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<document id="8ABE41FCA3CE4087E63411CE44811D3D" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58528FF9A5991FCE6503DFE5C" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) arturoi Silva, Louzada &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="478" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="477" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="AB855CDF386AF7C302BE1DBCB255A071">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="5B9E751C72A2CC68475E83E783DB6D28">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="82E901180519C7A1C8C3674A8EDED44C">Louzada, Júlio</mods:namePart>
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<mods:namePart id="354154CA729120A1B8F1BA5F41FB6649">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:title id="91E33AE74454B5339C464126D63D53F9">Zootaxa</mods:title>
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<mods:date id="4B2058E42112DBA5C18804200320708F">2015</mods:date>
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<treatment id="03DE87D58528FF9A5991FCE6503DFE5C" ID-DOI="http://doi.org/10.5281/zenodo.6115728" ID-GBIF-Taxon="119598945" ID-Zenodo-Dep="6115728" LSID="urn:lsid:plazi:treatment:03DE87D58528FF9A5991FCE6503DFE5C" httpUri="http://treatment.plazi.org/id/03DE87D58528FF9A5991FCE6503DFE5C" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
<subSubSection id="C36D65488528FF9B5991FCE65411FC8F" pageId="26" pageNumber="477" type="nomenclature">
<paragraph id="8BC836C38528FF9B5991FCE65793FCEE" blockId="26.[151,732,799,859]" box="[151,732,799,826]" pageId="26" pageNumber="477">
<heading id="D08081AF8528FF9B5991FCE65793FCEE" bold="true" box="[151,732,799,826]" fontSize="11" level="1" pageId="26" pageNumber="477" reason="1">
<emphasis id="B903EAD18528FF9B5991FCE65793FCEE" bold="true" box="[151,732,799,826]" pageId="26" pageNumber="477">
<taxonomicName id="4C774D408528FF9B5991FCE6570BFCED" ID-CoL="6CJ64" box="[151,580,799,826]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="arturoi" status="sp. nov." subGenus="Aganhyboma">
<emphasis id="B903EAD18528FF9B5991FCE65462FCED" bold="true" box="[151,301,799,825]" italics="true" pageId="26" pageNumber="477">Deltochilum</emphasis>
(
<emphasis id="B903EAD18528FF9B583BFCE65491FCED" bold="true" box="[317,478,799,825]" italics="true" pageId="26" pageNumber="477">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18528FF9B58F6FCE6570BFCED" bold="true" box="[496,580,799,825]" italics="true" pageId="26" pageNumber="477">arturoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057AA8528FF9B5B4AFCD95793FCEE" box="[588,732,800,826]" pageId="26" pageNumber="477" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38528FF9B5991FCBB5411FC8F" blockId="26.[151,732,799,859]" box="[151,350,834,859]" pageId="26" pageNumber="477">
(
<figureCitation id="134C2A468528FF9B5999FCBB55BAFC8F" box="[159,245,834,859]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="26" pageNumber="477">Figs 82</figureCitation>
,
<figureCitation id="134C2A468528FF9B5807FCBA5450FC8F" box="[257,287,835,859]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="26" pageNumber="477">91</figureCitation>
,
<figureCitation id="134C2A468528FF9B582DFCBA541AFC8F" box="[299,341,835,859]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="26" pageNumber="477">112</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488528FF9B5991FC735454FC13" pageId="26" pageNumber="477" type="etymology">
<paragraph id="8BC836C38528FF9B5991FC735454FC13" blockId="26.[151,1437,906,2011]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B5991FC735451FC77" bold="true" box="[151,286,906,931]" pageId="26" pageNumber="477">Etymology.</emphasis>
Named in honour of Fabio Arturo González Alvarado, friend of both authors and good dung beetle taxonomist.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488528FF9B59C1FC2B579CF8D7" pageId="26" pageNumber="477" type="description">
<paragraph id="8BC836C38528FF9B59C1FC2B579CF8D7" blockId="26.[151,1437,906,2011]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1FC2B54EAFC3F" bold="true" box="[199,421,978,1003]" pageId="26" pageNumber="477">Description. Body.</emphasis>
Head, pronotum, elytra and ventral surface black or dark brown, with an evident sheen. Length&gt;
<quantity id="4C8F9B268528FF9B5803FC0E5417FBDA" box="[261,344,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="26" pageNumber="477" unit="mm" value="14.0">14 mm</quantity>
.
<emphasis id="B903EAD18528FF9B5865FC0F54E5FBDB" bold="true" box="[355,426,1014,1039]" pageId="26" pageNumber="477">Head.</emphasis>
Punctures dense but not touching each other; foveiform punctures separated at the dorsal interocular surface by approximately half diameter. Punctures around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae, or with almost inconspicuous setae. Dorsal interocular distance approximately three times eye width (as in Fig. 61).
<emphasis id="B903EAD18528FF9B5991FB7F55B9FB4B" bold="true" box="[151,246,1158,1183]" pageId="26" pageNumber="477">Thorax.</emphasis>
Surface of pronotum with dense foveiform punctures. Punctures on the sides, anterior and posterior angles nearly touching. Discal punctures smaller and more dispersed than those of lateral portion, separated approximately by one diameter. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by at least one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles straight, with approximately the same length of margin between anterior and lateral angles. Hypomera with dense foveiform punctures on anterior portion and dispersed on posteromedial portion (
<figureCitation id="134C2A468528FF9B5B3DFAA757DDFAA3" box="[571,658,1374,1399]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="26" pageNumber="477">Fig. 82</figureCitation>
); anteromedial punctures separated by approximately half diameter. Diagonal band of hypomera from posteromedial portion to middle of lateral margin with few isolated punctures on otherwise smooth surface (
<figureCitation id="134C2A468528FF9B58C2FA5F5757FA6B" box="[452,536,1446,1471]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="26" pageNumber="477">Fig. 82</figureCitation>
).
<emphasis id="B903EAD18528FF9B5B2DFA5F5731FA6B" bold="true" box="[555,638,1446,1471]" pageId="26" pageNumber="477">Elytra.</emphasis>
Lateral margins slightly curved outward (
<figureCitation id="134C2A468528FF9B5D52FA5F51FDFA6B" box="[1108,1202,1446,1471]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="26" pageNumber="477">Fig. 112</figureCitation>
) and surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated; some punctures slightly elliptical, separated by regular spacing. Interstriae flattened, with foveiform punctures surrounded by approximately four well-defined shiny microtubercles. Foveiform punctures separated on elytral disc by at least one diameter. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length (as in Fig. 80). Apex of second interstria with carina or tubercle inconspicuous. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria approximately equal to the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina incomplete, effaced medially (as in Fig. 80).
</paragraph>
</subSubSection>
<subSubSection id="C36D65488528FF9B59C1F8F7510FF80F" pageId="26" pageNumber="477" type="materials_examined">
<paragraph id="8BC836C38528FF9B59C1F8F757C0F89F" blockId="26.[151,1437,906,2011]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1F8F757F6F8F3" bold="true" box="[199,697,1806,1831]" pageId="26" pageNumber="477">
Material studied.
<typeStatus id="54CC88618528FF9B58A4F8F7575CF8F3" box="[418,531,1806,1831]" pageId="26" pageNumber="477" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F36076538528FF9B5B25F8F757FAF8F3" box="[547,693,1806,1831]" name="Ecuador" pageId="26" pageNumber="477">ECUADOR</collectingCountry>
:
</emphasis>
PICHINCHA: B.P. Milpe,
<quantity id="4C8F9B268528FF9B5AFCF8F65105F8F3" box="[1018,1098,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="477" unit="m" value="1200.0">1200m</quantity>
,
<date id="FFC910038528FF9B5D5FF8F651FAF8F3" box="[1113,1205,1807,1831]" pageId="26" pageNumber="477" value="2003-04">IV.2003</date>
, Fumigación Dosel, P.Araujo &amp; William Chamorro (1 ♀
<collectionCode id="ED66AE068528FF9B5B36F8CA57CCF89F" box="[560,643,1843,1867]" pageId="26" pageNumber="477">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38528FF9B59C1F8AF5410F847" blockId="26.[151,1437,906,2011]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1F8AF5415F8BB" bold="true" box="[199,346,1878,1903]" pageId="26" pageNumber="477">
<typeStatus id="54CC88618528FF9B59C1F8AF547BF8BB" box="[199,308,1878,1903]" pageId="26" pageNumber="477" type="paratype">Paratype</typeStatus>
[1]
</emphasis>
:
<emphasis id="B903EAD18528FF9B586FF8AF54B9F8BB" bold="true" box="[361,502,1878,1903]" pageId="26" pageNumber="477">
<collectingCountry id="F36076538528FF9B586FF8AF54B9F8BB" box="[361,502,1878,1903]" name="Ecuador" pageId="26" pageNumber="477">ECUADOR</collectingCountry>
</emphasis>
:
<collectionCode id="ED66AE068528FF9B5B02F8AE572EF8BB" box="[516,609,1879,1903]" pageId="26" pageNumber="477">SANTO</collectionCode>
<collectingCountry id="F36076538528FF9B5B61F8AE57BCF8BB" box="[615,755,1879,1903]" name="Dominican Republic" pageId="26" pageNumber="477">DOMINGO</collectingCountry>
: Santo
<collectingCountry id="F36076538528FF9B5A45F8AF56E2F8BB" box="[835,941,1878,1903]" name="Dominican Republic" pageId="26" pageNumber="477">Domingo</collectingCountry>
(16 Km E), Tinalandia,
<date id="FFC910038528FF9B5DBDF8AF5009F8BB" box="[1211,1350,1878,1903]" pageId="26" pageNumber="477" value="1985-07-25">25.VII.1985</date>
, S. &amp; J. Peck (1 ♀
<collectionCode id="ED66AE068528FF9B5808F882541CF847" LSID="urn:lsid:biocol.org:col:34935" box="[270,339,1915,1939]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="26" pageNumber="477">CMN</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38528FF9B59C1F8675601F863" blockId="26.[151,1437,906,2011]" box="[199,846,1950,1975]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1F86754E3F863" bold="true" box="[199,428,1950,1975]" pageId="26" pageNumber="477">Sampling methods.</emphasis>
flight interception trap [
<specimenCount id="9D71FD4A8528FF9B5BB9F866560BF863" box="[703,836,1950,1975]" pageId="26" pageNumber="477" type="generic">1 specimen</specimenCount>
].
</paragraph>
<paragraph id="8BC836C38528FF9B59C1F83B510FF80F" blockId="26.[151,1437,906,2011]" box="[199,1088,1986,2011]" pageId="26" pageNumber="477">
<emphasis id="B903EAD18528FF9B59C1F83B5465F80F" bold="true" box="[199,298,1986,2011]" pageId="26" pageNumber="477">Habitat.</emphasis>
<collectingCountry id="F36076538528FF9B5837F83B54C1F80F" box="[305,398,1986,2011]" name="Ecuador" pageId="26" pageNumber="477">Ecuador</collectingCountry>
(
<figureCitation id="134C2A468528FF9B589BF83B54BFF80F" box="[413,496,1986,2011]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="26" pageNumber="477">Fig. 91</figureCitation>
), one specimen with label data “tropical rain forest”.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488529FF9A59C1FF6E503DFE5C" pageId="27" pageNumber="478" type="diagnosis">
<paragraph id="8BC836C38529FF9A59C1FF6E503DFE5C" blockId="27.[151,1436,151,392]" pageId="27" pageNumber="478">
<emphasis id="B903EAD18529FF9A59C1FF6E54ABFF64" bold="true" box="[199,484,151,176]" pageId="27" pageNumber="478">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408529FF9A5B7CFF6057AEFF64" box="[634,737,153,176]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD18529FF9A5B7CFF6057AEFF64" box="[634,737,153,176]" italics="true" pageId="27" pageNumber="478">acropyge</emphasis>
</taxonomicName>
complex): length&gt;
<quantity id="4C8F9B268529FF9A5ADEFF615160FF7B" box="[984,1071,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="478" unit="mm" value="14.0">14 mm</quantity>
; foveiform punctures of dorsal interocular surface separated by approximately half diameter; foveiform punctures of pronotal disc separated approximately by one diameter; posteromedial punctures of pronotum usually separated by at least one diameter; pronotal margin, between lateral and posterior angles, with approximately the same length of margin between anterior and lateral angles; some punctures of elytral striae slightly elliptical, separated by regular spacing; apical carina or tubercle of second elytral interstria inconspicuous; pseudepipleural carina incomplete, effaced medially (as in Fig. 80). The pattern of hypomera punctures on individuals of this species is also characteristic (
<figureCitation id="134C2A468529FF9A5C17FE96502AFE5C" box="[1297,1381,367,392]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="27" pageNumber="478">Fig. 82</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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