258 lines
26 KiB
XML
258 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1118.89593" ID-Pensoft-Pub="1313-2970-1118-1" ID-Pensoft-UUID="BAC22F9CBE275EE788FFC573D610E645" ID-ZooBank="B1B42B9E1CCA4E85A22E39710CFA755E" ModsDocID="1313-2970-1118-1" checkinTime="1660711579775" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Sittichaya, Wisut" docDate="2022" docId="4E5486C31E6D5230A5A310AB0C51AEF0" docLanguage="en" docName="ZooKeys 1118: 1-19" docOrigin="ZooKeys 1118" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/zookeys.1118.89593" docTitle="Malayorthomorpha hulutbeeda Likhitrakarn & Golovatch & Sittichaya 2022, sp. nov." docType="treatment" docUuid="49D76123-EB7C-4682-BD8F-48881036EDDC" docUuidSource="ZooBank" docVersion="1" id="BAC22F9CBE275EE788FFC573D610E645" lastPageNumber="1" masterDocId="BAC22F9CBE275EE788FFC573D610E645" masterDocTitle="Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1660711880571" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1306-317X</mods:nameIdentifier>
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<mods:affiliation>Program of Agriculture, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sittichaya, Wisut</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6200-1285</mods:nameIdentifier>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">wisut.s@psu.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1118</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>19</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1118.89593</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1118.89593</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1118-1</mods:identifier>
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<mods:identifier type="ZooBank">B1B42B9E1CCA4E85A22E39710CFA755E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BAC22F9CBE275EE788FFC573D610E645</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="1" type="taxonomy">
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<treatment LSID="urn:lsid:zoobank.org:act:49D76123-EB7C-4682-BD8F-48881036EDDC" httpUri="http://treatment.plazi.org/id/4E5486C31E6D5230A5A310AB0C51AEF0" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="https://zoobank.org/49D76123-EB7C-4682-BD8F-48881036EDDC" authority="Likhitrakarn & Golovatch & Sittichaya, 2022" authorityName="Likhitrakarn & Golovatch & Sittichaya" authorityYear="2022" class="Diplopoda" family="Paradoxosomatidae" genus="Malayorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malayorthomorpha hulutbeeda" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hulutbeeda" status="sp. nov.">Malayorthomorpha hulutbeeda</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">Figs 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A-D lateral, mesal, oral and caudal views, respectively E sternal cones between coxae 4, subcaudal view. Abbreviations: ll lamina lateralis, lm lamina medialis, sl solenomere, sph solenophore. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1118.89593.figure7" httpUri="https://binary.pensoft.net/fig/729949" pageId="0" pageNumber="1">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A, B mesal and lateral views, respectively C-F submesal, subsuperior, oral and subcaudal views, respectively. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1118.89593.figure8" httpUri="https://binary.pensoft.net/fig/729950" pageId="0" pageNumber="1">, 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2022-05-25" collectorName="Wisut Sittichaya" country="Thailand" elevation="1430" location="Malaya Phytochorion province" municipality="Betong District" specimenCount="♂" stateProvince="Yala Province" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Thailand">Thailand</collectingCountry>
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-
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<collectingRegion country="Thailand" name="Yala">Yala Province</collectingRegion>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingMunicipality>Betong District</collectingMunicipality>
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, elfin montane forest (
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<location LSID="urn:lsid:plazi:treatment:4E5486C31E6D5230A5A310AB0C51AEF0:DF2EE488ED28A6EA764806D069607D7D" country="Thailand" municipality="Betong District" name="Malaya Phytochorion province" stateProvince="Yala Province">Malaya Phytochorion province</location>
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);
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.43" unit="m" value="1430.0">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.43" unit="m" value="1430.0">1430 m</quantity>
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a.s.l.
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</elevation>
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;
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<collectingDate value="2022-05-25">25 May 2022</collectingDate>
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;
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<collectorName>Wisut Sittichaya</collectorName>
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leg.; CUMZ.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">This new species is distinguished from its two congeners in sternal process between male coxae 4 linguiform with a rounded tip, and lamina lateralis of gonopodal solenophore triangular, apically bifid and protruded laterally.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Length of holotype 31.5 mm, width of midbody pro- and metazonae 2.7 and 3.0 mm, respectively.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Colouration of alcohol material after one week of preservation dark red brown (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6A-F</figureCitation>
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); paraterga paler, head and antennae light brown to brown (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6A, B</figureCitation>
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), legs and venter contrasting light yellow to brown (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6</figureCitation>
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), antennae and legs increasingly darker brown distally (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6B, E, G</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Sittichaya" authorityYear="2022" class="Diplopoda" family="Paradoxosomatidae" genus="Malayorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malayorthomorpha hulutbeeda" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hulutbeeda">
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<emphasis italics="true" pageId="0" pageNumber="1">Malayorthomorpha hulutbeeda</emphasis>
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</taxonomicName>
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sp. nov., ♂ holotype
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<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
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anterior part of body, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
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segments 10 and 11, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="1">E-G</emphasis>
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posterior part of body, lateral, dorsal and subventral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="1">H, I</emphasis>
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sternal cones between coxae 4, subcaudal and sublateral views, respectively
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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All characters as in
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<taxonomicName genus="M." lsidName="M. halabala" pageId="0" pageNumber="1" rank="species" species="halabala">
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<emphasis italics="true" pageId="0" pageNumber="1">M. halabala</emphasis>
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</taxonomicName>
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sp. nov., except as follows.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Antennae rather long, extending caudally past metaterga 4 when stretched dorsally. Collum with three transverse rows of setae: 4+4 in anterior, 2+2 in intermediate, and 3+3 in posterior row; with a small lateral setigerous incision near midway (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6A, B</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Paraterga 2 broad, anterior edge angular and rounded, lateral edge with a small notch at about 1/4 in front of caudal corner (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6A</figureCitation>
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). Calluses on paraterga rather narrow, delimited by a sulcus fully on dorsal side and in posterior half on ventral side; on poreless rings narrower than on pore-bearing ones in dorsal view (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6B, D, E</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Transverse sulcus distinct (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6A, C, F</figureCitation>
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), complete on metaterga 5-17, narrow, line-shaped, rather deep, not reaching the bases of paraterga, smooth at bottom, incomplete and nearly wanting on ring 18. Stricture between pro- and metazona wide, deep, beaded at bottom down to base of paraterga starting with segment 5 (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6A-F</figureCitation>
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). Pleurosternal carinae complete crests with a sharp caudal tooth on rings 2-4, increasingly reduced and retaining a sharp caudal tooth on rings 5 and 6 thereafter, retaining a small caudal tooth on ring 7, missing further on (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6B, D, E</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Hypoproct roundly subtriangular, setigerous knobs at caudal edge very small and well-separated (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6G</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Sterna moderately setose, shining, cross-impressions shallow, without modifications; an entire, large, linguiform, sternal lobe between ♂ coxae 4, with a pair of small denticles laterally near base (Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, dorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively" figureDoi="10.3897/zookeys.1118.89593.figure6" httpUri="https://binary.pensoft.net/fig/729948" pageId="0" pageNumber="1">6H, I</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A-D lateral, mesal, oral and caudal views, respectively E sternal cones between coxae 4, subcaudal view. Abbreviations: ll lamina lateralis, lm lamina medialis, sl solenomere, sph solenophore. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1118.89593.figure7" httpUri="https://binary.pensoft.net/fig/729949" pageId="0" pageNumber="1">7E</figureCitation>
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). An inconspicuous and low ridge present in front of gonopod aperture. Legs long and slender, midbody ones ca. 1.6-1.9 times as long as body height, without modifications, ♂ tarsal brushes absent.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Gonopods (Figs
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A-D lateral, mesal, oral and caudal views, respectively E sternal cones between coxae 4, subcaudal view. Abbreviations: ll lamina lateralis, lm lamina medialis, sl solenomere, sph solenophore. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1118.89593.figure7" httpUri="https://binary.pensoft.net/fig/729949" pageId="0" pageNumber="1">7A-D</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A, B mesal and lateral views, respectively C-F submesal, subsuperior, oral and subcaudal views, respectively. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1118.89593.figure8" httpUri="https://binary.pensoft.net/fig/729950" pageId="0" pageNumber="1">8</figureCitation>
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) rather simple; coxa almost straight caudad, densely setose distoventrally. Prefemur as usual, densely setose, about 1/3 as long as femorite + postfemoral part. Femorite stout, suberect, showing a distinct mesal groove/hollow (
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<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
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), with a sulcus demarcating a postfemoral part; seminal groove running entirely mesally along fermorite, solenomere (
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<emphasis bold="true" pageId="0" pageNumber="1">sl</emphasis>
|
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) flagelliform, almost fully sheathed by solenophore (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">sph</emphasis>
|
||
). Lamina medialis (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">lm</emphasis>
|
||
) well developed, thick and large, unciform, terminal lobe sheathing the tip of solenomere. Lamina lateralis (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">ll</emphasis>
|
||
) triangular in shape, protruding laterally, tapered apically, bifid at tip (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A-D lateral, mesal, oral and caudal views, respectively E sternal cones between coxae 4, subcaudal view. Abbreviations: ll lamina lateralis, lm lamina medialis, sl solenomere, sph solenophore. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1118.89593.figure7" httpUri="https://binary.pensoft.net/fig/729949" pageId="0" pageNumber="1">7C, D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A, B mesal and lateral views, respectively C-F submesal, subsuperior, oral and subcaudal views, respectively. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1118.89593.figure8" httpUri="https://binary.pensoft.net/fig/729950" pageId="0" pageNumber="1">8A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Malayorthomorpha hulutbeeda sp. nov., ♂ holotype, right gonopod A, B mesal and lateral views, respectively C-F submesal, subsuperior, oral and subcaudal views, respectively. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1118.89593.figure8" httpUri="https://binary.pensoft.net/fig/729950" pageId="0" pageNumber="1">8C-D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1118.89593.figure7" httpUri="https://binary.pensoft.net/fig/729949" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
|
||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Sittichaya" authorityYear="2022" class="Diplopoda" family="Paradoxosomatidae" genus="Malayorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malayorthomorpha hulutbeeda" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hulutbeeda">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Malayorthomorpha hulutbeeda</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ holotype, right gonopod
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
|
||
lateral, mesal, oral and caudal views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
sternal cones between coxae 4, subcaudal view. Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="1">ll</emphasis>
|
||
lamina lateralis,
|
||
<emphasis bold="true" pageId="0" pageNumber="1">lm</emphasis>
|
||
lamina medialis,
|
||
<emphasis bold="true" pageId="0" pageNumber="1">sl</emphasis>
|
||
solenomere,
|
||
<emphasis bold="true" pageId="0" pageNumber="1">sph</emphasis>
|
||
solenophore. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1118.89593.figure8" httpUri="https://binary.pensoft.net/fig/729950" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
|
||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Sittichaya" authorityYear="2022" class="Diplopoda" family="Paradoxosomatidae" genus="Malayorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malayorthomorpha hulutbeeda" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hulutbeeda">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Malayorthomorpha hulutbeeda</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ holotype, right gonopod
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
mesal and lateral views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C-F</emphasis>
|
||
submesal, subsuperior, oral and subcaudal views, respectively. Scale bars: 0.2 mm
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
To emphasize "
|
||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Sittichaya" authorityYear="2022" class="Diplopoda" family="Paradoxosomatidae" genus="Malayorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malayorthomorpha hulutbeeda" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hulutbeeda">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Malayorthomorpha hulutbeeda</emphasis>
|
||
</taxonomicName>
|
||
" which means "flat-back millipede" in Malay dialect, a noun in apposition. A Malay dialect language is mainly used in three provinces of southern Thailand where the holotype was obtained.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remark.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species was found living together with
|
||
<taxonomicName genus="M." lsidName="M. halabala" pageId="0" pageNumber="1" rank="species" species="halabala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. halabala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Moreover, according to our observations, they may even occur syntopically, sharing the same habitat: leaf litter surface, branches of trees and tree trunks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |