142 lines
18 KiB
XML
142 lines
18 KiB
XML
<document id="C0F8EBC0A96A34DBB8052E8DADA81F46" ID-DOI="10.5281/zenodo.3674855" ID-GBIF-Dataset="ac53680f-6358-4d6b-8880-ccf3dd73297e" ID-ISSN="1942-1354" ID-Zenodo-Dep="3674855" ID-ZooBank="F9BABD1C-B18B-47FD-9B87-F44C49FD9D30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582111424063" checkinUser="felipe" docAuthor="F, Guillermo González, Hanley, Guy A. & Gordon, Robert D." docDate="2019" docId="4E5F8790C75095298D90D3E5265AF9A8" docLanguage="en" docName="InsectaMundi.0714.1-32.pdf.imf" docOrigin="Insecta Mundi 714 (714)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Cryptognatha karla Gonzalez and Hanley 2019, new species" docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="B266FFE8C75E95278D50D20C2369FF89" masterDocTitle="South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant" masterLastPageNumber="32" masterPageNumber="1" pageNumber="13" updateTime="1698719361767" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="840EB98B6864DE4D33C48F1C5717F554">South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant</mods:title>
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<mods:name id="E19F076A0506A4CF71317FF94C5DE82A" type="personal">
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<mods:roleTerm id="53BF826DE7CDB83A78C5CF2E62C9A15D">Author</mods:roleTerm>
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<mods:namePart id="07DAF6A2B5B47D9552A8844DCE27BFC9">F, Guillermo González</mods:namePart>
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<mods:roleTerm id="FC020222B6A6B32562DD10A3DC2D5DE0">Author</mods:roleTerm>
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<mods:namePart id="C47FC0AE2F3080315A6AC7EC702DB753">Hanley, Guy A.</mods:namePart>
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</mods:name>
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<mods:name id="89BB9D5B79471D271CDFE56E8FA99A3E" type="personal">
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<mods:roleTerm id="AA71E28C0E6332B354C42CBCBA61E725">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1EF4304DD7B489CA48B7A9F8000640FE">Gordon, Robert D.</mods:namePart>
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<mods:typeOfResource id="26A3DBFD2A7D4827D848C38EE12BB8D8">text</mods:typeOfResource>
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<mods:title id="DBA9E65A45B507B9085F9666763E3FCA">Insecta Mundi</mods:title>
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<mods:part id="7A4C020D7E2CCD95F4FFB9150E2838DE">
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<mods:date id="AD519B09AC2070F0F2240ED801D14418">2019</mods:date>
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<mods:number id="5378CF907B38C278CAE330DF1EA90B38">2019-06-28</mods:number>
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<mods:number id="723AACF5E66DE8853FCBEF4E6C316A6B">714</mods:number>
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<mods:number id="658D612C42CE1018EEFC79868B0216DC">714</mods:number>
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<mods:classification id="A64FCE8E918B27ADF4247AB6DA82CAF4">journal article</mods:classification>
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<mods:identifier id="FBAAC0EC25F0334E3F72341E2FC42149" type="DOI">10.5281/zenodo.3674855</mods:identifier>
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<treatment id="4E5F8790C75095298D90D3E5265AF9A8" ID-DOI="http://doi.org/10.5281/zenodo.3680660" ID-GBIF-Taxon="162173270" ID-Zenodo-Dep="3680660" LSID="urn:lsid:plazi:treatment:4E5F8790C75095298D90D3E5265AF9A8" httpUri="http://treatment.plazi.org/id/4E5F8790C75095298D90D3E5265AF9A8" lastPageNumber="13" pageId="14" pageNumber="13">
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<subSubSection id="8EEC650DC75095298D90D3E520B4FD88" box="[192,989,489,513]" pageId="14" pageNumber="13" type="nomenclature">
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<paragraph id="C6493686C75095298D90D3E520B4FD88" blockId="14.[192,989,489,513]" box="[192,989,489,513]" pageId="14" pageNumber="13">
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<heading id="9D0181EAC75095298D90D3E520B4FD88" bold="true" box="[192,989,489,513]" fontSize="10" level="2" pageId="14" pageNumber="13" reason="3">
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<emphasis id="F482EA94C75095298D90D3E520B4FD88" bold="true" box="[192,989,489,513]" pageId="14" pageNumber="13">
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10.
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<taxonomicName id="01F64D05C75095298DBDD3E52042FD88" authority="Gonzalez and Hanley" authorityName="Gonzalez and Hanley" authorityYear="2019" box="[237,811,489,513]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="karla" status="sp. nov.">
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<emphasis id="F482EA94C75095298DBDD3E52169FD88" bold="true" box="[237,512,489,513]" italics="true" pageId="14" pageNumber="13">Cryptognatha karla</emphasis>
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González and Hanley
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</taxonomicName>
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,
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<taxonomicNameLabel id="EFB157EFC75095298E69D3E520B4FD88" box="[825,989,489,513]" pageId="14" pageNumber="13" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8EEC650DC75095298D90D0252127FAE8" pageId="14" pageNumber="13" type="description">
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<paragraph id="C6493686C75095298D90D02520F8FB28" blockId="14.[192,1441,553,1185]" pageId="14" pageNumber="13">
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<emphasis id="F482EA94C75095298D90D025215DFDC8" bold="true" box="[192,564,553,577]" pageId="14" pageNumber="13">
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Description. Male
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<typeStatus id="194D8824C75095298CE5D0252147FDC8" box="[437,558,553,577]" pageId="14" pageNumber="13" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Length
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<quantity id="010E9B63C75095298FC5D0262198FDC8" box="[661,753,554,578]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="13" unit="mm" value="1.9">1.9 mm</quantity>
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, width
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<quantity id="010E9B63C75095298E16D02620CAFDC8" box="[838,931,554,578]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
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. Dorsal surface shiny, pronotum with faint trace of microsculpture. Color yellow; head slightly darkened yellow; pronotum yellow; elytron with large dark brown macula extended from just anterior to humeral callus posteriorly to midpoint of elytron, macula narrow at base, widened to irregularly truncate apex (
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<figureCitation id="5ECD2A03C75095298EE9D0852779FD28" box="[953,1040,649,673]" captionStart="Figures 55–72" captionStartId="28.[192,287,1816,1838]" captionTargetBox="[194,1426,307,1678]" captionTargetId="figure@28.[192,1438,292,1723]" captionTargetPageId="28" captionText="Figures 55–72. Cryptognatha spp. 55–61) Cryptognatha karla. 55) Habitus. 56) Frons. 57) Protibia. 58) Phallobase ventral. 59) Phallobase lateral. 60) Penis. 61) Spermathecal capsule. 62–66) Cryptognatha clarki. 62) Habitus. 63) Frons. 64) Phallobase ventral. 65) Phallobase lateral. 66) Penis. 67–72) Cryptognatha celia. 67) Habitus. 68) Frons. 69) Spermathecal capsule. 70) Phallobase ventral. 71) Phallobase lateral. 72) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674863" httpUri="https://zenodo.org/record/3674863/files/figure.png" pageId="14" pageNumber="13">Fig. 55</figureCitation>
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); venter with metasternum dark brown; basal abdominal ventrite brownish yellow medially. Head without punctures; pronotal punctures small, separated by a diameter or less; elytral punctures larger than on pronotum, separated by a diameter or less, becoming gradually larger toward lateral margin; prosternal punctures large, indistinct, separated by about a diameter; mesosternal punctures larger than on prosternum, separated by less than a diameter, contiguous, metasternal punctures large, separated by two or three times a diameter medially, entirely absent in lateral 1/3; basal abdominal ventrite with coarse punctures separated by about a diameter medially, lateral 1/3 of ventrite lacking punctation, ventrites 2–4 finely punctured, punctures separated by about a diameter, ventrite 5 with dense, fine punctation. Head with frons parallel-sided, medially depressed, three times as wide as eye, apex truncate medially, lateral 1/8 slightly retracted, angle acute (
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<figureCitation id="5ECD2A03C75095298C0CD1C522DAFC68" box="[348,435,969,993]" captionStart="Figures 55–72" captionStartId="28.[192,287,1816,1838]" captionTargetBox="[194,1426,307,1678]" captionTargetId="figure@28.[192,1438,292,1723]" captionTargetPageId="28" captionText="Figures 55–72. Cryptognatha spp. 55–61) Cryptognatha karla. 55) Habitus. 56) Frons. 57) Protibia. 58) Phallobase ventral. 59) Phallobase lateral. 60) Penis. 61) Spermathecal capsule. 62–66) Cryptognatha clarki. 62) Habitus. 63) Frons. 64) Phallobase ventral. 65) Phallobase lateral. 66) Penis. 67–72) Cryptognatha celia. 67) Habitus. 68) Frons. 69) Spermathecal capsule. 70) Phallobase ventral. 71) Phallobase lateral. 72) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674863" httpUri="https://zenodo.org/record/3674863/files/figure.png" pageId="14" pageNumber="13">Fig. 56</figureCitation>
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); eye canthus long, about 3/4 width of eye. Prosternum with lateral carina on each side extended from apex of intercoxal process 3/4 distance to apex of prosternum, strongly curved laterally. Epipleuron moderately descending externally. Postcoxal line on basal abdominal ventrite evenly curved, extended 3/4 distance to rear margin of ventrite. Genitalia short, penis guide about 3/4 as long as paramere, oval, with apical 1/8 narrowed to abruptly rounded apex; paramere long, slender, apex rounded (
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<figureCitation id="5ECD2A03C75095298C67D66522D7FB08" box="[311,446,1129,1153]" captionStart="Figures 55–72" captionStartId="28.[192,287,1816,1838]" captionTargetBox="[194,1426,307,1678]" captionTargetId="figure@28.[192,1438,292,1723]" captionTargetPageId="28" captionText="Figures 55–72. Cryptognatha spp. 55–61) Cryptognatha karla. 55) Habitus. 56) Frons. 57) Protibia. 58) Phallobase ventral. 59) Phallobase lateral. 60) Penis. 61) Spermathecal capsule. 62–66) Cryptognatha clarki. 62) Habitus. 63) Frons. 64) Phallobase ventral. 65) Phallobase lateral. 66) Penis. 67–72) Cryptognatha celia. 67) Habitus. 68) Frons. 69) Spermathecal capsule. 70) Phallobase ventral. 71) Phallobase lateral. 72) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674863" httpUri="https://zenodo.org/record/3674863/files/figure.png" pageId="14" pageNumber="13">Fig. 58, 59</figureCitation>
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); penis long, robust, apex shortly attenuate; penis capsule with inner arm long, tapered to abrupt apex, outer arm elongate, large (
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<figureCitation id="5ECD2A03C75095298E7DD68520ECFB28" box="[813,901,1161,1185]" captionStart="Figures 55–72" captionStartId="28.[192,287,1816,1838]" captionTargetBox="[194,1426,307,1678]" captionTargetId="figure@28.[192,1438,292,1723]" captionTargetPageId="28" captionText="Figures 55–72. Cryptognatha spp. 55–61) Cryptognatha karla. 55) Habitus. 56) Frons. 57) Protibia. 58) Phallobase ventral. 59) Phallobase lateral. 60) Penis. 61) Spermathecal capsule. 62–66) Cryptognatha clarki. 62) Habitus. 63) Frons. 64) Phallobase ventral. 65) Phallobase lateral. 66) Penis. 67–72) Cryptognatha celia. 67) Habitus. 68) Frons. 69) Spermathecal capsule. 70) Phallobase ventral. 71) Phallobase lateral. 72) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674863" httpUri="https://zenodo.org/record/3674863/files/figure.png" pageId="14" pageNumber="13">Fig. 60</figureCitation>
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).
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</paragraph>
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<paragraph id="C6493686C75095298D90D6B5210AFA98" blockId="14.[192,1441,1209,1297]" pageId="14" pageNumber="13">
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<emphasis id="F482EA94C75095298D90D6B52245FB58" bold="true" box="[192,300,1209,1233]" pageId="14" pageNumber="13">Female.</emphasis>
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Similar to male except head weakly depressed; genitalia with spermathecal capsule short, wide, tapered from wide ramus to abruptly rounded apex, portion of heavily sclerotized sperm duct present at base of ramus (
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<figureCitation id="5ECD2A03C75095298CAED6F5213CFA98" box="[510,597,1273,1297]" captionStart="Figures 55–72" captionStartId="28.[192,287,1816,1838]" captionTargetBox="[194,1426,307,1678]" captionTargetId="figure@28.[192,1438,292,1723]" captionTargetPageId="28" captionText="Figures 55–72. Cryptognatha spp. 55–61) Cryptognatha karla. 55) Habitus. 56) Frons. 57) Protibia. 58) Phallobase ventral. 59) Phallobase lateral. 60) Penis. 61) Spermathecal capsule. 62–66) Cryptognatha clarki. 62) Habitus. 63) Frons. 64) Phallobase ventral. 65) Phallobase lateral. 66) Penis. 67–72) Cryptognatha celia. 67) Habitus. 68) Frons. 69) Spermathecal capsule. 70) Phallobase ventral. 71) Phallobase lateral. 72) Penis." figureDoi="http://doi.org/10.5281/zenodo.3674863" httpUri="https://zenodo.org/record/3674863/files/figure.png" pageId="14" pageNumber="13">Fig. 61</figureCitation>
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).
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</paragraph>
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<paragraph id="C6493686C75095298D90D7252127FAE8" blockId="14.[192,1440,1321,1377]" pageId="14" pageNumber="13">
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<emphasis id="F482EA94C75095298D90D7252222FAC8" bold="true" box="[192,331,1321,1345]" pageId="14" pageNumber="13">Variation.</emphasis>
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Length
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<quantity id="010E9B63C75095298CE1D7262132FAC8" box="[433,603,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.1" metricValueMin="1.9" pageId="14" pageNumber="13" unit="mm" value="2.5" valueMax="3.1" valueMin="1.9">1.9 to 3.1 mm</quantity>
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, width to
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<quantity id="010E9B63C75095298F83D7262014FAC8" box="[723,893,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="14" pageNumber="13" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7 to 1.8 mm</quantity>
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. Pale areas on elytron may be slightly larger or smaller than described above.
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</paragraph>
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</subSubSection>
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<subSubSection id="8EEC650DC75095298D90D77520DFFA3B" pageId="14" pageNumber="13" type="materials_examined">
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<paragraph id="C6493686C75095298D90D77520DFFA3B" blockId="14.[192,1440,1401,1458]" pageId="14" pageNumber="13">
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<emphasis id="F482EA94C75095298D90D77522E3FA18" bold="true" box="[192,394,1401,1425]" pageId="14" pageNumber="13">Type material.</emphasis>
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<materialsCitation id="769E3CDBC75095298CC2D7752679FA1B" ID-GBIF-Occurrence="2569343952" box="[402,1296,1401,1426]" collectingDate="2001-01-22" collectionCode="INBP" country="Paraguay" location="Caaguazu" pageId="14" pageNumber="13" specimenCount="1" stateProvince="Caaguazu" typeStatus="holotype">
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<typeStatus id="194D8824C75095298CC2D7752169FA18" box="[402,512,1401,1425]" pageId="14" pageNumber="13" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="BEE17616C75095298F05D776219DFA1B" box="[597,756,1402,1426]" name="Paraguay" pageId="14" pageNumber="13">PARAGUAY</collectingCountry>
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,
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<collectingRegion id="0432F864C75095298E50D7762017FA1B" box="[768,894,1402,1426]" country="Paraguay" name="Caaguazu" pageId="14" pageNumber="13">Caaguazú</collectingRegion>
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, Repatriación.
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<date id="B2481046C75095298968D77627C4FA1B" box="[1080,1197,1402,1426]" pageId="14" pageNumber="13" value="2001-01-22">
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<collectingDate id="A20CE9AEC75095298968D77627C4FA1B" box="[1080,1197,1402,1426]" pageId="14" pageNumber="13" value="2001-01-22">22-I-2001</collectingDate>
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</date>
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(
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<collectionCode id="A0E7AE43C750952989EED776266EFA1B" box="[1214,1287,1402,1426]" country="Paraguay" httpUri="http://grbio.org/cool/nb6b-jwsh" name="Inventorio Biologico Nacional [Museo Nacional de Historia Natural del Paraguay]" pageId="14" pageNumber="13">INBP</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="769E3CDBC7509529884CD77620DFFA3B" ID-GBIF-Occurrence="2569343954" pageId="14" pageNumber="13" typeStatus="paratype">
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<typeStatus id="194D8824C7509529884CD77626F5FA1B" box="[1308,1436,1402,1426]" pageId="14" pageNumber="13" type="paratype">Paratypes</typeStatus>
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: 10, Chapada, Brazil, Acc.No. 2966, Aug., Nov., Sept. (
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<collectionCode id="A0E7AE43C75095298E1DD79620C3FA3B" box="[845,938,1434,1458]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="14" pageNumber="13" type="Museum">CMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8EEC650DC75095298D90D7C5265AF9A8" pageId="14" pageNumber="13" type="discussion">
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<paragraph id="C6493686C75095298D90D7C5265AF9A8" blockId="14.[192,1440,1481,1569]" pageId="14" pageNumber="13">
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<emphasis id="F482EA94C75095298D90D7C5222AFA68" bold="true" box="[192,323,1481,1505]" pageId="14" pageNumber="13">Remarks.</emphasis>
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Typical specimens of
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<taxonomicName id="01F64D05C75095298F03D7C5202FFA68" authorityName="Gonzalez and Hanley" authorityYear="2019" box="[595,838,1481,1505]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="karla">
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<emphasis id="F482EA94C75095298F03D7C5202FFA68" box="[595,838,1481,1505]" italics="true" pageId="14" pageNumber="13">Cryptognatha karla</emphasis>
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</taxonomicName>
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are recognized by the dark humeral macula and dark brown metasternum, those specimens lacking a dark macula on elytron are difficult to recognize but they also have a darkened metasternum, which is distinct on an otherwise yellow venter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |