id="4CD79B76FFC1FFB4FEC3FEE0FE827A14" box="[290,356,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="72" pageNumber="73" unit="mm" value="3.0">3 mm</quantity>
long and
<quantity id="4CD79B76FFC1FFB4FE39FEE1FDAD7A14" box="[472,587,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="72" pageNumber="73" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
in diameter, surrounded by a hard woody wall. Several galls may develop around the same acorn.
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFC1FFB4FF26FEA9FEF37AA4" pageId="72" pageNumber="73" type="biology_ecology">
<paragraph id="8B903693FFC1FFB4FF26FEA9FEF37AA4" blockId="72.[151,1437,150,429]" pageId="72" pageNumber="73">
<emphasis id="B95BEA81FFC1FFB4FF26FEA9FEC17A6D" bold="true" box="[199,295,330,356]" pageId="72" pageNumber="73">Biology.</emphasis>
Only a sexual generation is known, which induces acorn galls on
<taxonomicName id="4C2F4D10FFC1FFB4FBD4FEA8FB237A6D" box="[1077,1221,331,357]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="arizonica">
<emphasis id="B95BEA81FFC1FFB4FBD4FEA8FB237A6D" box="[1077,1221,331,357]" italics="true" pageId="72" pageNumber="73">Q. arizonica</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFBE4B62FFC1FFB4FB34FEA8FA837A6F" author="Weld, L. H." box="[1237,1381,331,358]" pageId="72" pageNumber="73" refId="ref70806" refString="Weld, L. H. (1960) Cynipid galls of the Southwest. Privately printed, Ann Arbor, Michigan, 50 pp." type="book" year="1960">Weld (1960)</bibRefCitation>
also records this gall
<typeStatus id="54948831FFC1FFB4FEB9FE8CFE6E7A80" box="[344,392,367,393]" pageId="72" pageNumber="73">type</typeStatus>
from
<taxonomicName id="4C2F4D10FFC1FFB4FE33FE8CFDBF7A80" box="[466,601,367,393]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="gambelii">
<emphasis id="B95BEA81FFC1FFB4FE33FE8CFDBF7A80" box="[466,601,367,393]" italics="true" pageId="72" pageNumber="73">Q. gambelii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D10FFC1FFB4FD88FE8CFD3A7A81" authorityName="Nee" authorityYear="1801" box="[617,732,367,393]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="rugosa">
<emphasis id="B95BEA81FFC1FFB4FD88FE8CFD3A7A81" box="[617,732,367,393]" italics="true" pageId="72" pageNumber="73">Q. rugosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F4D10FFC1FFB4FCF6FE8CFC4C7A80" box="[791,938,367,393]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="turbinella">
<emphasis id="B95BEA81FFC1FFB4FCF6FE8CFC4C7A80" box="[791,938,367,393]" italics="true" pageId="72" pageNumber="73">Q. turbinella</emphasis>
</taxonomicName>
. Galls mature in April, adults emerge soon afterwards.
</paragraph>
</subSubSection>
<caption id="DF50661BFFC1FFB4FF76F89DFD447CD7" ID-DOI="http://doi.org/10.5281/zenodo.5800856" ID-Zenodo-Dep="5800856" httpUri="https://zenodo.org/record/5800856/files/figure.png" pageId="72" pageNumber="73" startId="72.[151,264,1918,1942]" targetBox="[160,1417,470,1881]" targetPageId="72">
<paragraph id="8B903693FFC1FFB4FF76F89DFD447CD7" blockId="72.[151,1437,1918,2014]" pageId="72" pageNumber="73">
<emphasis id="B95BEA81FFC1FFB4FF76F89DFE897C9F" bold="true" box="[151,367,1918,1942]" pageId="72" pageNumber="73">FIGURES 265–269.</emphasis>
<taxonomicName id="4C2F4D10FFC1FFB4FE98F89DFCA17C9F" authority="Melika, Nicholls &amp; Stone" authorityName="Melika, Nicholls &amp; Stone" authorityYear="2021" box="[377,839,1918,1942]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
<emphasis id="B95BEA81FFC1FFB4FE98F89DFDD67C9F" box="[377,560,1918,1942]" italics="true" pageId="72" pageNumber="73">Andricus williami</emphasis>
Melika, Nicholls &amp; Stone
</taxonomicName>
,
<emphasis id="B95BEA81FFC1FFB4FCB5F89DFC417C9F" bold="true" box="[852,935,1918,1942]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="A26857FAFFC1FFB4FCB5F89DFC417C9F" box="[852,935,1918,1942]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, sexual generation, female. 265, mesosoma and propleura, frontal view, 266, mesosoma, lateral view, 267, mesoscutum, dorsal view, 268, mesoscutellum, dorsal view, 269, metascutellum and propodeum, posterodorsal view.
</paragraph>
</caption>
<subSubSection id="C3356518FFC0FFB5FF26FF75FCF37BB8" box="[199,789,150,177]" pageId="73" pageNumber="74" type="distribution">
<paragraph id="8B903693FFC0FFB5FF26FF75FCF37BB8" blockId="73.[151,1436,150,321]" box="[199,789,150,177]" pageId="73" pageNumber="74">
<emphasis id="B95BEA81FFC0FFB5FF26FF75FE877BB9" bold="true" box="[199,353,150,176]" pageId="73" pageNumber="74">Distribution.</emphasis>
<collectingCountry id="F3387603FFC0FFB5FE86FF74FE447BB8" box="[359,418,151,177]" name="United States of America" pageId="73" pageNumber="74">USA</collectingCountry>
,
<collectingRegion id="49EBF871FFC0FFB5FE4DFF74FDEF7BB8" box="[428,521,151,177]" country="United States of America" name="Arizona" pageId="73" pageNumber="74">Arizona</collectingRegion>
, Chiricahua Mountains.
</paragraph>
</subSubSection>
<subSubSection id="C3356518FFC0FFB5FF26FF59FB237A48" pageId="73" pageNumber="74" type="discussion">
<paragraph id="8B903693FFC0FFB5FF26FF59FB237A48" blockId="73.[151,1436,150,321]" pageId="73" pageNumber="74">
<emphasis id="B95BEA81FFC0FFB5FF26FF59FEA97BDD" bold="true" box="[199,335,186,212]" pageId="73" pageNumber="74">Comments.</emphasis>
All known
<taxonomicName id="4C2F4D10FFC0FFB5FE35FF58FDAD7BDC" authorityName="sensu Burks" authorityYear="1979" box="[468,587,187,213]" class="Insecta" family="Cynipidae" genus="Callirhytis" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEA81FFC0FFB5FE35FF58FDAD7BDC" box="[468,587,187,213]" italics="true" pageId="73" pageNumber="74">Callirhytis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F4D10FFC0FFB5FD63FF58FCA67BDC" box="[642,832,187,213]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95BEA81FFC0FFB5FD63FF58FD017BDC" box="[642,743,187,213]" italics="true" pageId="73" pageNumber="74">Andricus</emphasis>
species
</taxonomicName>
that induce acorn galls, whether pip galls (external to the acorn cup) or stone galls (within the acorn cup and/or also inside the cotyledons of the acorn), have their asexual generations making such galls. The only exception, other than
<taxonomicName id="4C2F4D10FFC0FFB5FC8FFEE0FC0A7A14" authorityName="Melika, Nicholls &amp; Stone" authorityYear="2021" box="[878,1004,259,285]" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="williami">
<emphasis id="B95BEA81FFC0FFB5FC8FFEE0FC0A7A14" box="[878,1004,259,285]" italics="true" pageId="73" pageNumber="74">A. williami</emphasis>
</taxonomicName>
, is the sexual generation of
<taxonomicName id="4C2F4D10FFC0FFB5FAD6FEE0FE567A48" authority="Weld, 1926" authorityName="Weld" authorityYear="1926" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="chrysobalani">
<emphasis id="B95BEA81FFC0FFB5FAD6FEE0FECF7A48" italics="true" pageId="73" pageNumber="74">Andricus chrysobalani</emphasis>
<bibRefCitation id="EFBE4B62FFC0FFB5FED1FEC4FE567A48" author="Weld, L. H." box="[304,432,295,321]" pageId="73" pageNumber="74" pagination="1 - 131" refId="ref70470" refString="Weld, L. H. (1926) Field Notes on Gall-inhabiting Cynipid Wasps with descriptions of new species. Proceedings of the United States National Museum, 68 (10), 1 - 131." type="journal article" year="1926">Weld, 1926</bibRefCitation>
</taxonomicName>
which develops on
<taxonomicName id="4C2F4D10FFC0FFB5FD71FEC4FCD47A48" authorityName="Liebmann" authorityYear="1854" box="[656,818,295,321]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="73" pageNumber="74" phylum="Tracheophyta" rank="species" species="chrysolepis">
<emphasis id="B95BEA81FFC0FFB5FD71FEC4FCD47A48" box="[656,818,295,321]" italics="true" pageId="73" pageNumber="74">Q. chrysolepis</emphasis>
</taxonomicName>
and is known only from
<collectingRegion id="49EBF871FFC0FFB5FBADFEC4FB277A48" box="[1100,1217,295,321]" country="United States of America" name="California" pageId="73" pageNumber="74">California</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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