209 lines
25 KiB
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209 lines
25 KiB
XML
<document id="8F8B18FD05C7086471CD9CB6F3C316D9" ID-CLB-Dataset="6276" ID-DOI="10.5281/zenodo.4906293" ID-GBIF-Dataset="3228103c-9c27-4ab0-9823-c31bbe817d96" ID-ISSN="1638-9395" ID-Zenodo-Dep="4906293" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623073274397" checkinUser="felipe" docAuthor="Steyer, Jean-Sébastien, Peecook, Brandon R., Arbez, Thomas, Nesbitt, Sterling J., Tolan, Steve, Stocker, Michelle R., Smith, Roger M. H., Angielczyk, Kenneth D. & Sidor, Christian A." docDate="2021" docId="03C8B219277A6A72AD5EFA10E06CD5FF" docLanguage="en" docName="Geodiversitas.43.12.365-376.pdf" docOrigin="Geodiversitas 43 (12)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Stereospondyli Zittel 1888" docType="treatment" docVersion="5" lastPageNumber="369" masterDocId="FFF1CA61277C6A74AC0DFFC3E51AD214" masterDocTitle="New data on the Triassic temnospondyls from the Karoo rift basins of Tanzania and Zambia" masterLastPageNumber="376" masterPageNumber="365" pageNumber="369" updateTime="1699000337765" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="41616D3238AF0D27E9782EC89D5B6F26">New data on the Triassic temnospondyls from the Karoo rift basins of Tanzania and Zambia</mods:title>
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<mods:namePart id="058FA69D19620965F73DF061525B1240">Steyer, Jean-Sébastien</mods:namePart>
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<mods:namePart id="907B32B1A14B5864A5973471EE03421C">Peecook, Brandon R.</mods:namePart>
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<mods:namePart id="01556D24E0513AF06447B7701A13B7AC">Arbez, Thomas</mods:namePart>
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<mods:namePart id="BF0FFD466339526D20F139F392245065">Nesbitt, Sterling J.</mods:namePart>
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<mods:namePart id="DA27F4FC33836CA7BB8A4DC8071E8531">Tolan, Steve</mods:namePart>
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<mods:namePart id="1B6F98C9CC45A0A7F7BE5C1747B24261">Stocker, Michelle R.</mods:namePart>
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<mods:namePart id="3FDB50C984023C816B5C648B2DD6A0A1">Smith, Roger M. H.</mods:namePart>
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<mods:namePart id="E14542B9EB860428E9C1F56DD13CFEE6">Angielczyk, Kenneth D.</mods:namePart>
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<mods:namePart id="8C3D14F4A1C04141EC3FDCC97ED0FD63">Sidor, Christian A.</mods:namePart>
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<mods:title id="82D15A791C9A23F477891290F52FD97B">Geodiversitas</mods:title>
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<mods:date id="193FDA604B28DAE51CC864553815CC9E">2021</mods:date>
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<mods:number id="AC5327ECE6D7F4BEFE1E5BB96621AAB3">2021-06-03</mods:number>
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<mods:number id="82A457C0FFB74A720CCD7721BC3C934A">43</mods:number>
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<treatment id="03C8B219277A6A72AD5EFA10E06CD5FF" ID-DOI="http://doi.org/10.5281/zenodo.4906871" ID-GBIF-Taxon="182597937" ID-Zenodo-Dep="4906871" LSID="urn:lsid:plazi:treatment:03C8B219277A6A72AD5EFA10E06CD5FF" httpUri="http://treatment.plazi.org/id/03C8B219277A6A72AD5EFA10E06CD5FF" lastPageNumber="369" pageId="6" pageNumber="369">
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<subSubSection id="C37B5084277A6A72AD5EFA10E722D7F9" box="[339,568,1491,1517]" pageId="6" pageNumber="369" type="nomenclature">
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<paragraph id="8BDE030F277A6A72AD5EFA10E722D7F9" blockId="6.[339,568,1491,1549]" box="[339,568,1491,1517]" pageId="6" pageNumber="369">
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<taxonomicName id="4C61788C277A6A72AD5EFA10E4E8D7F9" authorityName="Zittel" authorityYear="1888" box="[339,498,1491,1517]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stereospondyli" pageId="6" pageNumber="369" phylum="Chordata" rank="order">Stereospondyli</taxonomicName>
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indet.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B5084277A6A72AD93FA30E4F7D419" box="[414,493,1523,1549]" pageId="6" pageNumber="369" type="description">
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<paragraph id="8BDE030F277A6A72AD93FA30E4F7D419" blockId="6.[339,568,1491,1549]" box="[414,493,1523,1549]" pageId="6" pageNumber="369">
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(
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<figureCitation id="135A1F8A277A6A72ADAAFA30E4FED419" box="[423,484,1523,1549]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B5084277A6A72AC89F9EFE4C0D553" pageId="6" pageNumber="369" type="materials_examined">
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<paragraph id="8BDE030F277A6A72AC89F9EFE70BD4DC" blockId="6.[131,777,1580,1736]" pageId="6" pageNumber="369">
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REFERRED MATERIAL. —
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<materialsCitation id="3B090952277A6A72AD67F9EEE70BD4DC" ID-GBIF-Occurrence="3127475305" pageId="6" pageNumber="369">
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<emphasis id="B915DF1D277A6A72AD67F9EEE4AFD451" bold="true" box="[362,437,1581,1605]" pageId="6" pageNumber="369">
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<collectingCountry id="F376439F277A6A72AD67F9EEE4AFD451" box="[362,437,1581,1605]" name="Zambia" pageId="6" pageNumber="369">Zambia</collectingCountry>
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</emphasis>
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. NHCC LB678 (
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<figureCitation id="135A1F8A277A6A72AE57F9EFE795D450" box="[602,655,1580,1604]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4</figureCitation>
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and
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<bibRefCitation id="EFF07EFE277A6A72AEB5F9EFE5F3D44A" author="PEECOOK B. R. & STEYER J. S. & TABOR N. J. & SMITH R. M. H." pageId="6" pageNumber="369" pagination="8 - 38" refId="ref8849" refString="PEECOOK B. R., STEYER J. S., TABOR N. J. & SMITH R. M. H. 2018. - Updated geology and vertebrate paleontology of the Triassic Ntawere Formation of northeastern Zambia, with special emphasis on the archosauromorphs, in SIDOR C. A. & NESBITT S. J. (eds), Vertebrate and climatic evolution in the Triassic rift basins of Tanzania and Zambia. Society of Vertebrate Paleontology Memoir 17. Journal of Vertebrate Paleontology 37 (6): 8 - 38. https: // doi. org / 10.1080 / 02724634.2017.1410484" type="journal article" year="2018">
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Peecook
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<emphasis id="B915DF1D277A6A72AC89F984E5A8D44A" box="[132,178,1606,1630]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
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2018
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</bibRefCitation>
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: fig. 4A-C), a partial right mandible of a small-sized, but adult, individual based on the well-developed ornamentation visible on its labial side (e.g.,
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<bibRefCitation id="EFF07EFE277A6A72AD65F9B8E4C6D487" author="STEYER J. - S." box="[360,476,1659,1683]" pageId="6" pageNumber="369" pagination="449 - 467" refId="ref9793" refString="STEYER J. - S. 2000. - Ontogeny and phylogeny in temnospondyls: a new method of analysis. Zoological Journal of the Linnean Society 130: 449 - 467. https: // doi. org / 10.1111 / j. 1096 - 3642.2000. tb 01637. x" type="journal article" year="2000">Steyer 2000</bibRefCitation>
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) (
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<figureCitation id="135A1F8A277A6A72ADF9F9B8E758D487" box="[500,578,1659,1683]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4A</figureCitation>
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). The specimen was found by one of us (SJN) in 2014 together with small cylindrical vertebrae similar to NHCC LB690-728.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BDE030F277A6A72AC89F923E4C0D553" blockId="6.[130,776,1759,1863]" pageId="6" pageNumber="369">
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LOCALITY AND HORIZON. —
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<materialsCitation id="3B090952277A6A72AD9AF91CE4C0D553" ID-GBIF-Occurrence="3127475301" pageId="6" pageNumber="369">
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Locality L290 near the village of Sitwe, upper horizon of the Ntawere Formation, Middle-?Late Triassic (e.g.,
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<bibRefCitation id="EFF07EFE277A6A72AC8FF8D7E5E6D539" author="BATTAIL B." box="[130,252,1812,1837]" pageId="6" pageNumber="369" pagination="31 - 35" refId="ref6427" refString="BATTAIL B. 1993. - On the biostratigraphy of Triassic therapsid-bearing formation. New Mexico Museum of Natural History & Science Bulletin 3: 31 - 35." type="journal article" year="1993">Battail 1993</bibRefCitation>
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;
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<bibRefCitation id="EFF07EFE277A6A72AD06F8D6E488D539" author="HANCOX P. J." box="[267,402,1813,1837]" pageId="6" pageNumber="369" pagination="1285 - 1324" refId="ref7534" refString="HANCOX P. J. 2000. - The continental Triassic of South Africa. Zentralblatt fur Geologie und Palaontologie, Teil 1: 1285 - 1324." type="journal article" year="2000">Hancox 2000</bibRefCitation>
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;
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<bibRefCitation id="EFF07EFE277A6A72AD92F8D7E745D539" author="PEECOOK B. R. & STEYER J. S. & TABOR N. J. & SMITH R. M. H." box="[415,607,1812,1837]" pageId="6" pageNumber="369" pagination="8 - 38" refId="ref8849" refString="PEECOOK B. R., STEYER J. S., TABOR N. J. & SMITH R. M. H. 2018. - Updated geology and vertebrate paleontology of the Triassic Ntawere Formation of northeastern Zambia, with special emphasis on the archosauromorphs, in SIDOR C. A. & NESBITT S. J. (eds), Vertebrate and climatic evolution in the Triassic rift basins of Tanzania and Zambia. Society of Vertebrate Paleontology Memoir 17. Journal of Vertebrate Paleontology 37 (6): 8 - 38. https: // doi. org / 10.1080 / 02724634.2017.1410484" type="journal article" year="2018">
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Peecook
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<emphasis id="B915DF1D277A6A72ADFBF8D6E73CD538" box="[502,550,1812,1836]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
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2018
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</bibRefCitation>
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) of the Luangwa Basin (sensu
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<bibRefCitation id="EFF07EFE277A6A72AD0EF8ECE4D6D553" author="BARBOLINI N. & BAMFORD M. K. & TOLAN S." box="[259,460,1839,1863]" pageId="6" pageNumber="369" pagination="18 - 30" refId="ref6271" refString="BARBOLINI N., BAMFORD M. K. & TOLAN S. 2016. - Permo-Triassic palynology and palaeobotany of Zambia; a review. Palaeontologia Africana 50: 18 - 30." type="journal article" year="2016">
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Barbolini
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<emphasis id="B915DF1D277A6A72AD68F8F3E48ED553" box="[357,404,1839,1863]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
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2016
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</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B5084277A6A72AC89F8B1E06CD5FF" pageId="6" pageNumber="369" type="description">
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<paragraph id="8BDE030F277A6A72AC89F8B1E74DD598" blockId="6.[132,775,1905,2028]" box="[132,599,1905,1932]" pageId="6" pageNumber="369">
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DESCRIPTION (see also
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<bibRefCitation id="EFF07EFE277A6A72AD71F8B2E754D59F" author="PEECOOK B. R. & STEYER J. S. & TABOR N. J. & SMITH R. M. H." box="[380,590,1905,1931]" pageId="6" pageNumber="369" pagination="8 - 38" refId="ref8849" refString="PEECOOK B. R., STEYER J. S., TABOR N. J. & SMITH R. M. H. 2018. - Updated geology and vertebrate paleontology of the Triassic Ntawere Formation of northeastern Zambia, with special emphasis on the archosauromorphs, in SIDOR C. A. & NESBITT S. J. (eds), Vertebrate and climatic evolution in the Triassic rift basins of Tanzania and Zambia. Society of Vertebrate Paleontology Memoir 17. Journal of Vertebrate Paleontology 37 (6): 8 - 38. https: // doi. org / 10.1080 / 02724634.2017.1410484" type="journal article" year="2018">
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Peecook
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<emphasis id="B915DF1D277A6A72ADD1F8B1E70AD59F" box="[476,528,1905,1931]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
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2018
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</bibRefCitation>
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)
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</paragraph>
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<paragraph id="8BDE030F277A6A72AC89F852E166D438" blockId="6.[132,775,1905,2028]" lastBlockId="6.[811,1456,980,1581]" pageId="6" pageNumber="369">
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This small mandibular portion (
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<quantity id="4C99AEEA277A6A72ADF1F852E74BD5B8" box="[508,593,1937,1964]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="369" unit="mm" value="29.0">29 mm</quantity>
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in length) preserves three partial bones; a dentary labially and dorsally (
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<figureCitation id="135A1F8A277A6A72AC83F812E41FD5FF" box="[142,261,2000,2027]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4A, B</figureCitation>
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[d]), and two coronoids lingually and dorsally (
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<figureCitation id="135A1F8A277A6A72AF3BFC17E6B4D1FB" box="[822,942,980,1007]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4B, C</figureCitation>
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[cor1 and cor2]). The partial dentary is wellornamented, and its ornamentation pattern consists of interconnected pits of various sizes. It bears at least 12 marginal tooth sockets, which are slightly laterally compressed (a stereospondyl synapomorphy; e.g.,
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<bibRefCitation id="EFF07EFE277A6A72A884FB96E075D67A" author="YATES A. M. & WARREN A. A." box="[1161,1391,1108,1135]" pageId="6" pageNumber="369" pagination="77 - 121" refId="ref10229" refString="YATES A. M. & WARREN A. A. 2000. - The phylogeny of the ' higher' temnospondyls (Vertebrata: Choanata) and its implications for the monophyly and origins of the Stereospondyli. Zoological Journal of the Linnean Society 128: 77 - 121. https: // doi. org / 10.1111 / j. 1096 - 3642.2000. tb 00650. x" type="journal article" year="2000">Yates & Warren 2000</bibRefCitation>
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). The socket Nr. 8 still preserves a partial tooth with its medullar cavity. The two partial coronoids are covered by dense and well-separated denticle patches (
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<emphasis id="B915DF1D277A6A72A89CFB76E1B8D6DA" box="[1169,1186,1205,1230]" italics="true" pageId="6" pageNumber="369">c.</emphasis>
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3 teeth per mm
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<superScript id="7C14AE47277A6A72A95DFB76E040D6D2" attach="left" box="[1360,1370,1205,1222]" fontSize="7" pageId="6" pageNumber="369">2</superScript>
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). These denticles are conical, short (<
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<quantity id="4C99AEEA277A6A72A87AFB17E1A1D6FA" box="[1143,1211,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="369" unit="mm" value="1.0">1 mm</quantity>
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), and taper to a sharp point. These denticle patches are separated by a depression in the posterior region of the specimen that is visible lingually. The anterior patch is complete and covers most of the length of the specimen. It is supported by the betterpreserved coronoid (
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<figureCitation id="135A1F8A277A6A72A81EFAB0E193D799" box="[1043,1161,1394,1422]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4B, C</figureCitation>
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[cor1]). The other denticle patch and its corresponding coronoid (
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<figureCitation id="135A1F8A277A6A72A8E7FA50E078D7B9" box="[1258,1378,1426,1454]" captionStart="FIG" captionStartId="7.[132,143,1555,1572]" captionTargetBox="[141,1448,216,1512]" captionTargetId="figure-286@7.[698,1267,736,1277]" captionTargetPageId="7" captionText="FIG. 4. — Partial mandible of Stereospondyli (NHCC LB678) from the Triassic (?Anisian) of Zambia (Luangwa Basin) in labial (A), dorsal (B) and lingual (C) views. Abbreviations: cor1, coronoid 1; cor2, coronoid 2; d, dentary. Scale bar: 10 mm. Modified from Peecook et al. 2018." figureDoi="http://doi.org/10.5281/zenodo.4906303" httpUri="https://zenodo.org/record/4906303/files/figure.png" pageId="6" pageNumber="369">Fig. 4B, C</figureCitation>
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[cor2]) are partially preserved and only visible in the posterior extremity of the specimen. Due to the fragmentary nature of the specimen, it is not clear if these elements represent pre-, inter- or post-coronoids.
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</paragraph>
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<paragraph id="8BDE030F277A6A72AF20F990E6C6D47F" blockId="6.[811,1457,1619,2028]" box="[813,988,1619,1645]" pageId="6" pageNumber="369">IDENTIFICATION</paragraph>
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<paragraph id="8BDE030F277A6A72AF20F9B0E06CD5FF" blockId="6.[811,1457,1619,2028]" pageId="6" pageNumber="369">
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We identify this specimen as a stereospondyl based on the laterally compressed tooth sockets as well as the stereospondyl synapomorphy of a dentary that does not contribute substantially to the lingual side of the mandible (the dentary is clearly visible in lingual view in basal temnospondyls; e.g.,
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<bibRefCitation id="EFF07EFE277A6A72AF21F8D1E13BD538" author="SCHOCH R. R. & MILNER A. R." box="[812,1057,1810,1836]" pageId="6" pageNumber="369" refId="ref9290" refString="SCHOCH R. R. & MILNER A. R. 2000. - Handbuch der Palaoherpetologie / Encyclopedia of Paleoherpetology. Teil / Part 3 B. Stereospondyli. Friedrich Pfeil, Munchen, 219 p." type="book" year="2000">Schoch & Milner 2000</bibRefCitation>
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). It is difficult to assign a more precise identification because three stereospondyl clades have very dense denticle patches on at least two of their coronoids: the
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<taxonomicName id="4C61788C277A6A72AF21F8B2E6D7D59F" authorityName="Watson" authorityYear="1919" box="[812,973,1905,1931]" class="Amphibia" family="Rhinesuchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Temnospondyli" pageId="6" pageNumber="369" phylum="Chordata" rank="family">Rhinesuchidae</taxonomicName>
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, the
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<taxonomicName id="4C61788C277A6A72A80FF8B2E1B0D59F" authorityName="Watson" authorityYear="1919" box="[1026,1194,1905,1931]" class="Amphibia" family="Lydekkerinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Temnospondyli" pageId="6" pageNumber="369" phylum="Chordata" rank="family">Lydekkerinidae</taxonomicName>
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, and the
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<taxonomicName id="4C61788C277A6A72A900F8B2E0B4D59F" authorityName="von Huene" authorityYear="1920" box="[1293,1454,1905,1931]" class="Amphibia" family="Rhytidosteidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Temnospondyli" pageId="6" pageNumber="369" phylum="Chordata" rank="family">Rhytidosteidae</taxonomicName>
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(e.g.,
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<taxonomicName id="4C61788C277A6A72AF65F852E1E7D5BF" authority="Lehman, 1966" authorityName="Lehman" authorityYear="1966" box="[872,1277,1937,1963]" class="Amphibia" family="Rhytidosteidae" genus="Mahavisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Temnospondyli" pageId="6" pageNumber="369" phylum="Chordata" rank="species" species="dentatus">
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<emphasis id="B915DF1D277A6A72AF65F852E14DD5BF" box="[872,1111,1937,1963]" italics="true" pageId="6" pageNumber="369">Mahavisaurus dentatus</emphasis>
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<bibRefCitation id="EFF07EFE277A6A72A851F852E1E7D5BF" author="LEHMAN J. P." box="[1116,1277,1937,1963]" pageId="6" pageNumber="369" pagination="117 - 139" refId="ref8063" refString="LEHMAN J. P. 1966. - Nouveaux stegocephales de Madagascar. Annales de Paleontologie, Vertebres 52: 117 - 139." type="journal article" year="1966">Lehman, 1966</bibRefCitation>
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</taxonomicName>
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,
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<bibRefCitation id="EFF07EFE277A6A72A908F851E67FD5DF" author="MAGANUCO S. & PASINI G. & AUDITORE M." pageId="6" pageNumber="369" pagination="1 - 64" refId="ref8207" refString="MAGANUCO S., PASINI G. & AUDITORE M. 2014. - A revision of the short-faced stereospondyls Mahavisaurus dentatus and Lyrosaurus australis from the Lower Triassic of Madagascar: cranial anatomy, ontogenetic remarks, palaeoecology and rhytidosteids phylogeny. Memori della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 39: 1 - 64." type="journal article" year="2014">
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Maganuco
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<emphasis id="B915DF1D277A6A72A973F851E0AAD5BF" box="[1406,1456,1937,1963]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
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2014
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</bibRefCitation>
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: figs 8
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<emphasis id="B915DF1D277A6A72AFAAF872E6AAD5DF" bold="true" box="[935,944,1969,1995]" pageId="6" pageNumber="369">-</emphasis>
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9). Pending more discoveries, we therefore prefer to assign this fragmentary specimen to
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<taxonomicName id="4C61788C277A6A72A8DDF812E068D5FF" authorityName="Zittel" authorityYear="1888" box="[1232,1394,2001,2027]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stereospondyli" pageId="6" pageNumber="369" phylum="Chordata" rank="order">Stereospondyli</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |