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<document id="DA079D7D281A8C79F33A17571C77BC14" ID-DOI="10.5281/zenodo.195931" ID-GBIF-Dataset="346cce4c-6e5b-4bb9-8794-7270262cce50" ID-ISSN="1175-5326" ID-Zenodo-Dep="195931" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235905947" checkinUser="plazi" docAuthor="Li, Liang &amp; Zhou, Hong-Zhang" docDate="2010" docId="8709583BFFD81859FF036DA0606FFE43" docLanguage="en" docName="zt02506p025.pdf" docOrigin="Zootaxa 2506" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Craspedomerus zhangi Li &amp; Zhou, sp. n." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="7B302043FFD01853FF9469076102FFF4" masterDocTitle="Taxonomy of the genus Craspedomerus Bernhauer, 1911 (Coleoptera: Staphylinidae: Philonthina) from China" masterLastPageNumber="25" masterPageNumber="1" pageNumber="9" updateTime="1698239575847" updateUser="plazi">
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<mods:title id="3E59C9D1EC93DA2FE612812117FF7600">Taxonomy of the genus Craspedomerus Bernhauer, 1911 (Coleoptera: Staphylinidae: Philonthina) from China</mods:title>
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<mods:namePart id="0D64F40E2D80BD447D94F257906DC903">Li, Liang</mods:namePart>
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<mods:namePart id="E6CB2A685DCD5BCCF98A8C7EB7009FA6">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="F6E94A9134F49C608139B10B6D285BF0">2010</mods:date>
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<treatment id="8709583BFFD81859FF036DA0606FFE43" ID-DOI="http://doi.org/10.5281/zenodo.6195817" ID-GBIF-Taxon="119388208" ID-Zenodo-Dep="6195817" LSID="urn:lsid:plazi:treatment:8709583BFFD81859FF036DA0606FFE43" httpUri="http://treatment.plazi.org/id/8709583BFFD81859FF036DA0606FFE43" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="47BABAA6FFD8185BFF036DA063A2FB35" box="[151,672,1191,1217]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="0F1FE92DFFD8185BFF036DA063A2FB35" blockId="8.[151,672,1191,1252]" box="[151,672,1191,1217]" pageId="8" pageNumber="9">
<heading id="54575E41FFD8185BFF036DA063A2FB35" bold="true" box="[151,672,1191,1217]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="3DD4353FFFD8185BFF036DA063A2FB35" bold="true" box="[151,672,1191,1217]" pageId="8" pageNumber="9">
3.
<taxonomicName id="C8A092AEFFD8185BFF206DA06355FB35" ID-CoL="Z5RT" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" box="[180,599,1191,1217]" class="Insecta" family="Staphylinidae" genus="Craspedomerus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zhangi" status="sp. nov.">
<emphasis id="3DD4353FFFD8185BFF206DA060CBFB35" bold="true" box="[180,457,1191,1217]" italics="true" pageId="8" pageNumber="9">Craspedomerus zhangi</emphasis>
Li &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="26E78844FFD8185BFDCA6DA063A2FB35" box="[606,672,1191,1217]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="47BABAA6FFD8185BFF036DCD606CFB10" box="[151,366,1226,1252]" pageId="8" pageNumber="9" type="description">
<paragraph id="0F1FE92DFFD8185BFF036DCD606CFB10" blockId="8.[151,672,1191,1252]" box="[151,366,1226,1252]" pageId="8" pageNumber="9">
(
<figureCitation id="979BF5A8FFD8185BFF0B6DCD61ECFB10" box="[159,238,1226,1252]" captionStart="FIGURE 5" captionStartId="9.[151,255,1570,1594]" captionTargetBox="[275,1313,188,1536]" captionTargetId="figure@9.[268,1320,179,1548]" captionTargetPageId="9" captionText="FIGURE 5. Craspedomerus zhangi Li &amp; Zhou sp. n. A, aedeagus, ventral view; B, aedeagus, lateral view; C, sternite IX of male; D, sternite VIII of male; E, tergite X of male. Scale bars 1 mm." httpUri="https://zenodo.org/record/195936/files/figure.png" pageId="8" pageNumber="9">Figs. 5</figureCitation>
AE, 12E)
</paragraph>
</subSubSection>
<subSubSection id="47BABAA6FFD8185BFF036C0865D5FA80" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="0F1FE92DFFD8185BFF036C0865D5FA80" blockId="8.[151,1437,1295,2034]" pageId="8" pageNumber="9">
<emphasis id="3DD4353FFFD8185BFF036C0863AEFADD" bold="true" box="[151,684,1295,1321]" pageId="8" pageNumber="9">
<typeStatus id="D01B578FFFD8185BFF036C0861D4FADD" box="[151,214,1295,1321]" pageId="8" pageNumber="9">Type</typeStatus>
material.
<typeStatus id="D01B578FFFD8185BFEC96C0860D5FADD" box="[349,471,1295,1321]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="77B7A9BDFFD8185BFE716C08634CFADD" box="[485,590,1295,1321]" name="China" pageId="8" pageNumber="9">CHINA</collectingCountry>
: Tibet:
</emphasis>
ɗ, Mêdog (
<geoCoordinate id="6A948FEAFFD8185BFCAB6C1762AEFADE" box="[831,940,1296,1322]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="29.7">29°42´N</geoCoordinate>
,
<geoCoordinate id="6A948FEAFFD8185BFC2E6C176521FADE" box="[954,1059,1296,1322]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="95.933334">95°56´E</geoCoordinate>
),
<quantity id="C85844C8FFD8185BFBAE6C17659BFADE" box="[1082,1177,1296,1322]" metricMagnitude="3" metricUnit="m" metricValue="2.7649999999999997" pageId="8" pageNumber="9" unit="m" value="2765.0">2765 m</quantity>
,
<date id="7B1ECFEDFFD8185BFB326C17644AFADE" box="[1190,1352,1296,1322]" pageId="8" pageNumber="9" value="2006-08-26">26.VIII.2006</date>
, Liang Hongbin collected (IZ-CAS);
<emphasis id="3DD4353FFFD8185BFE6B6C326251FABB" bold="true" box="[511,851,1333,1359]" pageId="8" pageNumber="9">
<typeStatus id="D01B578FFFD8185BFE6B6C326381FABB" box="[511,643,1333,1359]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="77B7A9BDFFD8185BFD056C3263FAFABB" box="[657,760,1333,1359]" name="China" pageId="8" pageNumber="9">CHINA</collectingCountry>
: Tibet:
</emphasis>
2ΨΨ, same data as
<typeStatus id="D01B578FFFD8185BFBD66C3265A8FABB" box="[1090,1194,1333,1359]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(IZ-CAS); Ψ, Bomi (
<geoCoordinate id="6A948FEAFFD8185BFF0B6C5D600AFA80" box="[159,264,1370,1396]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="29.8">29°48´N</geoCoordinate>
,
<geoCoordinate id="6A948FEAFFD8185BFE806C5D607BFA80" box="[276,377,1370,1396]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="95.683334">95°41´E</geoCoordinate>
),
<quantity id="C85844C8FFD8185BFE1B6C5D60E8FA80" box="[399,490,1370,1396]" metricMagnitude="3" metricUnit="m" metricValue="3.65" pageId="8" pageNumber="9" unit="m" value="3650.0">3650 m</quantity>
,
<date id="7B1ECFEDFFD8185BFE626C5D6390FA80" box="[502,658,1370,1396]" pageId="8" pageNumber="9" value="2006-08-28">28.VIII.2006</date>
, Liang Hongbin &amp; Bai Ming collected (IZ-CAS).
</paragraph>
</subSubSection>
<subSubSection id="47BABAA6FFD81859FF526C85642FFF23" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="0F1FE92DFFD8185BFF526C856082F9C6" blockId="8.[151,1437,1295,2034]" pageId="8" pageNumber="9">
<emphasis id="3DD4353FFFD8185BFF526C856055FA68" bold="true" box="[198,343,1410,1436]" pageId="8" pageNumber="9">Description</emphasis>
. Head and pronotum black with slightly bronze-green reflex. Antennae black with 8th10th segments yellow, last segment black-brown. Elytra purple with shoulders, posterior margins and suture markedly reddish-yellow. Scutellum black. Abdomen black with strongly iridescent reflex, posterior half of tergite VII and entire tergites VIIIX reddish-yellow. Mandibles dark brown. Maxillary and labial palpi and legs reddish-brown.
</paragraph>
<paragraph id="0F1FE92DFFD8185BFF526F3A6524F8E6" blockId="8.[151,1437,1295,2034]" pageId="8" pageNumber="9">
Body moderately large,
<quantity id="C85844C8FFD8185BFE656F3A63A3F9A3" box="[497,673,1597,1623]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" metricValueMax="1.48" metricValueMin="1.08" pageId="8" pageNumber="9" unit="mm" value="12.8" valueMax="14.8" valueMin="10.8">10.814.8 mm</quantity>
long (HPL =
<quantity id="C85844C8FFD8185BFCCD6F3A650FF9A3" box="[857,1037,1597,1623]" metricMagnitude="-3" metricUnit="m" metricValue="4.535" metricValueMax="4.74" metricValueMin="4.33" pageId="8" pageNumber="9" unit="mm" value="4.535" valueMax="4.74" valueMin="4.33">4.334.74 mm</quantity>
). Head of rounded quadrangular shape,
<quantity id="C85844C8FFD8185BFF7F6F656098F988" box="[235,410,1634,1660]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="1.96" metricValueMin="1.88" pageId="8" pageNumber="9" unit="mm" value="1.92" valueMax="1.96" valueMin="1.88">1.881.96 mm</quantity>
long,
<quantity id="C85844C8FFD8185BFE7C6F656395F988" box="[488,663,1634,1660]" metricMagnitude="-3" metricUnit="m" metricValue="2.165" metricValueMax="2.29" metricValueMin="2.04" pageId="8" pageNumber="9" unit="mm" value="2.165" valueMax="2.29" valueMin="2.04">2.042.29 mm</quantity>
wide, slightly wider than long (HW:HL = 1.091.13). Tempora
<quantity id="C85844C8FFD8185BFF036F8F6043F956" box="[151,321,1672,1698]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="8.1" metricValueMin="6.5" pageId="8" pageNumber="9" unit="mm" value="0.73" valueMax="0.81" valueMin="0.65">0.650.81 mm</quantity>
long, almost evenly rounded, densely and coarsely punctate, setiferous. Eyes moderately large, slightly protruded,
<quantity id="C85844C8FFD8185BFE176FAA6336F933" box="[387,564,1709,1735]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="7.4" metricValueMin="5.699999999999999" pageId="8" pageNumber="9" unit="mm" value="0.655" valueMax="0.74" valueMin="0.57">0.570.74 mm</quantity>
long, 0.780.90 times as long as tempora. Dorsal surface of head with moderately numerous, large setiferous punctures, becoming sparser toward vertex, vertex largely impunctate; entire head with distinct and profound microsculpture of oblique, long waves.
</paragraph>
<paragraph id="0F1FE92DFFD8185BFF526E1A65B5F876" blockId="8.[151,1437,1295,2034]" pageId="8" pageNumber="9">
Pronotum subquadrate, slightly narrowed anteriad,
<quantity id="C85844C8FFD8185BFCAC6E1A62EAF8C3" box="[824,1000,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" metricValueMax="2.45" metricValueMin="2.29" pageId="8" pageNumber="9" unit="mm" value="2.37" valueMax="2.45" valueMin="2.29">2.292.45 mm</quantity>
long,
<quantity id="C85844C8FFD8185BFBA86E1A65E9F8C3" box="[1084,1259,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="2.205" metricValueMax="2.29" metricValueMin="2.12" pageId="8" pageNumber="9" unit="mm" value="2.205" valueMax="2.29" valueMin="2.12">2.122.29 mm</quantity>
wide, slightly wider than head (PW:HW = 1.001.10), sparsely and finely punctate, punctures separated by 47 times their diameter, narrowly impunctate along midline, microsculpture fine, similar to that on head.
</paragraph>
<paragraph id="0F1FE92DFFD8185BFF526E8A6000F806" blockId="8.[151,1437,1295,2034]" pageId="8" pageNumber="9">
Elytra
<quantity id="C85844C8FFD8185BFE816E8A60C2F853" box="[277,448,1933,1959]" metricMagnitude="-3" metricUnit="m" metricValue="2.855" metricValueMax="3.02" metricValueMin="2.69" pageId="8" pageNumber="9" unit="mm" value="2.855" valueMax="3.02" valueMin="2.69">2.693.02 mm</quantity>
long,
<quantity id="C85844C8FFD8185BFD9F6E8A63B7F853" box="[523,693,1933,1959]" metricMagnitude="-3" metricUnit="m" metricValue="3.185" metricValueMax="3.43" metricValueMin="2.94" pageId="8" pageNumber="9" unit="mm" value="3.185" valueMax="3.43" valueMin="2.94">2.943.43 mm</quantity>
wide, 1.181.30 times as long as pronotum, densely and finely punctate, punctures separated by 23 times their diameter. Scutellum large, triangular, densely and finely punctate.
</paragraph>
<caption id="5BDFB9A5FFD9185AFF036F256295F9AD" httpUri="https://zenodo.org/record/195936/files/figure.png" pageId="9" pageNumber="10" targetBox="[275,1313,188,1536]" targetPageId="9">
<paragraph id="0F1FE92DFFD9185AFF036F256295F9AD" blockId="9.[151,1436,1570,1625]" pageId="9" pageNumber="10">
<emphasis id="3DD4353FFFD9185AFF036F25601AF9CE" bold="true" box="[151,280,1570,1594]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
<taxonomicName id="C8A092AEFFD9185AFE8B6F25638EF9CD" authority="Li &amp; Zhou" authorityName="Li &amp; Zhou" box="[287,652,1570,1593]" class="Insecta" family="Staphylinidae" genus="Craspedomerus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zhangi" status="sp. nov.">
<emphasis id="3DD4353FFFD9185AFE8B6F256317F9CD" box="[287,533,1570,1593]" italics="true" pageId="9" pageNumber="10">Craspedomerus zhangi</emphasis>
Li &amp; Zhou
</taxonomicName>
<emphasis id="3DD4353FFFD9185AFD076F2563CCF9CE" bold="true" box="[659,718,1570,1594]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="26E78844FFD9185AFD076F2563CCF9CE" box="[659,718,1570,1594]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A, aedeagus, ventral view; B, aedeagus, lateral view; C, sternite IX of male; D, sternite VIII of male; E, tergite X of male. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="0F1FE92DFFD9185AFF526F8563BCF8F8" blockId="9.[151,1437,1666,2028]" pageId="9" pageNumber="10">
Abdomen a little narrowed posteriad, widest
<quantity id="C85844C8FFD9185AFD496F856289F968" box="[733,907,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.69" metricValueMin="2.45" pageId="9" pageNumber="10" unit="mm" value="2.57" valueMax="2.69" valueMin="2.45">2.452.69 mm</quantity>
; tergites densely and finely punctate, at base punctures separated by 23 times their diameter, gradually becoming sparser toward apex of each tergite, surface between punctures with dense and fine microsculpture of transverse striae; elevated area between basal lines of tergites IIIV almost impunctate.
</paragraph>
<paragraph id="0F1FE92DFFD9185AFF526E106585F855" blockId="9.[151,1437,1666,2028]" pageId="9" pageNumber="10">
Male. Sternite VIII with moderately wide, obtusely triangular medio-apical emargination (
<figureCitation id="979BF5A8FFD9185AFABD6E106474F8C5" box="[1321,1398,1815,1841]" captionStart="FIGURE 5" captionStartId="9.[151,255,1570,1594]" captionTargetBox="[275,1313,188,1536]" captionTargetId="figure@9.[268,1320,179,1548]" captionTargetPageId="9" captionText="FIGURE 5. Craspedomerus zhangi Li &amp; Zhou sp. n. A, aedeagus, ventral view; B, aedeagus, lateral view; C, sternite IX of male; D, sternite VIII of male; E, tergite X of male. Scale bars 1 mm." httpUri="https://zenodo.org/record/195936/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
D). Sternite IX with distinctly asymmetrical basal portion, apex deeply emarginate, each lobe bearing two long apical setae (
<figureCitation id="979BF5A8FFD9185AFEA36E656080F888" box="[311,386,1890,1916]" captionStart="FIGURE 5" captionStartId="9.[151,255,1570,1594]" captionTargetBox="[275,1313,188,1536]" captionTargetId="figure@9.[268,1320,179,1548]" captionTargetPageId="9" captionText="FIGURE 5. Craspedomerus zhangi Li &amp; Zhou sp. n. A, aedeagus, ventral view; B, aedeagus, lateral view; C, sternite IX of male; D, sternite VIII of male; E, tergite X of male. Scale bars 1 mm." httpUri="https://zenodo.org/record/195936/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
C). Genital segment with styli of tergite IX simple, densely setose apically. Tergite X simple, triangular, subrounded at apex, with variable number of apical setae (
<figureCitation id="979BF5A8FFD9185AFB8B6E806566F855" box="[1055,1124,1927,1953]" captionStart="FIGURE 5" captionStartId="9.[151,255,1570,1594]" captionTargetBox="[275,1313,188,1536]" captionTargetId="figure@9.[268,1320,179,1548]" captionTargetPageId="9" captionText="FIGURE 5. Craspedomerus zhangi Li &amp; Zhou sp. n. A, aedeagus, ventral view; B, aedeagus, lateral view; C, sternite IX of male; D, sternite VIII of male; E, tergite X of male. Scale bars 1 mm." httpUri="https://zenodo.org/record/195936/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
E).
</paragraph>
<paragraph id="0F1FE92DFFD9185AFF526EAA63B7F818" blockId="9.[151,1437,1666,2028]" pageId="9" pageNumber="10">
Aedeagus as (
<figureCitation id="979BF5A8FFD9185AFEFA6EAA60C2F833" box="[366,448,1965,1991]" captionStart="FIGURE 5" captionStartId="9.[151,255,1570,1594]" captionTargetBox="[275,1313,188,1536]" captionTargetId="figure@9.[268,1320,179,1548]" captionTargetPageId="9" captionText="FIGURE 5. Craspedomerus zhangi Li &amp; Zhou sp. n. A, aedeagus, ventral view; B, aedeagus, lateral view; C, sternite IX of male; D, sternite VIII of male; E, tergite X of male. Scale bars 1 mm." httpUri="https://zenodo.org/record/195936/files/figure.png" pageId="9" pageNumber="10">Figs. 5</figureCitation>
AB); underside of each branch of paramere with very small, pointed sensory peg setae arranged at subapical portion (
<figureCitation id="979BF5A8FFD9185AFDA96ED56391F818" box="[573,659,2002,2028]" captionStart="FIGURE 12" captionStartId="21.[151,255,1814,1838]" captionTargetBox="[151,1436,370,1791]" captionTargetId="figure@21.[151,1436,370,1793]" captionTargetPageId="21" captionText="FIGURE 12. Scanning electron micrographs of the underside of one of the branches of the paramere of Craspedomerus. A, C. sinetuber (Coiffait); B, C. glenoides (Schubert); C, C. beckeri Bernhauer; D, C. violaceipennis Cameron; E, C. zhangi Li &amp; Zhou sp. n.; F, C. giganteus Li &amp; Zhou sp. n.; G, C. ganeshensis Coiffait; H, C. gongshanus Li &amp; Zhou sp. n.; I, C. cyanipennis Scheerpeltz. Sensory peg setae are indicated with arrows." httpUri="https://zenodo.org/record/195943/files/figure.png" pageId="9" pageNumber="10">Fig. 12</figureCitation>
E).
</paragraph>
<paragraph id="0F1FE92DFFDA1859FF52699F642FFF23" blockId="10.[151,1436,152,439]" pageId="10" pageNumber="11">Female. Sternite VIII without medio-apical emargination. Gonocoxites moderately developed, second gonocoxites each with minute stylus bearing two long apical setae. Tergite X similar to that of male.</paragraph>
</subSubSection>
<subSubSection id="47BABAA6FFDA1859FF5269E561DBFEB3" pageId="10" pageNumber="11" type="discussion">
<paragraph id="0F1FE92DFFDA1859FF5269E561DBFEB3" blockId="10.[151,1436,152,439]" pageId="10" pageNumber="11">
<emphasis id="3DD4353FFFDA1859FF5269E5603DFF08" bold="true" box="[198,319,226,252]" pageId="10" pageNumber="11">Remarks.</emphasis>
<taxonomicName id="C8A092AEFFDA1859FEDE69E56359FF0F" box="[330,603,226,251]" class="Insecta" family="Staphylinidae" genus="Craspedomerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="3DD4353FFFDA1859FEDE69E56359FF0F" box="[330,603,226,251]" italics="true" pageId="10" pageNumber="11">Craspedomerus zhangi</emphasis>
</taxonomicName>
can be easily distinguished from all congeners by elytra purple with shoulders, posterior margins and suture markedly reddish-yellow and quite characteristic shape of sensory peg setae.
</paragraph>
</subSubSection>
<subSubSection id="47BABAA6FFDA1859FF5268556508FE98" box="[198,1034,338,364]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="0F1FE92DFFDA1859FF5268556508FE98" blockId="10.[151,1436,152,439]" box="[198,1034,338,364]" pageId="10" pageNumber="11">
<emphasis id="3DD4353FFFDA1859FF5268556065FE98" bold="true" box="[198,359,338,364]" pageId="10" pageNumber="11">Distribution.</emphasis>
The species is at present known only from
<collectingCountry id="77B7A9BDFFDA1859FCF0685562A8FE98" box="[868,938,338,364]" name="China" pageId="10" pageNumber="11">China</collectingCountry>
(Tibet).
</paragraph>
</subSubSection>
<subSubSection id="47BABAA6FFDA1859FF526870606FFE43" pageId="10" pageNumber="11" type="etymology">
<paragraph id="0F1FE92DFFDA1859FF526870606FFE43" blockId="10.[151,1436,152,439]" pageId="10" pageNumber="11">
<emphasis id="3DD4353FFFDA1859FF5268706051FE65" bold="true" box="[198,339,375,401]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species is named in honor of Dr. Luping Zhang, College of Life Sciences, Hebei Normal University,
<collectingCountry id="77B7A9BDFFDA1859FE8B689A606BFE43" box="[287,361,413,439]" name="China" pageId="10" pageNumber="11">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>