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<document id="722B6533FA79B9E2116E32CE4B6FF535" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58510FFA25991FB0D569FFA49" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) icariforme Paulian 1938" docType="treatment" docVersion="10" lastPageNumber="486" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="485" updateTime="1698565278120" updateUser="plazi">
<mods:mods id="B30C3C6B6CA57F6998743A48F8C6C283" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7C4B1AC0DA86632718B74DEDB0A9D9ED">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="B86B385B598B5787F333F7E804F70542">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="63807AA08DEF4E523FCED36C6E85B8DD">Louzada, Júlio</mods:namePart>
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<mods:namePart id="CFF2A7C06298E826635381031887BF40">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:typeOfResource id="45987CD6E33E181A2EE12D5F0FBA4FE1">text</mods:typeOfResource>
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<mods:title id="C0AF7E93A248B9F8C937CC132CA58E02">Zootaxa</mods:title>
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<mods:date id="2EA96526D18E7897FCC44BD10D51EC78">2015</mods:date>
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<mods:number id="C213CE3993995288A1255A9EF49FF067">3925</mods:number>
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<mods:classification id="72AFB90154740375A86EDD5B2134E962">journal article</mods:classification>
<mods:identifier id="23F46F435C3643657CEF71180F01EDF4" type="DOI">10.11646/zootaxa.3925.4.1</mods:identifier>
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<treatment id="03DE87D58510FFA25991FB0D569FFA49" ID-DOI="http://doi.org/10.5281/zenodo.6115750" ID-GBIF-Taxon="119598963" ID-Zenodo-Dep="6115750" LSID="urn:lsid:plazi:treatment:03DE87D58510FFA25991FB0D569FFA49" httpUri="http://treatment.plazi.org/id/03DE87D58510FFA25991FB0D569FFA49" lastPageId="35" lastPageNumber="486" pageId="34" pageNumber="485">
<subSubSection id="C36D65488510FFA35991FB0D5493FAFB" pageId="34" pageNumber="485" type="nomenclature">
<paragraph id="8BC836C38510FFA35991FB0D5126FADA" blockId="34.[151,1129,1268,1327]" box="[151,1129,1268,1294]" pageId="34" pageNumber="485">
<heading id="D08081AF8510FFA35991FB0D5126FADA" bold="true" box="[151,1129,1268,1294]" fontSize="11" level="1" pageId="34" pageNumber="485" reason="1">
<emphasis id="B903EAD18510FFA35991FB0D5126FADA" bold="true" box="[151,1129,1268,1294]" pageId="34" pageNumber="485">
<taxonomicName id="4C774D408510FFA35991FB0D5657FADA" ID-CoL="6CJ67" authority="Paulian 1938" authorityName="Paulian" authorityYear="1938" box="[151,792,1268,1294]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="icariforme" subGenus="Aganhyboma">
<emphasis id="B903EAD18510FFA35991FB0D5462FADA" bold="true" box="[151,301,1268,1294]" italics="true" pageId="34" pageNumber="485">Deltochilum</emphasis>
(
<emphasis id="B903EAD18510FFA3583BFB0D5491FADA" bold="true" box="[317,478,1268,1294]" italics="true" pageId="34" pageNumber="485">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18510FFA358F6FB0D5720FADA" bold="true" box="[496,623,1268,1294]" italics="true" pageId="34" pageNumber="485">icariforme</emphasis>
<bibRefCitation id="EFE64B328510FFA35B70FB0D5657FADA" author="Paulian" box="[630,792,1268,1294]" pageId="34" pageNumber="485" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
</taxonomicName>
new subgeneric placement
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38510FFA35991FAEF5493FAFB" blockId="34.[151,1129,1268,1327]" box="[151,476,1302,1327]" pageId="34" pageNumber="485">
(
<figureCitation id="134C2A468510FFA35999FAEF55BAFAFB" box="[159,245,1302,1327]" captionStart="FIGURES 20 – 23" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[169,1418,225,1833]" captionTargetId="figure@40.[151,1436,193,1836]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Aedeagus in Deltochilum (Aganhyboma). 20. D. paresi new species. 21. D. icariforme. 22. D. ritamourae new species. 23. D. subrubrum new species. Figure 24. Aedeagus illustration showing the angle between the phallobase and the parameres. Figures 25 – 27. Genital segment in Deltochilum (Aganhyboma). 25. D. cupreicolle. 26. D. viridescens new status. 27. D. trisignatum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244799/files/figure.png" pageId="34" pageNumber="485">Figs 21</figureCitation>
,
<figureCitation id="134C2A468510FFA35807FAEE546FFAFB" box="[257,288,1303,1327]" captionStart="FIGURES 28 – 32" captionStartId="41.[151,264,1863,1886]" captionTargetBox="[163,1427,199,1832]" captionTargetId="figure@41.[151,1435,193,1843]" captionTargetPageId="41" captionText="FIGURES 28 – 32. Genital segment in Deltochilum (Aganhyboma). 28. D. finestriatum new species. 29. D. schefflerorum new species 30. D. valgum. 31. D. icaroides. 32. D. icariforme. Figures 33 – 36. Lamella copulatrix sclerites in D. (Aganhyboma). 33. D. cupreicolle. 34. D. viridescens new status and D. viridicatum new species 35. D. trisignatum. 36. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244800/files/figure.png" pageId="34" pageNumber="485">32</figureCitation>
,
<figureCitation id="134C2A468510FFA3582DFAEE5405FAFB" box="[299,330,1303,1327]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="34" pageNumber="485">40</figureCitation>
,
<figureCitation id="134C2A468510FFA35853FAEF543CFAFB" box="[341,371,1302,1327]" captionStart="FIGURES 55 – 58" captionStartId="43.[151,264,1783,1806]" captionTargetBox="[205,1383,207,1725]" captionTargetId="figure@43.[174,1383,193,1728]" captionTargetPageId="43" captionText="FIGURES 55 – 58. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 55. D. valgum. 56. D. kolleri new species. 57. D. icariforme. 58. D. ritamourae new species. Figure 59. Complex of axial and subaxial sclerites (A + AS) in D. viridescens new status. Figures 60 – 61. Head in D. (Aganhyboma). 60. D. viridescens new status. 61. D. longiceps new status. Figures 62 – 63. Lateral view of the elytra in D. (Aganhyboma). 62. D. viridescens new status (arrow points the basal carina of ninth interstria). 63. D. finestriatum new species. (arrow points the basal carina of ninth interstria). Figures 64 – 69. Legs. 64. Metafemur in D. viridescens new status. 65. Metafemur in D. longiceps new status. 66. Protibia in D. trisignatum. 67. Metatibia in D. trisignatum. 68. Metatibia in D. viridescens new status. 69. Profemur in D. trisignatum (arrow indicates the anterior margin discontinuous). Figures scale = 1 mm." httpUri="https://zenodo.org/record/244802/files/figure.png" pageId="34" pageNumber="485">57</figureCitation>
,
<figureCitation id="134C2A468510FFA35879FAEE54D2FAFB" box="[383,413,1303,1327]" captionStart="FIGURES 93 – 94" captionStartId="47.[151,264,1007,1030]" captionTargetBox="[151,1436,193,984]" captionTargetId="figure@47.[151,1436,193,985]" captionTargetPageId="47" captionText="FIGURES 93 – 94. Known distribution of D. (Aganhyboma). 93. D. icaroides and D. valgum. 94. icariforme subcomplex." httpUri="https://zenodo.org/record/244806/files/figure.png" pageId="34" pageNumber="485">94</figureCitation>
,
<figureCitation id="134C2A468510FFA358AFFAEE549BFAFB" box="[425,468,1303,1327]" captionStart="FIGURES 119 – 123" captionStartId="51.[151,264,1596,1619]" captionTargetBox="[178,1436,193,1570]" captionTargetId="figure@51.[150,1436,193,1576]" captionTargetPageId="51" captionText="FIGURES 119 – 123. Species of D. (Aganhyboma). 119. D. ritamourae new species. 120. D. icariforme. 121. D. paresi new species. 122. D. kolleri new species. 123. D. subrubrum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244810/files/figure.png" pageId="34" pageNumber="485">120</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488510FFA35991FAA756E2F9DA" pageId="34" pageNumber="485" type="reference_group">
<paragraph id="8BC836C38510FFA35991FAA75755FA47" blockId="34.[151,1436,1374,1550]" pageId="34" pageNumber="485">
<treatmentCitationGroup id="AB6711ED8510FFA35991FAA75755FA47" pageId="34" pageNumber="485">
<taxonomicName id="4C774D408510FFA35991FAA757B3FAA0" ID-CoL="6CJ67" authority="Paulian 1938: 269" authorityName="Paulian" authorityPageNumber="269" authorityYear="1938" box="[151,764,1374,1396]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="icariforme" subGenus="Deltochilum">
<emphasis id="B903EAD18510FFA35991FAA7545BFAA0" box="[151,276,1374,1396]" italics="true" pageId="34" pageNumber="485">Deltochilum</emphasis>
