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<document id="8353B222107B1419BA4A4DC587116B5C" ID-DOI="10.5281/zenodo.180216" ID-GBIF-Dataset="f7c4a979-1dc7-400b-b5b0-eec6919fa931" ID-ISSN="1175-5326" ID-Zenodo-Dep="180216" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459801613191" checkinUser="plazi" docAuthor="Kehlmaier, Christian" docDate="2008" docId="03C1927DFFF14228FF3AF4A2FAECFE88" docLanguage="en" docName="zt01672p042.pdf" docOrigin="Zootaxa 1672" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pipunculus omissinervis Becker 1889" docType="treatment" docVersion="9" lastPageNumber="23" masterDocId="FFF8EA05FFE2423EFFADF06DFF98FFDA" masterDocTitle="Finnish Pipunculidae (Diptera) studies Part I: Taxonomic notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939" masterLastPageNumber="42" masterPageNumber="1" pageNumber="20" updateTime="1698220027910" updateUser="plazi">
<mods:mods id="183B48998A8ACF16F184CC8AF9C6DCFA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0C21F5B08BC24C445A930C67871ED2F0">
<mods:title id="56C0AEB077F8E2CE6991BF26CB250BA8">Finnish Pipunculidae (Diptera) studies Part I: Taxonomic notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939</mods:title>
</mods:titleInfo>
<mods:name id="00CB380D1886FABC49E5BB6687A4C0B6" type="personal">
<mods:role id="B85E45E495B9385594BCEE4B89687766">
<mods:roleTerm id="71043D8DB26023B0FC90D9DDC92A3C93">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5A4E886284A720C7FBF63866B9FE961">Kehlmaier, Christian</mods:namePart>
</mods:name>
<mods:typeOfResource id="03EB614B380268DC3965F25D6B8844F7">text</mods:typeOfResource>
<mods:relatedItem id="A21842F6D90D2C99D4F7613A87B49570" type="host">
<mods:titleInfo id="E134ACAF7B712FD1FE5A14B386617882">
<mods:title id="6725FA39F65E6101E56C998DFE5EAAAA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6D1E2B46132E2824113D8C359DD8B9CF">
<mods:date id="985DD3451F1ABF244C712E5255CE7EA9">2008</mods:date>
<mods:detail id="344013D3115DF2776980FB8AE8749E6B" type="volume">
<mods:number id="7E1294191E9DE1A1FADAC2F71814446B">1672</mods:number>
</mods:detail>
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<mods:start id="F33E979DFE2B7C18B6C0297D0A2E7FD5">1</mods:start>
<mods:end id="6DE0A93FF0344D90D264F664B10115F5">42</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B3C34D3890DBE4D55CE2736A90D1A6DF">journal article</mods:classification>
<mods:identifier id="AF156B2406CA54ECA546F751E1EA6905" type="DOI">10.5281/zenodo.180216</mods:identifier>
<mods:identifier id="F8001D39FEE99AD9708F6FC717E9A162" type="GBIF-Dataset">f7c4a979-1dc7-400b-b5b0-eec6919fa931</mods:identifier>
<mods:identifier id="3F54F810779ADFCF552EE91AC01FDB80" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5AB56AFA1583A954C72FDF2191C5188C" type="Zenodo-Dep">180216</mods:identifier>
</mods:mods>
<treatment id="03C1927DFFF14228FF3AF4A2FAECFE88" ID-DOI="http://doi.org/10.5281/zenodo.6232690" ID-GBIF-Taxon="119351850" ID-Zenodo-Dep="6232690" LSID="urn:lsid:plazi:treatment:03C1927DFFF14228FF3AF4A2FAECFE88" httpUri="http://treatment.plazi.org/id/03C1927DFFF14228FF3AF4A2FAECFE88" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20">
<subSubSection id="C37270E0FFF1422DFF3AF4A2FE50FAD6" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BD7236BFFF1422DFF3AF4A2FDFCFB33" blockId="19.[151,612,1231,1292]" box="[151,612,1231,1257]" pageId="19" pageNumber="20">
<heading id="D09F9407FFF1422DFF3AF4A2FDFCFB33" bold="true" box="[151,612,1231,1257]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="4C6858E8FFF1422DFF3AF4A2FDFCFB33" ID-CoL="4J7KY" authority="Becker, 1889" authorityName="Becker" authorityYear="1889" box="[151,612,1231,1257]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF1422DFF3AF4A2FDFCFB33" bold="true" box="[151,612,1231,1257]" pageId="19" pageNumber="20">
<emphasis id="B91CFF79FFF1422DFF3AF4A2FE27FB33" bold="true" box="[151,447,1231,1257]" italics="true" pageId="19" pageNumber="20">Pipunculus omissinervis</emphasis>
<bibRefCitation id="EFF95E9AFFF1422DFE68F4A2FDFCFB33" author="Becker" box="[453,612,1231,1257]" pageId="19" pageNumber="40" refString="Becker, T. (1889) Altes und Neues aus der Schweiz. Ein dipterologischer Beitrag. Wiener Entomologische Zeitung, 8, 73 - 84 + plate I." type="journal article" year="1889">Becker, 1889</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD7236BFFF1422DFF3AF49FFE50FAD6" blockId="19.[151,612,1231,1292]" box="[151,456,1266,1292]" pageId="19" pageNumber="20">
(
<figureCitation id="13533FEEFFF1422DFF32F49FFED7FAD6" box="[159,335,1266,1292]" captionStart="FIGURES 36 – 42" captionStartId="18.[151,269,1970,1994]" captionTargetBox="[272,1282,438,1945]" captionTargetId="figure@18.[264,1323,434,1946]" captionTargetPageId="18" captionText="FIGURES 36 – 42. Surstyli in strictly dorsal and outer surstylus in lateral view of male Pipunculus carlestolrai (36 &amp; 40), Pipunculus calceatus (37 &amp; 41) and Pipunculus omissinervis (38 – 39 &amp; 42). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/180222/files/figure.png" pageId="19" pageNumber="20">Figs 38–39, 42</figureCitation>
&amp;
<figureCitation id="13533FEEFFF1422DFED8F49FFE58FAD6" box="[373,448,1266,1292]" captionStart="FIGURES 67 – 73" captionStartId="24.[151,269,1542,1566]" captionTargetBox="[270,1308,485,1511]" captionTargetId="figure@24.[264,1323,474,1518]" captionTargetPageId="24" captionText="FIGURES 67 – 73. Ovipositor in lateral and dorsal view of female Pipunculus campestris agg. (67 – 71) and Pipunculus wolfi (72 – 73). Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/180225/files/figure.png" pageId="19" pageNumber="20">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37270E0FFF1422DFF3AF52FFDEEFA43" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="8BD7236BFFF1422DFF3AF52FFDC6FAA3" blockId="19.[151,746,1346,1433]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB780445FFF1422DFF3AF52FFDC6FAA3" pageId="19" pageNumber="20">
<taxonomicName id="4C6858E8FFF1422DFF3AF52FFD18FA83" ID-CoL="4J7KQ" authority="Kuznetzov, 1990: 378" authorityName="Kuznetzov" authorityPageNumber="378" authorityYear="1990" box="[151,640,1346,1369]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B91CFF79FFF1422DFF3AF52FFE13FA83" box="[151,395,1346,1369]" italics="true" pageId="19" pageNumber="20">Pipunculus nartshukae</emphasis>
<treatmentCitation id="0AC9057AFFF1422DFE3EF52FFD18FA83" author="Kuznetzov" box="[403,640,1346,1369]" page="378" pageId="19" pageNumber="20" year="1990">
<bibRefCitation id="EFF95E9AFFF1422DFE3EF52FFDD3FA83" author="Kuznetzov" box="[403,587,1346,1369]" pageId="19" pageNumber="41" refString="Kuznetzov, S. Y. (1990) New species of the family Pipunculidae from Mongolia. Insects of Mongolia, 11, 378 - 393." type="journal article" year="1990">Kuznetzov, 1990</bibRefCitation>
: 378
</treatmentCitation>
</taxonomicName>
. syn. nov.
<taxonomicName id="4C6858E8FFF1422DFF3AF50FFDC2FAA3" ID-CoL="4J7KY" authority="Becker, 1889: 83" authorityName="Becker" authorityPageNumber="83" authorityYear="1889" box="[151,602,1378,1401]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF1422DFF3AF50FFE02FAA3" box="[151,410,1378,1401]" italics="true" pageId="19" pageNumber="20">Pipunculus omissinervis</emphasis>
<treatmentCitation id="0AC9057AFFF1422DFE0CF50FFDC2FAA3" author="Becker" box="[417,602,1378,1401]" page="83" pageId="19" pageNumber="20" year="1889">
<bibRefCitation id="EFF95E9AFFF1422DFE0CF50FFDAAFAA3" author="Becker" box="[417,562,1378,1401]" pageId="19" pageNumber="40" refString="Becker, T. (1889) Altes und Neues aus der Schweiz. Ein dipterologischer Beitrag. Wiener Entomologische Zeitung, 8, 73 - 84 + plate I." type="journal article" year="1889">Becker, 1889</bibRefCitation>
: 83
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD7236BFFF1422DFF3AF5EFFDEEFA43" blockId="19.[151,746,1346,1433]" box="[151,630,1410,1433]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB780445FFF1422DFF3AF5EFFDEEFA43" box="[151,630,1410,1433]" pageId="19" pageNumber="20">
<taxonomicName id="4C6858E8FFF1422DFF3AF5EFFD93FA43" ID-CoL="4J7L5" authority="Coe, 1966: 49" authorityName="Coe" authorityPageNumber="49" authorityYear="1966" box="[151,523,1410,1433]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="phaeton">
<emphasis id="B91CFF79FFF1422DFF3AF5EFFEF3FA43" box="[151,363,1410,1433]" italics="true" pageId="19" pageNumber="20">Pipunculus phaeton</emphasis>
<treatmentCitation id="0AC9057AFFF1422DFEDFF5EFFD93FA43" author="Coe" box="[370,523,1410,1433]" page="49" pageId="19" pageNumber="20" year="1966">
<bibRefCitation id="EFF95E9AFFF1422DFEDFF5EFFE7BFA43" author="Coe" box="[370,483,1410,1433]" pageId="19" pageNumber="40" refString="Coe, R. L. (1966) Pipunculidae. Handbook for the Identification of British Insects. Vol. X, Part 2 (c). Royal Entomological Society of London, London, 83 pp." type="book" year="1966">Coe, 1966</bibRefCitation>
: 49
</treatmentCitation>
</taxonomicName>
. syn. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37270E0FFF1422DFF3AF5A7FBA7F926" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BD7236BFFF1422DFF3AF5A7FBA7F926" blockId="19.[151,1437,1482,2028]" pageId="19" pageNumber="20">
<emphasis id="B91CFF79FFF1422DFF3AF5A7FE96FA3E" bold="true" box="[151,270,1482,1508]" pageId="19" pageNumber="20">Material:</emphasis>
<collectingCountry id="F37F63FBFFF1422DFEBAF5A7FE3EFA3E" box="[279,422,1482,1508]" name="Switzerland" pageId="19" pageNumber="20">Switzerland</collectingCountry>
, Eggischhorn, Furka Pass, no. 19326: 1ɗ (
<typeStatus id="54D39DC9FFF1422DFC05F5A7FB8DFA3E" box="[936,1045,1482,1508]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
),
<date id="FFD605ABFFF1422DFB87F5A7FB1AFA3E" box="[1066,1154,1482,1508]" pageId="19" pageNumber="20">23.VII.</date>
, Becker (
<collectionCode id="ED79BBAEFFF1422DFB58F5A7FACDFA3E" box="[1269,1365,1482,1508]" pageId="19" pageNumber="20">MNHU</collectionCode>
);
<collectingCountry id="F37F63FBFFF1422DFAC6F5A7FF54F9D6" name="Finland" pageId="19" pageNumber="20">Finland</collectingCountry>
, N, Sibbo, Nevas, 6687:412: 2ɗ,
<date id="FFD605ABFFF1422DFDC3F59FFCBEF9D6" box="[622,806,1522,1548]" pageId="19" pageNumber="20" value="1985-07-07" valueMax="1985-07-13" valueMin="1985-07-07">7.–13.VII.1985</date>
; 1ɗ,
<date id="FFD605ABFFF1422DFCC6F59FFBA8F9D6" box="[875,1072,1522,1548]" pageId="19" pageNumber="20" value="1986-07-13" valueMax="1986-07-20" valueMin="1986-07-13">13.–20.VII.1986</date>
; 1ɗ,
<date id="FFD605ABFFF1422DFBDBF59FFAAAF9D6" box="[1142,1330,1522,1548]" pageId="19" pageNumber="20" value="1986-06-22" valueMax="1986-06-28" valueMin="1986-06-22">22.–28.VI.1986</date>
; 1ɗ,
<date id="FFD605ABFFF1422DFADAF59FFE83F9EE" pageId="19" pageNumber="20" value="1986-07-01" valueMax="1986-07-06" valueMin="1986-07-01">1.– 6.VII.1986</date>
; 4ɗ,
<date id="FFD605ABFFF1422DFEFBF677FD8AF9EE" box="[342,530,1562,1588]" pageId="19" pageNumber="20" value="1986-06-15" valueMax="1986-06-21" valueMin="1986-06-15">15.–21.VI.1986</date>
; 2ɗ,
<date id="FFD605ABFFF1422DFDE3F677FCA6F9EE" box="[590,830,1562,1588]" pageId="19" pageNumber="20" value="1986-07-28" valueMax="1986-08-04" valueMin="1986-07-28">28.VII.–4.VIII.1986</date>
; 1ɗ,
<date id="FFD605ABFFF1422DFCD7F677FBD2F9EE" box="[890,1098,1562,1588]" pageId="19" pageNumber="20" value="1984-08-22" valueMax="1984-08-26" valueMin="1984-08-22">22.–26.VIII.1984</date>
, Albrecht;
<collectingCountry id="F37F63FBFFF1422DFB67F677FABEF9EE" box="[1226,1318,1562,1588]" name="Finland" pageId="19" pageNumber="20">Finland</collectingCountry>
, Helsinki, Kallahdenniemi: 1ɗ,
<date id="FFD605ABFFF1422DFE3BF62FFD86F986" box="[406,542,1602,1628]" pageId="19" pageNumber="20" value="2003-06-18">18.VI.2003</date>
, Kahanpää (
<collectionCode id="ED79BBAEFFF1422DFD1EF62FFD66F986" box="[691,766,1602,1628]" pageId="19" pageNumber="20">PCJK</collectionCode>
);
<collectingCountry id="F37F63FBFFF1422DFCB9F62FFC1AF986" box="[788,898,1602,1628]" name="Germany" pageId="19" pageNumber="20">Germany</collectingCountry>
, Bavaria, Oberallgäu, Ofterschwang,
<quantity id="4C908E8EFFF1422DFAEEF62FFA0FF986" box="[1347,1431,1602,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1000.0">1000m</quantity>
,
<geoCoordinate id="EE5C45ACFFF1422DFF3AF607FEBFF95E" box="[151,295,1642,1668]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="15" value="10.2275">10°13’39’’E</geoCoordinate>
<geoCoordinate id="EE5C45ACFFF1422DFE9DF607FE50F95E" box="[304,456,1642,1668]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="15" value="47.49778">47°29’52’’N</geoCoordinate>
: 1ɗ,
<date id="FFD605ABFFF1422DFDABF607FD3BF95E" box="[518,675,1642,1668]" pageId="19" pageNumber="20" value="2004-08-12">12.VIII.2004</date>
, Kehlmaier (
<collectionCode id="ED79BBAEFFF1422DFC91F607FC17F95E" box="[828,911,1642,1668]" pageId="19" pageNumber="20">PCCK</collectionCode>
);
<collectingCountry id="F37F63FBFFF1422DFC09F607FB8AF95E" box="[932,1042,1642,1668]" name="Germany" pageId="19" pageNumber="20">Germany</collectingCountry>
, Bavaria, Oberallgäu, Sonthofen,
<quantity id="4C908E8EFFF1422DFF3AF6FFFF73F976" box="[151,235,1682,1708]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1000.0">1000m</quantity>
,
<geoCoordinate id="EE5C45ACFFF1422DFF53F6FFFE17F976" box="[254,399,1682,1708]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="15" value="10.320278">10°19’13’’E</geoCoordinate>
<geoCoordinate id="EE5C45ACFFF1422DFE30F6FFFDADF976" box="[413,565,1682,1708]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="15" value="47.519444">47°31’10’’N</geoCoordinate>
: 1ɗ,
<date id="FFD605ABFFF1422DFDD2F6FFFC82F976" box="[639,794,1682,1708]" pageId="19" pageNumber="20" value="2004-08-11">11.VIII.2004</date>
, Kehlmaier (
<collectionCode id="ED79BBAEFFF1422DFC12F6FFFB8AF976" box="[959,1042,1682,1708]" pageId="19" pageNumber="20">PCCK</collectionCode>
);
<collectingCountry id="F37F63FBFFF1422DFB80F6FFFB0DF976" box="[1069,1173,1682,1708]" name="Hungary" pageId="19" pageNumber="20">Hungary</collectingCountry>
, Nagy kovácsi, Júlia major: 1ɗ,
<date id="FFD605ABFFF1422DFE89F6D7FE43F90E" box="[292,475,1722,1748]" pageId="19" pageNumber="20" value="1983-04-25" valueMax="1983-04-27" valueMin="1983-04-25">25.–27.IV.1983</date>
, Visnyovszky (
<collectionCode id="ED79BBAEFFF1422DFD3BF6D7FD6FF90E" LSID="urn:lsid:biocol.org:col:34991" box="[662,759,1722,1748]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="19" pageNumber="20">HNHM</collectionCode>
);
<collectingCountry id="F37F63FBFFF1422DFCA2F6D7FCEFF90E" box="[783,887,1722,1748]" name="Hungary" pageId="19" pageNumber="20">Hungary</collectingCountry>
, Hárskut, Esztergáli-v.: 1ɗ,
<date id="FFD605ABFFF1422DFB7DF6D7FACCF90E" box="[1232,1364,1722,1748]" pageId="19" pageNumber="20" value="1977-04-27">27.IV.1977</date>
, Tóth (
<collectionCode id="ED79BBAEFFF1422DFF32F68FFF67F926" LSID="urn:lsid:biocol.org:col:34991" box="[159,255,1762,1788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="19" pageNumber="20">HNHM</collectionCode>
);
<collectingCountry id="F37F63FBFFF1422DFEBEF68FFEE3F926" box="[275,379,1762,1788]" name="Hungary" pageId="19" pageNumber="20">Hungary</collectingCountry>
,
<collectionCode id="ED79BBAEFFF1422DFE2AF68FFE58F926" box="[391,448,1762,1788]" httpUri="http://grbio.org/cool/vkvr-e0fw" name="Banff Park Museum" pageId="19" pageNumber="20">BNP</collectionCode>
Hármaskút: 1ɗ,
<date id="FFD605ABFFF1422DFD25F68FFCD5F926" box="[648,845,1762,1788]" pageId="19" pageNumber="20" value="1982-05-31" valueMax="1982-06-04" valueMin="1982-05-31">31.V.–4.VI.1982</date>
, Bessenyi (
<collectionCode id="ED79BBAEFFF1422DFC7FF68FFBABF926" LSID="urn:lsid:biocol.org:col:34991" box="[978,1075,1762,1788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="19" pageNumber="20">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37270E0FFF1422AFF6BF767FBFFF92E" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="8BD7236BFFF1422DFF6BF767FA04F836" blockId="19.[151,1437,1482,2028]" pageId="19" pageNumber="20">
<emphasis id="B91CFF79FFF1422DFF6BF767FE9EF8FE" bold="true" box="[198,262,1802,1828]" pageId="19" pageNumber="20">Male</emphasis>
: Body length:
<quantity id="4C908E8EFFF1422DFE19F767FDA5F8FE" box="[436,573,1802,1828]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.7" metricValueMin="3.8" pageId="19" pageNumber="20" unit="mm" value="4.25" valueMax="4.7" valueMin="3.8">3.8–4.7mm</quantity>
. Wing length:
<quantity id="4C908E8EFFF1422DFD4AF767FCE9F8FE" box="[743,881,1802,1828]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="19" pageNumber="20" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.1–5.1mm</quantity>
. Flagellum dark brown. Frons brown pollinose. Eyes meeting for length of frons or slightly more (13–16 facets). Occiput grey pollinose except posterior to ocellar triangle with brown pollinosity, which occasionally can be extended to as much as entire upper half. Postpronotal lobe dark brown. Prescutum and scutum entirely brown pollinose but can be greyish in anterior quarter. Pleura with anepimeron, katepimeron, posterior margin of anepisternum and upper margin of katepisternum light brown pollinose but this can be reduced to entirely absent, otherwise weakly grey pollinose.
</paragraph>
<paragraph id="8BD7236BFFF6422AFF3AF0F5FCFFFD98" blockId="20.[151,1437,152,2020]" pageId="20" pageNumber="21">
Scutellum with apical fringe of about 12 evenly spaced hairs. Subscutellum grey pollinose but occasionally some weak brown pollinosity can be present centrally or in upper lateral corners. LTC about 1.0–1.7 times LFC. Trochanter yellowish to brown. Femur with base (rarely) and apex narrowly yellow.
<collectingCountry id="F37F63FBFFF6422AFB15F085FB6BFED8" box="[1208,1267,232,258]" name="India" pageId="20" pageNumber="21">Hind</collectingCountry>
femur weakly concave in ventrobasal half. Tibia yellowish, ventrally darkened in apical two thirds. Tarsi yellowish to brown, with distitarsi darkest. Pulvilli of front leg slightly shorter than distitarsi. Tergite 1 anterodorsally brown, otherwise grey pollinose. Dorsally brown pollinosity covering entire tergite 2, anterior three quarters of tergite 3, anterior half to two thirds of tergite 4 and anterior quarter of tergite 5, otherwise shining. Laterally, tergites 2–5 entirely brown pollinose or with distinct triangular patches of greyish-brown to grey pollinosity in posterior half. Membranous area key-hole shaped. Surstyli elongated but can be slightly shorter than figured here (
<figureCitation id="13533FEEFFF6422AFE9AF26DFE5AFDC0" box="[311,450,512,538]" captionStart="FIGURES 36 – 42" captionStartId="18.[151,269,1970,1994]" captionTargetBox="[272,1282,438,1945]" captionTargetId="figure@18.[264,1323,434,1946]" captionTargetPageId="18" captionText="FIGURES 36 – 42. Surstyli in strictly dorsal and outer surstylus in lateral view of male Pipunculus carlestolrai (36 &amp; 40), Pipunculus calceatus (37 &amp; 41) and Pipunculus omissinervis (38 – 39 &amp; 42). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/180222/files/figure.png" pageId="20" pageNumber="21">Figs 38–39</figureCitation>
). Depending on the perspective, the shape of the surstyli can vary. Only two of the three ejaculatory ducts are distinctly sclerotized in distal half.
</paragraph>
<paragraph id="8BD7236BFFF6422AFF68F222FB81FDB0" blockId="20.[151,1437,152,2020]" box="[197,1049,591,618]" pageId="20" pageNumber="21">
<emphasis id="B91CFF79FFF6422AFF68F222FEB8FDB3" bold="true" box="[197,288,591,617]" pageId="20" pageNumber="21">Female</emphasis>
: Conspecific with
<emphasis id="B91CFF79FFF6422AFE53F23DFD0EFDB3" box="[510,662,592,617]" italics="true" pageId="20" pageNumber="21">P. c a m p e s t r i s</emphasis>
(see above for characterisation).
</paragraph>
<paragraph id="8BD7236BFFF6422AFF68F21AFF41FCD6" blockId="20.[151,1437,152,2020]" pageId="20" pageNumber="21">
<emphasis id="B91CFF79FFF6422AFF68F21AFEC6FD4B" bold="true" box="[197,350,631,657]" pageId="20" pageNumber="21">Annotations</emphasis>
:
<bibRefCitation id="EFF95E9AFFF6422AFEC2F215FD81FD48" author="Becker" box="[367,537,632,658]" pageId="20" pageNumber="40" refString="Becker, T. (1889) Altes und Neues aus der Schweiz. Ein dipterologischer Beitrag. Wiener Entomologische Zeitung, 8, 73 - 84 + plate I." type="journal article" year="1889">Becker (1889)</bibRefCitation>
collected a single male at Mount Eggischhorn. The specimen’s label states “Furka”, a pass about
<quantity id="4C908E8EFFF6422AFE0BF2CDFE72FD60" box="[422,490,672,698]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="21" unit="km" value="30.0">30km</quantity>
away according to De Meyer (1996). The female sex cannot be differentiated from
<taxonomicName id="4C6858E8FFF6422AFF75F2A5FEECFD3B" box="[216,372,712,737]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="B91CFF79FFF6422AFF75F2A5FEECFD3B" box="[216,372,712,737]" italics="true" pageId="20" pageNumber="21">P. campestris</emphasis>
</taxonomicName>
at present. The male from Hármaskút in
<collectingCountry id="F37F63FBFFF6422AFCF1F2A5FC5AFD38" box="[860,962,712,738]" name="Hungary" pageId="20" pageNumber="21">Hungary</collectingCountry>
has a short additional vein M
<quantity id="4C908E8EFFF6422AFA8FF2B5FAD1FD38" box="[1314,1353,712,742]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">2 in</quantity>
its left wing.
</paragraph>
<paragraph id="8BD7236BFFF6422AFF68F377FB9EFB46" blockId="20.[151,1437,152,2020]" pageId="20" pageNumber="21">
A comparison of European
<taxonomicName id="4C6858E8FFF6422AFDBFF376FD5CFCEE" box="[530,708,795,820]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFDBFF376FD5CFCEE" box="[530,708,795,820]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
and Nearctic
<taxonomicName id="4C6858E8FFF6422AFCC2F376FA9AFCEE" authority="Rapp, 1943" authorityName="Rapp" authorityYear="1943" box="[879,1282,794,820]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hertzogi">
<emphasis id="B91CFF79FFF6422AFCC2F376FBC4FCEE" box="[879,1116,795,820]" italics="true" pageId="20" pageNumber="21">Pipunculus hertzogi</emphasis>
(
<bibRefCitation id="EFF95E9AFFF6422AFBC3F377FB63FCEE" author="Rapp" box="[1134,1275,794,820]" pageId="20" pageNumber="41" refString="Rapp, W. F. (1943) A new Dorilaidae (Diptera). Entomological News, 54, 118." type="journal article" year="1943">Rapp, 1943</bibRefCitation>
)
</taxonomicName>
showed that male terminalia are conspecific. However, comparison of yet unpublished molecular data for a 658bp fragment of coxI suggests that both taxa have to be considered as distinct lineages (tested for one Nearctic male
<taxonomicName id="4C6858E8FFF6422AFA2BF306FF6FFC76" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hertzogi">
<emphasis id="B91CFF79FFF6422AFA2BF306FF6FFC76" italics="true" pageId="20" pageNumber="21">P. hertzogi</emphasis>
</taxonomicName>
(Skevington unpublished) and five Palaearctic male
<taxonomicName id="4C6858E8FFF6422AFCD5F3FEFBB3FC76" box="[888,1067,915,940]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFCD5F3FEFBB3FC76" box="[888,1067,915,940]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
specimens (Kehlmaier unpublished)). This is especially interesting because
<taxonomicName id="4C6858E8FFF6422AFD13F3D6FCA4FC0E" box="[702,828,955,980]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hertzogi">
<emphasis id="B91CFF79FFF6422AFD13F3D6FCA4FC0E" box="[702,828,955,980]" italics="true" pageId="20" pageNumber="21">P. hertzogi</emphasis>
</taxonomicName>
has lately been thoroughly reanalyzed by
<bibRefCitation id="EFF95E9AFFF6422AFA9FF3D7FE29FC26" author="Skevington" pageId="20" pageNumber="42" refString="Skevington, J. &amp; Marshall, S. A. (1998) Systematics of New World Pipunculus (Diptera: Pipunculidae). Thomas Say Publications in Entomology: Monographs, Lanham, 201 pp." type="book" year="1998">
Skevington &amp;
<collectingCountry id="F37F63FBFFF6422AFF40F38FFECDFC26" box="[237,341,994,1020]" name="Marshall Islands" pageId="20" pageNumber="21">Marshall</collectingCountry>
(1998)
</bibRefCitation>
who also established it as a senior synonym of
<taxonomicName id="4C6858E8FFF6422AFBB7F38EFA0FFC26" authority="Coe, 1966" authorityName="Coe" authorityYear="1966" box="[1050,1431,994,1020]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="phaeton">
<emphasis id="B91CFF79FFF6422AFBB7F38EFA92FC26" box="[1050,1290,995,1020]" italics="true" pageId="20" pageNumber="21">Pipunculus phaeton</emphasis>
<bibRefCitation id="EFF95E9AFFF6422AFABBF38FFA0FFC26" author="Coe" box="[1302,1431,994,1020]" pageId="20" pageNumber="40" refString="Coe, R. L. (1966) Pipunculidae. Handbook for the Identification of British Insects. Vol. X, Part 2 (c). Royal Entomological Society of London, London, 83 pp." type="book" year="1966">Coe, 1966</bibRefCitation>
</taxonomicName>
, originally described from 5ɗ and 6Ψ collected in the southern part of
<collectingCountry id="F37F63FBFFF6422AFC4DF467FB18FBFE" box="[992,1152,1034,1060]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
. Because of the present results it is proposed to regard
<taxonomicName id="4C6858E8FFF6422AFDABF45EFD1CFB96" box="[518,644,1075,1100]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hertzogi">
<emphasis id="B91CFF79FFF6422AFDABF45EFD1CFB96" box="[518,644,1075,1100]" italics="true" pageId="20" pageNumber="21">P. hertzogi</emphasis>
</taxonomicName>
as an exclusively Nearctic taxon and transfer
<taxonomicName id="4C6858E8FFF6422AFB05F45EFABCFB96" box="[1192,1316,1075,1100]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="phaeton">
<emphasis id="B91CFF79FFF6422AFB05F45EFABCFB96" box="[1192,1316,1075,1100]" italics="true" pageId="20" pageNumber="21">P. phaeton</emphasis>
</taxonomicName>
to synonymy with
<taxonomicName id="4C6858E8FFF6422AFEA0F436FE24FBAE" box="[269,444,1115,1140]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFEA0F436FE24FBAE" box="[269,444,1115,1140]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
instead (syn. nov.). The citings of
<taxonomicName id="4C6858E8FFF6422AFCE2F436FC54FBAE" box="[847,972,1115,1140]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hertzogi">
<emphasis id="B91CFF79FFF6422AFCE2F436FC54FBAE" box="[847,972,1115,1140]" italics="true" pageId="20" pageNumber="21">P. hertzogi</emphasis>
</taxonomicName>
from mainland
<collectingCountry id="F37F63FBFFF6422AFB2BF437FB24FBAE" box="[1158,1212,1114,1140]" name="Italy" pageId="20" pageNumber="21">Italy</collectingCountry>
and eastern
<collectingCountry id="F37F63FBFFF6422AFAE3F437FA04FBAE" box="[1358,1436,1114,1140]" name="Russia" pageId="20" pageNumber="21">Russia</collectingCountry>
(both
<bibRefCitation id="EFF95E9AFFF6422AFF71F4EFFDAAFB46" author="Skevington" box="[220,562,1154,1180]" pageId="20" pageNumber="42" refString="Skevington, J. &amp; Marshall, S. A. (1998) Systematics of New World Pipunculus (Diptera: Pipunculidae). Thomas Say Publications in Entomology: Monographs, Lanham, 201 pp." type="book" year="1998">
Skevington &amp;
<collectingCountry id="F37F63FBFFF6422AFE2AF4EFFE77FB46" box="[391,495,1154,1180]" name="Marshall Islands" pageId="20" pageNumber="21">Marshall</collectingCountry>
1998
</bibRefCitation>
) should be attributed to
<taxonomicName id="4C6858E8FFF6422AFCFDF4EEFC67FB46" box="[848,1023,1155,1180]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFCFDF4EEFC67FB46" box="[848,1023,1155,1180]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD7236BFFF6422AFF68F4C7FBFFF92E" blockId="20.[151,1437,152,2020]" pageId="20" pageNumber="21">
The male
<typeStatus id="54D39DC9FFF6422AFE93F4C7FE3DFB1E" box="[318,421,1194,1220]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
and additional two male and three female
<typeStatus id="54D39DC9FFF6422AFC05F4C7FB80FB1E" box="[936,1048,1194,1220]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C6858E8FFF6422AFBEEF4C6FB58FB1E" box="[1091,1216,1195,1220]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="phaeton">
<emphasis id="B91CFF79FFF6422AFBEEF4C6FB58FB1E" box="[1091,1216,1195,1220]" italics="true" pageId="20" pageNumber="21">P. phaeton</emphasis>
</taxonomicName>
were obtained for study but are not included in the listing above as they were not used for characterising the taxon due to the reasons stated below:
<collectingCountry id="F37F63FBFFF6422AFEDBF497FD8CFACE" box="[374,532,1274,1300]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
, Oxford: 1ɗ (
<typeStatus id="54D39DC9FFF6422AFD1BF497FCBBFACE" box="[694,803,1274,1300]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
),
<date id="FFD605ABFFF6422AFC9AF497FC27FACE" box="[823,959,1274,1300]" pageId="20" pageNumber="21" value="1909-06-27">27.VI.1909</date>
, Malloch (NHM);
<collectingCountry id="F37F63FBFFF6422AFB30F497FAA2FACE" box="[1181,1338,1274,1300]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
, Dorset, Hamworthy: 1ɗ (
<typeStatus id="54D39DC9FFF6422AFEC8F54FFE48FAE6" box="[357,464,1314,1340]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
),
<date id="FFD605ABFFF6422AFE49F54FFD18FAE6" box="[484,640,1314,1340]" pageId="20" pageNumber="21" value="1894-08-12">12.VIII.1894</date>
, Yerbury (NHM);
<collectingCountry id="F37F63FBFFF6422AFCF6F54FFC60FAE6" box="[859,1016,1314,1340]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
, Glamorgan, Port Talbot, Margam.: 1ɗ (
<typeStatus id="54D39DC9FFF6422AFF60F527FEA0FABE" box="[205,312,1354,1380]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
),
<date id="FFD605ABFFF6422AFEFCF527FE75FABE" box="[337,493,1354,1380]" pageId="20" pageNumber="21" value="1908-08-24">24.VIII.1908</date>
, Yerbury (NHM);
<collectingCountry id="F37F63FBFFF6422AFD78F527FCEFFABE" box="[725,887,1354,1380]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
, Cheshire, Rostherne: 1Ψ (
<typeStatus id="54D39DC9FFF6422AFB68F527FAA8FABE" box="[1221,1328,1354,1380]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
), Britton (NHM);
<collectingCountry id="F37F63FBFFF6422AFEADF51FFE06FA56" box="[256,414,1394,1420]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
, Hants., Woolmer Forest: 1Ψ (
<typeStatus id="54D39DC9FFF6422AFCA4F51FFCECFA56" box="[777,884,1394,1420]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
),
<date id="FFD605ABFFF6422AFC24F51FFB84FA56" box="[905,1052,1394,1420]" pageId="20" pageNumber="21" value="1903-07-12">12.VII.1903</date>
, Yerbury (NHM);
<collectingCountry id="F37F63FBFFF6422AFB54F51FFA0FFA56" box="[1273,1431,1394,1420]" name="United Kingdom" pageId="20" pageNumber="21">Great Britain</collectingCountry>
, Somerset, Sharpham: 1Ψ (
<typeStatus id="54D39DC9FFF6422AFE61F5F7FDAFFA6E" box="[460,567,1434,1460]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
),
<date id="FFD605ABFFF6422AFDE1F5F7FD70FA6E" box="[588,744,1434,1460]" pageId="20" pageNumber="21" value="1922-08-22">22.VIII.1922</date>
, Audcent (NHM). Male and presumably female terminalia are conspecific with
<emphasis id="B91CFF79FFF6422AFE21F5AEFDCFFA06" box="[396,599,1475,1500]" italics="true" pageId="20" pageNumber="21">P. omissinervis—</emphasis>
the male
<typeStatus id="54D39DC9FFF6422AFD69F5AFFCB2FA06" box="[708,810,1474,1500]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
was dissected—but certain outer morphological features concerning pollinosity stand in contrast: e.g. male with abdominal tergites 2–5 almost entirely shining, some weak brown pollinosity may be present laterally or dorsally along anterior margin, or female with frons shining in upper half at least and prescutum and scutum narrowly grey pollinose along anterior margin (up to anterior quarter), laterally almost or right down to wing base and otherwise shining. Despite this, it is proposed not to question the taxonomic status of
<taxonomicName id="4C6858E8FFF6422AFD01F6E6FCBFF97E" box="[684,807,1675,1700]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="phaeton">
<emphasis id="B91CFF79FFF6422AFD01F6E6FCBFF97E" box="[684,807,1675,1700]" italics="true" pageId="20" pageNumber="21">P. phaeton</emphasis>
</taxonomicName>
at present until fresh material for molecular work can be acquired and additional evidence generated. Also, the specimens recorded in the past (see
<bibRefCitation id="EFF95E9AFFF6422AFB7AF6DFFF4CF92E" author="Falk" pageId="20" pageNumber="40" refString="Falk, S. J. &amp; Chandler, P. J. (2005) A review of the scarce and threatened flies of Great Britain. Part 2: Nematocera and Aschiza not dealt with by Falk (1991). Species Status, 2, 1 - 189. Joint Nature Conservation Committee, Peterborough." type="journal article" year="2005">Falk &amp; Chandler 2005</bibRefCitation>
,
<bibRefCitation id="EFF95E9AFFF6422AFF4DF6B7FE3FF92E" author="De" box="[224,423,1754,1780]" pageId="20" pageNumber="40" refString="De Meyer, M. (1989 b) A synthesis of the present knowledge of Pipunculidae (Diptera) in Belgium. In: Comptes rendus du symposium ' invertebres de Belgique', KBIN, Brussels, pp. 373 - 378." type="book chapter" year="1989" yearSuffix="b">De Meyer 1989b</bibRefCitation>
and
<bibRefCitation id="EFF95E9AFFF6422AFE72F6B7FD64F92E" author="De" box="[479,764,1754,1780]" pageId="20" pageNumber="40" refString="De Meyer, M. &amp; Beuk, P. L. T. (2002) Family Pipunculidae. In: Beuk, P. L. T. (Ed.), Checklist of the Diptera of the Netherlands. KNNV Uitgeverij, Utrecht, pp. 196 - 199." type="book chapter" year="2002">De Meyer &amp; Beuk 2002</bibRefCitation>
) should be critically reviewed.
</paragraph>
</subSubSection>
<subSubSection id="C37270E0FFF6422BFF68F76EFA04F80D" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8BD7236BFFF6422BFF68F76EFB13FF00" blockId="20.[151,1437,152,2020]" lastBlockId="21.[151,1436,152,218]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<taxonomicName id="4C6858E8FFF6422AFF68F76EFD34F8C6" ID-CoL="4J7KQ" authority="Kuznetzov, 1990" authorityName="Kuznetzov" authorityYear="1990" box="[197,684,1794,1820]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B91CFF79FFF6422AFF68F76EFE4EF8C6" box="[197,470,1795,1820]" italics="true" pageId="20" pageNumber="21">Pipunculus nartshukae</emphasis>
<bibRefCitation id="EFF95E9AFFF6422AFE4CF76FFD34F8C6" author="Kuznetzov" box="[481,684,1794,1820]" pageId="20" pageNumber="41" refString="Kuznetzov, S. Y. (1990) New species of the family Pipunculidae from Mongolia. Insects of Mongolia, 11, 378 - 393." type="journal article" year="1990">Kuznetzov, 1990</bibRefCitation>
</taxonomicName>
was described from a single male collected in
<collectingCountry id="F37F63FBFFF6422AFB58F76FFAFFF8C6" box="[1269,1383,1794,1820]" name="Mongolia" pageId="20" pageNumber="21">Mongolia</collectingCountry>
and deposited at ZISP (not studied). The surstyli figured in
<bibRefCitation id="EFF95E9AFFF6422AFCB4F747FC74F89E" author="Kuznetzov" box="[793,1004,1834,1860]" pageId="20" pageNumber="41" refString="Kuznetzov, S. Y. (1990) New species of the family Pipunculidae from Mongolia. Insects of Mongolia, 11, 378 - 393." type="journal article" year="1990">Kuznetzov (1990)</bibRefCitation>
(his
<figureCitation id="13533FEEFFF6422AFB8BF747FB13F89E" box="[1062,1163,1834,1860]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,269,1735,1759]" captionTargetBox="[293,1306,203,1708]" captionTargetId="figure@6.[264,1323,197,1709]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Surstyli in strictly dorsal and outer surstylus in lateral view of male Pipunculus violovitshi (1 &amp; 5), Pipunculus elegans (2 &amp; 6), Pipunculus oldenbergi (3 &amp; 7) and Pipunculus zugmayeriae (4 &amp; 8). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/180217/files/figure.png" pageId="20" pageNumber="21">Figs 3–5</figureCitation>
&amp;
<figureCitation id="13533FEEFFF6422AFB02F747FB56F89E" box="[1199,1230,1834,1860]" captionStart="FIGURES 15 – 21" captionStartId="9.[151,269,1850,1874]" captionTargetBox="[278,1299,325,1825]" captionTargetId="figure@9.[264,1323,314,1826]" captionTargetPageId="9" captionText="FIGURES 15 – 21. Surstyli in strictly dorsal and outer surstylus in lateral view of male Pipunculus campestris (15 &amp; 19) and Pipunculus fonsecai (16 – 18 &amp; 20 – 21). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/180219/files/figure.png" pageId="20" pageNumber="21">20</figureCitation>
) seem to be identical to
<taxonomicName id="4C6858E8FFF6422AFF42F73EFE06F8B6" ID-CoL="4J7KY" box="[239,414,1875,1900]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFF42F73EFE06F8B6" box="[239,414,1875,1900]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
, as already pointed out by
<bibRefCitation id="EFF95E9AFFF6422AFD71F73FFBD1F8B6" author="Skevington" box="[732,1097,1874,1900]" pageId="20" pageNumber="42" refString="Skevington, J. &amp; Marshall, S. A. (1998) Systematics of New World Pipunculus (Diptera: Pipunculidae). Thomas Say Publications in Entomology: Monographs, Lanham, 201 pp." type="book" year="1998">
Skevington &amp;
<collectingCountry id="F37F63FBFFF6422AFC2AF73FFC77F8B6" box="[903,1007,1874,1900]" name="Marshall Islands" pageId="20" pageNumber="21">Marshall</collectingCountry>
(1998)
</bibRefCitation>
.
<bibRefCitation id="EFF95E9AFFF6422AFBF9F73FFAB0F8B6" author="Kuznetzov" box="[1108,1320,1874,1900]" pageId="20" pageNumber="41" refString="Kuznetzov, S. Y. (1990) New species of the family Pipunculidae from Mongolia. Insects of Mongolia, 11, 378 - 393." type="journal article" year="1990">Kuznetzov (1990)</bibRefCitation>
separates
<taxonomicName id="4C6858E8FFF6422AFF3AF716FEAFF84E" ID-CoL="4J7KQ" box="[151,311,1915,1940]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="B91CFF79FFF6422AFF3AF716FEAFF84E" box="[151,311,1915,1940]" italics="true" pageId="20" pageNumber="21">P. nartshukae</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6858E8FFF6422AFE2CF716FDAAF84E" ID-CoL="4J7KY" box="[385,562,1915,1940]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFE2CF716FDAAF84E" box="[385,562,1915,1940]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
through more strongly bent surstyli in lateral view and by less narrowing (presumably front) coxae in their apical half—in
<taxonomicName id="4C6858E8FFF6422AFD74F7CEFC10F866" ID-CoL="4J7KY" box="[729,904,1955,1980]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF6422AFD74F7CEFC10F866" box="[729,904,1955,1980]" italics="true" pageId="20" pageNumber="21">P. omissinervis</emphasis>
</taxonomicName>
the apex of the front coxa has approximately half its maximum width, what seems to be characteristic for all species of
<taxonomicName id="4C6858E8FFF6422AFC5AF7A6FBE3F83E" ID-CoL="6QSR" box="[1015,1147,1995,2020]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B91CFF79FFF6422AFC5AF7A6FBE3F83E" box="[1015,1147,1995,2020]" italics="true" pageId="20" pageNumber="21">Pipunculus</emphasis>
</taxonomicName>
studied. Both characters are considered as intraspecific variation and the names therefore synonymized (syn. nov.). Additionally, one male and one female of
<taxonomicName id="4C6858E8FFF7422BFE1CF0ADFDF9FF03" ID-CoL="4J7KY" box="[433,609,192,217]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF7422BFE1CF0ADFDF9FF03" box="[433,609,192,217]" italics="true" pageId="21" pageNumber="22">P. omissinervis</emphasis>
</taxonomicName>
were cited from
<collectingCountry id="F37F63FBFFF7422BFC84F0ADFC03FF00" box="[809,923,192,218]" name="Mongolia" pageId="21" pageNumber="22">Mongolia</collectingCountry>
by
<bibRefCitation id="EFF95E9AFFF7422BFC6BF0ADFB1FFF00" author="Kozanek" box="[966,1159,192,218]" pageId="21" pageNumber="41" refString="Kozanek, M. (1992) Contribution to the pipunculid fauna of Mongolia (Diptera: Pipunculidae). Akitu, New Series, 131, 1 - 16." type="journal article" year="1992">Kozánek (1992)</bibRefCitation>
.
</paragraph>
<caption id="DF1773E3FFF7422BFF3AF6B2FC53F8E6" httpUri="https://zenodo.org/record/180223/files/figure.png" pageId="21" pageNumber="22" targetBox="[275,1313,276,1728]" targetPageId="21">
<paragraph id="8BD7236BFFF7422BFF3AF6B2FC53F8E6" blockId="21.[151,1437,1759,1852]" pageId="21" pageNumber="22">
<emphasis id="B91CFF79FFF7422BFF3AF6B2FEC7F92D" bold="true" box="[151,351,1759,1783]" pageId="21" pageNumber="22">FIGURES 43–54.</emphasis>
Ovipositor in lateral and dorsal view of female
<taxonomicName id="4C6858E8FFF7422BFCC2F68DFBC6F92D" ID-CoL="4J7M7" box="[879,1118,1760,1783]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="violovitshi">
<emphasis id="B91CFF79FFF7422BFCC2F68DFBC6F92D" box="[879,1118,1760,1783]" italics="true" pageId="21" pageNumber="22">Pipunculus violovitshi</emphasis>
</taxonomicName>
(43–44),
<taxonomicName id="4C6858E8FFF7422BFB67F68DFA05F92D" ID-CoL="4J7J2" box="[1226,1437,1760,1783]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B91CFF79FFF7422BFB67F68DFA05F92D" box="[1226,1437,1760,1783]" italics="true" pageId="21" pageNumber="22">Pipunculus elegans</emphasis>
</taxonomicName>
(45–46),
<taxonomicName id="4C6858E8FFF7422BFF54F76FFE73F8C3" ID-CoL="4J7KX" box="[249,491,1794,1817]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="oldenbergi">
<emphasis id="B91CFF79FFF7422BFF54F76FFE73F8C3" box="[249,491,1794,1817]" italics="true" pageId="21" pageNumber="22">Pipunculus oldenbergi</emphasis>
</taxonomicName>
(47–48),
<taxonomicName id="4C6858E8FFF7422BFDF9F76FFCC4F8C3" ID-CoL="6VKRL" box="[596,860,1794,1817]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="zugmayeriae">
<emphasis id="B91CFF79FFF7422BFDF9F76FFCC4F8C3" box="[596,860,1794,1817]" italics="true" pageId="21" pageNumber="22">Pipunculus zugmayeriae</emphasis>
</taxonomicName>
(49–50),
<taxonomicName id="4C6858E8FFF7422BFC68F76FFB32F8C3" ID-CoL="4J7HB" box="[965,1194,1794,1817]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="calceatus">
<emphasis id="B91CFF79FFF7422BFC68F76FFB32F8C3" box="[965,1194,1794,1817]" italics="true" pageId="21" pageNumber="22">Pipunculus calceatus</emphasis>
</taxonomicName>
(51–52) and
<taxonomicName id="4C6858E8FFF7422BFA97F76FFEA5F8E6" ID-CoL="4J7LL" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="stackelbergi">
<emphasis id="B91CFF79FFF7422BFA97F76FFEA5F8E6" italics="true" pageId="21" pageNumber="22">Pipunculus stackelbergi</emphasis>
</taxonomicName>
(53–54, taken from Kuznetzov (1991)). Scale bars = 0.2mm.
</paragraph>
</caption>
<paragraph id="8BD7236BFFF7422BFF68F700FA04F80D" blockId="21.[151,1437,1901,2007]" pageId="21" pageNumber="22">
<bibRef id="52C8B79DFFF7422BFF68F700FA04F80D" pageId="21" pageNumber="22" type="book chapter">
<author id="53D5CB8EFFF7422BFF68F700FEF8F85C" box="[197,352,1901,1927]" fns="I" npo="LF" pageId="21" pageNumber="22">
In Europe,
<emphasis id="B91CFF79FFF7422BFEE7F700FEF8F85C" box="[330,352,1901,1926]" italics="true" pageId="21" pageNumber="22">P.</emphasis>
</author>
omissinervis shows a predominantly boreo-alpine distribution—the presence in
<collectingCountry id="F37F63FBFFF7422BFAB4F700FAC4F85D" box="[1305,1372,1901,1927]" name="Spain" pageId="21" pageNumber="22">Spain</collectingCountry>
indicated in De Meyer (
<year id="FFC06238FFF7422BFE29F7F8FE58F875" box="[388,448,1941,1967]" pageId="21" pageNumber="22">1996</year>
&amp;
<pagination id="B2A1DD9FFFF7422BFE45F7F8FDBCF875" box="[488,548,1941,1967]" pageId="21" pageNumber="22" type="pagination">2005</pagination>
)
<title id="F96CC85DFFF7422BFD98F7F8FC96F875" box="[565,782,1941,1967]" jopScore="0.07058009" pageId="21" pageNumber="22" type="_" typeScore="3">is questioned here</title>
.
<journalOrPublisher id="51A929A7FFF7422BFCB7F7F8FA00F80D" jopScore="0.16691603" pageId="21" pageNumber="22">The taxon should be readily collectable from southern Scandinavia up north and from the mountainous regions of western and central Europe to higher altitudes</journalOrPublisher>
.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="C37270E0FFF44228FF3AF0F5FAECFE88" pageId="22" pageNumber="23" type="description">
<paragraph id="8BD7236BFFF44228FF3AF0F5FAECFE88" blockId="22.[151,1437,152,338]" pageId="22" pageNumber="23">
Towards the more continental parts of central and eastern Europe, the taxon can already be found at lower altitudes. The specimens from
<collectingCountry id="F37F63FBFFF44228FE49F0ADFD19FF00" box="[484,641,192,218]" name="United Kingdom" pageId="22" pageNumber="23">Great Britain</collectingCountry>
(after
<bibRefCitation id="EFF95E9AFFF44228FD79F0ADFC6EFF00" author="Falk" box="[724,1014,192,218]" pageId="22" pageNumber="40" refString="Falk, S. J. &amp; Chandler, P. J. (2005) A review of the scarce and threatened flies of Great Britain. Part 2: Nematocera and Aschiza not dealt with by Falk (1991). Species Status, 2, 1 - 189. Joint Nature Conservation Committee, Peterborough." type="journal article" year="2005">Falk &amp; Chandler (2005)</bibRefCitation>
collected in the southern half incl.
<collectingCountry id="F37F63FBFFF44228FF3AF085FF79FED8" box="[151,225,232,258]" name="United Kingdom" pageId="22" pageNumber="23">Wales</collectingCountry>
, north to Cheshire and
<collectingCountry id="F37F63FBFFF44228FE59F085FDCEFED8" box="[500,598,232,258]" name="Norfolk Island" pageId="22" pageNumber="23">Norfolk</collectingCountry>
),
<collectingCountry id="F37F63FBFFF44228FDC7F085FD57FED8" box="[618,719,232,258]" name="Belgium" pageId="22" pageNumber="23">Belgium</collectingCountry>
(
<bibRefCitation id="EFF95E9AFFF44228FD72F085FC32FED8" author="De" box="[735,938,232,258]" pageId="22" pageNumber="40" refString="De Meyer, M. (1989 b) A synthesis of the present knowledge of Pipunculidae (Diptera) in Belgium. In: Comptes rendus du symposium ' invertebres de Belgique', KBIN, Brussels, pp. 373 - 378." type="book chapter" year="1989" yearSuffix="b">De Meyer 1989b</bibRefCitation>
) and
<collectingCountry id="F37F63FBFFF44228FC41F085FB36FED8" box="[1004,1198,232,258]" name="Netherlands" pageId="22" pageNumber="23">The Netherlands</collectingCountry>
(
<bibRefCitation id="EFF95E9AFFF44228FB12F085FF4EFEF0" author="De" pageId="22" pageNumber="40" refString="De Meyer, M. &amp; Beuk, P. L. T. (2002) Family Pipunculidae. In: Beuk, P. L. T. (Ed.), Checklist of the Diptera of the Netherlands. KNNV Uitgeverij, Utrecht, pp. 196 - 199." type="book chapter" year="2002">De Meyer &amp; Beuk 2002</bibRefCitation>
), cited as
<taxonomicName id="4C6858E8FFF44228FEE6F17DFE5EFEF3" box="[331,454,272,297]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="phaeton">
<emphasis id="B91CFF79FFF44228FEE6F17DFE5EFEF3" box="[331,454,272,297]" italics="true" pageId="22" pageNumber="23">P. phaeton</emphasis>
</taxonomicName>
in the past, do not fit in this pattern. With the present knowledge, they either represent an isolated population of
<taxonomicName id="4C6858E8FFF44228FE5AF155FD3FFE8B" box="[503,679,312,337]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="omissinervis">
<emphasis id="B91CFF79FFF44228FE5AF155FD3FFE8B" box="[503,679,312,337]" italics="true" pageId="22" pageNumber="23">P. omissinervis</emphasis>
</taxonomicName>
or might eventually be revealed to represent a distinct taxon.
</paragraph>
</subSubSection>
</treatment>
</document>
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