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<document id="74C3ACA063B9FCB0CA9FD9FC732AC767" ID-CLB-Dataset="7399" ID-DOI="10.11646/zootaxa.4951.1.6" ID-GBIF-Dataset="07eb2d3b-7c51-452d-a9be-34fa0ef8031d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4655824" ID-ZooBank="4769755A-E399-409D-B666-FEF0FF92574D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617267750445" checkinUser="plazi" docAuthor="Jiang, Ri-Xin, Liu, Xiao-Fan &amp; Wang, Shuo" docDate="2021" docId="03860A65FFDCFF8E2184CB23FE448DA8" docLanguage="en" docName="zootaxa.4951.1.6.pdf" docOrigin="Zootaxa 4951 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Broscosoma guoliangi Jiang, Liu &amp; Wang 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="139" masterDocId="FFBF721DFFDDFF8C2113C943FFC18A6B" masterDocTitle="New species and new distributional record of the genus Broscosoma Rosenhauer 1846 (Coleoptera, Carabidae, Broscinae) from China" masterLastPageNumber="146" masterPageNumber="137" pageNumber="138" updateTime="1698935843091" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DC4BE38E650604EB25454D685A81D141" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2F1731C370822D449E3E70A5D00F5E95">
<mods:title id="FC78CC186392D86C0AAE45D718B1E223">New species and new distributional record of the genus Broscosoma Rosenhauer 1846 (Coleoptera, Carabidae, Broscinae) from China</mods:title>
</mods:titleInfo>
<mods:name id="243924A40DB8643B00D364050DFBE6B3" type="personal">
<mods:role id="F0B3900ACAAFFC5EC2F8E9541B6DFA2C">
<mods:roleTerm id="907442DFA3CDD80207B61CE0B76D9E4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9B0B4B584F580CDB461E3CB0FFC3058">Jiang, Ri-Xin</mods:namePart>
<mods:nameIdentifier id="0112E48F82BE79B091CF3F71E868B5B9" type="ORCID">0000-0002-5339-853X</mods:nameIdentifier>
<mods:affiliation id="14B5F0B788A61C741EE124D25262BF14">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qingdao, 266042, P. R. China. &amp; maoshuwuyouzhi @ 163. com, https: // orcid. org / 0000 - 0002 - 5339 - 853 X</mods:affiliation>
</mods:name>
<mods:name id="1B4EF2390766CCFFE77F1FA6024B206E" type="personal">
<mods:role id="8ECC129564CBB1B931E85DAF7B9D1096">
<mods:roleTerm id="4902604FFFA4F8906783F13437B55E9B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C01FFD7893A85074599C53C43676D03A">Liu, Xiao-Fan</mods:namePart>
<mods:nameIdentifier id="1FFDE162AA5FA76C65BD749D1D7E6878" type="ORCID">0000-0002-0537-9596</mods:nameIdentifier>
<mods:affiliation id="94948D0B8F826D71887DE86427E649DE">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qingdao, 266042, P. R. China. &amp; 912210359 @ qq. com, https: // orcid. org / 0000 - 0002 - 0537 - 9596</mods:affiliation>
</mods:name>
<mods:name id="9919BFF6F04F94D1A100220CADC82401" type="personal">
<mods:role id="46957182E6BC48BD8289F705F80AA04B">
<mods:roleTerm id="6BFB10F4B2A5DB3BF4C713236A1B0E3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B10D54BFCC41BBB36ECE6AD9C8B0A395">Wang, Shuo</mods:namePart>
<mods:nameIdentifier id="248EAFC64C865FB62BE02CA389A10109" type="ORCID">0000-0003-0412-3799</mods:nameIdentifier>
<mods:affiliation id="5D9CD99FE14CE7D134712CCEA67A508F">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qingdao, 266042, P. R. China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C72CB514A454F273191248F8606B3E62">text</mods:typeOfResource>
<mods:relatedItem id="C3FD466A357C7F4D682BCAEAC18F4233" type="host">
<mods:titleInfo id="1884592969D1FDA05AAF64B26CD22A3B">
<mods:title id="6FA2EB6ABA3FA06ECF34FA177AE29E1C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DE2972F45E13DB826E1C0DD7B7E85BCE">
<mods:date id="007D969DA6927CB0BCAD5344930F44F8">2021</mods:date>
<mods:detail id="ACDB04B9C24F2CDCF854972275FDEF7A" type="pubDate">
<mods:number id="3C0AA5988813449FCE70ECF907CD5008">2021-04-01</mods:number>
</mods:detail>
<mods:detail id="198887574A72C88D6853D0201439421D" type="volume">
<mods:number id="A37D8681FE671183CE62B2446CED249D">4951</mods:number>
</mods:detail>
<mods:detail id="61F5E54130931D7C5880433120497EF7" type="issue">
<mods:number id="4433B9365DF6C5C9BD2D9810CDF8A69B">1</mods:number>
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<mods:start id="B74FA196B6FB1D324B7611DE65535304">137</mods:start>
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<mods:classification id="8587BBEBFDE365C4468566DFB6064B80">journal article</mods:classification>
<mods:identifier id="D27DE285BBD9FCAD1139725DB79A2902" type="CLB-Dataset">7399</mods:identifier>
<mods:identifier id="6433047B52089A82BD5DC87AD50D9BEE" type="DOI">10.11646/zootaxa.4951.1.6</mods:identifier>
<mods:identifier id="48448460350E01D2A885E684B36EC599" type="GBIF-Dataset">07eb2d3b-7c51-452d-a9be-34fa0ef8031d</mods:identifier>
<mods:identifier id="0080A33C9FB9D8E6E3504B2DEF3E42F8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="54D3B13373A3C2870CB74FE93D976056" type="Zenodo-Dep">4655824</mods:identifier>
<mods:identifier id="4DE12CAB1A996A21F4FD22D958C9C6A9" type="ZooBank">4769755A-E399-409D-B666-FEF0FF92574D</mods:identifier>
</mods:mods>
<treatment id="03860A65FFDCFF8E2184CB23FE448DA8" ID-DOI="http://doi.org/10.5281/zenodo.4683461" ID-GBIF-Taxon="181002232" ID-Zenodo-Dep="4683461" LSID="urn:lsid:plazi:treatment:03860A65FFDCFF8E2184CB23FE448DA8" httpUri="http://treatment.plazi.org/id/03860A65FFDCFF8E2184CB23FE448DA8" lastPageId="2" lastPageNumber="139" pageId="1" pageNumber="138">
<subSubSection id="C335E8F8FFDCFF8D2184CB23FC628813" box="[151,931,607,634]" pageId="1" pageNumber="138" type="nomenclature">
<paragraph id="8B90BB73FFDCFF8D2184CB23FC628813" blockId="1.[151,931,607,670]" box="[151,931,607,634]" pageId="1" pageNumber="138">
<heading id="D0D80C1FFFDCFF8D2184CB23FC628813" bold="true" box="[151,931,607,634]" fontSize="11" level="1" pageId="1" pageNumber="138" reason="1">
<emphasis id="B95B6761FFDCFF8D2184CB23FC628813" bold="true" box="[151,931,607,634]" pageId="1" pageNumber="138">
<taxonomicName id="4C2FC0F0FFDCFF8D2184CB23FD5E8811" ID-CoL="8MN69" authority="Jiang, Liu &amp; Wang" authorityName="Jiang, Liu &amp; Wang" authorityYear="2021" box="[151,671,607,634]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="guoliangi" status="sp. nov.">
<emphasis id="B95B6761FFDCFF8D2184CB23FE668811" bold="true" box="[151,423,607,634]" italics="true" pageId="1" pageNumber="138">Broscosoma guoliangi</emphasis>
Jiang, Liu &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="A268DA1AFFDCFF8D23BFCB23FCC88811" box="[684,777,608,634]" pageId="1" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
Ṟ亮ừ步m
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C335E8F8FFDCFF8D2184CBC7FF3E88F5" box="[151,255,644,670]" pageId="1" pageNumber="138" type="description">
<paragraph id="8B90BB73FFDCFF8D2184CBC7FF3E88F5" blockId="1.[151,931,607,670]" box="[151,255,644,670]" pageId="1" pageNumber="138">
<figureCitation id="1314A7F6FFDCFF8D2184CBC7FF3E88F5" box="[151,255,644,670]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1602,1626]" captionStartId-1="3.[151,250,1072,1096]" captionTargetBox-0="[177,1406,205,1561]" captionTargetBox-1="[158,1437,201,1049]" captionTargetId-0="figure-20@2.[151,1436,198,1576]" captionTargetId-1="figure-17@3.[151,1437,201,1049]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Dorsal habitus of Broscosoma guoliangi sp. nov. A. male; B. Female." captionText-1="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi-0="http://doi.org/10.5281/zenodo.4655828" figureDoi-1="http://doi.org/10.5281/zenodo.4655830" httpUri-0="https://zenodo.org/record/4655828/files/figure.png" httpUri-1="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Figs. 1–2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C335E8F8FFDCFF8D2184CB8FFF328939" pageId="1" pageNumber="138" type="materials_examined">
<paragraph id="8B90BB73FFDCFF8D2184CB8FFF328939" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D2184CB8FFEFE888D" bold="true" box="[151,319,716,742]" pageId="1" pageNumber="138">Type material</emphasis>
(
<specimenCount id="9D2970FAFFDCFF8D2043CB8FFE2C888D" box="[336,493,716,742]" pageId="1" pageNumber="138" type="generic">24 specimens</specimenCount>
,
<specimenCount id="9D2970FAFFDCFF8D20EACB8FFD8B888C" box="[505,586,716,743]" pageId="1" pageNumber="138" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="9D2970FAFFDCFF8D2344CB8EFD56888C" box="[599,663,717,743]" pageId="1" pageNumber="138" type="female">7 ♀♀</specimenCount>
).
<typeStatus id="549405D1FFDCFF8D23A3CB8FFCDC888D" box="[688,797,716,742]" pageId="1" pageNumber="138" type="holotype">Holotype</typeStatus>
:
<materialsCitation id="3B47B12EFFDCFF8D223BCB8FFBCF888D" ID-GBIF-Occurrence="3071255303" box="[808,1038,716,743]" country="China" location="CHINA" pageId="1" pageNumber="138" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F338FBE3FFDCFF8D223BCB8FFC44888D" box="[808,901,716,742]" name="China" pageId="1" pageNumber="138">CHINA</collectingCountry>
:
<specimenCount id="9D2970FAFFDCFF8D2282CB8EFC69888C" box="[913,936,717,743]" pageId="1" pageNumber="138" type="male">♂</specimenCount>
, labelled
</materialsCitation>
‘
<materialsCitation id="3B47B12EFFDCFF8D2508CB8FFA968961" ID-GBIF-Occurrence="3071255305" collectingDate="2020-05-18" collectionCode="SCAU" collectorName="Guo Liang" country="China" elevation="1450" location="Luobading" municipality="Sanming City" pageId="1" pageNumber="138" specimenCount="1" stateProvince="Fujian" typeStatus="holotype">
<collectingCountry id="F338FBE3FFDCFF8D2508CB8FFBA6888D" box="[1051,1127,716,742]" name="China" pageId="1" pageNumber="138">China</collectingCountry>
:
<collectingRegion id="49EB7591FFDCFF8D2560CB8FFAE6888D" box="[1139,1319,716,742]" country="China" name="Fujian" pageId="1" pageNumber="138">Fujian Province</collectingRegion>
(
<emphasis id="B95B6761FFDCFF8D242BCB8EFA4D888F" bold="true" box="[1336,1420,717,740]" pageId="1" pageNumber="138">AE建省</emphasis>
),
<collectingMunicipality id="6BF42109FFDCFF8D2184CBB3FEF38961" box="[151,306,752,778]" pageId="1" pageNumber="138">Sanming City</collectingMunicipality>
(
<emphasis id="B95B6761FFDCFF8D2051CBB2FE578963" bold="true" box="[322,406,753,776]" pageId="1" pageNumber="138">三DZ市</emphasis>
),
<location id="8EF0EDA8FFDCFF8D20BECBB3FDE78961" LSID="urn:lsid:plazi:treatment:03860A65FFDCFF8E2184CB23FE448DA8:8EF0EDA8FFDCFF8D20BECBB3FDE78961" box="[429,550,752,778]" country="China" municipality="Sanming City" name="Luobading" pageId="1" pageNumber="138" stateProvince="Fujian">Luobading</location>
(
<emphasis id="B95B6761FFDCFF8D2324CBB2FD4A8963" bold="true" box="[567,651,753,776]" pageId="1" pageNumber="138">ƤẘLj</emphasis>
),
<elevation id="00025C40FFDCFF8D23B1CBB3FC698960" box="[674,936,752,779]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="138" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
elevation
<quantity id="4CD71696FFDCFF8D2203CBB3FC698960" box="[784,936,752,779]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="138" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400-1500 m</quantity>
</elevation>
,
<date id="FF919DB3FFDCFF8D22A0CBB3FBF58960" box="[947,1076,752,779]" pageId="1" pageNumber="138" value="2020-05-18">
<collectingDate id="EFD5645BFFDCFF8D22A0CBB3FBF58960" box="[947,1076,752,779]" pageId="1" pageNumber="138" value="2020-05-18">2020.05.18</collectingDate>
</date>
,
<collectorName id="26DADEA5FFDCFF8D252CCBB3FB768961" box="[1087,1207,752,778]" pageId="1" pageNumber="138">Guo Liang</collectorName>
leg.’ (
<collectionCode id="ED3E23B6FFDCFF8D25EDCBB3FA8E8961" box="[1278,1359,752,778]" pageId="1" pageNumber="138">SCAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B47B12EFFDCFF8D2472CBB2FF2E8939" ID-GBIF-Occurrence="3071255302" collectingDate="2020-05-18" collectionCode="MYNU, NMPC, SCAU" collectorName="Guo Liang" country="China" elevation="1450" location="Luobading" municipality="Sanming City" pageId="1" pageNumber="138" specimenCount="46" specimenCount-female="14" specimenCount-male="32" stateProvince="Fujian" typeStatus="paratype">
<typeStatus id="549405D1FFDCFF8D2472CBB2FF0A8944" pageId="1" pageNumber="138" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F338FBE3FFDCFF8D21C5CA57FEF28945" box="[214,307,788,814]" name="China" pageId="1" pageNumber="138">CHINA</collectingCountry>
: (
<specimenCount id="9D2970FAFFDCFF8D2056CA57FE558944" box="[325,404,788,815]" pageId="1" pageNumber="138" type="male">16 ♂♂</specimenCount>
,
<specimenCount id="9D2970FAFFDCFF8D20B3CA56FE1C8944" box="[416,477,789,815]" pageId="1" pageNumber="138" type="female">7 ♀♀</specimenCount>
), same label data as the holotype (
<specimenCount id="9D2970FAFFDCFF8D2272CA57FC4A8944" box="[865,907,788,815]" pageId="1" pageNumber="138" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D2970FAFFDCFF8D2285CA57FC018944" box="[918,960,788,815]" pageId="1" pageNumber="138" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED3E23B6FFDCFF8D22DECA56FBE78944" box="[973,1062,789,815]" pageId="1" pageNumber="138">MYNU</collectionCode>
;
<specimenCount id="9D2970FAFFDCFF8D2522CA57FB9C8944" box="[1073,1117,788,815]" pageId="1" pageNumber="138" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D2970FAFFDCFF8D257BCA57FB538944" box="[1128,1170,788,815]" pageId="1" pageNumber="138" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED3E23B6FFDCFF8D258CCA57FB338945" box="[1183,1266,788,814]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="138">NMPC</collectionCode>
;
<specimenCount id="9D2970FAFFDCFF8D25EECA57FA8A8944" box="[1277,1355,788,815]" pageId="1" pageNumber="138" type="male">14 ♂♂</specimenCount>
,
<specimenCount id="9D2970FAFFDCFF8D2444CA57FA578944" box="[1367,1430,788,815]" pageId="1" pageNumber="138" type="female">5 ♀♀</specimenCount>
,
<collectionCode id="ED3E23B6FFDCFF8D2184CA7BFF268939" box="[151,231,824,850]" pageId="1" pageNumber="138">SCAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C335E8F8FFDCFF8D21D4CA1FFCB18988" pageId="1" pageNumber="138" type="diagnosis">
<paragraph id="8B90BB73FFDCFF8D21D4CA1FFCB18988" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D21D4CA1FFE83891D" bold="true" box="[199,322,860,886]" pageId="1" pageNumber="138">Diagnosis.</emphasis>
Body shiny, dark brown or black, legs, labrum, palpi and antennae brown. Frontal grooves irregular and impunctate, labrum with six setigerous punctures, clypeus with two setigerous punctures. Pronotum globose, with a pair of setigerous punctures. Elytra with eight distinct striae, shoulders largely rounded, each elytron with two setigerous punctures at base, two setigerous punctures near apex.
</paragraph>
</subSubSection>
<subSubSection id="C335E8F8FFDCFF8D21D4CAAFFE418DC4" pageId="1" pageNumber="138" type="description">
<paragraph id="8B90BB73FFDCFF8D21D4CAAFFF1B8E41" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D21D4CAAFFE988E6D" bold="true" box="[199,345,1004,1030]" pageId="1" pageNumber="138">Description.</emphasis>
Male. Body (
<figureCitation id="1314A7F6FFDCFF8D20E9CAAFFD938E6D" box="[506,594,1004,1030]" captionStart="FIGURE 1" captionStartId="2.[151,250,1602,1626]" captionTargetBox="[177,1406,205,1561]" captionTargetId="figure-20@2.[151,1436,198,1576]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Broscosoma guoliangi sp. nov. A. male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.4655828" httpUri="https://zenodo.org/record/4655828/files/figure.png" pageId="1" pageNumber="138">Fig. 1A</figureCitation>
) dark brown or black, slender and obviously convex, dorsal surface very shiny.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CD77FEAA8EB5" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
Head (
<figureCitation id="1314A7F6FFDCFF8D2006CD77FEB08E25" box="[277,369,1076,1102]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2A</figureCitation>
) wider than long, convex, dorsal surface shiny, lacking punctures, frontal grooves irregular and impunctate. Labrum nearly rectangular, with six setigerous punctures and emarginate at apical margin. Apical margin of clypeus slightly emarginate with two setigerous punctures. Eyes large, supraorbital setae level with rear margin of eyes. Antenna longer than the combination of head and pronotum. Antennomeres I–IV glabrous, V–XI densely pubescent.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CDAAFE0C8F04" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
Pronotum (
<figureCitation id="1314A7F6FFDCFF8D2055CDABFE638F69" box="[326,418,1256,1282]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2A</figureCitation>
), globose, distinctly convex, slightly longer than wide, widest slightly anterior of middle, equally constricted at anterior and basal margins, dorsal surface shiny without punctures. Median longitudinal impression and anterior impression shallow but distinct. One pair of setigerous punctures located at anterior middle of lateral margin of pronotum.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CC3BFD2D8C48" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
Elytra (
<figureCitation id="1314A7F6FFDCFF8D2008CC3BFEB08FF9" box="[283,369,1400,1426]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2B</figureCitation>
) longer than wide, strongly convex, dorsal surface shiny, each elytron with eight distinct striae. Scutellum nearly triangular, small. Shoulders largely rounded, narrow. Each elytron with four setigerous punctures, two pairs located at base and two pairs located near apex. The inner setigerous punctures of basal elytra located near base of stria II, outer punctures located at shoulders; the inner apical setigerous punctures located near apex of stria II, and the outer puncture located at apex of stria VII.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CF6EFB798C2D" blockId="1.[151,1437,716,2039]" box="[199,1208,1580,1607]" pageId="1" pageNumber="138">Legs simple, glabrous, only tibial apex with few thin setae, protarsomeres slightly dilated.</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CF13FD218CD8" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
Median lobe of aedeagus (
<figureCitation id="1314A7F6FFDCFF8D20E7CF13FD8B8C01" box="[500,586,1616,1642]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2C</figureCitation>
) simple, ending in a rounded tip; right paramere (
<figureCitation id="1314A7F6FFDCFF8D2591CF13FB1A8C01" box="[1154,1243,1616,1642]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2D</figureCitation>
) arcuate, acutely angular at apex, with dense long setae along apical margin; left paramere (
<figureCitation id="1314A7F6FFDCFF8D22C5CF37FBEB8CE5" box="[982,1066,1652,1678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2E</figureCitation>
) expanded at middle and tapered to narrow at apex, apical rounded and without setae.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CFFFFE418C91" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D21D4CFFFFE068CBD" bold="true" box="[199,455,1724,1750]" pageId="1" pageNumber="138">Measurements (mm).</emphasis>
BL 7.21–7.25; HL 1.13–1.20, HW 1.45–1.50; PL 2.13–2.25, PW 1.88–2.00; EL 3.80– 3.95, EW 2.70–2.80.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CE47FF168D0C" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
Female (
<figureCitation id="1314A7F6FFDCFF8D203ACE47FEBF8D75" box="[297,382,1796,1822]" captionStart="FIGURE 1" captionStartId="2.[151,250,1602,1626]" captionTargetBox="[177,1406,205,1561]" captionTargetId="figure-20@2.[151,1436,198,1576]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Broscosoma guoliangi sp. nov. A. male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.4655828" httpUri="https://zenodo.org/record/4655828/files/figure.png" pageId="1" pageNumber="138">Fig. 1B</figureCitation>
) similar to male, usually larger, with ovipositor smooth and arched, protarsomeres not dilated basally, basal segment of ovipositor (
<figureCitation id="1314A7F6FFDCFF8D2324CE6BFD4B8D29" box="[567,650,1832,1858]" captionStart="FIGURE 2" captionStartId="3.[151,250,1072,1096]" captionTargetBox="[158,1437,201,1049]" captionTargetId="figure-17@3.[151,1437,201,1049]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Broscosoma guoliangi sp. nov. A. Head and pronotum, dorsal view; B. Elytra; C. Aedeagus, lateral view; D. Right paramere; E. Left paramere; F. Left ovipositor. Features in red: sut, elytral suture; s1‒s8, elytral striae; red dots: setae. Red dots: setigerous punctures." figureDoi="http://doi.org/10.5281/zenodo.4655830" httpUri="https://zenodo.org/record/4655830/files/figure.png" pageId="1" pageNumber="138">Fig. 2F</figureCitation>
) with nine short setae and apical segment with two thinner and longer setae.
</paragraph>
<paragraph id="8B90BB73FFDCFF8D21D4CE33FE418DC4" blockId="1.[151,1437,716,2039]" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D21D4CE33FE068DE1" bold="true" box="[199,455,1904,1930]" pageId="1" pageNumber="138">Measurements (mm).</emphasis>
BL 7.48–8.03; HL 1.10–1.33, HW 1.48–1.53; PL 2.38–2.45, PW 2.00–2.23; EL 4.00– 4.25, EW 2.95–3.10.
</paragraph>
</subSubSection>
<subSubSection id="C335E8F8FFDCFF8E21D4CEFBFF1A8ADA" lastPageId="2" lastPageNumber="139" pageId="1" pageNumber="138" type="distribution">
<paragraph id="8B90BB73FFDCFF8D21D4CEFBFDB28DB9" blockId="1.[151,1437,716,2039]" box="[199,627,1976,2002]" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D21D4CEFBFEA08DB9" bold="true" box="[199,353,1976,2002]" pageId="1" pageNumber="138">Distribution.</emphasis>
<collectingCountry id="F338FBE3FFDCFF8D2074CEFBFE6F8DB9" box="[359,430,1976,2002]" name="China" pageId="1" pageNumber="138">China</collectingCountry>
:
<collectingRegion id="49EB7591FFDCFF8D20AACEFBFDAE8DB9" box="[441,623,1976,2002]" country="China" name="Fujian" pageId="1" pageNumber="138">Fujian Province</collectingRegion>
.
</paragraph>
<paragraph id="8B90BB73FFDCFF8E21D4CE9FFF1A8ADA" blockId="1.[151,1437,716,2039]" lastBlockId="2.[151,219,151,177]" lastPageId="2" lastPageNumber="139" pageId="1" pageNumber="138">
<emphasis id="B95B6761FFDCFF8D21D4CE9FFE488D9D" bold="true" box="[199,393,2012,2038]" pageId="1" pageNumber="138">Biological notes.</emphasis>
Both females andmales were collected on the lower surface of a damp cliff at night (
<figureCitation id="1314A7F6FFDCFF8D2425CE9FFA5D8D9C" box="[1334,1436,2012,2039]" captionStart="FIGURE 7" captionStartId="8.[151,250,1359,1383]" captionTargetBox="[151,1437,181,1335]" captionTargetId="figure-19@8.[151,1437,181,1335]" captionTargetPageId="8" captionText="FIGURE 7. Habitat of Broscosoma species. A. General environment of the type locality of B. qiului sp. nov.; B. Microenvironment of the type locality of B. guoliangi sp. nov.; C. Living adult of B. qiului sp. nov.; D–E. Living adult of B. guoliangi sp. nov. A &amp; C taken by Dr. Lu Qiu; B, D–E taken by Mr. Liang Guo." figureDoi="http://doi.org/10.5281/zenodo.4655842" httpUri="https://zenodo.org/record/4655842/files/figure.png" pageId="1" pageNumber="138">Figs. 7B,</figureCitation>
D–E).
</paragraph>
</subSubSection>
<caption id="DF50EBFBFFDFFF8E2184CF01FC1C8C31" ID-DOI="http://doi.org/10.5281/zenodo.4655828" ID-Zenodo-Dep="4655828" box="[151,989,1602,1627]" httpUri="https://zenodo.org/record/4655828/files/figure.png" pageId="2" pageNumber="139" startId="2.[151,250,1602,1626]" targetBox="[177,1406,205,1561]" targetPageId="2">
<paragraph id="8B90BB73FFDFFF8E2184CF01FC1C8C31" blockId="2.[151,989,1602,1627]" box="[151,989,1602,1627]" pageId="2" pageNumber="139">
<emphasis id="B95B6761FFDFFF8E2184CF01FED28C31" bold="true" box="[151,275,1602,1626]" pageId="2" pageNumber="139">FIGURE 1.</emphasis>
Dorsal habitus of
<taxonomicName id="4C2FC0F0FFDFFF8E20DDCF00FD738C31" authority="Jiang &amp; Liu &amp; Wang, 2021" authorityName="Jiang &amp; Liu &amp; Wang" authorityYear="2021" box="[462,690,1602,1626]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="guoliangi" status="sp. nov.">
<emphasis id="B95B6761FFDFFF8E20DDCF00FD738C31" box="[462,690,1602,1626]" italics="true" pageId="2" pageNumber="139">Broscosoma guoliangi</emphasis>
</taxonomicName>
<emphasis id="B95B6761FFDFFF8E23ABCF00FCE98C31" bold="true" box="[696,808,1602,1627]" pageId="2" pageNumber="139">
<taxonomicNameLabel id="A268DA1AFFDFFF8E23ABCF00FCC88C30" box="[696,777,1603,1627]" pageId="2" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
male;
<emphasis id="B95B6761FFDFFF8E227ECF00FC458C30" bold="true" box="[877,900,1603,1627]" pageId="2" pageNumber="139">B.</emphasis>
Female.
</paragraph>
</caption>
<subSubSection id="C335E8F8FFDFFF8E21D4CFCAFE258CAC" pageId="2" pageNumber="139" type="etymology">
<paragraph id="8B90BB73FFDFFF8E21D4CFCAFE258CAC" blockId="2.[151,1437,1673,1987]" pageId="2" pageNumber="139">
<emphasis id="B95B6761FFDFFF8E21D4CFCAFE8F8CC8" bold="true" box="[199,334,1673,1699]" pageId="2" pageNumber="139">Etymology.</emphasis>
This species is named in honor of Mr. Liang Guo (
<collectingRegion id="49EB7591FFDFFF8E2280CFCAFC1E8CC8" box="[915,991,1673,1699]" country="China" name="Fujian" pageId="2" pageNumber="139">Fujian</collectingRegion>
,
<collectingCountry id="F338FBE3FFDFFF8E22F8CFCAFBF38CCF" box="[1003,1074,1673,1700]" name="China" pageId="2" pageNumber="139">China</collectingCountry>
), who collected the new species and donated it to us for study.
</paragraph>
</subSubSection>
<subSubSection id="C335E8F8FFDFFF8E21D4CF92FE448DA8" pageId="2" pageNumber="139" type="discussion">
<paragraph id="8B90BB73FFDFFF8E21D4CF92FE448DA8" blockId="2.[151,1437,1673,1987]" pageId="2" pageNumber="139">
<emphasis id="B95B6761FFDFFF8E21D4CF92FEFA8C80" bold="true" box="[199,315,1745,1771]" pageId="2" pageNumber="139">Remarks.</emphasis>
The new species resembles
<taxonomicName id="4C2FC0F0FFDFFF8E2392CF92FAC68C80" authority="Jiang, Feng &amp; Wang, 2020" authorityName="Jiang, Feng &amp; Wang" authorityYear="2020" box="[641,1287,1745,1771]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="zhengyuandongi">
<emphasis id="B95B6761FFDFFF8E2392CF92FC0A8C80" box="[641,971,1745,1771]" italics="true" pageId="2" pageNumber="139">Broscosoma zhengyuandongi</emphasis>
<bibRefCitation id="EFBEC682FFDFFF8E22C0CF92FAC68C80" author="Jiang, R. - X. &amp; Feng, Q. &amp; Wang, S." box="[979,1287,1745,1771]" pageId="2" pageNumber="139" pagination="173 - 180" refId="ref3537" refString="Jiang, R. - X., Feng, Q. &amp; Wang, S. (2020) Two new species of the genus Broscosoma Rosenhauer, 1846 (Coleoptera, Carabidae, Broscinae) from China. Zootaxa, 4821 (1), 173 - 180. https: // doi. org / 10.11646 / zootaxa. 4821.1.10" type="journal article" year="2020">Jiang, Feng &amp; Wang, 2020</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C2FC0F0FFDFFF8E2456CF92FD978D64" authority="Jiang, Feng &amp; Wang, 2020" authorityName="Jiang, Feng &amp; Wang" authorityYear="2020" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="xuhaoi">
<emphasis id="B95B6761FFDFFF8E2456CF92FEE48D64" italics="true" pageId="2" pageNumber="139">Broscosoma xuhaoi</emphasis>
<bibRefCitation id="EFBEC682FFDFFF8E203FCFB6FD978D64" author="Jiang, R. - X. &amp; Feng, Q. &amp; Wang, S." box="[300,598,1781,1807]" pageId="2" pageNumber="139" pagination="173 - 180" refId="ref3537" refString="Jiang, R. - X., Feng, Q. &amp; Wang, S. (2020) Two new species of the genus Broscosoma Rosenhauer, 1846 (Coleoptera, Carabidae, Broscinae) from China. Zootaxa, 4821 (1), 173 - 180. https: // doi. org / 10.11646 / zootaxa. 4821.1.10" type="journal article" year="2020">
Jiang, Feng &amp; Wang,
<quantity id="4CD71696FFDFFF8E230DCFB6FD978D64" box="[542,598,1781,1807]" metricMagnitude="1" metricUnit="m" metricValue="5.1308" pageId="2" pageNumber="139" unit="in" value="2020.0">2020</quantity>
</bibRefCitation>
</taxonomicName>
in their habitus. However, it can be readily distinguished from
<taxonomicName id="4C2FC0F0FFDFFF8E240BCFB6FF328D58" authorityName="Jiang, Feng &amp; Wang" authorityYear="2020" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="zhengyuandongi">
<emphasis id="B95B6761FFDFFF8E240BCFB6FF328D58" italics="true" pageId="2" pageNumber="139">B. zhengyuandongi</emphasis>
</taxonomicName>
by the following characters: 1) striae on the elytra shallow (cf. deeper); 2) apical part of the left male paramere wide (cf. narrow); 3) the right male paramere covered with setae from middle to apex (cf. apical third); 4) basal segment of the female ovipositor with nine short setae (cf. seven). It can be easily distinguished from
<taxonomicName id="4C2FC0F0FFDFFF8E2497CE22FF228DF4" authorityName="Jiang, Feng &amp; Wang" authorityYear="2020" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="xuhaoi">
<emphasis id="B95B6761FFDFFF8E2497CE22FF228DF4" italics="true" pageId="2" pageNumber="139">B. xuhaoi</emphasis>
</taxonomicName>
by the apical margin of clypeus with two setigerous punctures (cf. four), and the left male paramere lacking setae (cf. with setae).
</paragraph>
</subSubSection>
</treatment>
</document>
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