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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1192.111275" ID-Pensoft-Pub="1313-2970-1192-111" ID-Pensoft-UUID="436A6870AC145BE79A56D786FB1C9716" ID-ZooBank="94FB491FB5A54514A1EF062B6A216D11" ModsDocID="1313-2970-1192-111" checkinTime="1708593701100" checkinUser="pensoft" docAuthor="Doan, Lydia M., Miller, James S., Brown, John W., Forister, Matthew L. &amp; Dyer, Lee A." docDate="2024" docId="E71494D918B25E2C9422686FD1FDF33B" docLanguage="en" docName="ZooKeys 1192: 111-140" docOrigin="ZooKeys 1192" docPubDate="2024-02-21" docSource="http://dx.doi.org/10.3897/zookeys.1192.111275" docTitle="Eois beebei Fletcher 1952, stat. rev." docType="treatment" docVersion="1" id="436A6870AC145BE79A56D786FB1C9716" lastPageNumber="111" masterDocId="436A6870AC145BE79A56D786FB1C9716" masterDocTitle="Two new species of the hyperdiverse geometrid moth genus Eois (Lepidoptera, Geometridae, Larentiinae) from Ecuador, with descriptions of early stages" masterLastPageNumber="140" masterPageNumber="111" pageNumber="111" updateTime="1708593701100" updateUser="pensoft">
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<mods:title id="0301BC62E230DB2C11E650BB90AB1979">Two new species of the hyperdiverse geometrid moth genus Eois (Lepidoptera, Geometridae, Larentiinae) from Ecuador, with descriptions of early stages</mods:title>
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<mods:namePart id="0CAD498F07ECB3681BCA0B19FD553076">Doan, Lydia M.</mods:namePart>
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<mods:namePart id="F98F8AFC659784208FD37708744552FC">Miller, James S.</mods:namePart>
<mods:affiliation id="C6ABA24DE79FB45A6B2A58F04178D7BF">Entomology Department, American Museum of Natural History, New York, NY, 10024, USA</mods:affiliation>
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<mods:namePart id="06664EC4E158D96C4F4A522167ECA72B">Brown, John W.</mods:namePart>
<mods:nameIdentifier id="F202425AF92201833EABF95ABA16CB73" type="ORCID">https://orcid.org/0000-0001-5610-9855</mods:nameIdentifier>
<mods:affiliation id="3483A28865FAAC20CD71AC995C4E5D07">Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington, DC, 20560, USA</mods:affiliation>
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<mods:namePart id="5A0567FAFB9D1869E453D81F4D88E8D1">Forister, Matthew L.</mods:namePart>
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<mods:affiliation id="B3907BB5341FA88F4CB3FE44494AA18D">Department of Biology, Ecology, Evolution and Conservation of Biology, University of Nevada, Reno, NV 89557, USA</mods:affiliation>
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<mods:namePart id="687A52A9914975E784AB9759BDCB5E71">Dyer, Lee A.</mods:namePart>
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<mods:affiliation id="C382ECD9A1DA5874410E411EBA7F1469">Department of Biology, Ecology, Evolution and Conservation of Biology, University of Nevada, Reno, NV 89557, USA</mods:affiliation>
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<mods:date id="003AB221DB36C2E86A1D39FF2FAE2EC4">2024</mods:date>
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<treatment id="E71494D918B25E2C9422686FD1FDF33B" LSID="urn:lsid:plazi:treatment:E71494D918B25E2C9422686FD1FDF33B" httpUri="http://treatment.plazi.org/id/E71494D918B25E2C9422686FD1FDF33B" lastPageNumber="111" pageId="0" pageNumber="111">
<subSubSection id="87C3F82BE9E7543BEDF2F92D49BF93E6" pageId="0" pageNumber="111" type="nomenclature">
<paragraph id="BAC686E05A3F6CE240397A585F10571D" pageId="0" pageNumber="111">
<taxonomicName id="2200C021610D3CC013938F950E2CAAA5" ID-CoL="98736" LSID="E71494D9-18B2-5E2C-9422-686FD1FDF33B" authority="Fletcher, 1952" authorityName="Fletcher" authorityYear="1952" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois beebei" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="beebei" status="stat. rev.">Eois beebei Fletcher, 1952</taxonomicName>
<taxonomicNameLabel id="BFE9313D42F2C34A8249A76486A66A18" pageId="0" pageNumber="111">stat. rev.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="9D0F16A2F0FCBB5B6889855AC8161CDA" pageId="0" pageNumber="111" type="description">
<paragraph id="392F09ECB050DF68345CBC5DEFDBB914" pageId="0" pageNumber="111">
<figureCitation id="3C36102FBA028EE5BE2E5118FD4B03CB" captionStart="Figures 15, 16" captionStartId="F5" captionText="Figures 15, 16. Eois beebei 15 holotype upperside (NHMUK) 16 male genitalia (NHMUK)." figureDoi="10.3897/zookeys.1192.111275.figures15-16" httpUri="https://binary.pensoft.net/fig/990478" pageId="0" pageNumber="111">Figs 15</figureCitation>
<figureCitation id="4B9EB7E89E5C23AA246E1D0F1F025034" captionStart="Figures 15, 16" captionStartId="F5" captionText="Figures 15, 16. Eois beebei 15 holotype upperside (NHMUK) 16 male genitalia (NHMUK)." figureDoi="10.3897/zookeys.1192.111275.figures15-16" httpUri="https://binary.pensoft.net/fig/990478" pageId="0" pageNumber="111">, 16</figureCitation>
</paragraph>
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<subSubSection id="9757771998C60D5F70B12869553F975B" pageId="0" pageNumber="111" type="reference_group">
<paragraph id="78D5F42B176AB56AF791228B9089E928" pageId="0" pageNumber="111">
<taxonomicName id="55337C5F087C2BB4EEFA88B6970F0BD0" authorityName="Doan &amp; Miller &amp; Brown &amp; Forister &amp; Dyer" authorityYear="2024" class="Insecta" family="Geometridae" genus="Racheospila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Racheospila beebei" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="beebei" status="stat. rev.">Racheospila beebei</taxonomicName>
Fletcher, 1952: 101.
</paragraph>
<paragraph id="3397BCC1C287C74BE73E5F4A005B5E1F" pageId="0" pageNumber="111">
<taxonomicName id="57DF14FA4EE932B8F4A4EDDDCEFE1F8F" authorityName="Doan &amp; Miller &amp; Brown &amp; Forister &amp; Dyer" authorityYear="2024" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois beebei" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="beebei" status="stat. rev.">Eois beebei</taxonomicName>
:
<bibRefCitation id="66DC234F2AF742A3E851A77DDBC37B5D" DOI="https://doi.org/10.1071/9780643101050" author="Parsons, M" journalOrPublisher="Systematic Entomology," pageId="0" pageNumber="111" refId="B23" refString="Parsons, M, Scoble, MJ, Honey, MR, Pitkin, LM, Pitkin, BR, 1999. The catalogue. In: Scoble MJ (Ed.) Geometrid moths of the world: catalogue. Commonwealth Scientific and Industrial Research Organization, Collingwood, Australia. https://doi.org/10.1071/9780643101050" title="The catalogue. In: Scoble MJ (Ed.) Geometrid moths of the world: catalogue. Commonwealth Scientific and Industrial Research Organization, Collingwood, Australia." url="https://doi.org/10.1071/9780643101050" year="1999">Parsons et al. 1999</bibRefCitation>
: 279;
<bibRefCitation id="0506BCFAEE1235C9B35A218414179EB8" DOI="https://doi.org/10.1603/AN10050" author="Brehm, G" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="111" pagination="1091 - 1107" refId="B2" refString="Brehm, G, Bodner, F, Strutzenberger, P, Huenefeld, F, Fiedler, K, 2011. Neotropical Eois (Lepidoptera: Geometridae): checklist, biogeography, diversity, and description patterns. Annals of the Entomological Society of America 104 (6): 1091 - 1107, DOI: https://doi.org/10.1603/AN10050" title="Neotropical Eois (Lepidoptera: Geometridae): checklist, biogeography, diversity, and description patterns." url="https://doi.org/10.1603/AN10050" volume="104" year="2011">Brehm et al. 2011</bibRefCitation>
: 1106.
</paragraph>
</subSubSection>
<subSubSection id="97A70B6D60D9CB3B18ECBD8A4F453203" pageId="0" pageNumber="111" type="type material">
<paragraph id="ACAB63A3A894C28559D9F76728CDDC03" pageId="0" pageNumber="111">Type material examined.</paragraph>
<paragraph id="62E8F1335C77DA9E8AD22901AC6E1B5B" pageId="0" pageNumber="111">
<emphasis id="3D8231C99A7F530548AC7FBFAD814BFB" bold="true" italics="true" pageId="0" pageNumber="111">Holotype</emphasis>
♂, Venezuela, Rancho Grande near Maracay, W. Beebe, No. 481604 (NHMUK).
</paragraph>
<caption id="C06EFB0EEBD8EEC9F8DF0B1D95A2D1E7" doi="10.3897/zookeys.1192.111275.figures15-16" httpUri="https://binary.pensoft.net/fig/990478" pageId="0" pageNumber="111" start="Figures 15, 16" startId="F5">
<paragraph id="DB648A051EAB2413B49D3C8D7BEA6889" pageId="0" pageNumber="111">
<emphasis id="58360F4E10B440E98E9D1A6C30EBC0C6" bold="true" pageId="0" pageNumber="111">Figures 15, 16.</emphasis>
<taxonomicName id="FEE7A250F697CAB1D7080B011626801C" authorityName="Doan &amp; Miller &amp; Brown &amp; Forister &amp; Dyer" authorityYear="2024" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois beebei" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="beebei">
<emphasis id="D399559378F53EEFDFD6D980CD5666E1" italics="true" pageId="0" pageNumber="111">Eois beebei</emphasis>
</taxonomicName>
<emphasis id="E6F7BF48E4D68BEB97A8F0497932F1FC" bold="true" pageId="0" pageNumber="111">15</emphasis>
holotype upperside (NHMUK)
<emphasis id="976E5D29AD878D0C85BDC1B63EA89B31" bold="true" pageId="0" pageNumber="111">16</emphasis>
male genitalia (NHMUK).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9E05C53107FEDBE17A79249CC6878A70" pageId="0" pageNumber="111" type="remarks">
<paragraph id="887DFFC02D12BE126C1484680A7791B3" pageId="0" pageNumber="111">Remarks and diagnosis.</paragraph>
<paragraph id="96979AC1A749E1926EE04E02D9B85A21" pageId="0" pageNumber="111">
<bibRefCitation id="42744CA8C2AECC4FFFDE0EF300715C2A" DOI="https://doi.org/10.5962/p.190383" author="Fletcher, DS" journalOrPublisher="Zoologica (New York)" pageId="0" pageNumber="111" pagination="101 - 105" refId="B10" refString="Fletcher, DS, 1952. Four new species of Geometridae (moths) from Rancho Grande, North-central Venezuela. Zoologica (New York) 37 (10): 101 - 105, DOI: https://doi.org/10.5962/p.190383" title="Four new species of Geometridae (moths) from Rancho Grande, North-central Venezuela." url="https://doi.org/10.5962/p.190383" volume="37" year="1952">Fletcher (1952)</bibRefCitation>
described this species from a single male collected by William Beebe at Rancho Grande (now known as Henri Pittier National Park), in the Venezuelan Costal Range, an historically well-known collecting locality.
<normalizedToken id="2E54F1778A9FAEA7F465C2E711D544A6" originalValue="Fletchers">Fletcher's</normalizedToken>
description is somewhat outdated, as is his rather crude drawing of the male genitalia. The species was treated as a synonym of
<taxonomicName id="ED63FD3FC89147172F856A4C1C18E9E8" lsidName="E. olivacea" pageId="0" pageNumber="111" rank="species" species="olivacea">
<emphasis id="66E8781D0FE7D975E418523AE7F0EB01" italics="true" pageId="0" pageNumber="111">E. olivacea</emphasis>
</taxonomicName>
by
<bibRefCitation id="0D49F0DB76FEE539720CA6B60C99BF40" DOI="https://doi.org/10.1071/9780643101050" author="Parsons, M" journalOrPublisher="Systematic Entomology," pageId="0" pageNumber="111" refId="B23" refString="Parsons, M, Scoble, MJ, Honey, MR, Pitkin, LM, Pitkin, BR, 1999. The catalogue. In: Scoble MJ (Ed.) Geometrid moths of the world: catalogue. Commonwealth Scientific and Industrial Research Organization, Collingwood, Australia. https://doi.org/10.1071/9780643101050" title="The catalogue. In: Scoble MJ (Ed.) Geometrid moths of the world: catalogue. Commonwealth Scientific and Industrial Research Organization, Collingwood, Australia." url="https://doi.org/10.1071/9780643101050" year="1999">Parsons et al. (1999)</bibRefCitation>
, without the benefit of a comparison of the genitalia with those of the latter.
</paragraph>
<paragraph id="26085BB1000BBF77188D7A21EA7A2C3E" pageId="0" pageNumber="111">
As in many members of Group I, the phallus of
<taxonomicName id="11D68F7D6E10715E5FDD70A89B8AEF4E" lsidName="E. beebei" pageId="0" pageNumber="111" rank="species" species="beebei">
<emphasis id="AB0E766102DF860534FC24C1CBB26094" italics="true" pageId="0" pageNumber="111">E. beebei</emphasis>
</taxonomicName>
has a large, conspicuous, scobinate plate with a saw-toothed edge situated near the distal end of the vesica, but lacks long cornuti (Fig.
<figureCitation id="026684B713C13B729DD4F6063E2D3945" captionStart="Figures 15, 16" captionStartId="F5" captionText="Figures 15, 16. Eois beebei 15 holotype upperside (NHMUK) 16 male genitalia (NHMUK)." figureDoi="10.3897/zookeys.1192.111275.figures15-16" httpUri="https://binary.pensoft.net/fig/990478" pageId="0" pageNumber="111">16</figureCitation>
). The species can be distinguished from
<taxonomicName id="B74E5BC42A562AA76274BCF2100D067B" lsidName="E. olivacea" pageId="0" pageNumber="111" rank="species" species="olivacea">
<emphasis id="0D6B7D273B97E96C90A550B768297FEB" italics="true" pageId="0" pageNumber="111">E. olivacea</emphasis>
</taxonomicName>
by the shorter and narrower valvae, and from
<taxonomicName id="0907002DD8BE340DFA836EA0C8A65BCD" lsidName="E. pseudolivacea" pageId="0" pageNumber="111" rank="species" species="pseudolivacea">
<emphasis id="21D61834CA71B71E94D24FCEED7276C0" italics="true" pageId="0" pageNumber="111">E. pseudolivacea</emphasis>
</taxonomicName>
by the shorter lacina.
</paragraph>
</subSubSection>
<subSubSection id="907F3519B043A826CE955F3ABDFDBA3A" pageId="0" pageNumber="111" type="redescription">
<paragraph id="C6D1DB7BFDA8DBCD56BFE8E1375FB166" pageId="0" pageNumber="111">Redescription.</paragraph>
<paragraph id="EF63381BCFB9312FA686037E6E7A3BBC" pageId="0" pageNumber="111">
<emphasis id="734406605E0899E080F9178027446DFA" bold="true" pageId="0" pageNumber="111">
Male.
<emphasis id="BFBE56006241315271109394DA8A250B" italics="true" pageId="0" pageNumber="111">Head</emphasis>
</emphasis>
: Frons and vertex pale pinkish buff with distinct white bar between bases of antennae; labial palpus pale pinkish buff, length~ 0.5 diameter of compound eye; pectinations of antenna ~ 4
<normalizedToken id="4AFB6780FDDEEE8AE5B4050B961D56CC" originalValue="×">x</normalizedToken>
as long as the diameter of the shaft.
<emphasis id="168E301D6B08222C9801E6236CD6338C" bold="true" italics="true" pageId="0" pageNumber="111">Thorax</emphasis>
: Pale olive; forewing ground color pale olive, anterior 0.5 irrorate with pale grayish brown, costa lightly irrorated with cream-brown, postmedial fascia white, discal spot fuscous. Fringe chalcedony yellow. Forewing undersurface white, glossy; discal spot minute.
<emphasis id="74D16C83004DDBEC916FA32F1833DE71" bold="true" italics="true" pageId="0" pageNumber="111">Abdomen</emphasis>
: Pale olive, each segment edged posteriorly with white. Male genitalia (Fig.
<figureCitation id="0E8006A9E6FE43522C6B1F619B974B9E" captionStart="Figures 15, 16" captionStartId="F5" captionText="Figures 15, 16. Eois beebei 15 holotype upperside (NHMUK) 16 male genitalia (NHMUK)." figureDoi="10.3897/zookeys.1192.111275.figures15-16" httpUri="https://binary.pensoft.net/fig/990478" pageId="0" pageNumber="111">16</figureCitation>
) with top of tegumen broadly rounded; lacina supporting long androconial scales; valva subrectangular with distinct sacculus along venter of basal 0.5. Phallus with weakly sclerotized patch near apex; vesica with two scobinate plates in apical 0.5, lacking elongate cornuti.
</paragraph>
<paragraph id="CC129396C154791B2D386C386450EE85" pageId="0" pageNumber="111">
<emphasis id="5FC9E5160C7EBACBD2428669B5EA7FF1" bold="true" pageId="0" pageNumber="111">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="8FA2FB2400F28D7E5DB43FEFF1D79950" pageId="0" pageNumber="111" type="distribution">
<paragraph id="120291A324D58927053DE2855F39758F" pageId="0" pageNumber="111">Distribution and biology.</paragraph>
<paragraph id="C6C5BB121551CD4DD439A0B73262340D" pageId="0" pageNumber="111">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</document>