(
<emphasis id="B903EAD18510FFA3582CFAA754E8FAA0" box="[298,423,1374,1396]" italics="true" pageId="34" pageNumber="485">Deltochilum</emphasis>
)
<emphasis id="B903EAD18510FFA358BAFAA75768FAA0" box="[444,551,1374,1396]" italics="true" pageId="34" pageNumber="485">icariforme</emphasis>
<treatmentCitation id="0AD610D28510FFA35B32FAA757B3FAA0" author="Paulian" box="[564,764,1374,1396]" page="269" pageId="34" pageNumber="485" year="1938">
<bibRefCitation id="EFE64B328510FFA35B32FAA7578CFAA0" author="Paulian" box="[564,707,1374,1396]" pageId="34" pageNumber="485" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
: 269
</treatmentCitation>
</taxonomicName>
, 273, 276,
<figureCitation id="134C2A468510FFA35A87FAA7569EFAA0" box="[897,977,1374,1396]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="34" pageNumber="485">Fig. 18</figureCitation>
, map n.5 (original description, taxonomic comments and identification key)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C38510FFA35991FA6557BFFA66" blockId="34.[151,1436,1374,1550]" box="[151,752,1436,1458]" pageId="34" pageNumber="485">
<treatmentCitationGroup id="AB6711ED8510FFA35991FA6557BFFA66" box="[151,752,1436,1458]" pageId="34" pageNumber="485">
<taxonomicName id="4C774D408510FFA35991FA655730FA66" ID-CoL="6CJ67" authority="Blackwelder 1944: 203" authorityName="Blackwelder" authorityPageNumber="203" authorityYear="1944" box="[151,639,1436,1458]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="icariforme">
<emphasis id="B903EAD18510FFA35991FA6554CAFA66" box="[151,389,1436,1458]" italics="true" pageId="34" pageNumber="485">Deltochilum icariforme</emphasis>
:
<treatmentCitation id="0AD610D28510FFA35895FA655730FA66" author="Blackwelder" box="[403,639,1436,1458]" page="203" pageId="34" pageNumber="485" year="1944">
<bibRefCitation id="EFE64B328510FFA35895FA655700FA66" author="Blackwelder" box="[403,591,1436,1458]" pageId="34" pageNumber="485" refString="Blackwelder, R. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 265." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 203
</treatmentCitation>
</taxonomicName>
(checklist)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C38510FFA35991FA43518BFA04" blockId="34.[151,1436,1374,1550]" box="[151,1220,1466,1488]" pageId="34" pageNumber="485">
<treatmentCitationGroup id="AB6711ED8510FFA35991FA43518BFA04" box="[151,1220,1466,1488]" pageId="34" pageNumber="485">
<taxonomicName id="4C774D408510FFA35991FA43564DFA04" authority="Martinez 1947: 274" authorityName="Martinez" authorityPageNumber="274" authorityYear="1947" box="[151,770,1466,1488]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="icariniforme" subGenus="Deltochilum">
<emphasis id="B903EAD18510FFA35991FA43545BFA04" box="[151,276,1466,1488]" italics="true" pageId="34" pageNumber="485">Deltochilum</emphasis>
(
<emphasis id="B903EAD18510FFA35825FA4354EFFA04" box="[291,416,1466,1488]" italics="true" pageId="34" pageNumber="485">Deltochilum</emphasis>
)
<emphasis id="B903EAD18510FFA358A8FA435763FA04" box="[430,556,1466,1488]" italics="true" pageId="34" pageNumber="485">icariniforme</emphasis>
:
<treatmentCitation id="0AD610D28510FFA35B3DFA43564DFA04" author="Martinez" box="[571,770,1466,1488]" page="274" pageId="34" pageNumber="485" year="1947">
<bibRefCitation id="EFE64B328510FFA35B3DFA43579DFA04" author="Martinez" box="[571,722,1466,1488]" pageId="34" pageNumber="485" refString="Martinez, A. (1947) Insectos nuevos o poco conocidos. V. El genero Pseudocanthon Bates, y algunas especies nuevas o poco conocidas de Scarabaeidae. Revista de la Sociedad Entomologica Argentina, 13 (1 - 5), pp. 263 - 280." type="journal article" year="1947">Martínez 1947</bibRefCitation>
: 274
</treatmentCitation>
</taxonomicName>
(comments about geographical distribution)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C38510FFA35991FA20562AFA3B" blockId="34.[151,1436,1374,1550]" box="[151,869,1497,1519]" pageId="34" pageNumber="485">
<treatmentCitationGroup id="AB6711ED8510FFA35991FA20562AFA3B" box="[151,869,1497,1519]" pageId="34" pageNumber="485">
<taxonomicName id="4C774D408510FFA35991FA2057A2FA3B" ID-CoL="6CJ67" authority="Martinez 1959: 53" authorityName="Martinez" authorityPageNumber="53" authorityYear="1959" box="[151,749,1497,1519]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="icariforme" subGenus="Deltohyboma">
<emphasis id="B903EAD18510FFA35991FA20576AFA3B" box="[151,549,1497,1519]" italics="true" pageId="34" pageNumber="485">Deltochilum (Deltohyboma) icariforme</emphasis>
:
<treatmentCitation id="0AD610D28510FFA35B34FA2057A2FA3B" author="Martinez" box="[562,749,1497,1519]" page="53" pageId="34" pageNumber="485" year="1959">
<bibRefCitation id="EFE64B328510FFA35B34FA205786FA3B" author="Martinez" box="[562,713,1497,1519]" pageId="34" pageNumber="485" refString="Martinez, A. (1959) Catalogo de los Scarabaeidae argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot;, 5, 1 - 126." type="journal article" year="1959">Martínez 1959</bibRefCitation>
: 53
</treatmentCitation>
</taxonomicName>
(catalogue)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C38510FFA35991FA0156E2F9DA" blockId="34.[151,1436,1374,1550]" box="[151,941,1528,1550]" pageId="34" pageNumber="485">
<bibRefCitation id="EFE64B328510FFA35991FA0154C0F9DA" author="Vulcano" box="[151,399,1528,1550]" pageId="34" pageNumber="485" refString="Vulcano, M. A. &amp; Pereira, F. S. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the western hemisphere. Entomologischen Arbeiten aus dem Museum G. Frey, 15, 569 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964</bibRefCitation>
: 655 (catalogue); Vaz-de-Mello 2000: 192 (checklist)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488510FFA25991F9C35631FE94" lastPageId="35" lastPageNumber="486" pageId="34" pageNumber="485" type="description">
<paragraph id="8BC836C38510FFA35991F9C350D2F80B" blockId="34.[151,1437,1594,2015]" pageId="34" pageNumber="485">
<emphasis id="B903EAD18510FFA35991F9C35438F987" bold="true" box="[151,375,1594,1619]" pageId="34" pageNumber="485">Description. Body.</emphasis>
Head, pronotum, elytra and ventral surface with copper coloration and a reddish metallic sheen. Length 14.5 to 15.8 mm.
<emphasis id="B903EAD18510FFA35B24F9A75726F9A3" bold="true" box="[546,617,1630,1655]" pageId="34" pageNumber="485">Head.</emphasis>
Foveiform punctures dense and deep, touching on dorsal interocular surface. Dorsal interocular distance approximately three times eye width. Lateral margin of gena strongly curved outward, with row of setae around eyes.
<emphasis id="B903EAD18510FFA35B60F95F5789F96B" bold="true" box="[614,710,1702,1727]" pageId="34" pageNumber="485">Thorax.</emphasis>
Surface of pronotum with deep and dense foveiform punctures, nearly touching on the sides and anterior angles. Anteromedial punctures more dispersed than those of surrounding surface. Margin between lateral and posterior angles straight. Posteromedial punctures separated by less than one diameter.
<emphasis id="B903EAD18510FFA3580EF8EB5413F8FF" bold="true" box="[264,348,1810,1835]" pageId="34" pageNumber="485">Elytra.</emphasis>
Lateral margins slightly curved outward, almost parallel (
<figureCitation id="134C2A468510FFA35D07F8EB5128F8FF" box="[1025,1127,1810,1835]" captionStart="FIGURES 119 – 123" captionStartId="51.[151,264,1596,1619]" captionTargetBox="[178,1436,193,1570]" captionTargetId="figure@51.[150,1436,193,1576]" captionTargetPageId="51" captionText="FIGURES 119 – 123. Species of D. (Aganhyboma). 119. D. ritamourae new species. 120. D. icariforme. 121. D. paresi new species. 122. D. kolleri new species. 123. D. subrubrum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244810/files/figure.png" pageId="34" pageNumber="485">Fig. 120</figureCitation>
). Surface slightly irregular (rugose). Elytral striae wider, carinate margins clearly separated (as in Fig. 75). Interstriae clearly convex, with its central portion nearly flat. Interstrial surface with foveiform punctures surrounded by microtubercles weakly defined. Interstriae surface usually more punctate on sides, close to striae. Foveiform punctures of third interstria denser laterally (punctures can be so scattered at center of interstria to produce smooth midlongitudinal strip) (as in Fig. 86). Basal carina of seventh interstria elongate, length about four or more times seventh interestria width.
</paragraph>
<paragraph id="8BC836C38511FFA25991FF6E5631FE94" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">Basal carina of ninth interstria present in three-quarters of elytra (not surpassing anterior limit of apical carina of seventh interstria). Apical carina of third interstria weaker than apical carinae of second and fourth interstriae. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria longer than the sum of lengths of apical carina of fifth and sixth interstriae. Pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63).</paragraph>
</subSubSection>
<subSubSection id="C36D65488511FFA259C1FEB5563EFBE1" pageId="35" pageNumber="486" type="materials_examined">
<paragraph id="8BC836C38511FFA259C1FEB554F5FE20" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
<emphasis id="B903EAD18511FFA259C1FEB5575DFEB1" bold="true" box="[199,530,332,357]" pageId="35" pageNumber="486">
Material studied.
<typeStatus id="54CC88618511FFA2589AFEB5575DFEB1" box="[412,530,332,357]" pageId="35" pageNumber="486" type="lectotype">Lectotype</typeStatus>
</emphasis>
(
<collectionCode id="ED66AE068511FFA25B27FEB55735FEB0" LSID="urn:lsid:biocol.org:col:34988" box="[545,634,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="486">MNHN</collectionCode>
)
<emphasis id="B903EAD18511FFA25B8EFEB55610FEB1" bold="true" box="[648,863,332,357]" pageId="35" pageNumber="486">[here designated]:</emphasis>
1. [old white label, bordered and printed in black]. Jatahy/ Etat de Goyaz/
<collectingCountry id="F36076538511FFA25899FE96548CFE5C" box="[415,451,367,392]" name="Switzerland" pageId="35" pageNumber="486">Ch</collectingCountry>
. Pujol
<date id="FFC910038511FFA25B15FE965737FE5C" box="[531,632,367,392]" pageId="35" pageNumber="486" value="1895" valueMax="1896">1895–96</date>
; 2. [old white label, handwritten].
<taxonomicName id="4C774D408511FFA25D04FE965138FE5C" box="[1026,1143,367,392]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="icariforme" status="sp. nov.">icariforme</taxonomicName>
<taxonomicNameLabel id="A23057AA8511FFA25D79FE8951F3FE53" box="[1151,1212,368,391]" pageId="35" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="54CC88618511FFA25DC2FE8951B0FE5C" box="[1220,1279,368,392]" pageId="35" pageNumber="486">Type</typeStatus>
; 3. [old white label, bordered and printed in black]. R. PAULIAN/ Vidit; 4. [red label, bordered and printed in black].
<typeStatus id="54CC88618511FFA25991FE415474FE04" box="[151,315,440,464]" pageId="35" pageNumber="486" type="lectotype">LECTOTYPE</typeStatus>
; 5. [white label, handwritten, bordered and printed in black].
<taxonomicName id="4C774D408511FFA25AE2FE4E5064FE04" authority="Paul." authorityName="Paul." box="[996,1323,439,464]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="icariforme">Deltochilum icariforme Paul.</taxonomicName>
LECTOT FVaz-de-Mello des. 2013.
</paragraph>
<paragraph id="8BC836C38511FFA259C1FDF9517BFDB4" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
In the original description,
<bibRefCitation id="EFE64B328511FFA258F3FE0657D5FDCC" author="Paulian" box="[501,666,511,536]" pageId="35" pageNumber="486" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian (1938)</bibRefCitation>
stated that the “
<typeStatus id="54CC88618511FFA25A4AFDF956CFFDCC" box="[844,896,512,536]" pageId="35" pageNumber="486">type</typeStatus>
” was deposited in the Oberthür collection. After examining the list of the specimens used by the author to describe the species, we found that only one individual from Jataí (Goiás) and one from Lacerda (Bahia) were deposited in this collection.
</paragraph>
<paragraph id="8BC836C38511FFA259C1FD9451D6FD24" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
We examined three of the five specimens listed in the original description. These were labeled by Paulian with the designation of &quot;
<typeStatus id="54CC88618511FFA25877FD6954E9FD7C" box="[369,422,656,680]" pageId="35" pageNumber="486">type</typeStatus>
&quot; and deposited in the Muséum National d'Histoire Naturelle (
<collectionCode id="ED66AE068511FFA25D5DFD6951F8FD7C" LSID="urn:lsid:biocol.org:col:34988" box="[1115,1207,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="486">MNHN</collectionCode>
), Paris,
<collectingCountry id="F36076538511FFA25C16FD69502FFD73" box="[1296,1376,656,679]" name="France" pageId="35" pageNumber="486">France</collectingCountry>
. Two of them correspond to the specimens from Jataí and Lacerda mentioned above, and the third is from
<collectingCountry id="F36076538511FFA25C29FD4C50D6FD19" box="[1327,1433,693,717]" name="Paraguay" pageId="35" pageNumber="486">Paraguay</collectingCountry>
. After careful analysis, we concluded that these individuals represent three different species.
</paragraph>
<paragraph id="8BC836C38511FFA259C1FD0551F2FC88" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
Thus, with the purpose of stabilizing nomenclature by choosing the name-bearing specimen, we designate the specimen from Jataí (Goiás) as the
<typeStatus id="54CC88618511FFA25B20FCE657C2FCEC" box="[550,653,799,824]" pageId="35" pageNumber="486" type="lectotype">lectotype</typeStatus>
of
<emphasis id="B903EAD18511FFA25BB2FCD8561BFCEC" box="[692,852,800,824]" italics="true" pageId="35" pageNumber="486">
<taxonomicName id="4C774D408511FFA25BB2FCD8561FFCEC" box="[692,848,800,824]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="icariforme">D. icariforme</taxonomicName>
.
</emphasis>
The specimen from Bahia represents a new species described here as
<taxonomicName id="4C774D408511FFA25858FCBC54B1FC88" box="[350,510,837,860]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="ritamourae" status="sp. nov.">
<emphasis id="B903EAD18511FFA25858FCBC54B1FC88" box="[350,510,837,860]" italics="true" pageId="35" pageNumber="486">D. ritamourae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057AA8511FFA25B00FCBD57C0FC89" box="[518,655,836,861]" pageId="35" pageNumber="486" rank="species">
<emphasis id="B903EAD18511FFA25B00FCBD57C0FC89" bold="true" box="[518,655,836,861]" pageId="35" pageNumber="486">new species</emphasis>
</taxonomicNameLabel>
, and the specimen from
<collectingCountry id="F36076538511FFA25AA2FCBC5143FC89" box="[932,1036,837,861]" name="Paraguay" pageId="35" pageNumber="486">Paraguay</collectingCountry>
is
<taxonomicName id="4C774D408511FFA25D2BFCBC51F9FC88" box="[1069,1206,836,860]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="icaroides">
<emphasis id="B903EAD18511FFA25D2BFCBC51F9FC88" box="[1069,1206,836,860]" italics="true" pageId="35" pageNumber="486">D. icaroides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC836C38511FFA259C1FC9E51C0FC1C" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
<emphasis id="B903EAD18511FFA259C1FC9E5717FC54" bold="true" box="[199,600,871,896]" pageId="35" pageNumber="486">
Non-type material [5]:
<collectingCountry id="F36076538511FFA258EBFC9E5717FC54" box="[493,600,871,896]" name="Brazil" pageId="35" pageNumber="486">BRAZIL</collectingCountry>
</emphasis>
:
<collectionCode id="ED66AE068511FFA25B68FC9157F2FC54" box="[622,701,872,896]" pageId="35" pageNumber="486">MATO</collectionCode>
GROSSO: Rio Verde,
<date id="FFC910038511FFA25ADCFC9E5178FC54" box="[986,1079,871,896]" pageId="35" pageNumber="486" value="1963-11">XI.1963</date>
(
<specimenCount id="9D71FD4A8511FFA25D56FC915131FCAB" box="[1104,1150,872,896]" pageId="35" pageNumber="486" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068511FFA25D8AFC915195FC54" box="[1164,1242,872,896]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="35" pageNumber="486">DZUP</collectionCode>
);
<collectionCode id="ED66AE068511FFA25DF2FC9E5063FC54" box="[1268,1324,871,896]" httpUri="http://grbio.org/cool/2pta-s8y1" name="Sammlung Oberli" pageId="35" pageNumber="486">SÃO</collectionCode>
<collectionCode id="ED66AE068511FFA25C3FFC9150D6FC54" box="[1337,1433,872,896]" pageId="35" pageNumber="486">PAULO</collectionCode>
: Itirapina,
<date id="FFC910038511FFA2580FFC7554CFFC70" box="[265,384,908,932]" pageId="35" pageNumber="486" value="2000-02-27">27.II.2000</date>
, G. Machado (1 ♀
<collectionCode id="ED66AE068511FFA25B76FC75578CFC70" box="[624,707,908,932]" pageId="35" pageNumber="486">CEMT</collectionCode>
); Itirapina, Estação Ecológica, (
<geoCoordinate id="EE4350048511FFA25D39FC7551D6FC70" box="[1087,1177,908,932]" direction="south" orientation="latitude" pageId="35" pageNumber="486" precision="925" value="-22.216667">22°13'S</geoCoordinate>
,
<geoCoordinate id="EE4350048511FFA25DACFC75505EFC71" box="[1194,1297,908,933]" direction="west" orientation="longitude" pageId="35" pageNumber="486" precision="925" value="-47.9">47°54'W</geoCoordinate>
),
<date id="FFC910038511FFA25C2CFC7550D7FC70" box="[1322,1432,908,932]" pageId="35" pageNumber="486" value="2009-01-30">30.I.2009</date>
, Fernando Silva (
<specimenCount id="9D71FD4A8511FFA25857FC495434FC13" box="[337,379,944,968]" pageId="35" pageNumber="486" type="generic">2 ♂</specimenCount>
<collectionCode id="ED66AE068511FFA25884FC49549AFC1C" box="[386,469,944,968]" pageId="35" pageNumber="486">CEMT</collectionCode>
); Pirassununga, E.E.C.P.,
<date id="FFC910038511FFA25BF1FC565620FC1C" box="[759,879,943,968]" pageId="35" pageNumber="486" value="1945-10-29">29.X.1945</date>
, Schubart (
<specimenCount id="9D71FD4A8511FFA25AF7FC495156FC13" box="[1009,1049,944,968]" pageId="35" pageNumber="486" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068511FFA25D26FC4951CCFC1C" box="[1056,1155,944,968]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="35" pageNumber="486">MZUSP</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38511FFA259C1FC2D516CFC39" blockId="35.[151,1437,151,1437]" box="[199,1059,980,1005]" pageId="35" pageNumber="486">
<emphasis id="B903EAD18511FFA259C1FC2D54E3FC39" bold="true" box="[199,428,980,1005]" pageId="35" pageNumber="486">Sampling methods.</emphasis>
baited pitfall trap with injured diplopods [
<specimenCount id="9D71FD4A8511FFA25A80FC2D5157FC39" box="[902,1048,980,1005]" pageId="35" pageNumber="486" type="generic">2 specimens</specimenCount>
].
</paragraph>
<paragraph id="8BC836C38511FFA259C1FC0E563EFBE1" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
<emphasis id="B903EAD18511FFA259C1FC0E5465FBC4" bold="true" box="[199,298,1015,1040]" pageId="35" pageNumber="486">Habitat.</emphasis>
<collectingCountry id="F36076538511FFA25833FC0E5436FBC4" box="[309,377,1015,1040]" name="Brazil" pageId="35" pageNumber="486">Brazil</collectingCountry>
(Goiás, Mato Grosso and São Paulo) (
<figureCitation id="134C2A468511FFA25A4BFC0E56E9FBC4" box="[845,934,1015,1040]" captionStart="FIGURES 93 – 94" captionStartId="47.[151,264,1007,1030]" captionTargetBox="[151,1436,193,984]" captionTargetId="figure@47.[151,1436,193,985]" captionTargetPageId="47" captionText="FIGURES 93 – 94. Known distribution of D. (Aganhyboma). 93. D. icaroides and D. valgum. 94. icariforme subcomplex." httpUri="https://zenodo.org/record/244806/files/figure.png" pageId="35" pageNumber="486">Fig. 94</figureCitation>
). From available data this species inhabits Cerrado. Two specimens with label data &quot;Campo Sujo&quot;, Cerrado.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488511FFA259C1FBC6569FFA49" pageId="35" pageNumber="486" type="diagnosis">
<paragraph id="8BC836C38511FFA259C1FBC6570CFAE4" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">
<emphasis id="B903EAD18511FFA259C1FBC65493FB8C" bold="true" box="[199,476,1087,1112]" pageId="35" pageNumber="486">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408511FFA25B61FBB95792FB8C" box="[615,733,1088,1112]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="icariforme">
<emphasis id="B903EAD18511FFA25B61FBB95792FB8C" box="[615,733,1088,1112]" italics="true" pageId="35" pageNumber="486">icariforme</emphasis>
</taxonomicName>
subcomplex): length 14.5 to 15.8 mm; length of apical carina of seventh interstria longer than the sum of lengths of apical carina of fifth and sixth interstriae; basal carina of ninth interstria present in three-quarters of elytra (not surpassing anterior limit of apical carina of seventh interstria). The following aedeagus characters are also diagnostic: parameres symmetric and elongate, in lateral view forming an angle of approximately 45° with phallobase; apex acuminate (
<figureCitation id="134C2A468511FFA25D09FB36512DFB3C" box="[1039,1122,1231,1256]" captionStart="FIGURES 20 – 23" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[169,1418,225,1833]" captionTargetId="figure@40.[151,1436,193,1836]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Aedeagus in Deltochilum (Aganhyboma). 20. D. paresi new species. 21. D. icariforme. 22. D. ritamourae new species. 23. D. subrubrum new species. Figure 24. Aedeagus illustration showing the angle between the phallobase and the parameres. Figures 25 – 27. Genital segment in Deltochilum (Aganhyboma). 25. D. cupreicolle. 26. D. viridescens new status. 27. D. trisignatum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244799/files/figure.png" pageId="35" pageNumber="486">Fig. 21</figureCitation>
). Genital segment as in
<figureCitation id="134C2A468511FFA25C76FB3655F9FAD8" captionStart="FIGURES 28 – 32" captionStartId="41.[151,264,1863,1886]" captionTargetBox="[163,1427,199,1832]" captionTargetId="figure@41.[151,1435,193,1843]" captionTargetPageId="41" captionText="FIGURES 28 – 32. Genital segment in Deltochilum (Aganhyboma). 28. D. finestriatum new species. 29. D. schefflerorum new species 30. D. valgum. 31. D. icaroides. 32. D. icariforme. Figures 33 – 36. Lamella copulatrix sclerites in D. (Aganhyboma). 33. D. cupreicolle. 34. D. viridescens new status and D. viridicatum new species 35. D. trisignatum. 36. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244800/files/figure.png" pageId="35" pageNumber="486">Fig. 32</figureCitation>
. Lamella copulatrix (LC): left lobe trapezoid (
<figureCitation id="134C2A468511FFA25BE2FB0D5672FAD8" box="[740,829,1268,1293]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="35" pageNumber="486">Fig. 40</figureCitation>
); right lobe subrectangular (
<figureCitation id="134C2A468511FFA25D89FB0D51A6FAD8" box="[1167,1257,1268,1293]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="35" pageNumber="486">Fig. 40</figureCitation>
). Fronto-lateral peripheral sclerite (FLP) as in
<figureCitation id="134C2A468511FFA258EAFAEE5771FAE4" box="[492,574,1303,1328]" captionStart="FIGURES 55 – 58" captionStartId="43.[151,264,1783,1806]" captionTargetBox="[205,1383,207,1725]" captionTargetId="figure@43.[174,1383,193,1728]" captionTargetPageId="43" captionText="FIGURES 55 – 58. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 55. D. valgum. 56. D. kolleri new species. 57. D. icariforme. 58. D. ritamourae new species. Figure 59. Complex of axial and subaxial sclerites (A + AS) in D. viridescens new status. Figures 60 – 61. Head in D. (Aganhyboma). 60. D. viridescens new status. 61. D. longiceps new status. Figures 62 – 63. Lateral view of the elytra in D. (Aganhyboma). 62. D. viridescens new status (arrow points the basal carina of ninth interstria). 63. D. finestriatum new species. (arrow points the basal carina of ninth interstria). Figures 64 – 69. Legs. 64. Metafemur in D. viridescens new status. 65. Metafemur in D. longiceps new status. 66. Protibia in D. trisignatum. 67. Metatibia in D. trisignatum. 68. Metatibia in D. viridescens new status. 69. Profemur in D. trisignatum (arrow indicates the anterior margin discontinuous). Figures scale = 1 mm." httpUri="https://zenodo.org/record/244802/files/figure.png" pageId="35" pageNumber="486">Fig. 57</figureCitation>
.
</paragraph>
<paragraph id="8BC836C38511FFA259C1FAC5569FFA49" blockId="35.[151,1437,151,1437]" pageId="35" pageNumber="486">The body coloration of some specimens can vary between dark brown, copper or red, with reddish or greenish sheen. In some specimens, apical carina of third interstria can be somewhat more prominent than apical carina of second and fourth interstriae, with a rounded form, resembling a tubercle.</paragraph>
</subSubSection>
</treatment>
</document>
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