281 lines
34 KiB
XML
281 lines
34 KiB
XML
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<mods:title id="3B852EE936A5C01BEA97EFD5261BB7FC">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="F89D5C679D919DF43D4ADA97B3B1A238">Silva, Fernando A. B.</mods:namePart>
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<treatment id="03DE87D5853AFF885991FEB051E3FDCC" ID-DOI="http://doi.org/10.5281/zenodo.6115686" ID-GBIF-Taxon="119598922" ID-Zenodo-Dep="6115686" LSID="urn:lsid:plazi:treatment:03DE87D5853AFF885991FEB051E3FDCC" httpUri="http://treatment.plazi.org/id/03DE87D5853AFF885991FEB051E3FDCC" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
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<paragraph id="8BC836C3853AFF895991FEB05733FEB7" blockId="8.[151,636,329,355]" box="[151,636,329,355]" pageId="8" pageNumber="459">
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<heading id="D08081AF853AFF895991FEB05733FEB7" bold="true" box="[151,636,329,355]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
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<emphasis id="B903EAD1853AFF895991FEB05733FEB7" bold="true" box="[151,636,329,355]" pageId="8" pageNumber="459">
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Subgenus
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<taxonomicName id="4C774D40853AFF895811FEB05733FEB7" authority="Kolbe 1893" authorityName="Kolbe" authorityYear="1893" box="[279,636,329,355]" class="Insecta" family="Scarabaeidae" genus="Aganhyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EAD1853AFF895811FEB054A8FEB7" bold="true" box="[279,487,329,355]" italics="true" pageId="8" pageNumber="459">AGANHYBOMA</emphasis>
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<bibRefCitation id="EFE64B32853AFF8958E8FEB05733FEB7" author="Kolbe" box="[494,636,329,355]" pageId="8" pageNumber="459" refString="Kolbe, H. J. (1893) Beitrage zur kenntniss der mistkafer, Lamellicornia onthophila. Stettiner entomologische Zeitung, 54, 188 - 202." type="journal article" year="1893">Kolbe 1893</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C36D6548853AFF895991FE6956A1FBFE" pageId="8" pageNumber="459" type="reference_group">
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<paragraph id="8BC836C3853AFF895991FE69578DFDF4" blockId="8.[151,1013,400,1066]" pageId="8" pageNumber="459">
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<treatmentCitationGroup id="AB6711ED853AFF895991FE69578DFDF4" pageId="8" pageNumber="459">
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<taxonomicName id="4C774D40853AFF895991FE69572EFE72" ID-CoL="8CVZ" authority="Kolbe 1893: 192" authorityName="Kolbe" authorityPageNumber="192" authorityYear="1893" box="[151,609,400,422]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="subGenus" subGenus="Aganhyboma">
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<emphasis id="B903EAD1853AFF895991FE69545BFE72" box="[151,276,400,422]" italics="true" pageId="8" pageNumber="459">Deltochilum</emphasis>
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(
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<emphasis id="B903EAD1853AFF895825FE6954E6FE72" box="[291,425,400,422]" italics="true" pageId="8" pageNumber="459">Aganhyboma</emphasis>
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)
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<treatmentCitation id="0AD610D2853AFF8958B1FE69572EFE72" author="Kolbe" box="[439,609,400,422]" page="192" pageId="8" pageNumber="459" year="1893">
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<bibRefCitation id="EFE64B32853AFF8958B1FE69577EFE72" author="Kolbe" box="[439,561,400,422]" pageId="8" pageNumber="459" refString="Kolbe, H. J. (1893) Beitrage zur kenntniss der mistkafer, Lamellicornia onthophila. Stettiner entomologische Zeitung, 54, 188 - 202." type="journal article" year="1893">Kolbe 1893</bibRefCitation>
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: 192
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</treatmentCitation>
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</taxonomicName>
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(original description)
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<treatmentCitation id="0AD610D2853AFF895991FE575471FE10" author="Shipp" box="[151,318,430,452]" page="195" pageId="8" pageNumber="459" year="1897">
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<bibRefCitation id="EFE64B32853AFF895991FE575441FE10" author="Shipp" box="[151,270,430,452]" pageId="8" pageNumber="459" refString="Shipp, J. W. (1897) On the synonymy of the genus Deltochilum. Entomologische Nachrichten, 23, 194 - 196." type="journal article" year="1897">Shipp 1897</bibRefCitation>
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: 195
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</treatmentCitation>
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||
(designation of
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<typeStatus id="54CC8861853AFF8958E5FE565740FE10" box="[483,527,431,452]" pageId="8" pageNumber="459">type</typeStatus>
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species; author treated
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<taxonomicName id="4C774D40853AFF895BF8FE5756CBFE10" box="[766,900,430,452]" class="Insecta" family="Scarabaeidae" genus="Aganhyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EAD1853AFF895BF8FE5756CBFE10" box="[766,900,430,452]" italics="true" pageId="8" pageNumber="459">Aganhyboma</emphasis>
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</taxonomicName>
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as genus)
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<treatmentCitation id="0AD610D2853AFF895991FE34541DFE37" author="Paulian" box="[151,338,461,483]" page="240" pageId="8" pageNumber="459" year="1938">
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<bibRefCitation id="EFE64B32853AFF895991FE345450FE37" author="Paulian" box="[151,287,461,483]" pageId="8" pageNumber="459" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
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: 240
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</treatmentCitation>
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, 243, 252 (identification keys, species descriptions)
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<treatmentCitation id="0AD610D2853AFF895991FE145477FDD6" author="Lane" box="[151,312,492,514]" page="172" pageId="8" pageNumber="459" year="1946">
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<bibRefCitation id="EFE64B32853AFF895991FE145449FDD6" author="Lane" box="[151,262,492,514]" pageId="8" pageNumber="459" refString="Lane, F. (1946) Sobre os tipos e a sinonimia de alguns Canthonini (Col. Scarabaeidae). Papeis Avulsos do Departamento de Zoologia, 7, 171 - 179." type="journal article" year="1946">Lane 1946</bibRefCitation>
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: 172
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||
</treatmentCitation>
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, 173, 174 (comment about history of names in
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<taxonomicName id="4C774D40853AFF895A10FE1556D4FDD6" box="[790,923,492,514]" class="Insecta" family="Scarabaeidae" genus="Aganhyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EAD1853AFF895A10FE1556D4FDD6" box="[790,923,492,514]" italics="true" pageId="8" pageNumber="459">Aganhyboma</emphasis>
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</taxonomicName>
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||
)
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<treatmentCitation id="0AD610D2853AFF895991FDF35486FDF4" author="Pereira" box="[151,457,522,544]" page="120" pageId="8" pageNumber="459" year="1956">
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||
<bibRefCitation id="EFE64B32853AFF895991FDF354D8FDF4" author="Pereira" box="[151,407,522,544]" pageId="8" pageNumber="459" refString="Pereira, F. S. & Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira & Martínez 1956</bibRefCitation>
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: 120
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||
</treatmentCitation>
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, 121 (identification key)
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BC836C3853AFF895991FDD05485FDEB" blockId="8.[151,1013,400,1066]" box="[151,458,553,575]" pageId="8" pageNumber="459">
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<bibRefCitation id="EFE64B32853AFF895991FDD05460FDEB" author="Martinez" box="[151,303,553,575]" pageId="8" pageNumber="459" refString="Martinez, A. (1959) Catalogo de los Scarabaeidae argentinos. Revista del Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", 5, 1 - 126." type="journal article" year="1959">Martínez 1959</bibRefCitation>
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: 51 (catalogue)
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</paragraph>
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<paragraph id="8BC836C3853AFF895991FDB15778FD8A" blockId="8.[151,1013,400,1066]" box="[151,567,584,606]" pageId="8" pageNumber="459">
|
||
<bibRefCitation id="EFE64B32853AFF895991FDB154C0FD8A" author="Vulcano" box="[151,399,584,606]" pageId="8" pageNumber="459" refString="Vulcano, M. A. & Pereira, F. S. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the western hemisphere. Entomologischen Arbeiten aus dem Museum G. Frey, 15, 569 - 685." type="journal article" year="1964">Vulcano & Pereira 1964</bibRefCitation>
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: 641 (catalogue)
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</paragraph>
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||
<paragraph id="8BC836C3853AFF895991FD9F578AFDA8" blockId="8.[151,1013,400,1066]" box="[151,709,614,636]" pageId="8" pageNumber="459">
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||
<bibRefCitation id="EFE64B32853AFF895991FD9F54C0FDA8" author="Vulcano" box="[151,399,614,636]" pageId="8" pageNumber="459" refString="Vulcano, M. A. & Pereira, F. S. (1967) Sinopse dos Passalidae e Scarabaeidae s. str. da regiao Amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica (Zoologia), 5, 533 - 603." type="journal article" year="1967">Vulcano & Pereira 1967</bibRefCitation>
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: 555, 556 (identification keys)
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</paragraph>
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||
<paragraph id="8BC836C3853AFF895991FD7C574FFD4F" blockId="8.[151,1013,400,1066]" box="[151,512,645,667]" pageId="8" pageNumber="459">Vaz-de-Mello 2000: 192 (checklist)</paragraph>
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||
<paragraph id="8BC836C3853AFF895991FD5D5760FCC2" blockId="8.[151,1013,400,1066]" pageId="8" pageNumber="459">
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||
Vaz-de-Mello
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<emphasis id="B903EAD1853AFF89582EFD5C541AFD6E" box="[296,341,676,698]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
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||
. 2011: 5, 11, 18, 26, 33, 41, 44 (checklist and identification keys).
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||
<taxonomicName id="4C774D40853AFF895991FD3B54E6FD0C" ID-CoL="42SM" box="[151,425,706,728]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EAD1853AFF895991FD3B545BFD0C" box="[151,276,706,728]" italics="true" pageId="8" pageNumber="459">Deltochilum</emphasis>
|
||
(
|
||
<emphasis id="B903EAD1853AFF895825FD3B54EFFD0C" box="[291,416,706,728]" italics="true" pageId="8" pageNumber="459">Deltochilum</emphasis>
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||
)
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||
</taxonomicName>
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||
in part
|
||
<bibRefCitation id="EFE64B32853AFF8958F0FD3B57E4FD0C" author="Eschscholtz" box="[502,683,706,728]" pageId="8" pageNumber="459" refString="Eschscholtz, J. F. (1822) Entomographien. Vol. I. G. Reimer, Berlin, 128 pp." type="book" year="1822">Eschscholtz 1822</bibRefCitation>
|
||
: 39 (original description)
|
||
<bibRefCitation id="EFE64B32853AFF895991FD18545EFD23" author="Kolbe" box="[151,273,737,759]" pageId="8" pageNumber="459" refString="Kolbe, H. J. (1893) Beitrage zur kenntniss der mistkafer, Lamellicornia onthophila. Stettiner entomologische Zeitung, 54, 188 - 202." type="journal article" year="1893">Kolbe 1893</bibRefCitation>
|
||
: 191 (establishement of seven subgenera under
|
||
<taxonomicName id="4C774D40853AFF895BF3FD18563DFD23" ID-CoL="42SM" box="[757,882,737,759]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EAD1853AFF895BF3FD18563DFD23" box="[757,882,737,759]" italics="true" pageId="8" pageNumber="459">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<bibRefCitation id="EFE64B32853AFF895991FCF95441FCC2" author="Shipp" box="[151,270,768,790]" pageId="8" pageNumber="459" refString="Shipp, J. W. (1897) On the synonymy of the genus Deltochilum. Entomologische Nachrichten, 23, 194 - 196." type="journal article" year="1897">Shipp 1897</bibRefCitation>
|
||
: 196 (taxonomic comments)
|
||
</paragraph>
|
||
<paragraph id="8BC836C3853AFF895991FCE754BFFC44" blockId="8.[151,1013,400,1066]" pageId="8" pageNumber="459">
|
||
<bibRefCitation id="EFE64B32853AFF895991FCE75450FCE0" author="Paulian" box="[151,287,798,820]" pageId="8" pageNumber="459" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
|
||
: 240, 243, 268, 269 (description, identification key and comments)
|
||
<bibRefCitation id="EFE64B32853AFF895991FCC75449FC87" author="Lane" box="[151,262,829,851]" pageId="8" pageNumber="459" refString="Lane, F. (1946) Sobre os tipos e a sinonimia de alguns Canthonini (Col. Scarabaeidae). Papeis Avulsos do Departamento de Zoologia, 7, 171 - 179." type="journal article" year="1946">Lane 1946</bibRefCitation>
|
||
: 172–175 (transfer of the
|
||
<taxonomicName id="4C774D40853AFF895B0DFCC45719FC87" ID-CoL="34KB6" box="[523,598,829,851]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="valgum">
|
||
<emphasis id="B903EAD1853AFF895B0DFCC45719FC87" box="[523,598,829,851]" italics="true" pageId="8" pageNumber="459">valgum</emphasis>
|
||
</taxonomicName>
|
||
group to
|
||
<taxonomicName id="4C774D40853AFF895BB1FCC45670FC87" box="[695,831,829,851]" class="Insecta" family="Scarabaeidae" genus="Deltohyboma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EAD1853AFF895BB1FCC45670FC87" box="[695,831,829,851]" italics="true" pageId="8" pageNumber="459">Deltohyboma</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C774D40853AFF895991FCA554FCFCA6" ID-CoL="42SM" authority="(Deltohyboma)" baseAuthorityName="Deltohyboma" box="[151,435,860,882]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EAD1853AFF895991FCA5545BFCA6" box="[151,276,860,882]" italics="true" pageId="8" pageNumber="459">Deltochilum</emphasis>
|
||
(
|
||
<emphasis id="B903EAD1853AFF895825FCA554E4FCA6" box="[291,427,860,882]" italics="true" pageId="8" pageNumber="459">Deltohyboma</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
in part
|
||
<bibRefCitation id="EFE64B32853AFF895B07FCA4573FFCA6" author="Lane" box="[513,624,860,882]" pageId="8" pageNumber="459" refString="Lane, F. (1946) Sobre os tipos e a sinonimia de alguns Canthonini (Col. Scarabaeidae). Papeis Avulsos do Departamento de Zoologia, 7, 171 - 179." type="journal article" year="1946">Lane 1946</bibRefCitation>
|
||
: 175 (designation of subgenus)
|
||
<bibRefCitation id="EFE64B32853AFF895991FC835460FC44" author="Martinez" box="[151,303,890,912]" pageId="8" pageNumber="459" refString="Martinez, A. (1959) Catalogo de los Scarabaeidae argentinos. Revista del Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", 5, 1 - 126." type="journal article" year="1959">Martínez 1959</bibRefCitation>
|
||
: 53–56 (catalogue)
|
||
</paragraph>
|
||
<paragraph id="8BC836C3853AFF895991FC60578DFC7B" blockId="8.[151,1013,400,1066]" box="[151,706,921,943]" pageId="8" pageNumber="459">
|
||
<bibRefCitation id="EFE64B32853AFF895991FC6054D8FC7B" author="Pereira" box="[151,407,921,943]" pageId="8" pageNumber="459" refString="Pereira, F. S. & Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira & Martínez 1956</bibRefCitation>
|
||
: 120, 121 (identification key)
|
||
</paragraph>
|
||
<paragraph id="8BC836C3853AFF895991FC415778FC1A" blockId="8.[151,1013,400,1066]" box="[151,567,952,974]" pageId="8" pageNumber="459">
|
||
<bibRefCitation id="EFE64B32853AFF895991FC4154C0FC1A" author="Vulcano" box="[151,399,952,974]" pageId="8" pageNumber="459" refString="Vulcano, M. A. & Pereira, F. S. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the western hemisphere. Entomologischen Arbeiten aus dem Museum G. Frey, 15, 569 - 685." type="journal article" year="1964">Vulcano & Pereira 1964</bibRefCitation>
|
||
: 641 (catalogue)
|
||
</paragraph>
|
||
<paragraph id="8BC836C3853AFF895991FC2F578AFC38" blockId="8.[151,1013,400,1066]" box="[151,709,982,1004]" pageId="8" pageNumber="459">
|
||
<bibRefCitation id="EFE64B32853AFF895991FC2F54C0FC38" author="Vulcano" box="[151,399,982,1004]" pageId="8" pageNumber="459" refString="Vulcano, M. A. & Pereira, F. S. (1967) Sinopse dos Passalidae e Scarabaeidae s. str. da regiao Amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica (Zoologia), 5, 533 - 603." type="journal article" year="1967">Vulcano & Pereira 1967</bibRefCitation>
|
||
: 555–560 (identification keys)
|
||
</paragraph>
|
||
<paragraph id="8BC836C3853AFF895991FC0C574FFBDF" blockId="8.[151,1013,400,1066]" box="[151,512,1013,1035]" pageId="8" pageNumber="459">Vaz-de-Mello 2000: 192 (checklist)</paragraph>
|
||
<paragraph id="8BC836C3853AFF895991FBED56A1FBFE" blockId="8.[151,1013,400,1066]" box="[151,1006,1044,1066]" pageId="8" pageNumber="459">
|
||
Vaz-de-Mello
|
||
<emphasis id="B903EAD1853AFF89582EFBEC541AFBFE" box="[296,341,1044,1066]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
|
||
. 2011: 5, 11, 18, 26, 33, 41, 44 (checklist and identification keys)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6548853AFF885991FBAF57C9FF2C" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="description">
|
||
<paragraph id="8BC836C3853AFF885991FBAF57C9FF2C" blockId="8.[151,1437,1110,2035]" lastBlockId="9.[151,1437,151,536]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
|
||
<emphasis id="B903EAD1853AFF895991FBAF543BFBBB" bold="true" box="[151,372,1110,1135]" pageId="8" pageNumber="459">Description. Body.</emphasis>
|
||
Lateral margins as well as dorsal surface vary from weakly to strongly convex (
|
||
<figureCitation id="134C2A46853AFF895DF8FBAF50C0FBBB" box="[1278,1423,1110,1135]" captionStart-0="FIGURES 95 – 102" captionStart-1="FIGURES 103 – 110" captionStart-2="FIGURES 111 – 118" captionStart-3="FIGURES 119 – 123" captionStartId-0="48.[151,264,1951,1974]" captionStartId-1="49.[151,264,1951,1974]" captionStartId-2="50.[151,264,1952,1975]" captionStartId-3="51.[151,264,1596,1619]" captionTargetBox-0="[181,1434,202,1924]" captionTargetBox-1="[197,1421,212,1926]" captionTargetBox-2="[201,1409,198,1924]" captionTargetBox-3="[178,1436,193,1570]" captionTargetId-0="figure@48.[152,1434,193,1930]" captionTargetId-1="figure@49.[165,1421,193,1931]" captionTargetId-2="figure@50.[175,1412,193,1931]" captionTargetId-3="figure@51.[150,1436,193,1576]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionTargetPageId-2="50" captionTargetPageId-3="51" captionText-0="FIGURES 95 – 102. Species of D. (Aganhyboma). 95. D. trisignatum. 96. D. kolbei. 97. D. violaceum. 98. D. amandaarcanjoae new species. 99. D. cupreicolle. 100. D. titovidaurrei new species. 101. D. viridescens new status. 102. D. viridescens new status. Figures scale = 1 mm." captionText-1="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." captionText-2="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." captionText-3="FIGURES 119 – 123. Species of D. (Aganhyboma). 119. D. ritamourae new species. 120. D. icariforme. 121. D. paresi new species. 122. D. kolleri new species. 123. D. subrubrum new species. Figures scale = 1 mm." httpUri-0="https://zenodo.org/record/244807/files/figure.png" httpUri-1="https://zenodo.org/record/244808/files/figure.png" httpUri-2="https://zenodo.org/record/244809/files/figure.png" httpUri-3="https://zenodo.org/record/244810/files/figure.png" pageId="8" pageNumber="459">Figs 95–123</figureCitation>
|
||
), with some species quite round in shape. Dorsal surface glabrous, with at most minute setae in some species. Length
|
||
<quantity id="4C8F9B26853AFF895991FB66547EFB62" box="[151,305,1183,1207]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="8" pageNumber="459" unit="mm" value="15.0" valueMax="19.0" valueMin="11.0">11 to 19 mm</quantity>
|
||
.
|
||
<emphasis id="B903EAD1853AFF895842FB6754C4FB63" bold="true" box="[324,395,1182,1207]" pageId="8" pageNumber="459">Head.</emphasis>
|
||
Surface with foveiform punctures, usually more conspicuous around eyes (Figs 60, 61). Anterior portion (between clypeogenal sutures) prolonged, similar to a small ‘muzzle’ (Figs 60, 61), bidentate. Margin of head, near clypeal teeth, straight and followed by a small tooth slightly pronounced (Fig. 61), or slightly curved and without teeth (Fig. 60). Lateral margin of gena curved outward until eyes, with a strong convex shape in the middle of this curvature, or with posterior half truncated, approximately straight. Clypeogenal suture well defined and shallow. Eye margin not emarginate. Eye rounded or comma-shaped in dorsal view. Dorsal interocular distance varies between three to ten times eye width (Figs 60, 61).
|
||
<emphasis id="B903EAD1853AFF895A94FA8F56BDFA5B" bold="true" box="[914,1010,1398,1423]" pageId="8" pageNumber="459">Thorax.</emphasis>
|
||
Pronotum with anterior angles acute, directed anteriorly. Lateral margin slightly angled. Pronotum punctures variable, dense or sparse, consisting of foveiform punctures. Sides of metasternum, anterior portion of hypomera, epimeron and episternum with a dense foveiform punctures. Hypomera not excavated to receive head margin (Figs 78, 79). Posterior portion of metasternum with a small, round, shallow fovea. Outer margin of mesocoxal cavity not reaching metaepimeron.
|
||
<emphasis id="B903EAD1853AFF895991F9D355A5F997" bold="true" box="[151,234,1578,1603]" pageId="8" pageNumber="459">Elytra.</emphasis>
|
||
Lateral margins ranging from semiparallel to outwardly curved. Elytral striae with foveiform punctures. Interstriae with foveiform punctures of different sizes. Basal carina of sixth elytral interstria absent. Basal carina of seventh interstria short or elongate, prominent or not (Figs 71, 72). Length of basal carina of ninth interstria reaches from mid-interstria, to apex (Figs 62, 63, 80). Apices of interstriae 2, 3, 4, 5, 6 and 7 with tubercle or carina (that of second interstria sometimes weak or absent). Length of apical carina of seventh interstria less than one-half that of interstria. Pseudepipleural carina complete, visible entire length of epipleuron in most species (Figs 62, 63). In some species this carina can be interrupted in the basal third, re-appearing halfway along elytral length (Fig. 80).
|
||
<emphasis id="B903EAD1853AFF895991F8DF5443F8EB" bold="true" box="[151,268,1830,1855]" pageId="8" pageNumber="459">Abdomen</emphasis>
|
||
. Abdominal ventrites shorter medially than laterally (Fig. 73). Punctures of ventrites most abundant and conspicuous at sides. Pygidium with rounded or elliptical foveiform punctures. Apex of pygidium acuminate or rounded.
|
||
<emphasis id="B903EAD1853AFF895806F8975470F853" bold="true" box="[256,319,1902,1927]" pageId="8" pageNumber="459">Legs.</emphasis>
|
||
Protibia abruptly expanded (Fig. 66) or not in basal third. External apical margin of protibia bearing three large prominent teeth. Medial lateral tooth can be equidistant between apical and basal tooth, or closer to apical tooth. Metatibia short, truncated and only slightly curved inward (Fig. 68), or elongate and apical one-third strongly curved inward (Fig. 67).
|
||
<emphasis id="B903EAD1853AFF895B48F8235787F827" bold="true" box="[590,712,2010,2035]" pageId="8" pageNumber="459">Aedeagus.</emphasis>
|
||
Parameres symmetric, short or elongate, simple, without ornamentation or setae (Fig. 24); endophallus: lobe in submedial portion; endophallic sclerites: lamella copulatrix sclerites (LC), superior right peripheral sclerite (SRP), fronto-lateral peripheral sclerite (FLP) and complex of axial and subaxial sclerites (A + SA) (
|
||
<figureCitation id="134C2A46853BFF885B03FF265704FF2C" box="[517,587,223,248]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1892,1915]" captionTargetBox="[168,1431,864,1864]" captionTargetId="figure@3.[151,1436,855,1871]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Male genital sclerites in Deltochilum (Aganhyboma). 1. Position and nomenclature of endophallic sclerites (modified from Medina et al. 2013). 2. Nomenclature of the main parts of the genital segment (D. amandaarcanjoae new species). 3. Nomenclature of the main parts of the superior right peripheral sclerite (SRP) (D. paresi new species). Figures scale = 1 mm." httpUri="https://zenodo.org/record/244796/files/figure.png" pageId="9" pageNumber="460">Figs 1</figureCitation>
|
||
, 59).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6548853BFF8859C1FEFD5461FE94" pageId="9" pageNumber="460" type="diagnosis">
|
||
<paragraph id="8BC836C3853BFF8859C1FEFD5461FE94" blockId="9.[151,1437,151,536]" pageId="9" pageNumber="460">
|
||
<emphasis id="B903EAD1853BFF8859C1FEFD540EFEC9" bold="true" box="[199,321,260,285]" pageId="9" pageNumber="460">Diagnosis.</emphasis>
|
||
Anterior portion of head (between clypeogenal sutures) prolonged, similar to a small ‘muzzle’ (Figs 60, 61).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6548853BFF8859C1FEB551E3FDCC" pageId="9" pageNumber="460" type="discussion">
|
||
<paragraph id="8BC836C3853BFF8859C1FEB551E3FDCC" blockId="9.[151,1437,151,536]" pageId="9" pageNumber="460">
|
||
<emphasis id="B903EAD1853BFF8859C1FEB55474FEB1" bold="true" box="[199,315,332,357]" pageId="9" pageNumber="460">Remarks.</emphasis>
|
||
The subgenus
|
||
<taxonomicName id="4C774D40853BFF8858E5FEB55738FEB0" box="[483,631,332,356]" class="Insecta" family="Scarabaeidae" genus="Aganhyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EAD1853BFF8858E5FEB55738FEB0" box="[483,631,332,356]" italics="true" pageId="9" pageNumber="460">Aganhyboma</emphasis>
|
||
</taxonomicName>
|
||
, as defined here, includes 26 species distributed in South
|
||
<collectingCountry id="F3607653853BFF885DFAFEB5502FFEB1" box="[1276,1376,332,357]" name="United States of America" pageId="9" pageNumber="460">America</collectingCountry>
|
||
, with one species (
|
||
<taxonomicName id="4C774D40853BFF88582AFE88574FFE5C" authority="Bates" authorityName="Bates" box="[300,512,368,392]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="acropyge">
|
||
<emphasis id="B903EAD1853BFF88582AFE8854F8FE5C" box="[300,439,369,392]" italics="true" pageId="9" pageNumber="460">D. acropyge</emphasis>
|
||
Bates
|
||
</taxonomicName>
|
||
) reaching as far North as
|
||
<collectingCountry id="F3607653853BFF885A2BFE9656C9FE5C" box="[813,902,367,392]" name="Mexico" pageId="9" pageNumber="460">Mexico</collectingCountry>
|
||
. The highest species richness occurs in
|
||
<collectingCountry id="F3607653853BFF885C56FE9650D7FE5C" box="[1360,1432,367,392]" name="Brazil" pageId="9" pageNumber="460">Brazil</collectingCountry>
|
||
. This work recognizes two major species groups within the subgenus, the
|
||
<taxonomicName id="4C774D40853BFF885AC9FE6C511FFE78" box="[975,1104,405,428]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="trisignatum">
|
||
<emphasis id="B903EAD1853BFF885AC9FE6C511FFE78" box="[975,1104,405,428]" italics="true" pageId="9" pageNumber="460">trisignatum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D40853BFF885D81FE6D5196FE78" box="[1159,1241,404,428]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="valgum">
|
||
<emphasis id="B903EAD1853BFF885D81FE6D5196FE78" box="[1159,1241,404,428]" italics="true" pageId="9" pageNumber="460">valgum</emphasis>
|
||
</taxonomicName>
|
||
groups. The first comprises species historically grouped in
|
||
<taxonomicName id="4C774D40853BFF885B7EFE415643FE04" box="[632,780,440,464]" class="Insecta" family="Scarabaeidae" genus="Aganhyboma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EAD1853BFF885B7EFE415643FE04" box="[632,780,440,464]" italics="true" pageId="9" pageNumber="460">Aganhyboma</emphasis>
|
||
</taxonomicName>
|
||
, including the genus
|
||
<typeStatus id="54CC8861853BFF885D00FE415175FE04" box="[1030,1082,440,464]" pageId="9" pageNumber="460">type</typeStatus>
|
||
. The
|
||
<taxonomicName id="4C774D40853BFF885D7AFE4151B2FE1B" box="[1148,1277,440,463]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="trisignatum">
|
||
<emphasis id="B903EAD1853BFF885D7AFE4151B2FE1B" box="[1148,1277,440,463]" italics="true" pageId="9" pageNumber="460">trisignatum</emphasis>
|
||
</taxonomicName>
|
||
group can be further split into two species complexes, the
|
||
<taxonomicName id="4C774D40853BFF885BAFFE245665FE20" box="[681,810,477,500]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="trisignatum">
|
||
<emphasis id="B903EAD1853BFF885BAFFE245665FE20" box="[681,810,477,500]" italics="true" pageId="9" pageNumber="460">trisignatum</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="4C774D40853BFF885A90FE25515CFE20" box="[918,1043,476,500]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="cupreicolle">
|
||
<emphasis id="B903EAD1853BFF885A90FE25515CFE20" box="[918,1043,476,500]" italics="true" pageId="9" pageNumber="460">cupreicolle</emphasis>
|
||
</taxonomicName>
|
||
complexes. The
|
||
<taxonomicName id="4C774D40853BFF885DD9FE25507EFE20" box="[1247,1329,476,500]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="valgum">
|
||
<emphasis id="B903EAD1853BFF885DD9FE25507EFE20" box="[1247,1329,476,500]" italics="true" pageId="9" pageNumber="460">valgum</emphasis>
|
||
</taxonomicName>
|
||
group is comprised of three species complexes, the
|
||
<emphasis id="B903EAD1853BFF885B70FDF8563CFDCC" box="[630,883,512,536]" italics="true" pageId="9" pageNumber="460">
|
||
<taxonomicName id="4C774D40853BFF885B70FDF857AFFDCC" box="[630,736,513,536]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="acropyge">acropyge</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D40853BFF885BEAFDF9563CFDCC" box="[748,883,512,536]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="finestriatum">finestriatum</taxonomicName>
|
||
</emphasis>
|
||
and the
|
||
<taxonomicName id="4C774D40853BFF885AD5FDF95169FDCC" box="[979,1062,512,536]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="valgum">
|
||
<emphasis id="B903EAD1853BFF885AD5FDF95169FDCC" box="[979,1062,512,536]" italics="true" pageId="9" pageNumber="460">valgum</emphasis>
|
||
</taxonomicName>
|
||
complexes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |