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<document ID-DOI="10.5252/zoosystema2023v45a3" ID-GBIF-Dataset="bc090a4d-4bce-42e0-bf6b-fb4bd45ca811" ID-ISSN="1638-9387" ID-Zenodo-Dep="7681295" ID-ZooBank="urn:lsid:zoobank.org:pub:62C2AC78-741C-4675-8C3A-67E359EB369E" checkinTime="1677503147187" checkinUser="felipe" docAuthor="Marchán, Daniel Fernández, Domínguez, Jorge, Hedde, Mickaël &amp; Decaëns, Thibaud" docDate="2023" docId="F03D0809FFC87451FE89C2AFA77BFB56" docLanguage="en" docName="Zoosystema.45.3.107-128.pdf" docOrigin="Zoosystema 45 (3)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.3:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="3" docTitle="Scherotheca qiui Marchan, Decaens &amp; Dominguez 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="117" masterDocId="0C047071FFC4745DFFE4C278A344FF98" masterDocTitle="The cradle of giants: insights into the origin of Scherotheca Bouché, 1972 (Lumbricidae, Crassiclitellata) with the descriptions of eight new species from Corsica, France" masterLastPageNumber="128" masterPageNumber="107" pageNumber="117" updateTime="1677510542406" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The cradle of giants: insights into the origin of Scherotheca Bouché, 1972 (Lumbricidae, Crassiclitellata) with the descriptions of eight new species from Corsica, France</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marchán, Daniel Fernández</mods:namePart>
<mods:affiliation>CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) danifermch @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">danifermch@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Domínguez, Jorge</mods:namePart>
<mods:affiliation>Universidade de Vigo, Grupo de Ecoloxía Animal (GEA), E- 36310 Vigo (Spain)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hedde, Mickaël</mods:namePart>
<mods:affiliation>Eco &amp; Sols, INRAE, IRD, CIRAD, SupAgro Montpellier, Campus SupAgro, Bâtiment 12, 2 place Viala, F- 34060 Montpellier cedex 2 (France) mickael. hedde @ inrae. fr</mods:affiliation>
<mods:nameIdentifier type="email">hedde@inrae.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Decaëns, Thibaud</mods:namePart>
<mods:affiliation>CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) thibaud. decaens @ cefe. cnrs. fr</mods:affiliation>
<mods:nameIdentifier type="email">decaens@cefe.cnrs.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoosystema</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>45</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>107</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/zoosystema2023v45a3</mods:identifier>
<mods:identifier type="GBIF-Dataset">bc090a4d-4bce-42e0-bf6b-fb4bd45ca811</mods:identifier>
<mods:identifier type="ISSN">1638-9387</mods:identifier>
<mods:identifier type="Zenodo-Dep">7681295</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:62C2AC78-741C-4675-8C3A-67E359EB369E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7681890" ID-Zenodo-Dep="7681890" LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56" httpUri="http://treatment.plazi.org/id/F03D0809FFC87451FE89C2AFA77BFB56" lastPageNumber="117" pageId="12" pageNumber="117">
<subSubSection pageId="12" pageNumber="117" type="nomenclature">
<paragraph blockId="12.[234,673,215,305]" pageId="12" pageNumber="117">
<heading centered="true" fontSize="11" level="2" pageId="12" pageNumber="117" reason="3">
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="qiui" status="n. sp.">
<emphasis bold="true" box="[365,542,215,242]" italics="true" pageId="12" pageNumber="117">Scherotheca qiui</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[614,673,248,274]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="description">
<paragraph blockId="12.[234,673,215,305]" box="[406,501,279,305]" pageId="12" pageNumber="117">
(
<figureCitation box="[413,494,279,305]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens &amp; Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens &amp; Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens &amp; Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens &amp; Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens &amp; Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens &amp; Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens &amp; Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens &amp; Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="12" pageNumber="117">Fig. 3E</figureCitation>
)
</paragraph>
<paragraph blockId="12.[161,745,339,357]" box="[161,745,339,357]" pageId="12" pageNumber="117">
<uri box="[161,745,339,357]" pageId="12" pageNumber="117">
urn:lsid:zoobank.org:act:
<uuid box="[364,745,339,357]" pageId="12" pageNumber="117">78F07B1F-8F87-4BC8-A454-5E7E3096EC24</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="materials_examined">
<paragraph blockId="12.[131,777,390,1129]" pageId="12" pageNumber="117">
<collectionCode box="[132,288,391,414]" pageId="12" pageNumber="117">TYPE MATERIAL</collectionCode>
. —
<materialsCitation collectingDate="2021-04-04" collectionCode="MNHN" collectorName="T. Decaens &amp; D. Fernandez Marchan" country="France" elevation="785" latitude="41.74278" location="Zonza" longLatPrecision="19" longitude="9.195" pageId="12" pageNumber="117" specimenCode="DFM-0235" specimenCount="1" specimenCount-adult="1" stateProvince="Corse du Sud" typeStatus="holotype">
<emphasis bold="true" box="[338,529,390,416]" pageId="12" pageNumber="117">
<typeStatus box="[338,438,390,414]" pageId="12" pageNumber="117">Holotype</typeStatus>
.
<collectingCountry box="[451,529,392,416]" name="France" pageId="12" pageNumber="117">Corsica</collectingCountry>
</emphasis>
•
<specimenCount box="[555,615,390,414]" pageId="12" pageNumber="117" type="adult">Adult</specimenCount>
;
<collectingRegion box="[628,771,390,414]" pageId="12" pageNumber="117">Corse du Sud</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FF62C3D9A381FE20" box="[134,197,417,440]" country="France" latitude="41.74278" longLatPrecision="19" longitude="9.195" name="Zonza" pageId="12" pageNumber="117" stateProvince="Corse du Sud">Zonza</location>
;
<geoCoordinate box="[203,323,417,441]" degrees="41" direction="north" minutes="44" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="34" value="41.74278">41°44’34”N</geoCoordinate>
,
<geoCoordinate box="[330,431,417,441]" degrees="9" direction="east" minutes="11" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="42" value="9.195">9°11’42”E</geoCoordinate>
;
<elevation box="[441,550,417,441]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="12" pageNumber="117" unit="m" value="785.0">
<quantity box="[441,502,417,441]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="12" pageNumber="117" unit="m" value="785.0">785 m</quantity>
a.s.l.
</elevation>
;
<date box="[559,658,417,441]" pageId="12" pageNumber="117" value="2021-04-04">
<collectingDate box="[559,658,417,441]" pageId="12" pageNumber="117" value="2021-04-04">4.IV.2021</collectingDate>
</date>
;
<collectorName box="[665,772,417,440]" pageId="12" pageNumber="117">T. Decaëns</collectorName>
,
<collectorName box="[131,343,443,467]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.; BOLD Sample ID:
<specimenCode box="[567,679,443,467]" pageId="12" pageNumber="117">DFM-0235</specimenCode>
;
<collectionCode box="[687,770,444,467]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph blockId="12.[131,777,390,1129]" pageId="12" pageNumber="117">
<materialsCitation collectingDate="2021-04-06" collectionCode="MNHN" collectorName="T. Decaens &amp; D. Fernandez Marchan" country="France" elevation="1178" latitude="41.836945" location="Olivese (D 69 road)" longLatPrecision="19" longitude="9.095833" municipality="Corse du Sud" pageId="12" pageNumber="117" specimenCode="DFM-0273" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[132,317,470,495]" pageId="12" pageNumber="117">
<typeStatus box="[132,230,470,494]" pageId="12" pageNumber="117">Paratypes</typeStatus>
.
<collectingCountry box="[242,317,471,495]" name="France" pageId="12" pageNumber="117">Corsica</collectingCountry>
</emphasis>
•
<specimenCount box="[343,542,470,494]" pageId="12" pageNumber="117">1 juvenile specimen</specimenCount>
;
<collectingMunicipality box="[553,691,470,494]" pageId="12" pageNumber="117">Corse du Sud</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FD5BC3AEA3BEFD90" country="France" latitude="41.836945" longLatPrecision="19" longitude="9.095833" municipality="Corse du Sud" name="Olivese (D 69 road)" pageId="12" pageNumber="117">Olivese (D69 road)</location>
;
<geoCoordinate box="[261,384,496,520]" degrees="41" direction="north" minutes="50" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="13" value="41.836945">41°50’13”N</geoCoordinate>
,
<geoCoordinate box="[396,485,496,520]" degrees="9" direction="east" minutes="5" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="45" value="9.095833">9°5’45”E</geoCoordinate>
;
<elevation box="[500,632,496,520]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="12" pageNumber="117" unit="m" value="1178.0">
<quantity box="[500,578,496,520]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="12" pageNumber="117" unit="m" value="1178.0">1178 m</quantity>
a.s.l.
</elevation>
;
<date box="[644,746,496,520]" pageId="12" pageNumber="117" value="2021-04-06">
<collectingDate box="[644,746,496,520]" pageId="12" pageNumber="117" value="2021-04-06">6.IV.2021</collectingDate>
</date>
;
<collectorName pageId="12" pageNumber="117">T. Decaëns</collectorName>
,
<collectorName box="[229,459,523,547]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.; BOLD Sample ID:
<specimenCode pageId="12" pageNumber="117">DFM-0273</specimenCode>
;
<collectionCode box="[196,279,550,573]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-03" collectionCode="T, BOLD, MNHN" collectorName="T. Decaens &amp; D. Fernandez Marchan" country="France" elevation="1248" latitude="41.87583" location="Ponte di Valpine" longLatPrecision="19" longitude="9.132778" pageId="12" pageNumber="117" specimenCount="2" specimenCount-adult="2" typeStatus="paratype">
<specimenCount box="[303,373,549,573]" pageId="12" pageNumber="117" type="adult">2 adult</specimenCount>
specimens;
<collectingCountry box="[492,549,549,573]" name="France" pageId="12" pageNumber="117">Corse</collectingCountry>
du Sud,
<collectorName box="[640,709,549,573]" pageId="12" pageNumber="117">Zicavo</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FD2BC05EA3AEFDCF" country="France" latitude="41.87583" longLatPrecision="19" longitude="9.132778" name="Ponte di Valpine" pageId="12" pageNumber="117">Ponte di Valpine</location>
;
<geoCoordinate box="[241,362,575,599]" degrees="41" direction="north" minutes="52" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="33" value="41.87583">41°52’33”N</geoCoordinate>
,
<geoCoordinate box="[371,460,575,599]" degrees="9" direction="east" minutes="7" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="58" value="9.132778">9°7’58”E</geoCoordinate>
;
<elevation box="[471,595,575,600]" metricMagnitude="3" metricUnit="m" metricValue="1.248" pageId="12" pageNumber="117" unit="m" value="1248.0">
<quantity box="[471,546,576,600]" metricMagnitude="3" metricUnit="m" metricValue="1.248" pageId="12" pageNumber="117" unit="m" value="1248.0">1248 m</quantity>
a.s.l.
</elevation>
;
<date box="[605,705,576,600]" pageId="12" pageNumber="117" value="2021-04-03">
<collectingDate box="[605,705,576,600]" pageId="12" pageNumber="117" value="2021-04-03">3.IV.2021</collectingDate>
</date>
;
<collectionCode box="[713,730,576,599]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="117">T</collectionCode>
.
<collectorName pageId="12" pageNumber="117">Decaëns</collectorName>
,
<collectorName box="[195,416,602,626]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.;
<collectionCode box="[466,535,602,626]" pageId="12" pageNumber="117">BOLD</collectionCode>
Sample ID: DFM-0441, DFM-0442;
<collectionCode box="[260,343,629,652]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-04" collectionCode="T, BOLD" country="France" county="Sample" elevation="785" latitude="41.74278" location="Sud" longLatPrecision="19" longitude="9.195" municipality="Zonza" pageId="12" pageNumber="117" specimenCount="3" specimenCount-adult="3" typeStatus="paratype">
<specimenCount box="[367,437,628,653]" pageId="12" pageNumber="117" type="adult">3 adult</specimenCount>
specimens;
<collectingCountry box="[557,614,629,653]" name="France" pageId="12" pageNumber="117">Corse</collectingCountry>
du
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FD6AC00CA1F2FD14" box="[654,694,628,652]" country="France" county="Sample" latitude="41.74278" longLatPrecision="19" longitude="9.195" municipality="Zonza" name="Sud" pageId="12" pageNumber="117">Sud</location>
,
<collectingMunicipality box="[705,769,629,652]" pageId="12" pageNumber="117">Zonza</collectingMunicipality>
;
<geoCoordinate box="[133,261,655,679]" degrees="41" direction="north" minutes="44" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="34" value="41.74278">41°44’34”N</geoCoordinate>
,
<geoCoordinate box="[274,381,655,679]" degrees="9" direction="east" minutes="11" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="42" value="9.195">9°11’42”E</geoCoordinate>
;
<elevation box="[397,522,655,679]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="12" pageNumber="117" unit="m" value="785.0">
<quantity box="[397,465,655,679]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="12" pageNumber="117" unit="m" value="785.0">785 m</quantity>
a.s.l.
</elevation>
;
<date box="[536,643,655,679]" pageId="12" pageNumber="117" value="2021-04-04">
<collectingDate box="[536,643,655,679]" pageId="12" pageNumber="117" value="2021-04-04">4.IV.2021</collectingDate>
</date>
;
<collectionCode box="[654,671,655,678]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="117">T</collectionCode>
.
<collectorName box="[684,772,655,678]" pageId="12" pageNumber="117">Decaëns</collectorName>
,
<collectorName box="[134,377,681,705]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.;
<collectionCode box="[439,511,681,705]" pageId="12" pageNumber="117">BOLD</collectionCode>
<collectingCounty box="[520,595,681,705]" pageId="12" pageNumber="117">Sample</collectingCounty>
ID: DFM-0230, DFM-0232, DFM-0233; Eco&amp;Sols
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-04" collectionCode="BOLD, MNHN" country="France" county="Sample" elevation="785" latitude="41.74278" location="Sud" longLatPrecision="19" longitude="9.195" municipality="Zonza" pageId="12" pageNumber="117" specimenCount="5" specimenCount-adult="4" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount box="[528,601,708,732]" pageId="12" pageNumber="117" type="adult">4 adult</specimenCount>
specimens,
<specimenCount pageId="12" pageNumber="117" type="juvenile">1 juvenile</specimenCount>
specimen; same data as for preceding;
<collectionCode box="[583,653,734,758]" pageId="12" pageNumber="117">BOLD</collectionCode>
Sample ID: DFM-0234, DFM-0236, DFM-0237, DFM-0238, DFM-0248;
<collectionCode box="[134,217,788,811]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-04" collectionCode="T, BOLD, MNHN" collectorName="Decaens, T. &amp; D. Fernandez Marchan" country="France" elevation="940" latitude="41.72389" location="Zonza" longLatPrecision="19" longitude="9.202778" municipality="Pacciunituli" pageId="12" pageNumber="117" specimenCount="11" specimenCount-adult="4" specimenCount-juvenile="7" typeStatus="paratype">
<specimenCount box="[239,308,787,811]" pageId="12" pageNumber="117" type="adult">4 adult</specimenCount>
specimens,
<specimenCount box="[425,520,787,811]" pageId="12" pageNumber="117" type="juvenile">7 juvenile</specimenCount>
specimens;
<collectingCountry box="[638,695,787,811]" name="France" pageId="12" pageNumber="117">Corse</collectingCountry>
du Sud,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FF60C156A380FCDD" box="[132,196,814,837]" country="France" latitude="41.72389" longLatPrecision="19" longitude="9.202778" municipality="Pacciunituli" name="Zonza" pageId="12" pageNumber="117">Zonza</location>
,
<collectingMunicipality box="[206,322,814,838]" pageId="12" pageNumber="117">Pacciunituli</collectingMunicipality>
(D368 road);
<geoCoordinate box="[464,586,814,838]" degrees="41" direction="north" minutes="43" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="26" value="41.72389">41°43’26”N</geoCoordinate>
,
<geoCoordinate box="[596,698,814,838]" degrees="9" direction="east" minutes="12" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="10" value="9.202778">9°12’10”E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="12" pageNumber="117" unit="m" value="940.0">
<quantity box="[711,775,814,838]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="12" pageNumber="117" unit="m" value="940.0">940 m</quantity>
a.s.l.
</elevation>
;
<date box="[189,290,840,864]" pageId="12" pageNumber="117" value="2021-04-04">
<collectingDate box="[189,290,840,864]" pageId="12" pageNumber="117" value="2021-04-04">4.IV.2021</collectingDate>
</date>
; T.
<collectorName box="[327,411,841,864]" pageId="12" pageNumber="117">Decaëns</collectorName>
,
<collectorName box="[422,651,840,864]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.;
<collectionCode box="[706,775,840,864]" pageId="12" pageNumber="117">BOLD</collectionCode>
Sample ID: DFM-0159, DFM-0160, DFM-0161, DFM-0162, DFM-0163, DFM-0164, DFM-0165, DFM-0166, DFM-0167, DFM-0168, DFM-0169;
<collectionCode box="[391,474,920,943]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-03" collectionCode="T, BOLD, MNHN" collectorName="Decaens, T. &amp; D. Fernandez Marchan" country="France" elevation="813" latitude="41.735832" location="Zonza" longLatPrecision="19" longitude="9.180834" municipality="Pacciunituli" pageId="12" pageNumber="117" specimenCount="2" specimenCount-adult="2" typeStatus="paratype">
<specimenCount box="[498,568,919,944]" pageId="12" pageNumber="117" type="adult">2 adult</specimenCount>
specimens;
<collectingCountry box="[687,744,920,944]" name="France" pageId="12" pageNumber="117">Corse</collectingCountry>
du Sud,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FF51C1CAA3B0FC51" box="[181,244,946,969]" country="France" latitude="41.735832" longLatPrecision="19" longitude="9.180834" municipality="Pacciunituli" name="Zonza" pageId="12" pageNumber="117">Zonza</location>
,
<collectingMunicipality box="[251,365,946,970]" pageId="12" pageNumber="117">Pacciunituli</collectingMunicipality>
(D67 road);
<geoCoordinate box="[488,595,946,970]" degrees="41" direction="north" minutes="44" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="9" value="41.735832">41°44’9”N</geoCoordinate>
,
<geoCoordinate box="[603,704,946,970]" degrees="9" direction="east" minutes="10" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="51" value="9.180834">9°10’51”E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.129999999999999" pageId="12" pageNumber="117" unit="m" value="813.0">
<quantity box="[714,776,946,970]" metricMagnitude="2" metricUnit="m" metricValue="8.129999999999999" pageId="12" pageNumber="117" unit="m" value="813.0">813 m</quantity>
a.s.l.
</elevation>
;
<date box="[186,300,972,996]" pageId="12" pageNumber="117" value="2021-04-03">
<collectingDate box="[186,300,972,996]" pageId="12" pageNumber="117" value="2021-04-03">03.IV.2021</collectingDate>
</date>
;
<collectionCode box="[308,325,973,996]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="117">T</collectionCode>
.
<collectorName box="[334,418,973,996]" pageId="12" pageNumber="117">Decaëns</collectorName>
,
<collectorName box="[427,653,972,996]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.;
<collectionCode box="[704,773,972,996]" pageId="12" pageNumber="117">BOLD</collectionCode>
Sample ID: DFM-0434, DFM-0435;
<collectionCode box="[494,576,999,1022]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-04" collectionCode="T, BOLD, MNHN" collectorName="Decaens, T. &amp; D. Fernandez Marchan" country="France" elevation="975" latitude="41.761944" location="Zonza" longLatPrecision="19" longitude="9.218888" municipality="Samulaghia" pageId="12" pageNumber="117" specimenCount="5" specimenCount-adult="5" typeStatus="paratype">
<specimenCount box="[600,666,999,1023]" pageId="12" pageNumber="117" type="adult">5 adult</specimenCount>
specimens;
<collectingCountry box="[131,188,1025,1049]" name="France" pageId="12" pageNumber="117">Corse</collectingCountry>
du Sud,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC87451FE89C2AFA77BFB56:7D4BEFC4FFC87451FEF0C67AA210FB81" box="[276,340,1026,1049]" country="France" latitude="41.761944" longLatPrecision="19" longitude="9.218888" municipality="Samulaghia" name="Zonza" pageId="12" pageNumber="117">Zonza</location>
,
<collectingMunicipality box="[349,465,1025,1049]" pageId="12" pageNumber="117">Samulaghia</collectingMunicipality>
;
<geoCoordinate box="[475,597,1025,1049]" degrees="41" direction="north" minutes="45" orientation="latitude" pageId="12" pageNumber="117" precision="15" seconds="43" value="41.761944">41°45’43”N</geoCoordinate>
,
<geoCoordinate box="[607,696,1025,1049]" degrees="9" direction="east" minutes="13" orientation="longitude" pageId="12" pageNumber="117" precision="15" seconds="8" value="9.218888">9°13’8”E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="12" pageNumber="117" unit="m" value="975.0">
<quantity box="[709,773,1025,1049]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="12" pageNumber="117" unit="m" value="975.0">975 m</quantity>
a.s.l.
</elevation>
;
<date box="[189,290,1052,1076]" pageId="12" pageNumber="117" value="2021-04-04">
<collectingDate box="[189,290,1052,1076]" pageId="12" pageNumber="117" value="2021-04-04">4.IV.2021</collectingDate>
</date>
;
<collectionCode box="[300,317,1052,1075]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="117">T</collectionCode>
.
<collectorName box="[327,411,1052,1075]" pageId="12" pageNumber="117">Decaëns</collectorName>
,
<collectorName box="[422,651,1052,1076]" pageId="12" pageNumber="117">D. Fernández Marchán</collectorName>
leg.;
<collectionCode box="[706,775,1052,1076]" pageId="12" pageNumber="117">BOLD</collectionCode>
Sample ID: DFM-0483, DFM-0484, DFM-0485, DFM-0486, DFM-0487);
<collectionCode box="[270,356,1105,1128]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="117">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="etymology">
<paragraph blockId="12.[132,775,1152,1230]" pageId="12" pageNumber="117">
ETYMOLOGY. — This species is named after Prof. Jianping Qiu, in recognition to its significant contribution to the taxonomic knowledge of the genus
<taxonomicName authorityName="Bouche" authorityYear="1972" box="[308,416,1205,1229]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="genus">
<emphasis box="[308,416,1205,1229]" italics="true" pageId="12" pageNumber="117">Scherotheca</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="diagnosis">
<paragraph blockId="12.[132,775,1253,1594]" pageId="12" pageNumber="117">
DIAGNOSIS. —
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[282,771,1253,1277]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="qiui" status="n. sp.">
<emphasis box="[282,434,1253,1277]" italics="true" pageId="12" pageNumber="117">Scherotheca qiui</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[132,183,1280,1303]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
can be distinguished from non-Corsican species of
<taxonomicName authorityName="Bouche" authorityYear="1972" box="[668,775,1279,1303]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="genus">
<emphasis box="[668,775,1279,1303]" italics="true" pageId="12" pageNumber="117">Scherotheca</emphasis>
</taxonomicName>
by the shorter and more anterior clitellum. It resembles
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="litoralis" status="n. sp.">
<emphasis italics="true" pageId="12" pageNumber="117">Scherotheca litoralis</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[560,615,1333,1356]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="mausoleana" status="n. sp.">
<emphasis italics="true" pageId="12" pageNumber="117">Scherotheca mausoleana</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[594,650,1359,1382]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
, from which it differs by the absence of pigmentation, the number of seminal vesicles (2 pairs instead of 4 pairs in its close relatives) and by the shorter
<emphasis box="[210,403,1438,1462]" italics="true" pageId="12" pageNumber="117">tubercula pubertatis</emphasis>
(30-33(
<superScript attach="none" box="[488,497,1438,1455]" fontSize="7" pageId="12" pageNumber="117">1</superScript>
/
<subScript attach="left" box="[505,514,1449,1465]" fontSize="7" pageId="12" pageNumber="117">n</subScript>
34) instead of 30-34(35) (
<tableCitation box="[140,215,1464,1489]" captionStart="TABLE" captionStartId="5.[131,142,219,237]" captionTargetPageId="5" captionText="TABLE 1. — Morphological characters of the newly described species of Corsican Scherotheca Bouché, 1972 and their closest relatives. Length in millimetres; weight in grams; 1ST D.P., position of first dorsal pore. Symbols: *, according to Pop (1947): composite between the different populations included in the original description; **, according to Bouché (1972)." httpUri="http://table.plazi.org/id/2CEBE997FFC17458FF67C2A3A2B6FE82" pageId="12" pageNumber="117" tableUuid="2CEBE997FFC17458FF67C2A3A2B6FE82">Table 1</tableCitation>
;
<tableCitation box="[227,241,1465,1489]" captionStart="TABLE" captionStartId="7.[132,143,219,237]" captionTargetPageId="7" captionText="TABLE 2. — Reproduction-related morphological characters of the newly described species of Corsican Scherotheca Bouché, 1972 and their closest relatives. Abbreviations: Sperm. pores, position of spermathecal pores; S.V., position of seminal vesicles; T.P., position of tubercula pubertatis. Symbols: *, according to Pop (1947): composite between the different populations included in the original description; **, according to Bouché (1972)." httpUri="http://table.plazi.org/id/2CEBE997FFC3745AFF60C2A3A7D6FE82" pageId="12" pageNumber="117" tableUuid="2CEBE997FFC3745AFF60C2A3A7D6FE82">2</tableCitation>
;
<figureCitation box="[253,312,1464,1489]" captionStart="FIG" captionStartId="8.[132,143,1935,1953]" captionTargetBox="[204,1366,211,1864]" captionTargetId="figure-101@8.[165,1136,207,1872]" captionTargetPageId="8" captionText="FIG. 2. — External and internal morphological differences between the newly described species of Corsican Scherotheca Bouché,1972 and their closest relatives. The three different shades represent absent, slight and moderate pigmentation. Open circles represent structures that only appear in some individuals. Structures shown in light and dark shades (genital papillae, clitellum and tubercula pubertatis) represent the maximum and minimum extension of these characters, respectively. Phylogenetic relationships are the same as those shown in Figure 1." figureDoi="http://doi.org/10.5281/zenodo.7681301" httpUri="https://zenodo.org/record/7681301/files/figure.png" pageId="12" pageNumber="117">Figs 2</figureCitation>
;
<figureCitation box="[323,353,1465,1489]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens &amp; Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens &amp; Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens &amp; Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens &amp; Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens &amp; Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens &amp; Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens &amp; Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens &amp; Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="12" pageNumber="117">3E</figureCitation>
).
</paragraph>
<paragraph blockId="12.[132,775,1253,1594]" pageId="12" pageNumber="117">
COI uncorrected average pairwise distances and topology of multilocus molecular phylogenetic trees support the status of
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="qiui" status="n. sp.">
<emphasis box="[691,774,1517,1541]" italics="true" pageId="12" pageNumber="117">Sch. qiui</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[488,544,1544,1567]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
as separate from other morphologically similar species.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="distribution">
<paragraph blockId="12.[130,776,1618,1748]" pageId="12" pageNumber="117">
DISTRIBUTION AND ECOLOGY. —
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="qiui" status="n. sp.">
<emphasis box="[434,576,1618,1642]" italics="true" pageId="12" pageNumber="117">Scherotheca qiui</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[259,314,1645,1668]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
was found in several localities in the region of Zonza and Zicavo,
<collectingCountry box="[316,372,1671,1695]" name="France" pageId="12" pageNumber="117">Corse</collectingCountry>
du Sud (
<figureCitation box="[455,548,1671,1695]" captionStart="FIG" captionStartId="14.[132,143,1613,1631]" captionTargetBox="[220,1354,216,1568]" captionTargetId="figure-223@14.[220,1354,216,1568]" captionTargetPageId="14" captionText="FIG. 4. — Distribution of the genus Scherotheca Bouché, 1972 in Corsica: A, localities documented in this work of the newly described species and other Scherotheca species sampled during this work; B, distribution of Corsican species of Scherotheca and Eumenescolex Qiu &amp; Bouché, 1998 according to Bouché (1972), Qiu &amp; Bouché (1998) and Szederjesi et al. (2021)." figureDoi="http://doi.org/10.5281/zenodo.7681307" httpUri="https://zenodo.org/record/7681307/files/figure.png" pageId="12" pageNumber="117">Figure 4a</figureCitation>
), mainly in
<taxonomicName box="[660,775,1671,1695]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="12" pageNumber="117" phylum="Tracheophyta" rank="species" species="laricio">
<emphasis box="[660,775,1671,1695]" italics="true" pageId="12" pageNumber="117">Pinus laricio</emphasis>
</taxonomicName>
forests at moderate elevations (
<quantity box="[421,566,1697,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.0165" metricValueMax="1.248" metricValueMin="0.7849999999999999" pageId="12" pageNumber="117" unit="m" value="1016.5" valueMax="1248.0" valueMin="785.0">785 to 1248 m</quantity>
), with some individuals found in
<taxonomicName box="[255,392,1724,1748]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="117" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis box="[255,392,1724,1748]" italics="true" pageId="12" pageNumber="117">Fagus sylvatica</emphasis>
</taxonomicName>
forests and shrublands.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="description">
<paragraph blockId="12.[130,778,1778,2028]" box="[132,277,1778,1804]" pageId="12" pageNumber="117">DESCRIPTION</paragraph>
<paragraph blockId="12.[130,778,1778,2028]" box="[132,346,1810,1836]" pageId="12" pageNumber="117">
<emphasis box="[132,346,1810,1836]" italics="true" pageId="12" pageNumber="117">External morphology</emphasis>
</paragraph>
<paragraph blockId="12.[130,778,1778,2028]" lastBlockId="12.[811,1458,215,497]" pageId="12" pageNumber="117">
Body unpigmented.White-beige in fixed specimens (
<figureCitation box="[683,762,1841,1868]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens &amp; Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens &amp; Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens &amp; Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens &amp; Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens &amp; Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens &amp; Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens &amp; Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens &amp; Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="12" pageNumber="117">Fig. 3E</figureCitation>
). Average length,
<quantity box="[309,390,1874,1900]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="117" unit="mm" value="74.0">74 mm</quantity>
(
<quantity box="[407,529,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="7.65" metricValueMax="8.9" metricValueMin="6.4" pageId="12" pageNumber="117" unit="mm" value="76.5" valueMax="89.0" valueMin="64.0">64-89 mm</quantity>
,
<emphasis box="[543,582,1874,1899]" italics="true" pageId="12" pageNumber="117">n =</emphasis>
<specimenCount box="[591,697,1873,1900]" pageId="12" pageNumber="117" type="adult">13 adults</specimenCount>
); body cylindrical in cross-section; average number of segments, 138 (119-152,
<emphasis box="[296,332,1938,1963]" italics="true" pageId="12" pageNumber="117">n =</emphasis>
<specimenCount box="[337,438,1937,1963]" pageId="12" pageNumber="117" type="adult">11 adults</specimenCount>
; 142 segments in the
<typeStatus box="[666,766,1937,1964]" pageId="12" pageNumber="117">holotype</typeStatus>
). Average weight (fixed specimens),
<quantity box="[513,585,1969,1996]" metricMagnitude="-4" metricUnit="kg" metricValue="9.2" pageId="12" pageNumber="117" unit="g" value="0.92">0.92 g</quantity>
(
<quantity box="[601,720,1969,1996]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" metricValueMax="1.4" metricValueMin="0.6" pageId="12" pageNumber="117" unit="g" value="1.0" valueMax="1.4" valueMin="0.6">0.60-1.4 g</quantity>
,
<emphasis box="[736,775,1970,1995]" italics="true" pageId="12" pageNumber="117">n =</emphasis>
<specimenCount box="[132,223,2001,2028]" pageId="12" pageNumber="117" type="adult">2 adults</specimenCount>
). Prostomium epilobous, closed. Longitudinal furrows in segments 1 and 2. First dorsal pore in intersegmental furrow (9/10)10/11. Nephridial pores “en solfège” (irregularly distributed). Spermathecal pores at intersegmental furrows 12/13 and 13/
<quantity box="[973,1031,311,337]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="12" pageNumber="117" unit="in" value="14.0">14 in</quantity>
setal line
<emphasis box="[1142,1151,312,337]" italics="true" pageId="12" pageNumber="117">c</emphasis>
. Male pores in segment 15, surrounded by a well-developed porophore. Female pores in segment 14. Clitellum saddle-shaped in segments (26)27- 35(36).
<emphasis box="[908,1135,406,432]" italics="true" pageId="12" pageNumber="117">Tubercula pubertatis</emphasis>
in segments 30-33(1/
<subScript attach="left" box="[1389,1399,420,436]" fontSize="7" pageId="12" pageNumber="117">n</subScript>
34). Chaetae small and closely paired. Genital papillae in segments 11-14,16, (26)27, 28, (29)34, 35, 36.
</paragraph>
<paragraph blockId="12.[811,1457,534,975]" box="[813,992,534,560]" pageId="12" pageNumber="117">
<emphasis box="[813,992,534,560]" italics="true" pageId="12" pageNumber="117">Internal anatomy</emphasis>
</paragraph>
<paragraph blockId="12.[811,1457,534,975]" pageId="12" pageNumber="117">
<collectingRegion box="[813,871,566,592]" country="Spain" name="Ceuta" pageId="12" pageNumber="117">Septa</collectingRegion>
5/6-10/11 thickened and muscular. Hearts in segments 6-11, oesophageal.Calciferous glands in segments 10-14, with diverticula in segment 10. Crop in segments 15-16, gizzard in segments 17-19. Typhlosole pinnate. Male sexual system holandric, testes and funnels (not enclosed in testes sacs, but with sperm present) located ventrally in segments 10 and 11. Two pairs of reniform seminal vesicles in segments 11 and 12, sometimes with one or two pairs of vestigial seminal vesicles in segments 9 and 10. Ovaries and female funnels in segment 13, ovarian receptacles (ovisacs) in segment 14. Two pairs of small globular spermathecae in segments 12 and 13 (intersegments 12/13, 13/14). Nephridial bladders U-shaped, reclinate in segment 30.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="117" type="discussion">
<paragraph blockId="12.[813,1457,1013,1230]" box="[813,913,1013,1039]" pageId="12" pageNumber="117">REMARKS</paragraph>
<paragraph blockId="12.[813,1457,1013,1230]" pageId="12" pageNumber="117">
This species corresponds to
<taxonomicName authorityName="* (Pop" authorityYear="1947" box="[1114,1245,1045,1071]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="corsicana">
<emphasis box="[1114,1245,1045,1071]" italics="true" pageId="12" pageNumber="117">Sc. corsicana</emphasis>
</taxonomicName>
L
<quantity box="[1270,1312,1044,1070]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="117" unit="in" value="2.0">2 in</quantity>
the checklist of Corsican earthworms of
<bibRefCitation author="MARCHAN D. F. &amp; GERARD S. &amp; HEDDE M. &amp; ROUGERIE R. &amp; DECAENS T." box="[1107,1357,1076,1103]" pageId="12" pageNumber="117" pagination="439 - 461" refId="ref16506" refString="MARCHAN D. F., GERARD S., HEDDE M., ROUGERIE R. &amp; DECAENS T. 2022 a. - An updated checklist and a DNA barcode library for the earthworms (Crassiclitellata, Oligochaeta) of Corsica, France. Zoosystema 44 (17): 439 - 461. https: // doi. org / 10.5252 / zoosystema 2022 v 44 a 17. http: // zoosystema. com / 44 / 17" type="journal article" year="2022">
Marchán
<emphasis box="[1212,1263,1076,1102]" italics="true" pageId="12" pageNumber="117">et al.</emphasis>
(2022a)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="12.[813,1457,1013,1230]" pageId="12" pageNumber="117">
<taxonomicName authority="Marchán &amp; Domínguez &amp; Hedde &amp; Decaëns, 2023" authorityName="Marchán &amp; Domínguez &amp; Hedde &amp; Decaëns" authorityYear="2023" box="[835,1023,1108,1134]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="117" phylum="Annelida" rank="species" species="quiui" status="n. sp.">
<emphasis box="[835,1023,1108,1134]" italics="true" pageId="12" pageNumber="117">Scherotheca quiui</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1032,1094,1109,1135]" pageId="12" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
displayed a level of intraspecific genetic divergence which was slightly above the expected threshold (around 9%), which could indicate the existence of cryptic genetic linages.
</paragraph>
</subSubSection>
</treatment>
</document>
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