393 lines
54 KiB
XML
393 lines
54 KiB
XML
<document id="FE4B8C9086699014DF58CA9F5667B292" ID-DOI="10.5852/ejt.2024.919.2407" ID-ISSN="2118-9773" ID-Zenodo-Dep="10559520" ID-ZooBank="72B7907F-F901-4DDD-B2DD-63D03253837E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706085863884" checkinUser="plazi" docAuthor="Constant, Jérôme & Pham, Hong Thai" docDate="2024" docId="03DE87B0FFC4FF83FDEFFCD5A37AFBA2" docLanguage="en" docName="EJT.2024.919.1-87.pdf" docOrigin="European Journal of Taxonomy 919 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2407/10617" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pusulissus quangninhensis Constant & Pham 2024, sp. nov." docType="treatment" docUuid="A5D5064C-EAB2-4C1A-809A-DA2074260167" docUuidSource="ZooBank" docVersion="2" lastPageNumber="56" masterDocId="FFE7FFC8FFF7FFB4FFABFFD4A706FFDA" masterDocTitle="Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)" masterLastPageNumber="87" masterPageNumber="1" pageNumber="52" updateTime="1706086549316" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="2E87D903C4DDDA43AD359672BE5C2CFF">
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<mods:title id="8C12F471CAAF2746F1BF21522D153491">Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)</mods:title>
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<mods:name id="C597618C30F3B7868347D115BA03174B" type="personal">
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<mods:role id="6D63CF7E6FA36E253872D2CEB4F4F715">
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<mods:roleTerm id="340DAD1667905632A51E024514518AC6">Author</mods:roleTerm>
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<mods:namePart id="C3016AF85DD07FBCAE14AC4D8259180F">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="D35F6BAEA993A852F59712AE6DD71FCF">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="BB02B819B84CFC10D171E67524E28EC8" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:name id="9494C33DE0FA0C4F4B6478F591397756" type="personal">
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<mods:namePart id="AE09CA2747B1486020B49D36A45D404E">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="B183207B951D255B2E6E6CFF376ABF13">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Street, Hanoi, Vietnam. & Graduate School of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam.</mods:affiliation>
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<mods:nameIdentifier id="E6BC99379993127974A0297650D13E8C" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="B375916433299D0F363B542172A3E323">European Journal of Taxonomy</mods:title>
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<mods:part id="B87D8E1A0F907E7A8B10B4E9BB20ABC5">
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<mods:date id="CBFE58B548FFF1B9178B3E9A2A26B1C4">2024</mods:date>
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<mods:number id="812E7ACAF5289758725E736052534FB1">2024-01-24</mods:number>
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<treatment id="03DE87B0FFC4FF83FDEFFCD5A37AFBA2" LSID="urn:lsid:zoobank.org:act:A5D5064C-EAB2-4C1A-809A-DA2074260167" httpUri="http://treatment.plazi.org/id/03DE87B0FFC4FF83FDEFFCD5A37AFBA2" lastPageId="55" lastPageNumber="56" pageId="51" pageNumber="52">
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<subSubSection id="C36D652DFFC4FF87FDEFFCD5A3CDFC9A" pageId="51" pageNumber="52" type="nomenclature">
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<paragraph id="8BC836A6FFC4FF87FDEFFCD5A4F6FCC7" blockId="51.[361,1227,769,866]" box="[580,1008,769,797]" pageId="51" pageNumber="52">
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<heading id="D08081CAFFC4FF87FDEFFCD5A4F6FCC7" box="[580,1008,769,797]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
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<taxonomicName id="4C774D25FFC4FF87FDEFFCD5A488FCC6" authority="Constant & Pham, 2024" authorityName="Constant & Pham" authorityYear="2024" box="[580,910,769,796]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="quangninhensis" status="sp. nov.">
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<emphasis id="B903EAB4FFC4FF87FDEFFCD5A488FCC6" bold="true" box="[580,910,769,796]" italics="true" pageId="51" pageNumber="52">Pusulissus quangninhensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057CFFFC4FF87FC3EFCD7A4F6FCC7" box="[917,1008,771,797]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BC836A6FFC4FF87FEC2FCF1A3CDFC9A" blockId="51.[361,1227,769,866]" box="[361,1227,805,832]" pageId="51" pageNumber="52">
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<uri id="FFE63AA4FFC4FF87FEC2FCF1A3CDFC9A" box="[361,1227,805,832]" pageId="51" pageNumber="52">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD10C73FFC4FF87FD2FFCF1A3CDFC9A" box="[644,1227,805,832]" pageId="51" pageNumber="52">A5D5064C-EAB2-4C1A-809A-DA2074260167</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D652DFFC4FF87FD73FC9CA45AFCB8" box="[728,860,840,866]" pageId="51" pageNumber="52" type="description">
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<paragraph id="8BC836A6FFC4FF87FD73FC9CA45AFCB8" blockId="51.[361,1227,769,866]" box="[728,860,840,866]" pageId="51" pageNumber="52">
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<figureCitation id="134C2A23FFC4FF87FD73FC9CA45AFCB8" box="[728,860,840,866]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="53.[189,232,1928,1954]" captionStartId-1="54.[189,232,1892,1918]" captionTargetBox-0="[248,1337,273,1877]" captionTargetBox-1="[264,1322,275,1843]" captionTargetId-0="figure-12@53.[244,1343,265,1881]" captionTargetId-1="figure-19@54.[257,1330,265,1843]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." captionText-1="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.10559593" figureDoi-1="http://doi.org/10.5281/zenodo.10559595" httpUri-0="https://zenodo.org/record/10559593/files/figure.png" httpUri-1="https://zenodo.org/record/10559595/files/figure.png" pageId="51" pageNumber="52">Figs 26–27</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D652DFFC4FF87FF16FC5EA57CFA1C" pageId="51" pageNumber="52" type="diagnosis">
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<paragraph id="8BC836A6FFC4FF87FF16FC5EA630FC7E" blockId="51.[189,310,906,932]" box="[189,310,906,932]" pageId="51" pageNumber="52">
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<heading id="D08081CAFFC4FF87FF16FC5EA630FC7E" bold="true" box="[189,310,906,932]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
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<emphasis id="B903EAB4FFC4FF87FF16FC5EA630FC7E" bold="true" box="[189,310,906,932]" pageId="51" pageNumber="52">Diagnosis</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="8BC836A6FFC4FF87FF16FC6CA45FFC08" blockId="51.[189,1399,952,1225]" box="[189,857,952,979]" pageId="51" pageNumber="52">
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<taxonomicName id="4C774D25FFC4FF87FF16FC6CA6FEFC08" authority="Constant & Pham, 2024" authorityName="Constant & Pham" authorityYear="2024" box="[189,504,952,978]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="quangninhensis" status="sp. nov.">
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<emphasis id="B903EAB4FFC4FF87FF16FC6CA6FEFC08" box="[189,504,952,978]" italics="true" pageId="51" pageNumber="52">Pusulissus quangninhensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057CFFFC4FF87FDABFC6DA55CFC09" box="[512,602,953,979]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
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can be recognized by
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</paragraph>
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<paragraph id="8BC836A6FFC4FF87FF16FC08A5BAFBE6" blockId="51.[189,1399,952,1225]" pageId="51" pageNumber="52">
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(1) the anal tube widely subrhomboidal as wide as long in midline with apical margin truncate and slightly emarginate in middle, with lateral flaps distinctly reflexed ventrad with lateral angles angularly rounded (
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<emphasis id="B903EAB4FFC4FF87FE4AFBF6A504FBE6" box="[481,514,1058,1084]" italics="true" pageId="51" pageNumber="52">An</emphasis>
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–
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<figureCitation id="134C2A23FFC4FF87FDB4FBF6A5A8FBE6" box="[543,686,1058,1084]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="51" pageNumber="52">Fig. 27A–B</figureCitation>
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);
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</paragraph>
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<paragraph id="8BC836A6FFC4FF87FF16FB91A329FB85" blockId="51.[189,1399,952,1225]" box="[189,1071,1093,1120]" pageId="51" pageNumber="52">
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(2) the neck of the capitulum of the gonostyli rather high (
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<emphasis id="B903EAB4FFC4FF87FCD0FB92A491FB85" box="[891,919,1094,1119]" italics="true" pageId="51" pageNumber="52">ca</emphasis>
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–
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<figureCitation id="134C2A23FFC4FF87FC1FFB91A324FB85" box="[948,1058,1093,1119]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="51" pageNumber="52">Fig. 27A</figureCitation>
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);
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</paragraph>
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<paragraph id="8BC836A6FFC4FF87FF16FBBDA21BFB13" blockId="51.[189,1399,952,1225]" pageId="51" pageNumber="52">
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(3) the shape of the lateroventral processes of the aedeagus directed mesocephalad in basal ⅓, then strongly upcurved and sinuate as to wrap around aedeagus, and with apical ⅓ indistinctly curved cephalad, but not surpassing dorsal margin of aedeagus in lateral view (
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<emphasis id="B903EAB4FFC4FF87FBE9FB7BA360FB13" box="[1090,1126,1199,1225]" italics="true" pageId="51" pageNumber="52">lvp</emphasis>
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–
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<figureCitation id="134C2A23FFC4FF87FB28FB7BA216FB13" box="[1155,1296,1199,1225]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="51" pageNumber="52">Fig. 27E–G</figureCitation>
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).
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</paragraph>
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<paragraph id="8BC836A6FFC4FF87FF16FB24A6CFFAD0" blockId="51.[189,457,1264,1290]" box="[189,457,1264,1290]" pageId="51" pageNumber="52">
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<heading id="D08081CAFFC4FF87FF16FB24A6CFFAD0" bold="true" box="[189,457,1264,1290]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
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<emphasis id="B903EAB4FFC4FF87FF16FB24A6CFFAD0" bold="true" box="[189,457,1264,1290]" pageId="51" pageNumber="52">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC836A6FFC4FF87FF16FACBA57CFA1C" blockId="51.[189,1399,1311,1478]" pageId="51" pageNumber="52">
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The closest species is
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<taxonomicName id="4C774D25FFC4FF87FE69FAF4A57EFAE3" authorityName="Bourgoin & Wang" authorityYear="2020" box="[450,632,1311,1337]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="phiaoacensis">
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<emphasis id="B903EAB4FFC4FF87FE69FAF4A57EFAE3" box="[450,632,1311,1337]" italics="true" pageId="51" pageNumber="52">P. phiaoacensis</emphasis>
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</taxonomicName>
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, which differs by a less wide anal tube (
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<bibRefCitation id="EFE64B57FFC4FF87FBF9FACBA275FAE3" author="Bourgoin T. & Wang M. - L." box="[1106,1395,1311,1337]" pageId="51" pageNumber="52" pagination="69 - 94" refId="ref32456" refString="Bourgoin T. & Wang M. - L. 2020. Parahiraciini (Hemiptera, Fulgoromorpha, Issidae): one new genus, two new species and three new subtribes. ZooKeys 997: 69 - 94. https: // doi. org / 10.3897 / zookeys. 997.52857" type="journal article" year="2020">Bourgoin & Wang 2020</bibRefCitation>
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||
: figs 8–9), a shorter neck of the capitulum of the gonostyli (
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<bibRefCitation id="EFE64B57FFC4FF87FC22FA96A3B6FA86" author="Bourgoin T. & Wang M. - L." box="[905,1200,1346,1373]" pageId="51" pageNumber="52" pagination="69 - 94" refId="ref32456" refString="Bourgoin T. & Wang M. - L. 2020. Parahiraciini (Hemiptera, Fulgoromorpha, Issidae): one new genus, two new species and three new subtribes. ZooKeys 997: 69 - 94. https: // doi. org / 10.3897 / zookeys. 997.52857" type="journal article" year="2020">Bourgoin & Wang 2020</bibRefCitation>
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: fig. 10) and the shape of the medioventral processes of the aedeagus, not directed mesad (subparallel) in basal portion, then strongly recurved with apex surpassing dorsal margin of aedeagus and directed posterodorsad (
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<bibRefCitation id="EFE64B57FFC4FF87FF6EFA7FA6E2FA1C" author="Bourgoin T. & Wang M. - L." box="[197,484,1451,1478]" pageId="51" pageNumber="52" pagination="69 - 94" refId="ref32456" refString="Bourgoin T. & Wang M. - L. 2020. Parahiraciini (Hemiptera, Fulgoromorpha, Issidae): one new genus, two new species and three new subtribes. ZooKeys 997: 69 - 94. https: // doi. org / 10.3897 / zookeys. 997.52857" type="journal article" year="2020">Bourgoin & Wang 2020</bibRefCitation>
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: figs 11–13).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36D652DFFC4FF87FF16FA39A6F9F983" pageId="51" pageNumber="52" type="etymology">
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<paragraph id="8BC836A6FFC4FF87FF16FA39A642F9DD" blockId="51.[189,324,1517,1543]" box="[189,324,1517,1543]" pageId="51" pageNumber="52">
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<heading id="D08081CAFFC4FF87FF16FA39A642F9DD" bold="true" box="[189,324,1517,1543]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
|
||
<emphasis id="B903EAB4FFC4FF87FF16FA39A642F9DD" bold="true" box="[189,324,1517,1543]" pageId="51" pageNumber="52">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC4FF87FF16F9CFA6F9F983" blockId="51.[189,1398,1563,1625]" pageId="51" pageNumber="52">
|
||
The species epithet refers to ‘Quang Ninh’, the province where Dong Son-Ky Thuong Nature Reserve, the
|
||
<typeStatus id="54CC8804FFC4FF87FF43F9EBA61DF983" box="[232,283,1599,1625]" pageId="51" pageNumber="52">type</typeStatus>
|
||
locality, is located.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC4FF87FF16F954A4B6F842" pageId="51" pageNumber="52" type="materials_examined">
|
||
<paragraph id="8BC836A6FFC4FF87FF16F954A66AF940" blockId="51.[189,364,1664,1690]" box="[189,364,1664,1690]" pageId="51" pageNumber="52">
|
||
<heading id="D08081CAFFC4FF87FF16F954A66AF940" bold="true" box="[189,364,1664,1690]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
|
||
<emphasis id="B903EAB4FFC4FF87FF16F954A66AF940" bold="true" box="[189,364,1664,1690]" pageId="51" pageNumber="52">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC4FF87FF41F97BA65AF913" blockId="51.[189,1398,1711,1843]" box="[234,348,1711,1737]" pageId="51" pageNumber="52">
|
||
<emphasis id="B903EAB4FFC4FF87FF41F97BA65AF913" bold="true" box="[234,348,1711,1737]" pageId="51" pageNumber="52">
|
||
<typeStatus id="54CC8804FFC4FF87FF41F97BA65AF913" box="[234,348,1711,1737]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC4FF87FF16F906A444F8E8" blockId="51.[189,1398,1711,1843]" pageId="51" pageNumber="52">
|
||
<materialsCitation id="3B1F3CFBFFC4FF87FF16F906A444F8E8" collectingDate="2022-08-27" collectionCode="RBINS" collectorName="Project & J. Constant & J. Bresseel & L. Semeraro" country="Vietnam" elevation="550" latitude="21.141388" location="Thuong Nature Reserve" longLatPrecision="21" longitude="107.08139" municipality="Dong" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Quang Ninh" typeStatus="holotype">
|
||
<collectingCountry id="F3607636FFC4FF87FF16F906A642F936" box="[189,324,1746,1772]" name="Vietnam" pageId="51" pageNumber="52">VIETNAM</collectingCountry>
|
||
•
|
||
<specimenCount id="9D71FD2FFFC4FF87FEF0F906A677F936" box="[347,369,1746,1772]" count="1" pageId="51" pageNumber="52" type="male">♂</specimenCount>
|
||
(dissected –
|
||
<figureCitation id="134C2A23FFC4FF87FDA2F906A589F936" box="[521,655,1746,1772]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="53.[189,232,1928,1954]" captionStartId-1="54.[189,232,1892,1918]" captionTargetBox-0="[248,1337,273,1877]" captionTargetBox-1="[264,1322,275,1843]" captionTargetId-0="figure-12@53.[244,1343,265,1881]" captionTargetId-1="figure-19@54.[257,1330,265,1843]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." captionText-1="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.10559593" figureDoi-1="http://doi.org/10.5281/zenodo.10559595" httpUri-0="https://zenodo.org/record/10559593/files/figure.png" httpUri-1="https://zenodo.org/record/10559595/files/figure.png" pageId="51" pageNumber="52">Figs 26–27</figureCitation>
|
||
);
|
||
<collectingRegion id="49B3F844FFC4FF87FD08F906A4A2F936" box="[675,932,1746,1772]" country="Vietnam" name="Quang Ninh" pageId="51" pageNumber="52">Quang Ninh Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BACACDCFFC4FF87FC04F906A4F6F936" box="[943,1008,1746,1772]" pageId="51" pageNumber="52">Dong</collectingMunicipality>
|
||
Son-Ky
|
||
<location id="8EA8607DFFC4FF87FBF3F906A274F936" LSID="urn:lsid:plazi:treatment:03DE87B0FFC4FF83FDEFFCD5A37AFBA2:8EA8607DFFC4FF87FBF3F906A274F936" box="[1112,1394,1746,1772]" country="Vietnam" latitude="21.141388" longLatPrecision="21" longitude="107.08139" municipality="Dong" name="Thuong Nature Reserve" pageId="51" pageNumber="52" stateProvince="Quang Ninh">Thuong Nature Reserve</location>
|
||
;
|
||
<geoCoordinate id="EE435061FFC4FF87FF16F920A650F8CA" box="[189,342,1780,1808]" degrees="21" direction="north" minutes="08" orientation="latitude" pageId="51" pageNumber="52" precision="15" seconds="29" value="21.141388">21°08′29″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE435061FFC4FF87FECEF920A50FF8CA" box="[357,521,1780,1808]" degrees="107" direction="east" minutes="04" orientation="longitude" pageId="51" pageNumber="52" precision="15" seconds="53" value="107.08139">107°04′53″ E</geoCoordinate>
|
||
;
|
||
<date id="FFC91066FFC4FF87FDB2F921A5C7F8D5" box="[537,705,1781,1807]" pageId="51" pageNumber="52" value="2022-08-27">
|
||
<collectingDate id="EF8DE98EFFC4FF87FDB2F921A5C7F8D5" box="[537,705,1781,1807]" pageId="51" pageNumber="52" value="2022-08-27">27 Aug. 2022</collectingDate>
|
||
</date>
|
||
;
|
||
<elevation id="005AD195FFC4FF87FD7AF921A467F8D5" box="[721,865,1781,1808]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="51" pageNumber="52" unit="m" value="550.0">
|
||
<quantity id="4C8F9B43FFC4FF87FD7AF921A418F8CA" box="[721,798,1781,1808]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="51" pageNumber="52" unit="m" value="550.0">550 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
; secondary forest; GTI
|
||
<collectorName id="26825370FFC4FF87FB2FF921A3DDF8D5" box="[1156,1243,1781,1807]" pageId="51" pageNumber="52">Project</collectorName>
|
||
;
|
||
<collectorName id="26825370FFC4FF87FB41F922A274F8D5" box="[1258,1394,1781,1808]" pageId="51" pageNumber="52">J. Constant</collectorName>
|
||
,
|
||
<collectorName id="26825370FFC4FF87FF16F8CDA63CF8E8" box="[189,314,1816,1843]" pageId="51" pageNumber="52">J. Bresseel</collectorName>
|
||
and
|
||
<collectorName id="26825370FFC4FF87FED8F8CDA502F8E8" box="[371,516,1816,1843]" pageId="51" pageNumber="52">L. Semeraro</collectorName>
|
||
leg.; I.G.: 34.518;
|
||
<collectionCode id="ED66AE63FFC4FF87FD4FF8CCA438F8E8" box="[740,830,1816,1842]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="51" pageNumber="52" type="Museum">RBINS</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC4FF87FF41F88EA65AF8AE" blockId="51.[189,944,1882,1944]" box="[234,348,1882,1908]" pageId="51" pageNumber="52">
|
||
<emphasis id="B903EAB4FFC4FF87FF41F88EA65AF8AE" bold="true" box="[234,348,1882,1908]" pageId="51" pageNumber="52">
|
||
<typeStatus id="54CC8804FFC4FF87FF41F88EA65AF8AE" box="[234,348,1882,1908]" pageId="51" pageNumber="52" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC4FF87FF16F8A9A4B6F842" blockId="51.[189,944,1882,1944]" box="[189,944,1917,1944]" pageId="51" pageNumber="52">
|
||
<materialsCitation id="3B1F3CFBFFC4FF87FF16F8A9A4AAF842" box="[189,940,1917,1944]" collectingDate="2022-08-27" collectionCode="VNMN" collectorName="Project & J. Constant & J. Bresseel & L. Semeraro" country="Vietnam" elevation="550" latitude="21.141388" location="Thuong Nature Reserve" longLatPrecision="21" longitude="107.08139" municipality="Dong" pageId="51" pageNumber="52" specimenCount="1" specimenCount-female="1" stateProvince="Quang Ninh" typeStatus="paratype">
|
||
<collectingCountry id="F3607636FFC4FF87FF16F8A9A642F84D" box="[189,324,1917,1943]" name="Vietnam" pageId="51" pageNumber="52">VIETNAM</collectingCountry>
|
||
•
|
||
<specimenCount id="9D71FD2FFFC4FF87FEF6F8A9A68CF842" box="[349,394,1917,1944]" count="1" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED66AE63FFC4FF87FCE5F8AAA4AAF842" box="[846,940,1918,1944]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="51" pageNumber="52">VNMN</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC3FF83FF16FEDCA596FCAF" lastPageId="55" lastPageNumber="56" pageId="52" pageNumber="53" type="description">
|
||
<paragraph id="8BC836A6FFC3FF80FF16FEDCA648FEF8" blockId="52.[189,334,264,290]" box="[189,334,264,290]" pageId="52" pageNumber="53">
|
||
<heading id="D08081CAFFC3FF80FF16FEDCA648FEF8" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
|
||
<emphasis id="B903EAB4FFC3FF80FF16FEDCA648FEF8" bold="true" box="[189,334,264,290]" pageId="52" pageNumber="53">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FF80FF16FEECA5E6FEAE" blockId="52.[189,1399,311,373]" pageId="52" pageNumber="53">
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FF16FEECA668FE8A" baselines="331,331" box="[189,366,312,338]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="52" pageNumber="53">MEASUREMENTS</smallCapsWord>
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FEDDFEE9A6A5FE8A" baselines="331" box="[374,419,317,336]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="52" pageNumber="53">AND</smallCapsWord>
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FE07FEE9A6F3FE8A" baselines="331" box="[428,501,317,336]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="ratios" pageId="52" pageNumber="53">RATIOS</smallCapsWord>
|
||
. LT:
|
||
<specimenCount id="9D71FD2FFFC3FF80FD9FFEECA54CFE88" box="[564,586,312,338]" count="1" pageId="52" pageNumber="53" type="male">♂</specimenCount>
|
||
(n = 1):
|
||
<quantity id="4C8F9B43FFC3FF80FD1FFEE3A414FE88" box="[692,786,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="52" pageNumber="53" unit="mm" value="6.9">6.9 mm</quantity>
|
||
,
|
||
<specimenCount id="9D71FD2FFFC3FF80FCB5FEECA432FE88" box="[798,820,312,338]" count="1" pageId="52" pageNumber="53" type="female">♀</specimenCount>
|
||
(n = 1):
|
||
<quantity id="4C8F9B43FFC3FF80FC35FEE3A4FDFE88" box="[926,1019,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="52" pageNumber="53" unit="mm" value="7.1">7.1 mm</quantity>
|
||
; LT/BB = 1.44; LTg/BTg = 2.0; LW/BW = 1.30; BV/LV = 2.15; LF/BF = 1.20.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FF80FF16FE48A40DFCE3" blockId="52.[189,1400,412,825]" pageId="52" pageNumber="53">
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FF16FE48A7FAFE6E" baselines="431,431" box="[189,252,412,438]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="52" pageNumber="53">HEAD</smallCapsWord>
|
||
(
|
||
<figureCitation id="134C2A23FFC3FF80FEBBFE48A6A5FE6C" box="[272,419,412,438]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[248,1337,273,1877]" captionTargetId="figure-12@53.[244,1343,265,1881]" captionTargetPageId="53" captionText="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="52" pageNumber="53">Fig. 26A–D</figureCitation>
|
||
). Vertex variegated brown, distinctly broader than long in midline and with lateral margins longer than midline, with all margins carinate; anterior margin weakly convex, posterior one deeply angularly concave and lateral ones subparallel; disc shallowly excavate with obsolete median carina. Side of head yellowish with triangular black marking at anterodorsal angle; anteroventral angle projecting anteriorly in a small round lobe. Frons elongate and generally smooth, brown with band along dorsal margin and narrower band subparallel to lateral margins, blackish, distinct yellowish spot in middle on each side of median carina and yellowish band along frontoclypeal suture; median carina extending from dorsal margin down nearly to rounded frontoclypeal suture, orangish on disc; row of irregular yellowish tubercles along dorsal and lateral margins (in black bands); dorsal margin distinctly concave, lateral margins distinctly sinuate. Clypeus variegated black-brown, weakly elevated medially. Labium brown with last segment longer than broad, and shorter than penultimate. Scape short, ring-shaped, brown; pedicel bulbous, yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FF80FF16FCB5A68DFB6D" blockId="52.[188,1400,864,1207]" pageId="52" pageNumber="53">
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FF16FCB5A61FFCA3" baselines="884,884" box="[189,281,865,891]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="52" pageNumber="53">THORAX</smallCapsWord>
|
||
(
|
||
<figureCitation id="134C2A23FFC3FF80FE82FCB4A6DAFCA0" box="[297,476,864,891]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[248,1337,273,1877]" captionTargetId="figure-12@53.[244,1343,265,1881]" captionTargetPageId="53" captionText="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="52" pageNumber="53">Fig. 26A, C–D</figureCitation>
|
||
). Pronotum variegated yellowish brown; about 0.6 times as long as mesonotum in midline; anterior margin carinate, strongly sinuate and strongly, angularly protruding anteriorly between eyes, with carinae directed obliquely posteriorly, not reaching hind margin of pronotum; posterior margin weakly bisinuate; no median carina but with impressed point on each side of median line; lateral portion behind eye very narrow, laminate; blunt, pale yellowish tubercles along posterior margin; paranotal lobes (lateral view) broad, brown in outer portion, yellowish in inner portion and with large black marking behind lower margin of eye, and with posteroventral angle straight. Mesonotum variegated yellowish brown with longitudinal carinae obsolete but marked by dark brown lines; smooth, slightly depressed before scutellum; nearly complete transverse carina along anterior margin. Tegulae yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FF80FF16FB0BA629F982" blockId="52.[188,1401,1246,1624]" pageId="52" pageNumber="53">
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FF16FB0BA624FB2D" baselines="1266,1266" box="[189,290,1247,1273]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Tegmina" pageId="52" pageNumber="53">TEGMINA</smallCapsWord>
|
||
(
|
||
<figureCitation id="134C2A23FFC3FF80FE9DFB0AA6E9FB23" box="[310,495,1246,1273]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[248,1337,273,1877]" captionTargetId="figure-12@53.[244,1343,265,1881]" captionTargetPageId="53" captionText="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="52" pageNumber="53">Fig. 26A–B, D</figureCitation>
|
||
). Variegated yellowish brown (greener in live specimens) with darker area along costal margin, sometimes darker basal marking between ScP+RA and RP and more or less distinct transverse band at midlength; subcoriaceous with longitudinal veins yellow, elevated and with dense reticulum of pale yellow veinlets; shape elongate and convex with sides broadly rounded, widest at basal ⅓, about 2.4 times as long as broad; narrowly rounded apically. Postclaval margin weakly rounded on distal half and weakly notched at apex of clavus. Clavus closed, reaching about midlength of tegmen. Venation: ScP+R forking close to base after short common stem, ScP+RA and RP running more or less parallel to costal margin and not forked; MP forked close to base after short common stem, resulting veins both forked again further; CuA simple, sinuate, more or less parallel to claval joint, then to sutural margin towards apex of tegmen; Pcu and A
|
||
<subScript id="17F334E3FFC3FF80FD18F9FAA5BAF9E6" attach="left" box="[691,700,1582,1596]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
|
||
fused at apical ⅓ of clavus, resulting Pcu+A
|
||
<subScript id="17F334E3FFC3FF80FB6FF9FAA3CBF9E6" attach="left" box="[1220,1229,1582,1596]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
|
||
reaching apex of clavus.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FF80FF16F955A3A0F890" blockId="52.[188,1399,1664,1873]" pageId="52" pageNumber="53">
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FF16F955A7F1F943" baselines="1684,1684" box="[189,247,1665,1691]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="52" pageNumber="53">HIND</smallCapsWord>
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FEA9F952A640F943" baselines="1684" box="[258,326,1670,1689]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="wings" pageId="52" pageNumber="53">WINGS</smallCapsWord>
|
||
(
|
||
<figureCitation id="134C2A23FFC3FF80FEF1F954A6C1F940" box="[346,455,1664,1691]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[248,1337,273,1877]" captionTargetId="figure-12@53.[244,1343,265,1881]" captionTargetPageId="53" captionText="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="52" pageNumber="53">Fig. 26E</figureCitation>
|
||
). Brown with large basicostal portion of lobe Sc-R-MP-CuA and base of lobe CuP-Pcu-A 1 yellow-brown; venation slightly darker than corresponding background; wing broader than tegmen and deeply bilobed at CuP; costal margin moderately sinuate; CuP-Pcu-A
|
||
<subScript id="17F334E3FFC3FF80FBC4F90DA37EF93D" attach="left" box="[1135,1144,1753,1767]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
|
||
lobe broadly rounded along postclaval margin and about 1.4 times as wide as Sc-R-MP-CuA lobe, the latter slightly shorter and with apical margin obliquely rounded; A 2 lobe reduced and narrow. Venation: longitudinal veins ScP-R-MP-Cu well distinct; Pcu and A
|
||
<subScript id="17F334E3FFC3FF80FD23F897A597F88B" attach="left" box="[648,657,1859,1873]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
|
||
separated; numerous cross-veins; A
|
||
<subScript id="17F334E3FFC3FF80FB93F897A347F88B" attach="left" box="[1080,1089,1859,1873]" fontSize="6" pageId="52" pageNumber="53">2</subScript>
|
||
distinct.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FF80FF16F8A6A3ADF808" blockId="52.[189,1399,1905,2002]" pageId="52" pageNumber="53">
|
||
<smallCapsWord id="8D2EA07AFFC3FF80FF16F8A6A7F0F850" baselines="1925,1925" box="[189,246,1906,1932]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="52" pageNumber="53">LEGS</smallCapsWord>
|
||
(
|
||
<figureCitation id="134C2A23FFC3FF80FEA2F8A5A69AF851" box="[265,412,1905,1932]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[248,1337,273,1877]" captionTargetId="figure-12@53.[244,1343,265,1881]" captionTargetPageId="53" captionText="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="52" pageNumber="53">Fig. 26A–D</figureCitation>
|
||
). Moderately elongate and slender, brown femora darker than corresponding tibiae; posterior margin of pro- and mesofemora with row of acute irregular small teeth. Metatibiae with 2 lateral spines in distal half and 8 apical spines. Metatibiotarsal formula: (2) 8 / 8 / 2.
|
||
</paragraph>
|
||
<caption id="DF08662EFFC2FF81FF16F85CA63DF832" ID-DOI="http://doi.org/10.5281/zenodo.10559593" ID-Zenodo-Dep="10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="53" pageNumber="54" startId="53.[189,232,1928,1954]" targetBox="[248,1337,273,1877]" targetPageId="53">
|
||
<paragraph id="8BC836A6FFC2FF81FF16F85CA63DF832" blockId="53.[189,1399,1928,2024]" pageId="53" pageNumber="54">
|
||
<emphasis id="B903EAB4FFC2FF81FF16F85CA61DF878" bold="true" box="[189,283,1928,1954]" pageId="53" pageNumber="54">Fig. 26.</emphasis>
|
||
<taxonomicName id="4C774D25FFC2FF81FE8FF85CA567F878" authority="Constant & Pham, 2024" authorityName="Constant & Pham" authorityYear="2024" box="[292,609,1928,1954]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="quangninhensis" status="sp. nov.">
|
||
<emphasis id="B903EAB4FFC2FF81FE8FF85CA567F878" box="[292,609,1928,1954]" italics="true" pageId="53" pageNumber="54">Pusulissus quangninhensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23057CFFFC2FF81FDC1F85CA5C1F878" box="[618,711,1928,1954]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, dissected holotype, ♂ (RBINS).
|
||
<emphasis id="B903EAB4FFC2FF81FBFCF85CA36AF878" bold="true" box="[1111,1132,1928,1954]" pageId="53" pageNumber="54">A</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B903EAB4FFC2FF81FF15F87FA7D6F81F" bold="true" box="[190,208,1963,1989]" pageId="53" pageNumber="54">B</emphasis>
|
||
. Habitus, ventral view.
|
||
<emphasis id="B903EAB4FFC2FF81FE59F87FA501F81F" bold="true" box="[498,519,1963,1989]" pageId="53" pageNumber="54">C</emphasis>
|
||
. Habitus, perpendicular view of frons.
|
||
<emphasis id="B903EAB4FFC2FF81FC49F87FA4F1F81F" bold="true" box="[994,1015,1963,1989]" pageId="53" pageNumber="54">D</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="B903EAB4FFC2FF81FABBF87FA222F81F" bold="true" box="[1296,1316,1963,1989]" pageId="53" pageNumber="54">E</emphasis>
|
||
. Right hind wing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF08662EFFC1FF82FF16F8B0A35DF832" ID-DOI="http://doi.org/10.5281/zenodo.10559595" ID-Zenodo-Dep="10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="54" pageNumber="55" startId="54.[189,232,1892,1918]" targetBox="[264,1322,275,1843]" targetPageId="54">
|
||
<paragraph id="8BC836A6FFC1FF82FF16F8B0A35DF832" blockId="54.[189,1399,1892,2025]" pageId="54" pageNumber="55">
|
||
<emphasis id="B903EAB4FFC1FF82FF16F8B0A61EF8A5" bold="true" box="[189,280,1892,1919]" pageId="54" pageNumber="55">Fig. 27.</emphasis>
|
||
<taxonomicName id="4C774D25FFC1FF82FE8BF8B1A55DF8A5" authority="Constant & Pham, 2024" authorityName="Constant & Pham" authorityYear="2024" box="[288,603,1893,1919]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="quangninhensis" status="sp. nov.">
|
||
<emphasis id="B903EAB4FFC1FF82FE8BF8B1A55DF8A5" box="[288,603,1893,1919]" italics="true" pageId="54" pageNumber="55">Pusulissus quangninhensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23057CFFFC1FF82FDC9F8B1A5B9F8A5" box="[610,703,1893,1919]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (RBINS), terminalia.
|
||
<emphasis id="B903EAB4FFC1FF82FBF1F8B0A392F8A4" bold="true" box="[1114,1172,1892,1918]" pageId="54" pageNumber="55">A–D</emphasis>
|
||
. Pygofer, anal tube and gonostyli.
|
||
<emphasis id="B903EAB4FFC1FF82FEC7F85CA687F878" bold="true" box="[364,385,1928,1954]" pageId="54" pageNumber="55">A</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="B903EAB4FFC1FF82FD99F85CA543F878" bold="true" box="[562,581,1928,1954]" pageId="54" pageNumber="55">B</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="B903EAB4FFC1FF82FD59F85CA401F878" bold="true" box="[754,775,1928,1954]" pageId="54" pageNumber="55">C</emphasis>
|
||
. Laterocaudal view.
|
||
<emphasis id="B903EAB4FFC1FF82FC56F85CA314F878" bold="true" box="[1021,1042,1928,1954]" pageId="54" pageNumber="55">D</emphasis>
|
||
. Caudal view.
|
||
<emphasis id="B903EAB4FFC1FF82FB68F85CA3F5F878" bold="true" box="[1219,1267,1928,1954]" pageId="54" pageNumber="55">E–J</emphasis>
|
||
. Aedeagus and connective.
|
||
<emphasis id="B903EAB4FFC1FF82FE2DF87FA69CF81F" bold="true" box="[390,410,1963,1989]" pageId="54" pageNumber="55">E</emphasis>
|
||
. Left lateral view.
|
||
<emphasis id="B903EAB4FFC1FF82FD20F87FA59BF81F" bold="true" box="[651,669,1963,1989]" pageId="54" pageNumber="55">F</emphasis>
|
||
. Left ventrolateral view.
|
||
<emphasis id="B903EAB4FFC1FF82FC72F87FA4F6F81F" bold="true" box="[985,1008,1963,1989]" pageId="54" pageNumber="55">G</emphasis>
|
||
. Caudal view.
|
||
<emphasis id="B903EAB4FFC1FF82FB06F87FA3C2F81F" bold="true" box="[1197,1220,1963,1989]" pageId="54" pageNumber="55">H</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="B903EAB4FFC1FF82FF16F81AA7CEF832" bold="true" box="[189,200,1998,2024]" pageId="54" pageNumber="55">I</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="B903EAB4FFC1FF82FED9F81AA687F832" bold="true" box="[370,385,1998,2024]" pageId="54" pageNumber="55">J</emphasis>
|
||
. Anterodorsal view.Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC836A6FFC0FF83FF16FEDDA271FEF9" blockId="55.[189,1399,265,291]" box="[189,1399,265,291]" pageId="55" pageNumber="56">
|
||
<smallCapsWord id="8D2EA07AFFC0FF83FF16FEDDA62AFEFB" baselines="284,284" box="[189,300,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="55" pageNumber="56">ABDOMEN</smallCapsWord>
|
||
(
|
||
<figureCitation id="134C2A23FFC0FF83FE97FEDDA6A1FEF9" box="[316,423,265,291]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[248,1337,273,1877]" captionTargetId="figure-12@53.[244,1343,265,1881]" captionTargetPageId="53" captionText="Fig. 26. Pusulissus quangninhensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, perpendicular view of frons. D. Habitus, lateral view. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559593" httpUri="https://zenodo.org/record/10559593/files/figure.png" pageId="55" pageNumber="56">Fig. 26B</figureCitation>
|
||
). Black brown, each tergite with a yellowish, crescent-shaped marking on each side.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC0FF83FF16FE9FA596FCAF" blockId="55.[189,1399,330,885]" pageId="55" pageNumber="56">
|
||
<smallCapsWord id="8D2EA07AFFC0FF83FF16FE9FA7F9FEB9" baselines="350,350" box="[189,255,331,357]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="55" pageNumber="56">MALE</smallCapsWord>
|
||
<smallCapsWord id="8D2EA07AFFC0FF83FEA0FE84A68AFEB9" baselines="350" box="[267,396,336,355]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="55" pageNumber="56">TERMINALIA</smallCapsWord>
|
||
. Pygofer (
|
||
<emphasis id="B903EAB4FFC0FF83FDBBFE9FA529FEBE" box="[528,559,331,356]" italics="true" pageId="55" pageNumber="56">Py</emphasis>
|
||
–
|
||
<figureCitation id="134C2A23FFC0FF83FDF8FE9EA5E3FEBF" box="[595,741,330,357]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="55" pageNumber="56">Fig. 27A–D</figureCitation>
|
||
) about 2.5 times as high as long in lateral view, with anterior margin weakly concave and posterior margin more or less straight on both sides of an obtuse angle under half height. Gonostyli (
|
||
<emphasis id="B903EAB4FFC0FF83FDC1FE45A579FE71" box="[618,639,401,427]" italics="true" pageId="55" pageNumber="56">G</emphasis>
|
||
–
|
||
<figureCitation id="134C2A23FFC0FF83FD0AFE45A452FE71" box="[673,852,401,427]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="55" pageNumber="56">Fig. 27A, C–D</figureCitation>
|
||
) (in lateral view) about as long as high (with dorsal capitulum), projecting posteriorly in distinct lobe rounded apically, and concave; capitulum (
|
||
<emphasis id="B903EAB4FFC0FF83FAFAFE61A26BFE14" box="[1361,1389,437,462]" italics="true" pageId="55" pageNumber="56">ca</emphasis>
|
||
) with wide neck, directed dorsad, with distal portion anteroposteriorly flattened, with apical hook directed cephalad and with lateral laminate projection in dorsal half with outer margin under it strongly concave in caudal view. Aedeagus (
|
||
<figureCitation id="134C2A23FFC0FF83FE50FDCAA578FDE2" box="[507,638,542,568]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="55" pageNumber="56">Fig. 27E–J</figureCitation>
|
||
) symmetrical, strongly curved dorsad (in lateral view), but with distal ⅓ more or less straight; pair of elongate lateroventral processes (
|
||
<emphasis id="B903EAB4FFC0FF83FBBAFD95A333FD81" box="[1041,1077,577,603]" italics="true" pageId="55" pageNumber="56">lvp</emphasis>
|
||
) directed mesocephalad in basal ⅓, then strongly upcurved and sinuate as to wrap around aedeagus and with apical ⅓ weakly curved cephalad, but not surpassing dorsal margin of aedeagus in lateral view; ventral lobe of periandrium (
|
||
<emphasis id="B903EAB4FFC0FF83FAFCFD53A26AFD7B" box="[1367,1388,647,673]" italics="true" pageId="55" pageNumber="56">vl</emphasis>
|
||
) evenly widening posterior to medioventral processes, then more or less parallel-sided, apical margin truncate with small round projection in middle. Connective (
|
||
<emphasis id="B903EAB4FFC0FF83FC3BFD1AA4AAFD3D" box="[912,940,718,743]" italics="true" pageId="55" pageNumber="56">co</emphasis>
|
||
) elongate, curved, with teciductus (
|
||
<emphasis id="B903EAB4FFC0FF83FAFCFD1AA26AFD3D" box="[1367,1388,718,743]" italics="true" pageId="55" pageNumber="56">te</emphasis>
|
||
) well developed. Anal tube (
|
||
<emphasis id="B903EAB4FFC0FF83FDA3FD25A52FFCD1" box="[520,553,753,779]" italics="true" pageId="55" pageNumber="56">An</emphasis>
|
||
–
|
||
<figureCitation id="134C2A23FFC0FF83FDE1FD25A5DFFCD1" box="[586,729,753,779]" captionStart="Fig" captionStartId="54.[189,232,1892,1918]" captionTargetBox="[264,1322,275,1843]" captionTargetId="figure-19@54.[257,1330,265,1843]" captionTargetPageId="54" captionText="Fig. 27. Pusulissus quangninhensis sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Lateral view. B. Dorsal view. C. Laterocaudal view. D. Caudal view. E–J. Aedeagus and connective. E. Left lateral view. F. Left ventrolateral view. G. Caudal view. H. Ventral view. I. Dorsal view. J. Anterodorsal view.Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559595" httpUri="https://zenodo.org/record/10559595/files/figure.png" pageId="55" pageNumber="56">Fig. 27A–D</figureCitation>
|
||
) dorsoventrally flattened, in dorsal view subrhomboidal with rather short parallel-sided base, and apical margin truncate and slightly emarginate in middle, as wide as long in midline, with lateral flaps distinctly reflexed ventrad with lateral angles angularly rounded; anal opening at about basal ¼.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC0FF83FF16FC48A48DFBD2" pageId="55" pageNumber="56" type="biology_ecology">
|
||
<paragraph id="8BC836A6FFC0FF83FF16FC48A61AFC6C" blockId="55.[189,284,924,950]" box="[189,284,924,950]" pageId="55" pageNumber="56">
|
||
<heading id="D08081CAFFC0FF83FF16FC48A61AFC6C" bold="true" box="[189,284,924,950]" fontSize="11" level="3" pageId="55" pageNumber="56" reason="3">
|
||
<emphasis id="B903EAB4FFC0FF83FF16FC48A61AFC6C" bold="true" box="[189,284,924,950]" pageId="55" pageNumber="56">Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC0FF83FF16FC1EA48DFBD2" blockId="55.[189,1398,970,1032]" pageId="55" pageNumber="56">
|
||
<taxonomicName id="4C774D25FFC0FF83FF16FC1EA6FCFC3E" authority="Constant & Pham, 2024" authorityName="Constant & Pham" authorityYear="2024" box="[189,506,970,996]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="quangninhensis" status="sp. nov.">
|
||
<emphasis id="B903EAB4FFC0FF83FF16FC1EA6FCFC3E" box="[189,506,970,996]" italics="true" pageId="55" pageNumber="56">Pusulissus quangninhensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23057CFFFC0FF83FDA8FC1FA559FC3F" box="[515,607,971,997]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was collected in August on lower vegetation and bushes, in moist evergreen tropical forest at about
|
||
<quantity id="4C8F9B43FFC0FF83FDE2FC39A595FBD2" box="[585,659,1005,1032]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="55" pageNumber="56" unit="m" value="550.0">550 m</quantity>
|
||
in altitude (
|
||
<figureCitation id="134C2A23FFC0FF83FC8AFC39A47AFBD2" box="[801,892,1005,1032]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="55" pageNumber="56">Fig. 1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC0FF83FF16FBFBA37AFBA2" box="[189,1148,1071,1144]" pageId="55" pageNumber="56" type="distribution">
|
||
<paragraph id="8BC836A6FFC0FF83FF16FBFBA650FB93" blockId="55.[189,342,1071,1097]" box="[189,342,1071,1097]" pageId="55" pageNumber="56">
|
||
<heading id="D08081CAFFC0FF83FF16FBFBA650FB93" bold="true" box="[189,342,1071,1097]" fontSize="11" level="3" pageId="55" pageNumber="56" reason="3">
|
||
<emphasis id="B903EAB4FFC0FF83FF16FBFBA650FB93" bold="true" box="[189,342,1071,1097]" pageId="55" pageNumber="56">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC0FF83FF16FB89A37AFBA2" blockId="55.[189,1148,1117,1144]" box="[189,1148,1117,1144]" pageId="55" pageNumber="56">
|
||
<collectingCountry id="F3607636FFC0FF83FF16FB89A623FBAD" box="[189,293,1117,1143]" name="Vietnam" pageId="55" pageNumber="56">Vietnam</collectingCountry>
|
||
:
|
||
<collectingRegion id="49B3F844FFC0FF83FE9BFB8AA534FBAD" box="[304,562,1117,1144]" country="Vietnam" name="Quang Ninh" pageId="55" pageNumber="56">Quang Ninh Province</collectingRegion>
|
||
, Dong Son-Ky Thuong Nature Reserve (
|
||
<figureCitation id="134C2A23FFC0FF83FBBAFB89A368FBA2" box="[1041,1134,1117,1144]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="55" pageNumber="56">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |