233 lines
29 KiB
XML
233 lines
29 KiB
XML
<document id="BB61C1978C40FCA209E5D2545E79DED3" ID-DOI="10.11646/zootaxa.4750.3.7" ID-GBIF-Dataset="e8a4de57-4e3e-47f6-93f2-20f4426cfe0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3707533" ID-ZooBank="4AA5B325-AB4E-4275-8F89-E1DA27EE6F07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584002421674" checkinUser="plazi" docAuthor="Alekseev, Vitalii I., Bukejs, Andris & Sontag, Elżbieta" docDate="2020" docId="1A7087EEFFCF8B60FF31FE5195F8FEC3" docLanguage="en" docName="zootaxa.4750.3.7.pdf" docOrigin="Zootaxa 4750 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eledonoprius incoronatus Alekseev & Bukejs & Sontag 2020, sp. nov." docType="treatment" docUuid="2F4128E7-17BD-468E-8109-920B84A52BB9" docUuidSource="ZooBank" docVersion="7" lastPageNumber="423" masterDocId="E649FF96FFCE8B65FFA6FFF49743FF80" masterDocTitle="A new fossil species of Bolitophagini (Coleoptera: Tenebrionidae) from Baltic amber suggests the genus Eledonoprius Reitter is persistent in the Western Palaearctic since the Tertiary" masterLastPageNumber="424" masterPageNumber="418" pageNumber="419" updateTime="1698727250520" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="22D80E75BFB160684B055E8DE64CF2F9" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="44B959794B7AC3793F1A85FC171AB95B">
|
||
<mods:title id="84AEC6AA4563A48652557F37C0BCAA34">A new fossil species of Bolitophagini (Coleoptera: Tenebrionidae) from Baltic amber suggests the genus Eledonoprius Reitter is persistent in the Western Palaearctic since the Tertiary</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0E2B72C1971C3CA6DAB229103141903E" type="personal">
|
||
<mods:role id="3A244F64908E35958E69E6303735EC72">
|
||
<mods:roleTerm id="DFABA2E97317D2FF8CE8214957BA3795">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AA82DD4023B8BE4899D363940BF6760D">Alekseev, Vitalii I.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="45A045F237656084D0DC9DB5F2C8D2D6" type="personal">
|
||
<mods:role id="0340E148F9FCBA9168808124E4C5FEAD">
|
||
<mods:roleTerm id="AEFDCD09E58BF78610EA8EC2C1F2E0C0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="61FE3F693CF989E41B8FCD41A1C0BE72">Bukejs, Andris</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8452450F222F804ADF44F06AC728600F" type="personal">
|
||
<mods:role id="A1E755518D53E283BBDF188B5C0EE06E">
|
||
<mods:roleTerm id="5CC9D60CFFC1543EB7B0377F8035C5F3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="918EB2B4AAC3EECEF04D9723A961BB5B">Sontag, Elżbieta</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B61E3C82073A3FA6FF8CF325AB8FBF4B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="963EA3E888C09D9504AA168F0146E997" type="host">
|
||
<mods:titleInfo id="D4887D9F2BEC92D338F2F3651573F5A2">
|
||
<mods:title id="01BDEFF81CBFEE74D662B94761549966">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="AF96575E976B2BDB9E2385EC2A5A50F2">
|
||
<mods:date id="9F836B0C72FB432A7BD9E8E08D21AE76">2020</mods:date>
|
||
<mods:detail id="10FF2D837A83A1DA9A1899E84BBD6F2C" type="pubDate">
|
||
<mods:number id="3E653E39D6E1FEF9BB7D6FE1871C8DAA">2020-03-12</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4A6A74805047FAB0D34EDC1224324A81" type="volume">
|
||
<mods:number id="3BB405BA1C02FD0DA787EC381F29AC21">4750</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2726DB3A40379448BDC41BCF77D84770" type="issue">
|
||
<mods:number id="54D4268454A1A6E1F3750B52B4223A8B">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FED4B4B48C6F2A63E9308A802E1E239B" unit="page">
|
||
<mods:start id="443DFC4982D3B3A9D83740DC851B85BE">418</mods:start>
|
||
<mods:end id="0E1162A138F421E28060A5F391D87E5D">424</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="EEFBC0D51133D9C49C0EE9242F8DE589">journal article</mods:classification>
|
||
<mods:identifier id="7A00C1D35913829CAED9DEF404A7F420" type="DOI">10.11646/zootaxa.4750.3.7</mods:identifier>
|
||
<mods:identifier id="279B4DD32028FB311321081F374052D9" type="GBIF-Dataset">e8a4de57-4e3e-47f6-93f2-20f4426cfe0d</mods:identifier>
|
||
<mods:identifier id="A3DED995AFD348AFE1A34D06A2BC0583" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="4B9219F9F4C49603ADC61613B51CFE12" type="Zenodo-Dep">3707533</mods:identifier>
|
||
<mods:identifier id="B1774EE59077F65F184B004093E94338" type="ZooBank">4AA5B325-AB4E-4275-8F89-E1DA27EE6F07</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="1A7087EEFFCF8B60FF31FE5195F8FEC3" ID-DOI="http://doi.org/10.5281/zenodo.3717916" ID-GBIF-Taxon="162484014" ID-Zenodo-Dep="3717916" LSID="urn:lsid:zoobank.org:act:2F4128E7-17BD-468E-8109-920B84A52BB9" httpUri="http://treatment.plazi.org/id/1A7087EEFFCF8B60FF31FE5195F8FEC3" lastPageId="5" lastPageNumber="423" pageId="1" pageNumber="419">
|
||
<subSubSection id="DAC36573FFCF8B64FF31FE5195E2FE3E" box="[151,673,420,447]" pageId="1" pageNumber="419" type="nomenclature">
|
||
<paragraph id="926636F8FFCF8B64FF31FE5195E2FE3E" blockId="1.[151,673,420,483]" box="[151,673,420,447]" pageId="1" pageNumber="419">
|
||
<heading id="C92E8194FFCF8B64FF31FE5195E2FE3E" bold="true" box="[151,673,420,447]" fontSize="11" level="1" pageId="1" pageNumber="419" reason="1">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF31FE5195E2FE3E" bold="true" box="[151,673,420,447]" pageId="1" pageNumber="419">
|
||
<taxonomicName id="55D94D7BFFCF8B64FF31FE51957FFE3E" authority="Alekseev & Bukejs" authorityName="Alekseev & Bukejs & Sontag" authorityYear="2020" box="[151,572,420,447]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="species" species="incoronatus" status="sp. nov.">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF31FE519609FE3E" bold="true" box="[151,330,420,447]" italics="true" pageId="1" pageNumber="419">E. incoronatus</emphasis>
|
||
Alekseev & Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BB9E5791FFCF8B64FDE2FE5095E2FE3E" box="[580,673,420,446]" pageId="1" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAC36573FFCF8B64FF31FE3C9651FE63" box="[151,274,456,483]" pageId="1" pageNumber="419" type="description">
|
||
<paragraph id="926636F8FFCF8B64FF31FE3C9651FE63" blockId="1.[151,673,420,483]" box="[151,274,456,483]" pageId="1" pageNumber="419">
|
||
(
|
||
<figureCitation id="0AE22A7DFFCF8B64FF39FE3C964AFE63" box="[159,265,456,483]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStart-2="FIGURES 5–6" captionStartId-0="2.[151,264,1953,1978]" captionStartId-1="3.[151,264,1963,1988]" captionStartId-2="4.[151,264,1955,1980]" captionTargetBox-0="[154,1433,181,1928]" captionTargetBox-1="[158,1429,199,1938]" captionTargetBox-2="[155,1433,181,1929]" captionTargetId-0="figure@2.[154,1433,181,1928]" captionTargetId-1="figure@3.[158,1429,199,1938]" captionTargetId-2="figure@4.[155,1433,181,1929]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–2. Eledonoprius incoronatus sp. nov., holotype: 1 – habitus, dorsal view; 2 – habitus, lateral view. Scale bars represent 0.5 mm." captionText-1="FIGURES 3–4. Eledonoprius incoronatus sp. nov., holotype: 3 – habitus, ventral view; 4 – forebody, ventral view. Scale bars represent 0.5 mm (Fig. 3) and 0.2 mm (Fig.4)." captionText-2="FIGURES 5–6. Eledonoprius incoronatus sp. nov., holotype: 5 – genitalia, lateral view; 6 – forebody, lateral view. Scale bars represent 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3707535" figureDoi-1="http://doi.org/10.5281/zenodo.3707537" figureDoi-2="http://doi.org/10.5281/zenodo.3707539" httpUri-0="https://zenodo.org/record/3707535/files/figure.png" httpUri-1="https://zenodo.org/record/3707537/files/figure.png" httpUri-2="https://zenodo.org/record/3707539/files/figure.png" pageId="1" pageNumber="419">Figs. 1–6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAC36573FFCF8B64FF31FDFF94CAFD10" pageId="1" pageNumber="419" type="materials_examined">
|
||
<paragraph id="926636F8FFCF8B64FF31FDFF97F8FDC8" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF31FDFF9606FDA5" bold="true" box="[151,325,522,549]" pageId="1" pageNumber="419">Type material.</emphasis>
|
||
<materialsCitation id="22B13CA5FFCF8B64FEEBFDFF97F8FDC8" ID-GBIF-Occurrence="2575868881" collectionCode="MAIG" pageId="1" pageNumber="419" specimenCode="MAIG 6012" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="4D62885AFFCF8B64FEEBFDFF96F9FDA5" box="[333,442,523,549]" pageId="1" pageNumber="419" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="C27F9E83FFCF8B64FE62FDFF9511FDA5" box="[452,594,523,549]" collectionCode="MAIG" pageId="1" pageNumber="419">MAIG 6012</specimenCode>
|
||
, adult, male, complete beetle inclusion with partially exposed male genitalia.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61FDA694CAFD10" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">
|
||
<materialsCitation id="22B13CA5FFCF8B64FF61FDA694C6FD10" ID-GBIF-Occurrence="2575868880" country="Russia" county="Mid-Eocene" location="Yantarny" municipality="Baltic" pageId="1" pageNumber="419" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF61FDA69544FDEC" bold="true" box="[199,519,593,620]" pageId="1" pageNumber="419">
|
||
<typeStatus id="4D62885AFFCF8B64FF61FDA69641FDEC" box="[199,258,594,620]" pageId="1" pageNumber="419">Type</typeStatus>
|
||
locality and horizon.
|
||
</emphasis>
|
||
<location id="97066023FFCF8B64FDB4FDA69534FDEC" LSID="urn:lsid:plazi:treatment:1A7087EEFFCF8B60FF31FE5195F8FEC3:97066023FFCF8B64FDB4FDA69534FDEC" box="[530,631,594,620]" country="Russia" county="Mid-Eocene" municipality="Baltic" name="Yantarny" pageId="1" pageNumber="419" stateProvince="Kaliningrad">Yantarny</location>
|
||
settlement (formerly Palmnicken),
|
||
<location id="97066023FFCF8B64FBB5FDA69336FDEC" LSID="urn:lsid:plazi:treatment:1A7087EEFFCF8B60FF31FE5195F8FEC3:97066023FFCF8B64FBB5FDA69336FDEC" box="[1043,1141,594,620]" country="Russia" county="Mid-Eocene" municipality="Baltic" name="Sambian" pageId="1" pageNumber="419" stateProvince="Kaliningrad">Sambian</location>
|
||
(Samland) peninsula, the
|
||
<collectingRegion id="501DF81AFFCF8B64FF31FD81962DFD0F" box="[151,366,629,655]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="419">Kaliningrad region</collectingRegion>
|
||
,
|
||
<collectingCountry id="EACE7668FFCF8B64FEDFFD819684FD0F" box="[377,455,629,655]" name="Russia" pageId="1" pageNumber="419">Russia</collectingCountry>
|
||
.
|
||
<collectingMunicipality id="7202AC82FFCF8B64FE74FD819556FD0F" box="[466,533,629,655]" pageId="1" pageNumber="419">Baltic</collectingMunicipality>
|
||
amber.
|
||
<collectingCounty id="7B074E74FFCF8B64FDC8FD8195BBFD0F" box="[622,760,629,655]" pageId="1" pageNumber="419">Mid-Eocene</collectingCounty>
|
||
(Bartonian)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAC36573FFCF8B64FF61FD6C9577F9F4" pageId="1" pageNumber="419" type="description">
|
||
<paragraph id="926636F8FFCF8B64FF61FD6C94A9FD7B" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF61FD6C961AFD33" bold="true" box="[199,345,664,691]" pageId="1" pageNumber="419">Description.</emphasis>
|
||
Body oblong, subparallel-sided, moderately convex dorsally; uniformly black, coarsely punctate, bare. Total body length about
|
||
<quantity id="55219B1DFFCF8B64FE4EFD499513FD57" box="[488,592,701,727]" metricMagnitude="-3" metricUnit="m" metricValue="2.14" pageId="1" pageNumber="419" unit="mm" value="2.14">2.14 mm</quantity>
|
||
; maximum length of pronotum
|
||
<quantity id="55219B1DFFCF8B64FC13FD48934DFD57" box="[949,1038,700,727]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="419" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; pronotal maximum width
|
||
<quantity id="55219B1DFFCF8B64FA98FD4992DBFD57" box="[1342,1432,701,727]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="419" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; length of an elytron along suture
|
||
<quantity id="55219B1DFFCF8B64FDADFD149527FD7B" box="[523,612,736,763]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="419" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; width of elytra at humeri
|
||
<quantity id="55219B1DFFCF8B64FC2BFD1494A5FD7B" box="[909,998,736,763]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="419" unit="mm" value="1.1">1.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61FCF09554FBBB" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">Head transverse; frontoclypeus widely rounded and non-denticulate anteriorly, with two short straight sharp horns and inconspicuous sparse short setae; genae broader than eyes, non-denticulate; frons almost flat, with four distinct sharpened tubercles located in concave arcuate line between posterior margins of eyes; vertex slightly convex. Punctation of head dense and rounded. Compound eyes oval, with vertical diameter about 2× transverse diameter, very deeply emarginate by genae anteriorly (almost divided in two parts: dorsal and ventral); without interfacetal setae. Terminal maxillary palpomere cylindrical, about 4.0× as long as wide, longer than two previous palpomeres combined. Antennae 11-segmented, comparatively short; antennomeres 6–8 almost quadrate; antennomeres 9–10 transverse; antennomere 11 longest, oval, about 1.8× as long as antennomere 10, slightly shorter than antennomeres 9 and 10 combined.</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61FBB19457FA91" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">Pronotum strongly transverse, about 2.2× as wide as long, with maximum width in posterior one-third, as wide as elytral base; pronotal disc moderately convex, with two shallow, indistinct oblique latero-basal impressions; lateral margins widely explanate, slightly rounded, with 5–6 sharp triangular denticles; posterior margin bisinuate; anterior margin deflexed, arcuate; anterior pronotal angles distinct, triangular, produced forwards; posterior angles distinct, almost orthogonal. Pronotal surface with rough, rounded, dense punctation, interspersed with sharp granules. Procoxae narrowly separated by prosternal process.</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61FAEE92D3FA42" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">Elytra elongate, slightly widened postmedially; disc moderately convex; humeral angles well-developed, rounded. Each elytron with 9 deep striae; elytral punctation very shallow and discernible in basal one-quarter, obliterate apically. Interstriae convex, with distinct sharp granules in posterior one-third and obtuse granules on the disc. Humeral interval with sharp granules along the entire length. Scutellar shield as long as wide, rounded apically. Epipleura wide (epipleuron in basal part distinctly wider than metanepisternum), gradually narrowing posteriorly.</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61FA38958BF9AE" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">Legs slender. Femora and tibiae moderately flattened; tibia with one very short spine apically. Tarsal formula 5-5-4. All tarsomeres simple (non-lobed); tarsomere 1 longer than all previous tarsomeres combined; tarsal claws long, falcate, about 0.5× as long as last tarsomere.</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61F9C39577F9F4" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">Abdomen with five ventrites, coarsely and densely punctate, bare. Anterior process of ventrite 1 triangular. Distal parts of male genitalia visible.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAC36573FFCF8B64FF61F98A95A8F93C" pageId="1" pageNumber="419" type="etymology">
|
||
<paragraph id="926636F8FFCF8B64FF61F98A95A8F93C" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF61F98A960DF918" bold="true" box="[199,334,1662,1688]" pageId="1" pageNumber="419">Etymology.</emphasis>
|
||
The epithet of this new species [Latin adjective “
|
||
<taxonomicName id="55D94D7BFFCF8B64FCCBF98B94B1F918" authorityName="Alekseev & Bukejs & Sontag" authorityYear="2020" box="[877,1010,1663,1688]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="species" species="incoronatus">
|
||
<emphasis id="A0ADEAEAFFCF8B64FCCBF98B94B1F918" box="[877,1010,1663,1688]" italics="true" pageId="1" pageNumber="419">incoronatus</emphasis>
|
||
</taxonomicName>
|
||
” – uncrowned, deprived of the crown] refers to its non-denticulate frontoclypeus and genae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAC36573FFCF8B64FF61F93093C4F87D" pageId="1" pageNumber="419" type="diagnosis">
|
||
<paragraph id="926636F8FFCF8B64FF61F9309247F8ED" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF61F930968EF95F" bold="true" box="[199,461,1732,1759]" pageId="1" pageNumber="419">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="55D94D7BFFCF8B64FE72F93195B4F95F" authority="Alekseev & Bukejs & Sontag, 2020" authorityName="Alekseev & Bukejs & Sontag" authorityYear="2020" box="[468,759,1733,1759]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="species" species="incoronatus" status="sp. nov.">
|
||
<emphasis id="A0ADEAEAFFCF8B64FE72F93195B4F95F" box="[468,759,1733,1759]" italics="true" pageId="1" pageNumber="419">Eledonoprius incoronatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0ADEAEAFFCF8B64FD5BF9319415F95F" bold="true" box="[765,854,1733,1759]" pageId="1" pageNumber="419">
|
||
<taxonomicNameLabel id="BB9E5791FFCF8B64FD5BF9319415F95F" box="[765,854,1733,1759]" pageId="1" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is placed in the genus
|
||
<taxonomicName id="55D94D7BFFCF8B64FBF2F93193A8F95F" authorityName="Reitter" authorityYear="1911" box="[1108,1259,1733,1759]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0ADEAEAFFCF8B64FBF2F93193A8F95F" box="[1108,1259,1733,1759]" italics="true" pageId="1" pageNumber="419">Eledonoprius</emphasis>
|
||
</taxonomicName>
|
||
within the tribe
|
||
<taxonomicName id="55D94D7BFFCF8B64FF31F91D9672F883" authorityName="Kirby" authorityYear="1837" box="[151,305,1769,1795]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="tribe" tribe="Bolitophagini">Bolitophagini</taxonomicName>
|
||
on the basis of the following morphological characters: small size (slightly larger than
|
||
<quantity id="55219B1DFFCF8B64FAB5F91D921AF883" box="[1299,1369,1769,1795]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="419" unit="mm" value="2.0">2 mm</quantity>
|
||
); pronotal lateral sides widely explanate, denticulate and not-margined; anterior pronotal angles distinct but not strongly produced forwards; elytral intervals and pronotal surface with sharp granules; eyes distinctly and deeply emarginate by genae anteriorly; head with pair of short horns; epipleuron distinctly wider than metanepisternum.
|
||
</paragraph>
|
||
<paragraph id="926636F8FFCF8B64FF61F88393C4F87D" blockId="1.[151,1437,522,2045]" pageId="1" pageNumber="419">
|
||
The newly described fossil species clearly differs from the extant European
|
||
<taxonomicName id="55D94D7BFFCF8B64FB8EF88392DEF812" authority="(Reitter)" baseAuthorityName="Reitter" baseAuthorityYear="1890" box="[1064,1437,1911,1938]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="species" species="serrifrons">
|
||
<emphasis id="A0ADEAEAFFCF8B64FB8EF8839276F811" box="[1064,1333,1911,1937]" italics="true" pageId="1" pageNumber="419">Eledonoprius serrifrons</emphasis>
|
||
(Reitter)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="55D94D7BFFCF8B64FF63F86896E7F835" authority="(Panzer)" baseAuthorityName="Panzer" baseAuthorityYear="1799" box="[197,420,1947,1973]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="A0ADEAEAFFCF8B64FF63F868967CF835" box="[197,319,1948,1973]" italics="true" pageId="1" pageNumber="419">E. armatus</emphasis>
|
||
(Panzer)
|
||
</taxonomicName>
|
||
in having the frontoclypeus and genae without denticulation (in contrast both extant species have distinct, well-developed denticulation on the head anteriorly); less numerous lateral denticles of pronotum; apical antennomere shorter than antennomeres 9 and 10 combined; and smaller body size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C6A66670FFCC8B67FF31F855960CF85D" ID-DOI="http://doi.org/10.5281/zenodo.3707535" ID-Zenodo-Dep="3707535" httpUri="https://zenodo.org/record/3707535/files/figure.png" pageId="2" pageNumber="420" startId="2.[151,264,1953,1978]" targetBox="[154,1433,181,1928]" targetPageId="2">
|
||
<paragraph id="926636F8FFCC8B67FF31F855960CF85D" blockId="2.[151,1436,1953,2014]" pageId="2" pageNumber="420">
|
||
<emphasis id="A0ADEAEAFFCC8B67FF31F855967EF83A" bold="true" box="[151,317,1953,1978]" pageId="2" pageNumber="420">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="55D94D7BFFCC8B67FEE3F856950DF83A" authority="Alekseev & Bukejs & Sontag, 2020" authorityName="Alekseev & Bukejs & Sontag" authorityYear="2020" box="[325,590,1954,1978]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="420" phylum="Arthropoda" rank="species" species="incoronatus" status="sp. nov.">
|
||
<emphasis id="A0ADEAEAFFCC8B67FEE3F856950DF83A" box="[325,590,1954,1978]" italics="true" pageId="2" pageNumber="420">Eledonoprius incoronatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0ADEAEAFFCC8B67FDF1F85695EAF83A" bold="true" box="[599,681,1954,1978]" pageId="2" pageNumber="420">
|
||
<taxonomicNameLabel id="BB9E5791FFCC8B67FDF1F85695EAF83A" box="[599,681,1954,1978]" pageId="2" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype: 1 – habitus, dorsal view; 2 – habitus, lateral view. Scale bars represent 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C6A66670FFCD8B66FF31F85F952FF868" ID-DOI="http://doi.org/10.5281/zenodo.3707537" ID-Zenodo-Dep="3707537" httpUri="https://zenodo.org/record/3707537/files/figure.png" pageId="3" pageNumber="421" startId="3.[151,264,1963,1988]" targetBox="[158,1429,199,1938]" targetPageId="3">
|
||
<paragraph id="926636F8FFCD8B66FF31F85F952FF868" blockId="3.[151,1436,1963,2024]" pageId="3" pageNumber="421">
|
||
<emphasis id="A0ADEAEAFFCD8B66FF31F85F9678F844" bold="true" box="[151,315,1963,1988]" pageId="3" pageNumber="421">FIGURES 3–4.</emphasis>
|
||
<taxonomicName id="55D94D7BFFCD8B66FEE4F85F950AF843" authority="Alekseev & Bukejs & Sontag, 2020" authorityName="Alekseev & Bukejs & Sontag" authorityYear="2020" box="[322,585,1963,1987]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="421" phylum="Arthropoda" rank="species" species="incoronatus" status="sp. nov.">
|
||
<emphasis id="A0ADEAEAFFCD8B66FEE4F85F950AF843" box="[322,585,1963,1987]" italics="true" pageId="3" pageNumber="421">Eledonoprius incoronatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0ADEAEAFFCD8B66FDF6F85F95E3F843" bold="true" box="[592,672,1963,1987]" pageId="3" pageNumber="421">
|
||
<taxonomicNameLabel id="BB9E5791FFCD8B66FDF6F85F95E3F843" box="[592,672,1963,1987]" pageId="3" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype: 3 – habitus, ventral view; 4 – forebody, ventral view. Scale bars represent 0.5 mm (Fig. 3) and 0.2 mm (Fig.4).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C6A66670FFCA8B61FF31F857960CF85F" ID-DOI="http://doi.org/10.5281/zenodo.3707539" ID-Zenodo-Dep="3707539" httpUri="https://zenodo.org/record/3707539/files/figure.png" pageId="4" pageNumber="422" startId="4.[151,264,1955,1980]" targetBox="[155,1433,181,1929]" targetPageId="4">
|
||
<paragraph id="926636F8FFCA8B61FF31F857960CF85F" blockId="4.[151,1436,1955,2016]" pageId="4" pageNumber="422">
|
||
<emphasis id="A0ADEAEAFFCA8B61FF31F8579678F83C" bold="true" box="[151,315,1955,1980]" pageId="4" pageNumber="422">FIGURES 5–6.</emphasis>
|
||
<taxonomicName id="55D94D7BFFCA8B61FEE4F850950AF83B" authority="Alekseev & Bukejs & Sontag, 2020" authorityName="Alekseev & Bukejs & Sontag" authorityYear="2020" box="[322,585,1956,1980]" class="Insecta" family="Tenebrionidae" genus="Eledonoprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="422" phylum="Arthropoda" rank="species" species="incoronatus" status="sp. nov.">
|
||
<emphasis id="A0ADEAEAFFCA8B61FEE4F850950AF83B" box="[322,585,1956,1980]" italics="true" pageId="4" pageNumber="422">Eledonoprius incoronatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0ADEAEAFFCA8B61FDF6F85095E3F83C" bold="true" box="[592,672,1956,1980]" pageId="4" pageNumber="422">
|
||
<taxonomicNameLabel id="BB9E5791FFCA8B61FDF6F85095E3F83C" box="[592,672,1956,1980]" pageId="4" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype: 5 – genitalia, lateral view; 6 – forebody, lateral view. Scale bars represent 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DAC36573FFCB8B60FF61FF6D95F8FEC3" pageId="5" pageNumber="423" type="discussion">
|
||
<paragraph id="926636F8FFCB8B60FF61FF6D95F8FEC3" blockId="5.[151,1437,152,323]" pageId="5" pageNumber="423">
|
||
<emphasis id="A0ADEAEAFFCB8B60FF61FF6D9677FF33" bold="true" box="[199,308,153,179]" pageId="5" pageNumber="423">Remarks</emphasis>
|
||
. The new species superficially resembles (possibly, due to evolutionary parallelism and adaptation to similar ecological niche) the zopherid species
|
||
<taxonomicName id="55D94D7BFFCB8B60FD1DFF48938BFF56" authority="Alekseev & Bukejs, 2016" authorityName="Alekseev & Bukejs" authorityYear="2016" box="[699,1224,188,215]" class="Insecta" family="Tenebrionidae" genus="Endophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="423" phylum="Arthropoda" rank="species" species="gorskii">
|
||
<emphasis id="A0ADEAEAFFCB8B60FD1DFF4894E1FF56" box="[699,930,188,214]" italics="true" pageId="5" pageNumber="423">Endophloeus gorskii</emphasis>
|
||
<bibRefCitation id="F6484B09FFCB8B60FC01FF48938BFF56" author="Alekseev, V. I. & Bukejs, A." box="[935,1224,188,215]" pageId="5" pageNumber="423" pagination="409 - 427" refId="ref2220" refString="Alekseev, V. I. & Bukejs, A. (2016) New Zopheridae (Coleoptera: Tenebrionoidea) from Baltic amber. Zootaxa, 4178 (3), 409 - 427. https: // doi. org / 10.11646 / zootaxa. 4178.3.6" type="journal article" year="2016">Alekseev & Bukejs, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from Baltic amber (
|
||
<bibRefCitation id="F6484B09FFCB8B60FF39FF149682FF7B" author="Alekseev, V. I. & Bukejs, A." box="[159,449,224,251]" pageId="5" pageNumber="423" pagination="409 - 427" refId="ref2220" refString="Alekseev, V. I. & Bukejs, A. (2016) New Zopheridae (Coleoptera: Tenebrionoidea) from Baltic amber. Zootaxa, 4178 (3), 409 - 427. https: // doi. org / 10.11646 / zootaxa. 4178.3.6" type="journal article" year="2016">Alekseev & Bukejs 2016</bibRefCitation>
|
||
), but differs in is glabrous body (elytra with apparent scale-like setae in
|
||
<taxonomicName id="55D94D7BFFCB8B60FB5CFF1492CFFF7A" box="[1274,1420,224,250]" class="Insecta" family="Tenebrionidae" genus="Endophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="423" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0ADEAEAFFCB8B60FB5CFF1492CFFF7A" box="[1274,1420,224,250]" italics="true" pageId="5" pageNumber="423">Endophloeus</emphasis>
|
||
</taxonomicName>
|
||
), tarsal formula 5-5-4 (4-4-
|
||
<quantity id="55219B1DFFCB8B60FE14FEF1969EFE9E" box="[434,477,260,287]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="423" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="55D94D7BFFCB8B60FE45FEF09536FE9E" box="[483,629,260,286]" class="Insecta" family="Tenebrionidae" genus="Endophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="423" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0ADEAEAFFCB8B60FE45FEF09536FE9E" box="[483,629,260,286]" italics="true" pageId="5" pageNumber="423">Endophloeus</emphasis>
|
||
</taxonomicName>
|
||
), antennae without distinct club (in contrast to distinct 2-segmented club in
|
||
<taxonomicName id="55D94D7BFFCB8B60FF12FEDC9605FEC2" box="[180,326,296,322]" class="Insecta" family="Tenebrionidae" genus="Endophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="423" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0ADEAEAFFCB8B60FF12FEDC9605FEC2" box="[180,326,296,322]" italics="true" pageId="5" pageNumber="423">Endophloeus</emphasis>
|
||
</taxonomicName>
|
||
), and more transverse pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |