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<document id="0F9056E4CC58A80CB1874301389D8CE5" ID-CLB-Dataset="7931" ID-DOI="10.11646/zootaxa.4878.3.2" ID-GBIF-Dataset="288c6fc9-269b-468f-8811-d79d4350f6e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425132" ID-ZooBank="4F59F5B2-CFDB-4819-AF83-0EF232328F8D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610039812786" checkinUser="plazi" docAuthor="Gil, Marta, Ramil, Fran &amp; Agís, José Ansín" docDate="2020" docId="03DE879D1C0A336F46B2C654A1446C67" docLanguage="en" docName="zootaxa.4878.3.2.pdf" docOrigin="Zootaxa 4878 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemertesia caboverdensis Gil &amp; Ramil &amp; Agís 2020, n. sp." docType="treatment" docVersion="10" lastPageNumber="441" masterDocId="FFE7FFE51C1133724625C569A205683E" masterDocTitle="Hydroids (Cnidaria, Hydrozoa) from Mauritanian Coral Mounds" masterLastPageNumber="466" masterPageNumber="412" pageNumber="439" updateTime="1698923825618" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AF68D6A652D6C75263C6B053D508F8AB">Hydroids (Cnidaria, Hydrozoa) from Mauritanian Coral Mounds</mods:title>
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<mods:namePart id="650B4BD346D4F223ADE3879622910F4C">Gil, Marta</mods:namePart>
<mods:affiliation id="9CCC4E8085074F0967BFF94549BC57FE">Departamento de Ecoloxía e Bioloxía Animal, Facultade de Ciencias do Mar e Centro de Investigación Mariña (CIM-UVigo), Universidade de Vigo. Campus Lagoas-Marcosende. 36310 Vigo. Spain. &amp; Instituto Español de Oceanografía, Centro Oceanográfico de Vigo. Subida Radio Faro 50. 36390 Vigo. Spain.</mods:affiliation>
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<mods:namePart id="61FC7EE8BA6A76BB02B7BD80807D7166">Ramil, Fran</mods:namePart>
<mods:affiliation id="338989D7ED913751290F670B86C3F2D6">Departamento de Ecoloxía e Bioloxía Animal, Facultade de Ciencias do Mar e Centro de Investigación Mariña (CIM-UVigo), Universidade de Vigo. Campus Lagoas-Marcosende. 36310 Vigo. Spain. &amp; framil @ uvigo. es</mods:affiliation>
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<mods:namePart id="B59B02386BB81DC19BD63B4C0A673CB6">Agís, José Ansín</mods:namePart>
<mods:affiliation id="3D85624A8E511569253C23926E86DC9A">Departamento de Ecoloxía e Bioloxía Animal, Facultade de Ciencias do Mar e Centro de Investigación Mariña (CIM-UVigo), Universidade de Vigo. Campus Lagoas-Marcosende. 36310 Vigo. Spain. &amp; ansinjose @ gmail. com</mods:affiliation>
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<mods:title id="1640F119EAC6FE9C0B16AB078B9C7260">Zootaxa</mods:title>
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<mods:date id="C2A16BC7FB7FAA7E0359DCBFE85C7E07">2020</mods:date>
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<mods:number id="38E27FD9FD170EDF4DCF9C0E3CE3201F">2020-11-16</mods:number>
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<mods:number id="4525ACDFF2C17B226F087252838987D5">4878</mods:number>
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<treatment id="03DE879D1C0A336F46B2C654A1446C67" ID-DOI="http://doi.org/10.5281/zenodo.4426156" ID-GBIF-Taxon="180323462" ID-Zenodo-Dep="4426156" LSID="urn:lsid:plazi:treatment:03DE879D1C0A336F46B2C654A1446C67" httpUri="http://treatment.plazi.org/id/03DE879D1C0A336F46B2C654A1446C67" lastPageId="29" lastPageNumber="441" pageId="27" pageNumber="439">
<subSubSection id="C36D65001C0A336946B2C654A01E6B66" box="[151,539,829,856]" pageId="27" pageNumber="439" type="nomenclature">
<paragraph id="8BC8368B1C0A336946B2C654A01E6B66" blockId="27.[151,539,829,892]" box="[151,539,829,856]" pageId="27" pageNumber="439">
<heading id="D08081E71C0A336946B2C654A01E6B66" bold="true" box="[151,539,829,856]" fontSize="11" level="1" pageId="27" pageNumber="439" reason="1">
<emphasis id="B903EA991C0A336946B2C654A01E6B66" bold="true" box="[151,539,829,856]" pageId="27" pageNumber="439">
<taxonomicName id="4C774D081C0A336946B2C654A3D56B66" ID-CoL="84BD5" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[151,464,829,856]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="439" phylum="Cnidaria" rank="species" species="caboverdensis" status="sp. nov.">
<emphasis id="B903EA991C0A336946B2C654A3D56B66" bold="true" box="[151,464,829,856]" italics="true" pageId="27" pageNumber="439">Nemertesia caboverdensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057E21C0A336947FDC657A01E6B66" box="[472,539,830,856]" pageId="27" pageNumber="439" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336946B2C608A3486B42" box="[151,333,865,892]" pageId="27" pageNumber="439" type="description">
<paragraph id="8BC8368B1C0A336946B2C608A3486B42" blockId="27.[151,539,829,892]" box="[151,333,865,892]" pageId="27" pageNumber="439">
(
<figureCitation id="134C2A0E1C0A33694685C608A2E16B42" box="[160,228,865,892]" captionStart="FIGURE 5" captionStartId="28.[151,250,1979,2003]" captionTargetBox="[186,1399,193,1946]" captionTargetId="figure-17@28.[182,1405,181,1955]" captionTargetPageId="28" captionText="FIGURE 5. AE, Nemertesia caboverdensis n. sp., holotype (stn 6.078; A), colony. B, detail of colony, frontal view. C, hydrocladium, lateral view. D, gonotheca, frontal view. E, branched hydrocladium. Scales: a, 1 cm; be, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4425142" httpUri="https://zenodo.org/record/4425142/files/figure.png" pageId="27" pageNumber="439">Fig. 5</figureCitation>
;
<tableCitation id="C6F503301C0A336946D5C608A3416B42" box="[240,324,865,892]" captionStart="TABLE 2" captionStartId="29.[151,239,1160,1184]" captionTargetPageId="29" captionText="TABLE 2. Measurements of Nemertesia caboverdensis n. sp. in µm." httpUri="http://table.plazi.org/id/DF0866031C0C336F46B2C1E1A17D6C9F" pageId="27" pageNumber="439" tableUuid="DF0866031C0C336F46B2C1E1A17D6C9F">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336946B2C6CEA3566BE2" pageId="27" pageNumber="439" type="reference_group">
<paragraph id="8BC8368B1C0A336946B2C6CEA3566BE2" blockId="27.[151,1437,934,988]" pageId="27" pageNumber="439">
<treatmentCitationGroup id="AB6711A51C0A336946B2C6CEA3566BE2" pageId="27" pageNumber="439">
<treatmentCitation id="0AD6109A1C0A336946B2C6CEA0686B80" author="Ansin Agis, J." box="[151,621,934,958]" page="460" pageId="27" pageNumber="439" year="1998">
<taxonomicName id="4C774D081C0A336946B2C6CEA0686B80" ID-CoL="467LG" authority=": Ansin Agis, 1998: 457 - 460" authorityName="Ansin Agis" authorityPageNumber="457 - 460" authorityYear="1998" box="[151,621,934,958]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="439" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0A336946B2C6CEA34F6B80" box="[151,330,934,958]" italics="true" pageId="27" pageNumber="439">Nemertesia belini</emphasis>
:
<bibRefCitation id="EFE64B7A1C0A33694773C6CFA0686B80" author="Ansin Agis, J." box="[342,621,934,958]" pageId="27" pageNumber="439" refId="ref36934" refString="Ansin Agis, J. (1998) Plumularioidea (Cnidaria, Hydrozoa) recolectados por las expediciones holandesas &quot; CANCAP &quot; en el Atlantico nor-oriental. Tesis Doctoral, Universidade de Vigo, Vigo, Galicia, 675 pp." type="book" year="1998">Ansín Agís, 1998: 457460</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; figs. 6263;
<treatmentCitation id="0AD6109A1C0A336944D4C6CFA6386B80" author="Ansin Agis, J. &amp; Ramil, F. &amp; Vervoort, W." box="[753,1085,934,958]" page="200" pageId="27" pageNumber="439" year="2001">
<bibRefCitation id="EFE64B7A1C0A336944D4C6CFA6386B80" author="Ansin Agis, J. &amp; Ramil, F. &amp; Vervoort, W." box="[753,1085,934,958]" pageId="27" pageNumber="439" pagination="1 - 268" refId="ref36975" refString="Ansin Agis, J., Ramil, F. &amp; Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische verhandelingen, Leiden, 333, 1 - 268." type="journal article" year="2001">
Ansín Agís
<emphasis id="B903EA991C0A33694543C6CEA1946B80" box="[870,913,934,958]" italics="true" pageId="27" pageNumber="439">et al</emphasis>
., 2001: 200204
</bibRefCitation>
</treatmentCitation>
, figs. 7879 (not
<taxonomicName id="4C774D081C0A336942CFC6CEA3426BE2" ID-CoL="467LG" authority="Bedot, 1916" authorityName="Bedot" authorityYear="1916" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="439" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0A336942CFC6CEA7986B80" box="[1258,1437,934,958]" italics="true" pageId="27" pageNumber="439">Nemertesia belini</emphasis>
<bibRefCitation id="EFE64B7A1C0A336946E2C6ADA3426BE2" author="Bedot, M." box="[199,327,964,988]" pageId="27" pageNumber="439" pagination="637 - 648" refId="ref37282" refString="Bedot, M. (1916) Sur le genre Kirchenpaueria. Revue Suisse de Zoologie, 24 (11), 637 - 648." type="journal article" year="1916">Bedot, 1916</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336946B2C163A0046D1F" pageId="27" pageNumber="439" type="materials_examined">
<paragraph id="8BC8368B1C0A336946B2C163A6FA6C77" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
<emphasis id="B903EA991C0A336946B2C163A3846C1A" bold="true" box="[151,385,1034,1060]" pageId="27" pageNumber="439">
<collectingRegion id="49B3F8691C0A336946B2C163A2FA6C1A" box="[151,255,1034,1060]" pageId="27" pageNumber="439">Material</collectingRegion>
examined.
</emphasis>
<materialsCitation id="3B1F3CD61C0A336947ACC163A6FF6C77" ID-GBIF-Occurrence="3044195330" collectingDate="1982-06-13" collectionCode="R" country="Cape Verde" elevation="110" latitude="15.9" location="Islands." longLatPrecision="1284" longitude="-23.1" pageId="27" pageNumber="439" specimenCount="1" stateProvince="Islands." typeStatus="paratype">
<emphasis id="B903EA991C0A336947ACC163A07C6C1B" bold="true" box="[393,633,1034,1061]" pageId="27" pageNumber="439">
<collectingCountry id="F360761B1C0A336947ACC163A0106C1B" box="[393,533,1034,1061]" name="Cape Verde" pageId="27" pageNumber="439">Cape Verde</collectingCountry>
<collectingRegion id="49B3F8691C0A33694439C162A07C6C1B" box="[540,633,1035,1061]" pageId="27" pageNumber="439">Islands.</collectingRegion>
</emphasis>
CANCAP, stn 6.072,
<geoCoordinate id="EE43504C1C0A33694551C162A1D06C18" box="[884,981,1035,1062]" degrees="15" direction="north" minutes="54" orientation="latitude" pageId="27" pageNumber="439" precision="925" value="15.9">15º54´N</geoCoordinate>
,
<geoCoordinate id="EE43504C1C0A336945C4C162A6406C1B" box="[993,1093,1035,1061]" degrees="23" direction="west" minutes="06" orientation="longitude" pageId="27" pageNumber="439" precision="925" value="-23.1">23º06´W</geoCoordinate>
,
<quantity id="4C8F9B6E1C0A33694277C162A69F6C1B" box="[1106,1178,1035,1061]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="439" unit="m" value="110.0">
<elevation id="005AD1B81C0A33694277C162A69F6C1B" box="[1106,1178,1035,1061]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="439" unit="m" value="110.0">110 m</elevation>
</quantity>
,
<date id="FFC9104B1C0A33694282C162A72A6C1B" box="[1191,1327,1035,1061]" pageId="27" pageNumber="439" value="1982-06-13">
<collectingDate id="EF8DE9A31C0A33694282C162A72A6C1B" box="[1191,1327,1035,1061]" pageId="27" pageNumber="439" value="1982-06-13">13-VI-1982</collectingDate>
</date>
: six colonies
<quantity id="4C8F9B6E1C0A336946EEC146A3576C77" box="[203,338,1071,1097]" metricMagnitude="-2" metricUnit="m" metricValue="9.85" metricValueMax="17.1" metricValueMin="2.6" pageId="27" pageNumber="439" unit="mm" value="98.5" valueMax="171.0" valueMin="26.0">26171 mm</quantity>
high, without gonothecae;
<typeStatus id="54CC88291C0A336944A1C146A0EA6C77" box="[644,751,1071,1097]" pageId="27" pageNumber="439" type="paratype">paratypes</typeStatus>
(RMNH-Coel. 28754; LZM-UV slide
<collectionCode id="ED66AE4E1C0A33694286C146A6BC6C77" box="[1187,1209,1071,1097]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="439">R</collectionCode>
. 332)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC8368B1C0A336946E2C13AA39D6CAC" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
CANCAP, stn 6.074,
<geoCoordinate id="EE43504C1C0A3369479CC13AA01F6C50" box="[441,538,1107,1134]" degrees="15" direction="north" minutes="55" orientation="latitude" pageId="27" pageNumber="439" precision="925" value="15.916667">15º55´N</geoCoordinate>
,
<geoCoordinate id="EE43504C1C0A33694403C13AA08F6C53" box="[550,650,1107,1133]" degrees="23" direction="west" minutes="04" orientation="longitude" pageId="27" pageNumber="439" precision="925" value="-23.066668">23º04´W</geoCoordinate>
,
<quantity id="4C8F9B6E1C0A336944B3C13AA0D46C53" box="[662,721,1107,1133]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="27" pageNumber="439" unit="m" value="91.0">91 m</quantity>
,
<date id="FFC9104B1C0A336944F8C13AA1636C53" box="[733,870,1107,1133]" pageId="27" pageNumber="439" value="1982-06-13">13-VI-1982</date>
: one colony
<quantity id="4C8F9B6E1C0A336945D1C13AA6466C53" box="[1012,1091,1107,1134]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="27" pageNumber="439" unit="mm" value="53.0">53 mm</quantity>
high, no gonothecae;
<typeStatus id="54CC88291C0A33694319C13AA7996C53" box="[1340,1436,1107,1133]" pageId="27" pageNumber="439" type="paratype">paratype</typeStatus>
(RMNH-Coel. 28762).
</paragraph>
<paragraph id="8BC8368B1C0A336946E2C1F2A3CA6CE7" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
CANCAP, stn 6.076,
<geoCoordinate id="EE43504C1C0A33694792C1F2A01D6C88" box="[439,536,1179,1206]" degrees="15" direction="north" minutes="55" orientation="latitude" pageId="27" pageNumber="439" precision="925" value="15.916667">15º55´N</geoCoordinate>
,
<geoCoordinate id="EE43504C1C0A33694406C1F2A08D6C88" box="[547,648,1179,1206]" degrees="23" direction="west" minutes="05" orientation="longitude" pageId="27" pageNumber="439" precision="925" value="-23.083334">23º05´W</geoCoordinate>
,
<quantity id="4C8F9B6E1C0A336944B7C1F2A0C86C8B" box="[658,717,1179,1205]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="27" pageNumber="439" unit="m" value="92.0">92 m</quantity>
,
<date id="FFC9104B1C0A336944FDC1F2A15C6C8B" box="[728,857,1179,1205]" pageId="27" pageNumber="439" value="1982-06-13">13-vi-1982</date>
: one colony
<quantity id="4C8F9B6E1C0A336945C0C1F2A6366C8B" box="[997,1075,1179,1205]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="27" pageNumber="439" unit="mm" value="69.0">69 mm</quantity>
high, without gonothecae;
<typeStatus id="54CC88291C0A33694346C1F2A2C26CE7" pageId="27" pageNumber="439" type="paratype">paratype</typeStatus>
(RMNH-Coel. 29101).
</paragraph>
<paragraph id="8BC8368B1C0A336946E2C18AA0046D1F" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
CANCAP, stn 6.078,
<geoCoordinate id="EE43504C1C0A33694798C18AA01B6CC0" box="[445,542,1251,1278]" degrees="15" direction="north" minutes="55" orientation="latitude" pageId="27" pageNumber="439" precision="925" value="15.916667">15º55´N</geoCoordinate>
,
<geoCoordinate id="EE43504C1C0A3369440EC18AA0956CC3" box="[555,656,1251,1277]" degrees="23" direction="west" minutes="06" orientation="longitude" pageId="27" pageNumber="439" precision="925" value="-23.1">23º06´W</geoCoordinate>
,
<quantity id="4C8F9B6E1C0A336944B8C18AA1256CC3" box="[669,800,1251,1278]" metricMagnitude="2" metricUnit="m" metricValue="1.875" metricValueMax="1.9" metricValueMin="1.85" pageId="27" pageNumber="439" unit="m" value="187.5" valueMax="190.0" valueMin="185.0">185190 m</quantity>
,
<date id="FFC9104B1C0A33694508C18AA1AB6CC3" box="[813,942,1251,1277]" pageId="27" pageNumber="439" value="1982-06-13">13-vi-1982</date>
: one colony
<quantity id="4C8F9B6E1C0A33694265C18AA6946CC3" box="[1088,1169,1251,1277]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="27" pageNumber="439" unit="mm" value="92.0">92 mm</quantity>
high, with gonothecae;
<typeStatus id="54CC88291C0A336946B2C06EA2FC6D1F" box="[151,249,1287,1313]" pageId="27" pageNumber="439" type="holotype">holotype</typeStatus>
(RMNH-Coel. 28788).
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336946E2C042A3B56D57" pageId="27" pageNumber="439" type="etymology">
<paragraph id="8BC8368B1C0A336946E2C042A3B56D57" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
<emphasis id="B903EA991C0A336946E2C042A34B6D7B" bold="true" box="[199,334,1323,1349]" pageId="27" pageNumber="439">Etymology.</emphasis>
The specific name,
<taxonomicName id="4C774D081C0A3369440BC042A0C86D7B" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[558,717,1323,1349]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="439" phylum="Cnidaria" rank="species" species="caboverdensis">
<emphasis id="B903EA991C0A3369440BC042A0C86D7B" box="[558,717,1323,1349]" italics="true" pageId="27" pageNumber="439">caboverdensis</emphasis>
</taxonomicName>
, refers to the locality from which this new species was obtained:
<collectingCountry id="F360761B1C0A336946B2C026A31D6D57" box="[151,280,1359,1385]" name="Cape Verde" pageId="27" pageNumber="439">Cape Verde</collectingCountry>
Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336946E2C01BA6736DB3" box="[199,1142,1394,1421]" pageId="27" pageNumber="439" type="biology_ecology">
<paragraph id="8BC8368B1C0A336946E2C01BA6736DB3" blockId="27.[151,1437,1034,2033]" box="[199,1142,1394,1421]" pageId="27" pageNumber="439">
<emphasis id="B903EA991C0A336946E2C01BA3226DB2" bold="true" box="[199,295,1394,1420]" pageId="27" pageNumber="439">Biology.</emphasis>
Gonothecae were found in June (
<bibRefCitation id="EFE64B7A1C0A336944BAC01AA1656DB3" author="Ansin Agis, J." box="[671,864,1395,1421]" pageId="27" pageNumber="439" refId="ref36934" refString="Ansin Agis, J. (1998) Plumularioidea (Cnidaria, Hydrozoa) recolectados por las expediciones holandesas &quot; CANCAP &quot; en el Atlantico nor-oriental. Tesis Doctoral, Universidade de Vigo, Vigo, Galicia, 675 pp." type="book" year="1998">Ansín Agís 1998</bibRefCitation>
;
<bibRefCitation id="EFE64B7A1C0A3369454FC01AA66C6DB3" author="Ansin Agis, J. &amp; Ramil, F. &amp; Vervoort, W." box="[874,1129,1395,1421]" pageId="27" pageNumber="439" pagination="1 - 268" refId="ref36975" refString="Ansin Agis, J., Ramil, F. &amp; Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische verhandelingen, Leiden, 333, 1 - 268." type="journal article" year="2001">
Ansín Agís
<emphasis id="B903EA991C0A336945CBC01DA6256DB3" box="[1006,1056,1395,1421]" italics="true" pageId="27" pageNumber="439">et al</emphasis>
. 2001
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336946E2C0FFA0216DEB" pageId="27" pageNumber="439" type="distribution">
<paragraph id="8BC8368B1C0A336946E2C0FFA0216DEB" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
<emphasis id="B903EA991C0A336946E2C0FFA3646D8E" bold="true" box="[199,353,1430,1456]" pageId="27" pageNumber="439">Distribution.</emphasis>
Only recorded in four localities of
<collectingCountry id="F360761B1C0A336944D4C0FEA1776D8F" box="[753,882,1431,1457]" name="Cape Verde" pageId="27" pageNumber="439">Cape Verde</collectingCountry>
Islands, between depths of 91 and
<quantity id="4C8F9B6E1C0A33694324C0FEA74C6D8F" box="[1281,1353,1431,1457]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="439" unit="m" value="190.0">190 m</quantity>
(
<bibRefCitation id="EFE64B7A1C0A3369437DC0FEA30B6DEB" author="Ansin Agis, J." pageId="27" pageNumber="439" refId="ref36934" refString="Ansin Agis, J. (1998) Plumularioidea (Cnidaria, Hydrozoa) recolectados por las expediciones holandesas &quot; CANCAP &quot; en el Atlantico nor-oriental. Tesis Doctoral, Universidade de Vigo, Vigo, Galicia, 675 pp." type="book" year="1998">Ansín Agís 1998</bibRefCitation>
;
<bibRefCitation id="EFE64B7A1C0A3369473DC0D2A01D6DEB" author="Ansin Agis, J. &amp; Ramil, F. &amp; Vervoort, W." box="[280,536,1467,1493]" pageId="27" pageNumber="439" pagination="1 - 268" refId="ref36975" refString="Ansin Agis, J., Ramil, F. &amp; Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische verhandelingen, Leiden, 333, 1 - 268." type="journal article" year="2001">
Ansín Agís
<emphasis id="B903EA991C0A336947B8C0D5A3CB6DEB" box="[413,462,1467,1493]" italics="true" pageId="27" pageNumber="439">et al</emphasis>
. 2001
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0A336F46E2C0B7A1A5695B" lastPageId="29" lastPageNumber="441" pageId="27" pageNumber="439" type="description">
<paragraph id="8BC8368B1C0A336946E2C0B7A67E6EEF" blockId="27.[151,1437,1034,2033]" pageId="27" pageNumber="439">
<emphasis id="B903EA991C0A336946E2C0B7A35C6DC6" bold="true" box="[199,345,1502,1528]" pageId="27" pageNumber="439">Description.</emphasis>
Hydrorhiza tubular, adhering to substrate, giving rise to several monosiphonic, unbranched hydrocauli, divided into internodes by straight nodes, distinctly visible along the entire stem, especially in distal part. Each internode provided with two opposite apophyses in basal part of stem and a verticil of four apophyses in distal part, with decussate disposition; moreover, each axial internode carries several nematothecae distributed under the apophyses. Length of apophyses varies, short basally and long distally; each apophysis with two axillar nematothecae, a mamelon on its upper surface and an unpaired distal nematotheca. Long apophyses may have, after the mamelon, a pair of proximal nematothecae and up to three unpaired distal nematothecae.
</paragraph>
<paragraph id="8BC8368B1C0A336F46E2C3B2A0A168EB" blockId="27.[151,1437,1034,2033]" lastBlockId="29.[151,1437,151,1113]" lastPageId="29" lastPageNumber="441" pageId="27" pageNumber="439">In basal half of stem, the apophyses are short and each hydrocladium starts with a short ahydrothecate internode, provided with a single nematotheca. In distal half, with long apophyses, first internode of hydrocladium is hydrothecate because of the fusion of the first ahydrothecate internode with the apophysis. Hydrocladia composed of succession of hydrothecate internodes with slightly oblique nodes, each with one hydrotheca on distal third, two to four mesial inferior nematothecae, a pair of lateral nematothecae and occasionally a supracalycine nematotheca. Hydrothecae cup-shaped, adcauline wall fully adnate, abcauline wall straight, with the aperture perpendicular to longitudinal axis of hydrocladia; rim smooth.Ahydrothecate internodes occur but distributed without any regularity, bearing one to three nematothecae, usually as the result of node formation in basal part of succeeding hydrothecate internode; in this case, the hydrotheca occupies a central position on the internode. All nematothecae bithalamic and movable, with long and narrow basal chamber.</paragraph>
<caption id="DF0866031C0D336E46B2C2D2A6D06FC9" ID-DOI="http://doi.org/10.5281/zenodo.4425142" ID-Zenodo-Dep="4425142" httpUri="https://zenodo.org/record/4425142/files/figure.png" pageId="28" pageNumber="440" startId="28.[151,250,1979,2003]" targetBox="[186,1399,193,1946]" targetPageId="28">
<paragraph id="8BC8368B1C0D336E46B2C2D2A6D06FC9" blockId="28.[151,1436,1979,2040]" pageId="28" pageNumber="440">
<emphasis id="B903EA991C0D336E46B2C2D2A3166FEA" bold="true" box="[151,275,1979,2004]" pageId="28" pageNumber="440">FIGURE 5.</emphasis>
AE,
<taxonomicName id="4C774D081C0D336E4771C2D5A0586FEA" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[340,605,1980,2004]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="28" pageNumber="440" phylum="Cnidaria" rank="species" species="caboverdensis" status="sp. nov.">
<emphasis id="B903EA991C0D336E4771C2D5A0586FEA" box="[340,605,1980,2004]" italics="true" pageId="28" pageNumber="440">Nemertesia caboverdensis</emphasis>
</taxonomicName>
<emphasis id="B903EA991C0D336E4446C2D5A0996FEA" bold="true" box="[611,668,1980,2004]" pageId="28" pageNumber="440">
<taxonomicNameLabel id="A23057E21C0D336E4446C2D5A0996FEA" box="[611,668,1980,2004]" pageId="28" pageNumber="440" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (stn 6.078; A), colony. B, detail of colony, frontal view. C, hydrocladium, lateral view. D, gonotheca, frontal view. E, branched hydrocladium. Scales: a, 1 cm; be, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC8368B1C0C336F46E2C5B6A1886923" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="441">
Gonothecae inserted on apophyses, ovoid, narrowing basally into a short pedicel. Aperture latero-terminal, broadly oval. Only one
<typeStatus id="54CC88291C0C336F4785C46AA3D56923" box="[416,464,259,285]" pageId="29" pageNumber="441">type</typeStatus>
of gonothecae observed; sex unknown.
</paragraph>
<paragraph id="8BC8368B1C0C336F46E2C44EA1A5695B" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="441">A colony from Stn 6.078 showed two secondary hydrocladia arising from the interior of a hydrotheca or a small apophysis below the hydrotheca of a primary hydrocladium (fig. 5E).</paragraph>
</subSubSection>
<subSubSection id="C36D65001C0C336F46E2C406A1446C67" pageId="29" pageNumber="441" type="discussion">
<paragraph id="8BC8368B1C0C336F46E2C406A74969EF" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="441">
<emphasis id="B903EA991C0C336F46E2C406A33E69B7" bold="true" box="[199,315,367,393]" pageId="29" pageNumber="441">Remarks.</emphasis>
Review of the
<typeStatus id="54CC88291C0C336F47C5C406A01569B7" box="[480,528,367,393]" pageId="29" pageNumber="441">type</typeStatus>
material of
<taxonomicName id="4C774D081C0C336F44B1C406A0F169B7" authorityName="Bedot" authorityYear="1916" box="[660,756,367,393]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F44B1C406A0F169B7" box="[660,756,367,393]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
and study of the samples identified as
<taxonomicName id="4C774D081C0C336F42B9C406A6FE69B7" authorityName="Bedot" authorityYear="1916" box="[1180,1275,367,393]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F42B9C406A6FE69B7" box="[1180,1275,367,393]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE64B7A1C0C336F4305C406A2E76993" author="Ansin Agis, J." pageId="29" pageNumber="441" refId="ref36934" refString="Ansin Agis, J. (1998) Plumularioidea (Cnidaria, Hydrozoa) recolectados por las expediciones holandesas &quot; CANCAP &quot; en el Atlantico nor-oriental. Tesis Doctoral, Universidade de Vigo, Vigo, Galicia, 675 pp." type="book" year="1998">Ansín Agís (1998)</bibRefCitation>
and
<bibRefCitation id="EFE64B7A1C0C336F4731C4FAA0186993" author="Ansin Agis, J. &amp; Ramil, F. &amp; Vervoort, W." box="[276,541,403,429]" pageId="29" pageNumber="441" pagination="1 - 268" refId="ref36975" refString="Ansin Agis, J., Ramil, F. &amp; Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische verhandelingen, Leiden, 333, 1 - 268." type="journal article" year="2001">
Ansín Agís
<emphasis id="B903EA991C0C336F47B3C4FAA3C86993" box="[406,461,403,429]" italics="true" pageId="29" pageNumber="441">et al.</emphasis>
(2001)
</bibRefCitation>
led us to the conclusion that they are different species.Although this species was not collected in our surveys, for comparative purposes, we have also included a description of this material.
</paragraph>
<paragraph id="8BC8368B1C0C336F46E2C4B2A6796A5F" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="441">
Comparison of these colonies with the
<typeStatus id="54CC88291C0C336F44ABC4B2A0BB69CB" box="[654,702,475,501]" pageId="29" pageNumber="441">type</typeStatus>
material of
<taxonomicName id="4C774D081C0C336F4575C4B2A1B669CB" authorityName="Bedot" authorityYear="1916" box="[848,947,475,501]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F4575C4B2A1B669CB" box="[848,947,475,501]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
showed a fair similarity in morphological traits, both with respect to trophosome and gonosome. Moreover, this material also shows wide variations related to the length of apophyses, segmentation of the stem and hydrocladia, and number of nematothecae per hydrocladial internode. This variability is consistent with that described by
<bibRefCitation id="EFE64B7A1C0C336F4574C72EA1E36A5C" author="Bedot, M." box="[849,998,583,610]" pageId="29" pageNumber="441" pagination="637 - 648" refId="ref37282" refString="Bedot, M. (1916) Sur le genre Kirchenpaueria. Revue Suisse de Zoologie, 24 (11), 637 - 648." type="journal article" year="1916">Bedot (1916)</bibRefCitation>
for
<taxonomicName id="4C774D081C0C336F4231C72EA6706A5F" authorityName="Bedot" authorityYear="1916" box="[1044,1141,583,609]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F4231C72EA6706A5F" box="[1044,1141,583,609]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC8368B1C0C336F46E2C702A0CF6B9B" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="441">
Nevertheless, we found differences that, in our opinion, justify its specific separation. In
<taxonomicName id="4C774D081C0C336F4299C702A7786ABB" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[1212,1405,619,645]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="caboverdensis" status="sp. nov.">
<emphasis id="B903EA991C0C336F4299C702A7786ABB" box="[1212,1405,619,645]" italics="true" pageId="29" pageNumber="441">N. caboverdensis</emphasis>
</taxonomicName>
<emphasis id="B903EA991C0C336F43A3C702A2BD6A97" bold="true" pageId="29" pageNumber="441">
<taxonomicNameLabel id="A23057E21C0C336F43A3C702A2BD6A97" pageId="29" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, hydrothecae are disposed in distal third of internodes and its aperture is almost perpendicular to the longitudinal axis of hydrocladia, whereas in
<taxonomicName id="4C774D081C0C336F4402C7DAA08D6AF3" authorityName="Bedot" authorityYear="1916" box="[551,648,691,717]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F4402C7DAA08D6AF3" box="[551,648,691,717]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
, hydrothecae are located in basal half of internodes and its aperture is clearly tilted to abcauline side (adcauline wall longer than abcauline one) (fig. 4F, G). The number of supracalycine nematothecae is also different: one to three in
<taxonomicName id="4C774D081C0C336F44BFC792A0FE6B2B" authorityName="Bedot" authorityYear="1916" box="[666,763,763,789]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F44BFC792A0FE6B2B" box="[666,763,763,789]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
, two being the more common number (
<bibRefCitation id="EFE64B7A1C0C336F4296C792A7436B28" author="Bedot, M." box="[1203,1350,763,790]" pageId="29" pageNumber="441" pagination="1 - 73" refId="ref37386" refString="Bedot, M. (1921 b) Hydroides provenant des campagnes des yachts Hirondelle et Princesse-Alice (1887 - 1912). I. Plumulariidae. Resultats des Campagnes scientifiques accomplies sur son yacht par le Prince Albert Ier de Monaco, 60, 1 - 73." type="journal article" year="1921">Bedot 1921b</bibRefCitation>
) versus zero to one in the
<taxonomicName id="4C774D081C0C336F4740C676A0206B07" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[357,549,799,825]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="caboverdensis" status="sp. nov.">
<emphasis id="B903EA991C0C336F4740C676A0206B07" box="[357,549,799,825]" italics="true" pageId="29" pageNumber="441">N. caboverdensis</emphasis>
</taxonomicName>
<emphasis id="B903EA991C0C336F4408C676A0686B07" bold="true" box="[557,621,799,825]" pageId="29" pageNumber="441">
<taxonomicNameLabel id="A23057E21C0C336F4408C676A0686B07" box="[557,621,799,825]" pageId="29" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, one being the most common; moreover, in
<taxonomicName id="4C774D081C0C336F4246C676A7266B07" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[1123,1315,799,825]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="caboverdensis" status="sp. nov.">
<emphasis id="B903EA991C0C336F4246C676A7266B07" box="[1123,1315,799,825]" italics="true" pageId="29" pageNumber="441">N. caboverdensis</emphasis>
</taxonomicName>
<emphasis id="B903EA991C0C336F430EC676A76E6B07" bold="true" box="[1323,1387,799,825]" pageId="29" pageNumber="441">
<taxonomicNameLabel id="A23057E21C0C336F430EC676A76E6B07" box="[1323,1387,799,825]" pageId="29" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the nematothecae are longer (see
<tableCitation id="C6F503301C0C336F47C4C62AA0336B63" box="[481,566,835,861]" captionStart="TABLE 2" captionStartId="29.[151,239,1160,1184]" captionTargetPageId="29" captionText="TABLE 2. Measurements of Nemertesia caboverdensis n. sp. in µm." httpUri="http://table.plazi.org/id/DF0866031C0C336F46B2C1E1A17D6C9F" pageId="29" pageNumber="441" tableUuid="DF0866031C0C336F46B2C1E1A17D6C9F">Table 2</tableCitation>
). In addition,
<taxonomicName id="4C774D081C0C336F44F4C62AA1956B63" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[721,912,835,861]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="caboverdensis" status="sp. nov.">
<emphasis id="B903EA991C0C336F44F4C62AA1956B63" box="[721,912,835,861]" italics="true" pageId="29" pageNumber="441">N. caboverdensis</emphasis>
</taxonomicName>
<emphasis id="B903EA991C0C336F45B2C62AA1D06B63" bold="true" box="[919,981,835,861]" pageId="29" pageNumber="441">
<taxonomicNameLabel id="A23057E21C0C336F45B2C62AA1D06B63" box="[919,981,835,861]" pageId="29" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has ahydrothecate internodes irregularly distributed along hydrocladia, whereas in
<taxonomicName id="4C774D081C0C336F444AC60EA0D56BBF" authorityName="Bedot" authorityYear="1916" box="[623,720,871,897]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F444AC60EA0D56BBF" box="[623,720,871,897]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
, hydrocladia are composed of a succession of hydrothecate and ahydrothecate internodes as standard morphology.
</paragraph>
<paragraph id="8BC8368B1C0C336F46E2C6C6A0586C0B" blockId="29.[151,1437,151,1113]" pageId="29" pageNumber="441">
Material from CANCAP stn 5010, collected in the Azores area and reported by
<bibRefCitation id="EFE64B7A1C0C336F4274C6C6A7276BF7" author="Ansin Agis, J." box="[1105,1314,943,969]" pageId="29" pageNumber="441" refId="ref36934" refString="Ansin Agis, J. (1998) Plumularioidea (Cnidaria, Hydrozoa) recolectados por las expediciones holandesas &quot; CANCAP &quot; en el Atlantico nor-oriental. Tesis Doctoral, Universidade de Vigo, Vigo, Galicia, 675 pp." type="book" year="1998">Ansín Agís (1998)</bibRefCitation>
and
<bibRefCitation id="EFE64B7A1C0C336F437CC6C6A3596BD3" author="Ansin Agis, J. &amp; Ramil, F. &amp; Vervoort, W." pageId="29" pageNumber="441" pagination="1 - 268" refId="ref36975" refString="Ansin Agis, J., Ramil, F. &amp; Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische verhandelingen, Leiden, 333, 1 - 268." type="journal article" year="2001">
Ansín Agís
<emphasis id="B903EA991C0C336F46F6C6BAA30E6BD3" box="[211,267,979,1005]" italics="true" pageId="29" pageNumber="441">et al.</emphasis>
(2001)
</bibRefCitation>
within
<taxonomicName id="4C774D081C0C336F4794C6BAA0176BD3" authorityName="Bedot" authorityYear="1916" box="[433,530,979,1005]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="belini">
<emphasis id="B903EA991C0C336F4794C6BAA0176BD3" box="[433,530,979,1005]" italics="true" pageId="29" pageNumber="441">N. belini</emphasis>
</taxonomicName>
, must be reviewed before its definitive identification. Some features of this colony, such as the hydrothecal aperture tilted in abcauline direction and the length of its nematothecae clearly smaller, suggest that could be a different species.
</paragraph>
<paragraph id="8BC8368B1C0C336F46E2C156A1446C67" blockId="29.[151,1437,151,1113]" box="[199,833,1087,1113]" pageId="29" pageNumber="441">
Differences with
<taxonomicName id="4C774D081C0C336F47ACC156A00B6C67" authority="Gil &amp; Ramil &amp; Agís, 2020" authorityName="Gil &amp; Ramil &amp; Agís" authorityYear="2020" box="[393,526,1087,1113]" class="Hydrozoa" family="Plumulariidae" genus="Nemertesia" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="441" phylum="Cnidaria" rank="species" species="freiwaldi" status="sp. nov.">
<emphasis id="B903EA991C0C336F47ACC156A00B6C67" box="[393,526,1087,1113]" italics="true" pageId="29" pageNumber="441">N. freiwaldi</emphasis>
</taxonomicName>
<emphasis id="B903EA991C0C336F4430C156A0566C67" bold="true" box="[533,595,1087,1113]" pageId="29" pageNumber="441">
<taxonomicNameLabel id="A23057E21C0C336F4430C156A0566C67" box="[533,595,1087,1113]" pageId="29" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are discussed below.
</paragraph>
</subSubSection>
</treatment>
</document>