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<document id="1E8AE8413ECA738D4CC993B02441BC25" ID-DOI="10.11646/zootaxa.5002.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5122655" ID-ZooBank="5DC2A66A-3F04-42D0-8A8C-F0686054E556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626888359662" checkinUser="plazi" docAuthor="Motamedinia, Behnam, Skevington, Jeffrey H. &amp; Kelso, Scott" docDate="2021" docId="03A887D05F5AFF91FF3EEA98FD914819" docLanguage="en" docName="zootaxa.5002.1.1.pdf" docOrigin="Zootaxa 5002 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomosvaryella docta De Meyer 1995" docType="treatment" docVersion="3" lastPageNumber="25" masterDocId="FF91FFA85F42FF89FFA9EB42FFAF4F30" masterDocTitle="Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species" masterLastPageNumber="103" masterPageNumber="1" pageNumber="25" updateTime="1699012478263" updateUser="plazi">
<mods:mods id="6361DCB040BFCD2697D2DFDD5119D8B4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4F06E0FFBC97F870DF56E1B0749D51B5">
<mods:title id="3841F40BF5626925780F2AEAE1E660CE">Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species</mods:title>
</mods:titleInfo>
<mods:name id="63E174A0B4178E42BA5CB1B5855CBF97" type="personal">
<mods:role id="3338E537FC67E04E21A101BCB8CDF4DF">
<mods:roleTerm id="84280BD1FC96967F6DE8AA3204E4FBD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="30D6C4263E67F7CF4C582130DDC13946">Motamedinia, Behnam</mods:namePart>
</mods:name>
<mods:name id="943FA0B308700D5D36713F01A59A475B" type="personal">
<mods:role id="984FC560B084139E8B436645E5B3FC98">
<mods:roleTerm id="192B3CC958F2683D60FA680ABB1E36FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58CB676333D0DAB130713543E9F86091">Skevington, Jeffrey H.</mods:namePart>
<mods:nameIdentifier id="D5200A08F09235E347B47939E7255699" type="ORCID">0000-0002-1445-9870</mods:nameIdentifier>
<mods:nameIdentifier id="1261EFA469959175E3A34AD46DC85981" type="email">jhskevington@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="D235080C007A7971EC0B3CEDE2DDEF0E" type="personal">
<mods:role id="BFD26C375962438A90FA37F8DED12D46">
<mods:roleTerm id="D057B66D4E69F9237325B1BAB1EAB86C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75EBAB1505FB15C6949009E09DDCDAC2">Kelso, Scott</mods:namePart>
<mods:nameIdentifier id="4B68453E0FE0F9D409E10C9BCBF38105" type="ORCID">0000-0001-5559-9477</mods:nameIdentifier>
<mods:nameIdentifier id="0DA4D80E4E0F5979480CDEADBC8892DA" type="email">scott.kelso@canada.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8E971CAD37AE2045F53CE07D09CB128E">text</mods:typeOfResource>
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<mods:titleInfo id="89ECCCC572CA7126D4358C3C6326DE42">
<mods:title id="6A4A8ACFF64F1D5080259CECE143B90C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5F00B6575181A8E5581B4FF5C9E2390C">
<mods:date id="EAD67BB86FC11192A5401A845557AD22">2021</mods:date>
<mods:detail id="D9F8FE0859F7643FF85AAAC8AFEC30AE" type="pubDate">
<mods:number id="53678B59DB7EDEE6540EF70BC4123460">2021-07-16</mods:number>
</mods:detail>
<mods:detail id="4BA49A7D0101D29A66B2E1F404B242F7" type="volume">
<mods:number id="1E12AA0DDAEE984B27D405B560D8ABA2">5002</mods:number>
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<mods:number id="DE83BD0C4033C366ABCCCDD3B4E35F8A">1</mods:number>
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<mods:start id="A30B26D72DBB70A5A268B5D33C84FC1E">1</mods:start>
<mods:end id="921622FEA902B8016C6982C742E0D79F">103</mods:end>
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<mods:classification id="E8557E456999DCA4D680231C218055E2">journal article</mods:classification>
<mods:identifier id="DEF391A66C247A1E34D8B5F48E136FA2" type="DOI">10.11646/zootaxa.5002.1.1</mods:identifier>
<mods:identifier id="34DB653481B373D6DB798FCE7F7EF57B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FA412EE118A20F71A8FCC49C0EF76E72" type="Zenodo-Dep">5122655</mods:identifier>
<mods:identifier id="B10BAFE39DCC516E615CD771DF870ED6" type="ZooBank">5DC2A66A-3F04-42D0-8A8C-F0686054E556</mods:identifier>
</mods:mods>
<treatment id="03A887D05F5AFF91FF3EEA98FD914819" LSID="urn:lsid:plazi:treatment:03A887D05F5AFF91FF3EEA98FD914819" httpUri="http://treatment.plazi.org/id/03A887D05F5AFF91FF3EEA98FD914819" lastPageNumber="25" pageId="24" pageNumber="25">
<subSubSection id="C31B654D5F5AFF91FF3EEA98FDF74EC4" box="[151,600,474,501]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BBE36C65F5AFF91FF3EEA98FDF74EC4" blockId="24.[151,600,474,537]" box="[151,600,474,501]" pageId="24" pageNumber="25">
<heading id="D0F681AA5F5AFF91FF3EEA98FDF74EC4" bold="true" box="[151,600,474,501]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="4">
<taxonomicName id="4C014D455F5AFF91FF3EEA98FDF74EC4" ID-CoL="7CH4N" authority="De Meyer, 1995" authorityName="De Meyer" authorityYear="1995" box="[151,600,474,501]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="docta">
<emphasis id="B975EAD45F5AFF91FF3EEA98FDF74EC4" bold="true" box="[151,600,474,501]" pageId="24" pageNumber="25">
<emphasis id="B975EAD45F5AFF91FF3EEA98FE214EC5" bold="true" box="[151,398,474,501]" italics="true" pageId="24" pageNumber="25">Tomosvaryella docta</emphasis>
<bibRefCitation id="EF904B375F5AFF91FE3FEA99FDF74EC4" author="De Meyer, M." box="[406,600,474,501]" pageId="24" pageNumber="25" pagination="283 - 319" refId="ref56694" refString="De Meyer, M. (1995) The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae). Spixiana, 18, 283 - 319." type="journal article" year="1995">De Meyer, 1995</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B654D5F5AFF91FF3EEABDFEEC4D29" box="[151,323,510,537]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BBE36C65F5AFF91FF3EEABDFEEC4D29" blockId="24.[151,600,474,537]" box="[151,323,510,537]" pageId="24" pageNumber="25">
<figureCitation id="133A2A435F5AFF91FF3EEABDFEB44D29" box="[151,283,511,537]" captionStart="FIGURE 11" captionStartId="25.[151,250,1880,1904]" captionTargetBox="[170,1412,185,1845]" captionTargetId="figure-21@25.[151,1436,181,1857]" captionTargetPageId="25" captionText="FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122695" httpUri="https://zenodo.org/record/5122695/files/figure.png" pageId="24" pageNumber="25">Figs 11A–F</figureCitation>
,
<figureCitation id="133A2A435F5AFF91FE8EEABCFEEC4D29" box="[295,323,510,537]" captionStart="FIGURE 52" captionStartId="86.[151,250,1598,1622]" captionTargetBox="[151,1436,932,1574]" captionTargetId="figure-68@86.[151,1436,932,1574]" captionTargetPageId="86" captionText="FIGURE 52. Distribution of Tomosvaryella coquilletti, T. debruyni, T. docta in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122775" httpUri="https://zenodo.org/record/5122775/files/figure.png" pageId="24" pageNumber="25">52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B654D5F5AFF91FF3EE904FCA5498D" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BBE36C65F5AFF91FF3EE904FD964DC1" blockId="24.[151,1437,582,1834]" pageId="24" pageNumber="25">
<emphasis id="B975EAD45F5AFF91FF3EE904FEBB4D50" bold="true" box="[151,276,582,608]" pageId="24" pageNumber="25">Diagnosis:</emphasis>
This species can be recognized by the shape of surstyli in dorsal view, both broadened in basal half; right surstylus narrowed moderately in in apical half, left surstylus narrowed abruptly in apical half (
<figureCitation id="133A2A435F5AFF91FA8CE929FA204DB5" box="[1317,1423,619,645]" captionStart="FIGURE 11" captionStartId="25.[151,250,1880,1904]" captionTargetBox="[170,1412,185,1845]" captionTargetId="figure-21@25.[151,1436,181,1857]" captionTargetPageId="25" captionText="FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122695" httpUri="https://zenodo.org/record/5122695/files/figure.png" pageId="24" pageNumber="25">Fig. 11A</figureCitation>
); epandrium longer than wide (
<figureCitation id="133A2A435F5AFF91FE49E9CDFDE94D99" box="[480,582,655,681]" captionStart="FIGURE 11" captionStartId="25.[151,250,1880,1904]" captionTargetBox="[170,1412,185,1845]" captionTargetId="figure-21@25.[151,1436,181,1857]" captionTargetPageId="25" captionText="FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122695" httpUri="https://zenodo.org/record/5122695/files/figure.png" pageId="24" pageNumber="25">Fig. 11A</figureCitation>
); subepandrial sclerite wide at the base in ventral view (
<figureCitation id="133A2A435F5AFF91FB1FE9CDFAB44D99" box="[1206,1307,655,681]" captionStart="FIGURE 11" captionStartId="25.[151,250,1880,1904]" captionTargetBox="[170,1412,185,1845]" captionTargetId="figure-21@25.[151,1436,181,1857]" captionTargetPageId="25" captionText="FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122695" httpUri="https://zenodo.org/record/5122695/files/figure.png" pageId="24" pageNumber="25">Fig. 11B</figureCitation>
); gonopods with an unequal projection, right is farther than left one (
<figureCitation id="133A2A435F5AFF91FC88E9F1FC274DFD" box="[801,904,691,717]" captionStart="FIGURE 11" captionStartId="25.[151,250,1880,1904]" captionTargetBox="[170,1412,185,1845]" captionTargetId="figure-21@25.[151,1436,181,1857]" captionTargetPageId="25" captionText="FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122695" httpUri="https://zenodo.org/record/5122695/files/figure.png" pageId="24" pageNumber="25">Fig. 11B</figureCitation>
); phallus trifid, one with some small teeth-like projections in lateral view (
<figureCitation id="133A2A435F5AFF91FE63E995FD824DC1" box="[458,557,727,753]" captionStart="FIGURE 11" captionStartId="25.[151,250,1880,1904]" captionTargetBox="[170,1412,185,1845]" captionTargetId="figure-21@25.[151,1436,181,1857]" captionTargetPageId="25" captionText="FIGURE 11. Male genitalia of Tomosvaryella docta (CNCD137579, EAD) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view, F) phallus and phallic guide in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122695" httpUri="https://zenodo.org/record/5122695/files/figure.png" pageId="24" pageNumber="25">Fig. 11F</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C65F5AFF91FF6EE9B8FCA5498D" blockId="24.[151,1437,582,1834]" pageId="24" pageNumber="25">
<emphasis id="B975EAD45F5AFF91FF6EE9B8FE654C24" bold="true" box="[199,458,762,788]" pageId="24" pageNumber="25">Specimens examined:</emphasis>
<collectingCountry id="F31676565F5AFF91FE78E9B9FDB54C25" box="[465,538,763,789]" name="Iran" pageId="24" pageNumber="25">IRAN</collectingCountry>
:
<collectingRegion id="49C5F8245F5AFF91FD8CE9B9FD0F4C25" box="[549,672,763,789]" country="Iran" name="Khuzestan" pageId="24" pageNumber="25">Khuzestan</collectingRegion>
: Shush,
<geoCoordinate id="EE3550015F5AFF91FD57E9B9FCCD4C25" box="[766,866,763,789]" degrees="32" direction="north" minutes="06" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="32.1">32°06’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FCC7E9B9FC614C25" box="[878,974,763,789]" degrees="48" direction="east" minutes="26" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="48.433334">48°26’E</geoCoordinate>
,
<quantity id="4CF99B235F5AFF91FC73E9B8FBBB4C25" box="[986,1044,762,789]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="24" pageNumber="25" unit="m" value="55.0">55 m</quantity>
a.s.l.,
<date id="FFBF10065F5AFF91FBF6E9B9FABD4C25" box="[1119,1298,763,789]" pageId="24" pageNumber="25" value="2013-08-29" valueMax="2013-08-31" valueMin="2013-08-29">29–31.viii.2013</date>
, M. Parchami-Araghi, Malaise trap, JSS52189 (
<specimenCount id="9D07FD4F5F5AFF91FD90E85DFDF54C09" box="[569,602,799,825]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
, CNC); Sistan &amp; Baluchestan: Daman,
<geoCoordinate id="EE3550015F5AFF91FBB7E85DFB2E4C09" box="[1054,1153,799,825]" degrees="27" direction="north" minutes="24" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="27.4">27°24’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FB24E85DFB424C09" box="[1165,1261,799,825]" degrees="60" direction="east" minutes="49" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="60.816666">60°49’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FB53E85DFA374C09" box="[1274,1432,799,825]" pageId="24" pageNumber="25" value="2016-07-02" valueMax="2016-07-12" valueMin="2016-07-02">2–12.vii.2016</date>
, H. Davari, Malaise trap, JSS51822 (
<specimenCount id="9D07FD4F5F5AFF91FD95E801FDCF4C6D" box="[572,608,835,861]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC); Saravan,
<geoCoordinate id="EE3550015F5AFF91FC87E800FC3E4C6D" box="[814,913,834,861]" degrees="27" direction="north" minutes="25" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="27.416666">27°25’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FC09E801FBAF4C6D" box="[928,1024,835,861]" degrees="62" direction="east" minutes="17" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="62.283333">62°17’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FBA7E801FB2E4C6D" box="[1038,1153,835,861]" pageId="24" pageNumber="25" value="2016-10-17">17.x.2016</date>
, F. Hamzavi, sweep net, JSS51827 (
<specimenCount id="9D07FD4F5F5AFF91FEB4E825FE914CB1" box="[285,318,871,897]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
, CNC); Zabol, Dostmohammad,
<geoCoordinate id="EE3550015F5AFF91FD16E825FC8C4CB1" box="[703,803,871,897]" degrees="31" direction="north" minutes="09" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="31.15">31°09’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FC98E825FC3E4CB1" box="[817,913,871,897]" degrees="61" direction="east" minutes="43" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="61.716667">61°43’E</geoCoordinate>
,
<quantity id="4CF99B235F5AFF91FC09E825FC454CB1" box="[928,1002,871,897]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="24" pageNumber="25" unit="m" value="470.0">470 m</quantity>
a.s.l.,
<date id="FFBF10065F5AFF91FB90E825FB1C4CB1" box="[1081,1203,871,897]" pageId="24" pageNumber="25" value="2016-06-29">29.vi.2016</date>
, M. Enaiatnia, Mal- aise trap, JSS51933 (
<specimenCount id="9D07FD4F5F5AFF91FE23E8C9FE034C95" box="[394,428,907,933]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
, CNC); Zabol, 31°
<geoCoordinate id="EE3550015F5AFF91FD2DE8C9FD664C95" box="[644,713,907,933]" degrees="0" direction="north" minutes="3" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="0.05">0 3’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FD7EE8C9FC984C95" box="[727,823,907,933]" degrees="61" direction="east" minutes="22" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="61.366665">61°22’E</geoCoordinate>
,
<quantity id="4CF99B235F5AFF91FCEDE8C9FC234C95" box="[836,908,907,933]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="24" pageNumber="25" unit="m" value="480.0">480 m</quantity>
a.s.l.,
<date id="FFBF10065F5AFF91FC71E8C9FBF54C95" box="[984,1114,907,933]" pageId="24" pageNumber="25" value="2016-07-10">10.vii.2016</date>
, M. Enaiatnia, Malaise trap, JSS51860 (
<specimenCount id="9D07FD4F5F5AFF91FEB4E8EDFEEE4CF9" box="[285,321,943,969]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<geoCoordinate id="EE3550015F5AFF91FE0EE8EDFDA44CF9" box="[423,523,943,969]" degrees="31" direction="north" minutes="02" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="31.033333">31°02’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FDB2E8EDFDD44CF9" box="[539,635,943,969]" degrees="61" direction="east" minutes="32" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="61.533333">61°32’E</geoCoordinate>
,
<quantity id="4CF99B235F5AFF91FD22E8EDFD794CF9" box="[651,726,943,969]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="24" pageNumber="25" unit="m" value="470.0">470 m</quantity>
a.s.l.,
<date id="FFBF10065F5AFF91FC81E8EDFC3B4CF9" box="[808,916,943,969]" pageId="24" pageNumber="25" value="2016-09-02">2.ix.2016</date>
, M. Enaiatnia, Malaise trap, JSS51907 (
<specimenCount id="9D07FD4F5F5AFF91FADCE8EDFA394CF9" box="[1397,1430,943,969]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
, CNC);
<collectingRegion id="49C5F8245F5AFF91FF42E891FEE54CDD" box="[235,330,979,1005]" country="Iran" name="Kerman" pageId="24" pageNumber="25">Kerman</collectingRegion>
: Fariab, Horesfandi,
<geoCoordinate id="EE3550015F5AFF91FD93E891FD324CDD" box="[570,669,979,1005]" degrees="28" direction="north" minutes="08" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="28.133333">28°08’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FD02E890FCA54CDD" box="[683,778,978,1005]" degrees="57" direction="east" minutes="16" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="57.266666">57°16’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FCB1E890FC3A4CDD" box="[792,917,978,1005]" pageId="24" pageNumber="25" value="2016-08-05">5.viii.2016</date>
, M. Arabzadeh, Malaise trap, JSS51922 (
<specimenCount id="9D07FD4F5F5AFF91FADAE891FA384CDD" box="[1395,1431,979,1005]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<collectingCountry id="F31676565F5AFF91FF41E8B5FEE14B21" box="[232,334,1015,1041]" name="Israel" pageId="24" pageNumber="25">ISRAEL</collectingCountry>
: Hazeva Field School,
<geoCoordinate id="EE3550015F5AFF91FDE5E8B5FD004B21" box="[588,687,1015,1041]" degrees="30" direction="north" minutes="43" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="30.716667">30°43’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FD13E8B4FCB64B21" box="[698,793,1014,1041]" degrees="35" direction="east" minutes="15" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="35.25">35°15’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FC8DE8B5FC264B21" box="[804,905,1015,1041]" pageId="24" pageNumber="25" value="1993-10-02">2.x.1993</date>
, S. Plotkin, Malaise trap, JSS51704 (
<specimenCount id="9D07FD4F5F5AFF91FA8CE8B5FAE64B21" box="[1317,1353,1015,1041]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, TAU);
<date id="FFBF10065F5AFF91FF3EEF58FF544B05" box="[151,251,1050,1077]" pageId="24" pageNumber="25" value="1997-10-05">5.x.1997</date>
, S. Plotkin, Malaise trap, JSS52080 (
<specimenCount id="9D07FD4F5F5AFF91FD1DEF59FD764B05" box="[692,729,1051,1077]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, TAU); Hazeva, Field School,
<geoCoordinate id="EE3550015F5AFF91FBEDEF59FB084B05" box="[1092,1191,1051,1077]" degrees="30" direction="north" minutes="43" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="30.716667">30°43’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FB1EEF58FAB94B05" box="[1207,1302,1050,1077]" degrees="35" direction="east" minutes="15" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="35.25">35°15’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FA8FEF59FA384B05" box="[1318,1431,1051,1077]" pageId="24" pageNumber="25" value="1997-10-11">11.x.1997</date>
, A. Maklakov, Malaise trap, JSS51769–70 (
<specimenCount id="9D07FD4F5F5AFF91FD2DEF7DFD054B69" box="[644,682,1087,1113]" pageId="24" pageNumber="25" type="male">2♂</specimenCount>
, TAU);
<collectingCountry id="F31676565F5AFF91FCA0EF7DFCD54B69" box="[777,890,1087,1113]" name="Cyprus" pageId="24" pageNumber="25">CYPRUS</collectingCountry>
:
<collectingRegion id="49C5F8245F5AFF91FC2EEF7DFC4B4B69" box="[903,996,1087,1113]" country="Cyprus" name="Kyrenia" pageId="24" pageNumber="25">Kyrenia</collectingRegion>
,
<geoCoordinate id="EE3550015F5AFF91FC58EF7CFBFB4B69" box="[1009,1108,1086,1113]" degrees="35" direction="north" minutes="21" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="35.35">35°21’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FBCBEF7DFB6D4B69" box="[1122,1218,1087,1113]" degrees="33" direction="east" minutes="09" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="33.15">33°09’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FB66EF7DFADC4B69" box="[1231,1395,1087,1113]" pageId="24" pageNumber="25" value="2017-09-10" valueMax="2017-09-17" valueMin="2017-09-10">10–17.ix.2017</date>
, O. Ozden, Malaise trap, JSS52344 (
<specimenCount id="9D07FD4F5F5AFF91FDBBEF21FD994B4D" box="[530,566,1123,1149]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FD30EF21FCEA4B4D" box="[665,837,1123,1149]" pageId="24" pageNumber="25" value="2017-07-16" valueMax="2017-07-30" valueMin="2017-07-16">16–30.vii.2017</date>
, O. Ozden, Malaise trap, JSS52325, JSS52355 (
<specimenCount id="9D07FD4F5F5AFF91FADBEF21FA374B4D" box="[1394,1432,1123,1149]" pageId="24" pageNumber="25" type="male">2♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FF44EFC5FE374B91" box="[237,408,1159,1185]" pageId="24" pageNumber="25" value="2017-09-24" valueMax="2017-10-01" valueMin="2017-09-24">24.ix–1.x.2017</date>
, O. Ozden, Malaise trap, JSS52354 (
<specimenCount id="9D07FD4F5F5AFF91FCF9EFC5FCDB4B91" box="[848,884,1159,1185]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FC73EFC5FB1D4B91" box="[986,1202,1159,1185]" pageId="24" pageNumber="25" value="2017-07-30" valueMax="2017-08-13" valueMin="2017-07-30">30.vii–13.viii.2017</date>
, O. Ozden, Malaise trap, JSS52366 (
<specimenCount id="9D07FD4F5F5AFF91FEF8EFE9FEDA4BF5" box="[337,373,1195,1221]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FE78EFE9FDDF4BF5" box="[465,624,1195,1221]" pageId="24" pageNumber="25" value="2017-07-09" valueMax="2017-07-16" valueMin="2017-07-09">9–16.vii.2017</date>
, O. Ozden, Malaise trap, JSS52330 (
<specimenCount id="9D07FD4F5F5AFF91FBA2EFE9FB804BF5" box="[1035,1071,1195,1221]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC); UNITED ARAB EMIR- ATES: Abu Dhabi: Al Wathba Wetland Reserve,
<geoCoordinate id="EE3550015F5AFF91FD7FEF8CFC964BD9" box="[726,825,1230,1257]" degrees="24" direction="north" minutes="15" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="24.25">24°15’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FCE0EF8CFC064BD9" box="[841,937,1230,1257]" degrees="54" direction="east" minutes="37" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="54.616665">54°37’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FC11EF8CFBAD4BD9" box="[952,1026,1230,1257]" pageId="24" pageNumber="25" value="2015-01">i.2015</date>
, A. Saji &amp; A. van Harten, Malaise trap, CNC470746 (
<specimenCount id="9D07FD4F5F5AFF91FEDEEFB1FE344A3D" box="[375,411,1267,1293]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FDA9EFB0FDFE4A3D" box="[512,593,1266,1293]" pageId="24" pageNumber="25" value="2015-02">ii.2015</date>
, A. Saji &amp; A. van Harten, Malaise trap, CNC470512, CNC470572 (
<specimenCount id="9D07FD4F5F5AFF91FADBEFB1FA374A3D" box="[1394,1432,1267,1293]" pageId="24" pageNumber="25" type="male">2♂</specimenCount>
, USNM);
<date id="FFBF10065F5AFF91FEA8EE54FEF64A01" box="[257,345,1302,1329]" pageId="24" pageNumber="25" value="2015-03">iii.2015</date>
, A. Saji &amp; A. van Harten, Malaise trap, CNC470645, CNC470654, CNC470679–80 (
<specimenCount id="9D07FD4F5F5AFF91FAB6EE55FAEA4A01" box="[1311,1349,1303,1329]" pageId="24" pageNumber="25" type="male">4♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FF3EEE79FF434A65" box="[151,236,1339,1365]" pageId="24" pageNumber="25" value="2013-04">iv.2013</date>
, A. Saji &amp; A. van Harten, Malaise trap, CNC470729 (
<specimenCount id="9D07FD4F5F5AFF91FCF3EE79FCD14A65" box="[858,894,1339,1365]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FC76EE78FB9A4A65" box="[991,1077,1338,1365]" pageId="24" pageNumber="25" value="2015-04">iv.2015</date>
, A. Saji &amp; A. van Harten, Mal- aise trap, CNC470808 (
<specimenCount id="9D07FD4F5F5AFF91FE0DEE1DFE674A49" box="[420,456,1375,1401]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FD8FEE1DFDD24A49" box="[550,637,1375,1401]" pageId="24" pageNumber="25" value="2013-09">ix.2013</date>
, A. Saji &amp; A. van Harten, Malaise trap, CNC465503, CNC470708–10, CNC470713, CNC470716 (
<specimenCount id="9D07FD4F5F5AFF91FE78EEC1FE594AAD" box="[465,502,1411,1437]" pageId="24" pageNumber="25" type="male">6♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FDFDEEC1FD034AAD" box="[596,684,1411,1437]" pageId="24" pageNumber="25" value="2013-11">xi.2013</date>
, A. Saji &amp; A. van Harten, Malaise trap, CNC470721, CNC470725, CNC470728 (
<specimenCount id="9D07FD4F5F5AFF91FE9DEEE5FEF64AF1" box="[308,345,1447,1473]" pageId="24" pageNumber="25" type="male">3♂</specimenCount>
, CNC); Al Bida’a Protected Area,
<geoCoordinate id="EE3550015F5AFF91FD70EEE5FC944AF1" box="[729,827,1447,1473]" degrees="23" direction="north" minutes="36" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="23.6">23°36’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FCEFEEE4FC094AF1" box="[838,934,1446,1473]" degrees="53" direction="east" minutes="58" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="53.966667">53°58’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FC18EEE5FB8B4AF1" box="[945,1060,1447,1473]" pageId="24" pageNumber="25" value="2018-10-29">29.x.2018</date>
, A. Saji &amp; A. van Harten, Malaise trap, JSS52267–70 (
<specimenCount id="9D07FD4F5F5AFF91FED6EE89FE0A4AD5" box="[383,421,1483,1509]" pageId="24" pageNumber="25" type="male">4♂</specimenCount>
, CNC); Al Tawi Protected Area,
<geoCoordinate id="EE3550015F5AFF91FC8DEE88FC284AD5" box="[804,903,1482,1509]" degrees="23" direction="north" minutes="50" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="23.833334">23°50’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FC3CEE88FC5A4AD5" box="[917,1013,1482,1509]" degrees="53" direction="east" minutes="10" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="53.166668">53°10’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FBAAEE89FBC04AD5" box="[1027,1135,1483,1509]" pageId="24" pageNumber="25" value="2018-11-01">1.xi.2018</date>
, A. Saji &amp; A. van Harten, Malaise trap, JSS52295–98 (
<specimenCount id="9D07FD4F5F5AFF91FE49EEADFDA94939" box="[480,518,1519,1545]" pageId="24" pageNumber="25" type="male">4♂</specimenCount>
, CNC); Barqa Al Suqoor Protected Area,
<geoCoordinate id="EE3550015F5AFF91FC4EEEADFBE54939" box="[999,1098,1519,1545]" degrees="23" direction="north" minutes="60" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="24.0">23°60’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FBFEEEACFB174939" box="[1111,1208,1518,1545]" degrees="52" direction="east" minutes="41" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="52.683334">52°41’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FB6CEEADFAEE4939" box="[1221,1345,1519,1545]" pageId="24" pageNumber="25" value="2018-03-20">20.iii.2018</date>
, A. Saji &amp; A. van Harten, Malaise trap, JSS52260 (
<specimenCount id="9D07FD4F5F5AFF91FDD0ED51FD32491D" box="[633,669,1555,1581]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FD52ED51FCC2491D" box="[763,877,1555,1581]" pageId="24" pageNumber="25" value="2018-10-29">29.x.2018</date>
, A. Saji &amp; A. van Harten, Malaise trap, JSS52282 (
<specimenCount id="9D07FD4F5F5AFF91FF08ED75FF6A4961" box="[161,197,1591,1617]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC); Wadi Wurayah farm,
<geoCoordinate id="EE3550015F5AFF91FDB4ED74FD2F4961" box="[541,640,1590,1617]" degrees="25" direction="north" minutes="14" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="25.233334">25°14’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FD24ED74FD434961" box="[653,748,1590,1617]" degrees="56" direction="east" minutes="11" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="56.183334">56°11’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FD53ED75FC0F4961" box="[762,928,1591,1617]" pageId="24" pageNumber="25" value="2009-03-17" valueMax="2009-03-24" valueMin="2009-03-17">17–24.iii.2009</date>
, A. van Harten, Malaise trap, CNCD137123 (
<specimenCount id="9D07FD4F5F5AFF91FF08ED19FF6A4945" box="[161,197,1627,1653]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FE82ED19FE4D4945" box="[299,482,1627,1653]" pageId="24" pageNumber="25" value="2009-04-19" valueMax="2009-05-19" valueMin="2009-04-19">19.iv–19.v.2009</date>
, A. van Harten, Malaise trap, CNCD137544 (
<specimenCount id="9D07FD4F5F5AFF91FC57ED19FB8D4945" box="[1022,1058,1627,1653]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, CNC);
<date id="FFBF10065F5AFF91FB2EED19FAEF4945" box="[1159,1344,1627,1653]" pageId="24" pageNumber="25" value="2009-05-31" valueMax="2009-06-14" valueMin="2009-05-31">31.v–14.vi.2009</date>
, A. van Harten, Malaise trap, CNCD137579, CNCD137582 (
<specimenCount id="9D07FD4F5F5AFF91FD40ED3DFCA049A9" box="[745,783,1663,1689]" pageId="24" pageNumber="25" type="male">2♂</specimenCount>
, EAD); Wadi Wurayah,
<geoCoordinate id="EE3550015F5AFF91FBB4ED3CFB2F49A9" box="[1053,1152,1662,1689]" degrees="25" direction="north" minutes="14" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="25.233334">25°14’N</geoCoordinate>
,
<geoCoordinate id="EE3550015F5AFF91FB22ED3CFB4549A9" box="[1163,1258,1662,1689]" degrees="56" direction="east" minutes="11" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="56.183334">56°11’E</geoCoordinate>
,
<date id="FFBF10065F5AFF91FB5CED3DFA3749A9" box="[1269,1432,1663,1689]" pageId="24" pageNumber="25" value="2009-04-19" valueMax="2009-04-26" valueMin="2009-04-19">19–26.iv.2009</date>
, A. van Harten, Malaise trap, CNCD160581 (
<specimenCount id="9D07FD4F5F5AFF91FD26EDE1FD1C498D" box="[655,691,1699,1725]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
, EAD).
</paragraph>
</subSubSection>
<subSubSection id="C31B654D5F5AFF91FF6EED84FD914819" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BBE36C65F5AFF91FF6EED84FA8449D1" blockId="24.[151,1437,582,1834]" box="[199,1323,1734,1761]" pageId="24" pageNumber="25">
<emphasis id="B975EAD45F5AFF91FF6EED84FECC49D0" bold="true" box="[199,355,1734,1760]" pageId="24" pageNumber="25">Distribution:</emphasis>
<collectingCountry id="F31676565F5AFF91FEC3ED85FE1149D1" box="[362,446,1735,1761]" name="Cyprus" pageId="24" pageNumber="25">Cyprus</collectingCountry>
,
<collectingCountry id="F31676565F5AFF91FE60ED85FDA049D1" box="[457,527,1735,1761]" name="Egypt" pageId="24" pageNumber="25">Egypt</collectingCountry>
,
<collectingCountry id="F31676565F5AFF91FDB3ED85FDE549D1" box="[538,586,1735,1761]" name="Iran" pageId="24" pageNumber="25">Iran</collectingCountry>
,
<collectingCountry id="F31676565F5AFF91FDFCED85FD3949D1" box="[597,662,1735,1761]" name="Israel" pageId="24" pageNumber="25">Israel</collectingCountry>
,
<collectingCountry id="F31676565F5AFF91FD08ED85FD4B49D1" box="[673,740,1735,1761]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
,
<collectingCountry id="F31676565F5AFF91FD46ED85FC4C49D1" box="[751,995,1735,1761]" name="United Arab Emirates" pageId="24" pageNumber="25">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="EF904B375F5AFF91FC58ED85FB1B49D1" author="Skevington, J. H." box="[1009,1204,1735,1761]" pageId="24" pageNumber="25" refId="ref58350" refString="Skevington, J. H. (2020) &quot; Pipunculidae &quot;. Canadian National Collection of Insects, Arachnids and Nematodes database. Available from: https: // cnc. agr. gc. ca / taxonomy / Taxonomy. php? id = 18557 (accessed 18 October 2020) [public version of Pipunculidae catalogue maintained by J. H. Skevington]" type="url" year="2020">Skevington 2020</bibRefCitation>
) (
<figureCitation id="133A2A435F5AFF91FB63ED85FAB049D1" box="[1226,1311,1734,1761]" captionStart="FIGURE 52" captionStartId="86.[151,250,1598,1622]" captionTargetBox="[151,1436,932,1574]" captionTargetId="figure-68@86.[151,1436,932,1574]" captionTargetPageId="86" captionText="FIGURE 52. Distribution of Tomosvaryella coquilletti, T. debruyni, T. docta in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122775" httpUri="https://zenodo.org/record/5122775/files/figure.png" pageId="24" pageNumber="25">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C65F5AFF91FF6EEDA9FD914819" blockId="24.[151,1437,582,1834]" pageId="24" pageNumber="25">
<emphasis id="B975EAD45F5AFF91FF6EEDA9FEA74835" bold="true" box="[199,264,1771,1797]" pageId="24" pageNumber="25">Note:</emphasis>
DNA barcodes of this species are genetically most similar to
<taxonomicName id="4C014D455F5AFF91FC60EDA9FBFB4835" authorityName="Kuznetzov" authorityYear="1994" box="[969,1108,1771,1797]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gazliensis">
<emphasis id="B975EAD45F5AFF91FC60EDA9FBFB4835" box="[969,1108,1771,1797]" italics="true" pageId="24" pageNumber="25">T. gazliensis</emphasis>
</taxonomicName>
, differing by 7.4% (pairwise divergence) (Supplementary
<tableCitation id="C683037D5F5AFF91FE72EC4DFD824819" box="[475,557,1807,1833]" captionStart="TABLE 1" captionStartId="5.[151,239,810,836]" captionText="TABLE 1. Cytochrome c oxidase subunit I (COI) and internal transcribed spacer (ITS) primers." pageId="24" pageNumber="25">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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