441 lines
52 KiB
XML
441 lines
52 KiB
XML
<document id="1FE165DD38BB831496DFAECAB5F937A0" ID-CLB-Dataset="8579" ID-DOI="10.11646/zootaxa.4850.1.1" ID-GBIF-Dataset="686337a2-1528-43d0-ae32-68ca023d8df3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407072" ID-ZooBank="18AA0411-0E54-4922-84C7-608EAC68D281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609428297648" checkinUser="plazi" docAuthor="Guéorguiev, Borislav & Liang, Hongbin" docDate="2020" docId="03BC5E5B2948FFB5FF4BF972ED88FD54" docLanguage="en" docName="zootaxa.4850.1.1.pdf" docOrigin="Zootaxa 4850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudoodes Guéorguiev & Liang 2020, gen. n." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FF8526232940FFBFFFDCFFB4EC2EFFFD" masterDocTitle="Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="9" updateTime="1698852686469" updateUser="ExternalLinkService">
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<mods:mods id="9CAC63B048850613D3E53AEEF8556818" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FBC3C3BBF254500373D754E8457ACD42">
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<mods:title id="D0D8F1496B440567C3E2FC41CE29E576">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
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<mods:name id="52D3683DDF2EB1513642DE629F614669" type="personal">
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<mods:role id="EE97F8644C640AAE49A3660A7556E656">
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<mods:roleTerm id="1BE3051F888A9DC78785F72DA8367D9C">Author</mods:roleTerm>
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<mods:namePart id="6ED76A0F952DA58872B78A17E8E6B381">Guéorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier id="E7EA43D2E3E800627A6392F55139AD16" type="ORCID">0000-0002-8532-0657</mods:nameIdentifier>
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<mods:nameIdentifier id="3354FA168B3E14D66951E52A805CC15D" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:name id="E03E8E77B67029FC54460C0D19688A3E" type="personal">
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<mods:roleTerm id="FEFA590BDFAD39B27FFE20C417138032">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4CB6847B05F3DF3E136D35152C8649B3">Liang, Hongbin</mods:namePart>
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<mods:nameIdentifier id="769D3BA9089BB514EEF8301FD70CD24D" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
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<mods:nameIdentifier id="9BBCE2E7CCB56ED317DCD3BADC191E9F" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="230133F0A561CEB03EEFCEC76E08F949">Zootaxa</mods:title>
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<mods:part id="97862E7A1273DEF822992CF1D54BF80E">
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<mods:date id="CCDF6BA448D196BA36AE42843C52A6CE">2020</mods:date>
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<mods:number id="EDAC1F5F501E2FFA429A6444F71F305A">2020-09-08</mods:number>
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<mods:number id="075156BF4F243848C654219986B4305F">4850</mods:number>
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<mods:identifier id="F77883D83FDDFEA136EEDDED64B4FFDB" type="DOI">10.11646/zootaxa.4850.1.1</mods:identifier>
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<treatment id="03BC5E5B2948FFB5FF4BF972ED88FD54" ID-DOI="http://doi.org/10.5281/zenodo.4479977" ID-GBIF-Taxon="177139743" ID-Zenodo-Dep="4479977" LSID="urn:lsid:plazi:treatment:03BC5E5B2948FFB5FF4BF972ED88FD54" httpUri="http://treatment.plazi.org/id/03BC5E5B2948FFB5FF4BF972ED88FD54" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="C30FBCC62948FFB7FF4BF972ED51F91C" box="[151,383,1734,1761]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BAAEF4D2948FFB7FF4BF972ED51F91C" blockId="8.[151,383,1734,1761]" box="[151,383,1734,1761]" pageId="8" pageNumber="9">
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<heading id="D0E258212948FFB7FF4BF972ED51F91C" bold="true" box="[151,383,1734,1761]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
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<emphasis id="B961335F2948FFB7FF4BF972ED51F91C" bold="true" box="[151,383,1734,1761]" pageId="8" pageNumber="9">
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<taxonomicName id="4C1594CE2948FFB7FF4BF972ED0BF91C" ID-CoL="8LYNX" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[151,293,1734,1761]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B961335F2948FFB7FF4BF972ED0BF91C" bold="true" box="[151,293,1734,1761]" italics="true" pageId="8" pageNumber="9">Pseudoodes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2528E242948FFB7FEF0F973ED51F91C" box="[300,383,1735,1761]" pageId="8" pageNumber="9" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FBCC62948FFB7FF4BF8BBEF85F8D4" box="[151,939,1807,1833]" pageId="8" pageNumber="9" type="type_taxon">
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<paragraph id="8BAAEF4D2948FFB7FF4BF8BBEF85F8D4" blockId="8.[151,939,1807,1833]" box="[151,939,1807,1833]" pageId="8" pageNumber="9">
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<typeStatus id="54AE51EF2948FFB7FF4BF8BBECE1F8D4" box="[151,207,1807,1833]" pageId="8" pageNumber="9">Type</typeStatus>
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species:
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<taxonomicName id="4C1594CE2948FFB7FEE9F8BBEEB3F8D4" ID-CoL="8LL28" authority="Bates, 1892" authorityName="Bates. Consists" authorityYear="1892" box="[309,669,1807,1833]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cribristernis">
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<emphasis id="B961335F2948FFB7FEE9F8BBEE20F8D4" box="[309,526,1807,1833]" italics="true" pageId="8" pageNumber="9">Oodes cribristernis</emphasis>
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<bibRefCitation id="EF8492BC2948FFB7FDC9F8BBEEB3F8D4" author="Bates, H. W." box="[533,669,1807,1833]" pageId="8" pageNumber="9" pagination="267 - 428" refId="ref59903" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, 32, 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
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</taxonomicName>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FBCC62948FFB6FF4BF8E2ED95FE50" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BAAEF4D2948FFB7FF4BF8E2EFC5F88C" blockId="8.[151,1003,1878,1905]" box="[151,1003,1878,1905]" pageId="8" pageNumber="9">
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<emphasis id="B961335F2948FFB7FF4BF8E2ED24F88D" bold="true" box="[151,266,1878,1904]" pageId="8" pageNumber="9">Diagnosis</emphasis>
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. The species from this genus share the following character states:
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</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BF82BEECCF844" blockId="8.[151,837,1951,2013]" box="[151,738,1951,1977]" pageId="8" pageNumber="9">
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(1)
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<collectingRegion id="49D121AF2948FFB7FF1AF82BED0BF844" box="[198,293,1951,1977]" country="Myanmar" name="Chin" pageId="8" pageNumber="9">Mentum</collectingRegion>
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tooth with apex truncate to emarginate;
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</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BF877EF6BF820" blockId="8.[151,837,1951,2013]" box="[151,837,1987,2013]" pageId="8" pageNumber="9">(2) Submentum with medial and lateral setiferous punctures;</paragraph>
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<paragraph id="8BAAEF4D2949FFB6FF4BFF23EE0FFF4C" blockId="9.[151,1437,151,429]" box="[151,545,151,177]" pageId="9" pageNumber="10">(3) Prosternal process unbordered;</paragraph>
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<paragraph id="8BAAEF4D2949FFB6FF4BFF0FEEF1FF28" blockId="9.[151,1437,151,429]" box="[151,735,187,213]" pageId="9" pageNumber="10">(4) Abdominal ventrite 3 without ambulatory setae;</paragraph>
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<paragraph id="8BAAEF4D2949FFB6FF4BFF6AEF95FF04" blockId="9.[151,1437,151,429]" box="[151,955,222,249]" pageId="9" pageNumber="10">(5) Male protarsomere 3 subtrapezoidal, with sides constricted distally;</paragraph>
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<paragraph id="8BAAEF4D2949FFB6FF4BFEB7E822FEE0" blockId="9.[151,1437,151,429]" box="[151,1036,259,286]" pageId="9" pageNumber="10">(6) Basal bulb of median lobe of aedeagus closed dorsally, with apical aileron;</paragraph>
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<paragraph id="8BAAEF4D2949FFB6FF4BFE93ED17FE98" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="10">(7) Sclerite of internal sac of median lobe very long, spiral, formed by broader proximal part and narrower distal extension;</paragraph>
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<paragraph id="8BAAEF4D2949FFB6FF4BFEDBED95FE50" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="10">
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(8) Spermatheca differentiated to receptaculum and seminal canal (reverse state occurs in several taxa of “
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<taxonomicName id="4C1594CE2949FFB6FF0EFE27ED7BFE50" authorityName="Jedlicka. Consists" authorityYear="1931" box="[210,341,403,429]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rambouseki">
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<emphasis id="B961335F2949FFB6FF0EFE27ED7BFE50" box="[210,341,403,429]" italics="true" pageId="9" pageNumber="10">rambouseki</emphasis>
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</taxonomicName>
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” group).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FBCC62949FFB6FF1BFE60E909F8DF" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BAAEF4D2949FFB6FF1BFE60E909F8DF" blockId="9.[151,1437,467,2043]" pageId="9" pageNumber="10">
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<emphasis id="B961335F2949FFB6FF1BFE60ED7FFE13" bold="true" box="[199,337,468,494]" pageId="9" pageNumber="10">Description</emphasis>
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.
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<emphasis id="B961335F2949FFB6FE82FE67EE38FE13" bold="true" box="[350,534,467,494]" italics="true" pageId="9" pageNumber="10">Color and luster</emphasis>
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. Dorsal part of head and ventral surface dark-brown to black; pronotum and elytra dark brown to black (most species) or bluish (
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<taxonomicName id="4C1594CE2949FFB6FD43FE4DEF14FDEF" authorityName="Chaudoir" authorityYear="1882" box="[671,826,504,531]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
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<emphasis id="B961335F2949FFB6FD43FE4DEE9EFDEF" box="[671,688,505,530]" italics="true" pageId="9" pageNumber="10">P</emphasis>
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.
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<emphasis id="B961335F2949FFB6FD62FE4CEF14FDEF" box="[702,826,504,530]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
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</taxonomicName>
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); appendages piceous to rufopiceous.
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<emphasis id="B961335F2949FFB6FB37FE43EDB9FDCB" bold="true" italics="true" pageId="9" pageNumber="10">Microsculpture and micropunctuation</emphasis>
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. Dorsal surface of head, pronotum and elytra with regular micropunctuation and isodiametric meshes well impressed which together form “punctate rosettes” microreticulation (like
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<taxonomicName id="4C1594CE2949FFB6FBB5FDF4E8F0FDA7" box="[1129,1246,576,602]" class="Insecta" family="Carabidae" genus="Anatrichis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2949FFB6FBB5FDF4E8F0FDA7" box="[1129,1246,576,602]" italics="true" pageId="9" pageNumber="10">Anatrichis</emphasis>
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</taxonomicName>
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spp., see
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<bibRefCitation id="EF8492BC2949FFB6FA97FDF4ECFCFD83" author="Spence, J. R." pageId="9" pageNumber="10" pagination="567 - 580" refId="ref64265" refString="Spence, J. R. (1983) Taxonomic status, relationships, and biogeography of Anatrichis LeConte and Oodinus Motschulsky (Carabidae: Oodini). The Coleopterist Bulletin, 36 (4), 567 - 580." type="journal article" year="1983">Spence 1983</bibRefCitation>
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;
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<bibRefCitation id="EF8492BC2949FFB6FF3CFDD0EE33FD83" author="Gueorguiev, B. & Morita, S." box="[224,541,612,638]" pageId="9" pageNumber="10" pagination="249 - 263" refId="ref61579" refString="Gueorguiev, B. & Morita, S. (2017) Two new species of Anatrichis LeConte (Coleoptera: Carabidae: Oodini) from Japan, with notes on the taxonomic position of Oodes infimus Andrewes. Zootaxa, 4226 (2), 249 - 263. https: // doi. org / 10.11646 / zootaxa. 4226.2.5" type="journal article" year="2017">Guéorguiev & Morita 2017</bibRefCitation>
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), micropunctuation larger, denser and more regular on head than on pronotum and elytra; ventral surface without regular micropunctuation, with reduced sculpticells (sometimes meso- and metacoxae, and metasternum with microreticulation).
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<emphasis id="B961335F2949FFB6FD2BFD1FEF54FD3B" bold="true" box="[759,890,683,710]" italics="true" pageId="9" pageNumber="10">Chaetotaxy</emphasis>
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. Labrum with six more or less equidistant setae, the two lateral shorter and slightly remote from medial four. Clypeal setae present. Anterior supraorbital seta absent. Ventral seta of antennomere 2 located in apical half. Anterior seta of stipes present. Setae on apical margin of ligula distant. Penultimate labial palpomere glabrous. Pronotum with basal setiferous punctures (except
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<taxonomicName id="4C1594CE2949FFB6FB33FCADE9A5FCCF" authorityName="Chaudoir" authorityYear="1882" box="[1263,1419,792,819]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
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<emphasis id="B961335F2949FFB6FB33FCADE92EFCCF" box="[1263,1280,793,818]" italics="true" pageId="9" pageNumber="10">P</emphasis>
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.
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<emphasis id="B961335F2949FFB6FAD3FCACE9A5FCCF" box="[1295,1419,792,818]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
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</taxonomicName>
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). Parascutellar seta of elytron present. Elytral interval 3 with two discal setiferous punctures near stria 2, first situated in medial third, second in posterior third. Mesocoxa with lateral and posterior setae; mesotrochanter with one seta. Hind femur without posterior setae. Last visible sternite with two apical punctures in male, four in female. Basal gonocoxite with lateroapical setae, without medioapical setae. Apical gonocoxite with two, well-developed or reduced, dorsolateral ensiform setae; dorsomedial ensiform seta present, located near base or equidistant from base and middle, rarely near middle; nematiform setae present, located near apex or equidistant from apex and middle.
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<emphasis id="B961335F2949FFB6FF28FBA7ED1CFBD3" bold="true" box="[244,306,1043,1070]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
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. Eye large, its vertical diameter greater than length of antennomere 1 (in most species) or subequal to it (in “
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<taxonomicName id="4C1594CE2949FFB6FEDEFB8CEDABFBAF" authorityName="Jedlicka. Consists" authorityYear="1931" box="[258,389,1080,1106]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rambouseki">
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<emphasis id="B961335F2949FFB6FEDEFB8CEDABFBAF" box="[258,389,1080,1106]" italics="true" pageId="9" pageNumber="10">rambouseki</emphasis>
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</taxonomicName>
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” group). Anterior margin of labrum straight. Antenna long, with last two or three segments exceeding pronotal base. Mandibles large, convex. Last two maxillary palpomeres of subequal length. Paraglossa long, not expanded.
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<collectingRegion id="49D121AF2949FFB6FEA6FB35EDF7FB66" box="[378,473,1153,1179]" country="Myanmar" name="Chin" pageId="9" pageNumber="10">Mentum</collectingRegion>
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||
tooth with apex truncate (most species), more rarely emarginate (in
|
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<taxonomicName id="4C1594CE2949FFB6FB09FB35E960FB67" authorityName="Guéorguiev & Liang" authorityYear="2020" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1237,1358,1153,1179]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vicarius">
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||
<emphasis id="B961335F2949FFB6FB09FB35E8C8FB67" box="[1237,1254,1153,1178]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FB28FB35E960FB67" box="[1268,1358,1153,1178]" italics="true" pageId="9" pageNumber="10">vicarius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE2949FFB6FA58FB35ED34FB43" authorityName="Jedlicka. Consists" authorityYear="1931" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rambouseki">
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<emphasis id="B961335F2949FFB6FA58FB35E9BBFB67" box="[1412,1429,1153,1178]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FF4BFB10ED34FB43" box="[151,282,1188,1214]" italics="true" pageId="9" pageNumber="10">rambouseki</emphasis>
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</taxonomicName>
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||
).
|
||
<emphasis id="B961335F2949FFB6FEEFFB17EDA7FB43" bold="true" box="[307,393,1187,1214]" italics="true" pageId="9" pageNumber="10">Thorax</emphasis>
|
||
. Pronotum with anterior angles slightly to moderately projected. Prosternal process elliptic (most species), rarely ovate (in
|
||
<taxonomicName id="4C1594CE2949FFB6FDD3FB7DEEA3FB1F" authorityName="Guéorguiev & Liang" authorityYear="2020" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[527,653,1225,1251]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="B961335F2949FFB6FDD3FB7DEE0EFB1F" box="[527,544,1225,1250]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FDEFFB7DEEA3FB1F" box="[563,653,1225,1250]" italics="true" pageId="9" pageNumber="10">vicarius</emphasis>
|
||
</taxonomicName>
|
||
), with apex rounded or subpointed. Mesosternum deeply concave; mesepisternum without apodemal pit; mesocoxal rim entire.
|
||
<emphasis id="B961335F2949FFB6FCB7FB5FEF9DFAFB" bold="true" box="[875,947,1259,1286]" italics="true" pageId="9" pageNumber="10">Elytra</emphasis>
|
||
. Parascutellar striola distinct or reduced, situated between suture and stria 1; stria 7 as distinct as medial striae. Granulation in marginal furrow continuous.
|
||
<emphasis id="B961335F2949FFB6FF4BFA80ECE3FAB3" bold="true" box="[151,205,1332,1358]" italics="true" pageId="9" pageNumber="10">Legs</emphasis>
|
||
. Metacoxal basal sulcus well developed, extending to lateral fourth (in “
|
||
<taxonomicName id="4C1594CE2949FFB6FBF8FA80E880FAB3" authorityName="Bates. Consists" authorityYear="1892" box="[1060,1198,1332,1358]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cribristernis">
|
||
<emphasis id="B961335F2949FFB6FBF8FA80E880FAB3" box="[1060,1198,1332,1358]" italics="true" pageId="9" pageNumber="10">cribristernis</emphasis>
|
||
</taxonomicName>
|
||
” group) or reduced, reaching lateral third of coxa (in “
|
||
<taxonomicName id="4C1594CE2949FFB6FDFFFAECEE99FA8F" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[547,695,1368,1394]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ampliusculus">
|
||
<emphasis id="B961335F2949FFB6FDFFFAECEE99FA8F" box="[547,695,1368,1394]" italics="true" pageId="9" pageNumber="10">ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
” and “
|
||
<taxonomicName id="4C1594CE2949FFB6FCD7FAECEFA0FA8F" authorityName="Jedlicka. Consists" authorityYear="1931" box="[779,910,1368,1394]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F2949FFB6FCD7FAECEFA0FA8F" box="[779,910,1368,1394]" italics="true" pageId="9" pageNumber="10">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
” groups), rarely ending at medial third (in
|
||
<taxonomicName id="4C1594CE2949FFB6FA58FAEDED3DFA6B" authorityName="Chaudoir" authorityYear="1882" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F2949FFB6FA58FAEDE9BBFA8F" box="[1412,1429,1369,1394]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FF4BFAC8ED3DFA6B" box="[151,275,1404,1430]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
|
||
</taxonomicName>
|
||
). Male mesotibia slightly dilated apically (most species), rarely markedly swollen in apical two-thirds (in
|
||
<taxonomicName id="4C1594CE2949FFB6FF62FA15ED4FFA47" authorityName="Jedlicka. Consists" authorityYear="1931" box="[190,353,1440,1467]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F2949FFB6FF62FA15ECE1FA47" box="[190,207,1441,1466]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FF02FA14ED4FFA47" box="[222,353,1440,1466]" italics="true" pageId="9" pageNumber="10">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
) or greatly curved in basal third and then widened apically (in
|
||
<taxonomicName id="4C1594CE2949FFB6FBECFA15E8E2FA47" authorityName="Chaudoir" authorityYear="1882" box="[1072,1228,1440,1467]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F2949FFB6FBECFA15E86FFA47" box="[1072,1089,1441,1466]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FB8CFA14E8E2FA47" box="[1104,1228,1440,1466]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
|
||
</taxonomicName>
|
||
). Mesotarsomeres 2–4 and metatarsomeres
|
||
<quantity id="4CED42A82949FFB6FE77FA70EDDFFA23" box="[427,497,1476,1502]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="9" pageNumber="10" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
|
||
both sexes as well as protarsomeres
|
||
<quantity id="4CED42A82949FFB6FC5AFA70EFE2FA23" box="[902,972,1476,1502]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="9" pageNumber="10" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
|
||
female with (in
|
||
<taxonomicName id="4C1594CE2949FFB6FB5CFA71E934FA23" authorityName="Chaudoir" authorityYear="1882" box="[1152,1306,1476,1503]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F2949FFB6FB5CFA71E8BFFA23" box="[1152,1169,1477,1502]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FB42FA70E934FA23" box="[1182,1306,1476,1502]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
|
||
</taxonomicName>
|
||
) or without (in other species) long and dense, yellowish setae on ventral surface. Male front tarsomeres 1–3 with numerous adhesive setae beneath; adhesive setae covering apical three-fourths of tarsomere 1.
|
||
<emphasis id="B961335F2949FFB6FBBBF9BFE93FF9DB" bold="true" box="[1127,1297,1547,1574]" italics="true" pageId="9" pageNumber="10">Male genitalia</emphasis>
|
||
. Basal bulb of median lobe broad. Ostium reaching (in
|
||
<taxonomicName id="4C1594CE2949FFB6FD56F985EF06F9B7" authorityName="Chaudoir" authorityYear="1882" box="[650,808,1584,1611]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F2949FFB6FD56F985EEB5F9B7" box="[650,667,1585,1610]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FD70F984EF06F9B7" box="[684,808,1584,1610]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
|
||
</taxonomicName>
|
||
) or not reaching basal bulb (in other species). Internal sac with elongate sclerite occupying greater length of shaft.
|
||
<emphasis id="B961335F2949FFB6FC96F9E7E821F993" bold="true" box="[842,1039,1619,1646]" italics="true" pageId="9" pageNumber="10">Female genitalia</emphasis>
|
||
. Apical gonocoxite subelongate or subtriangular.
|
||
<collectingRegion id="49D121AF2949FFB6FEE6F9CDED55F96E" box="[314,379,1657,1683]" country="Turkey" name="Bursa" pageId="9" pageNumber="10">Bursa</collectingRegion>
|
||
copulatrix with or without dorsal sclerite; spermatheca moderately long to very long, more or less differentiated (most species) or undifferentiated (
|
||
<taxonomicName id="4C1594CE2949FFB6FCC3F929EF90F94B" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[799,958,1692,1719]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F2949FFB6FCC3F929EF1EF94B" box="[799,816,1693,1718]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FC9CF928EF90F94B" box="[832,958,1692,1718]" italics="true" pageId="9" pageNumber="10">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE2949FFB6FC05F929E89EF94B" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[985,1200,1692,1719]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leigongshanicus">
|
||
<emphasis id="B961335F2949FFB6FC05F929EFC4F94B" box="[985,1002,1693,1718]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FC26F928E89EF94B" box="[1018,1200,1692,1718]" italics="true" pageId="9" pageNumber="10">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
), with receptaculum twisted (most species) or coiled apically (
|
||
<taxonomicName id="4C1594CE2949FFB6FD5DF975EF0FF927" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[641,801,1728,1755]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F2949FFB6FD5DF975EEBCF927" box="[641,658,1729,1754]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FD7FF974EF0FF927" box="[675,801,1728,1754]" italics="true" pageId="9" pageNumber="10">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE2949FFB6FCEFF975E822F927" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[819,1036,1728,1755]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leigongshanicus">
|
||
<emphasis id="B961335F2949FFB6FCEFF975EF6AF927" box="[819,836,1729,1754]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FC8AF974E822F927" box="[854,1036,1728,1754]" italics="true" pageId="9" pageNumber="10">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
); spermathecal gland with atrium, connected in apical fifth (“
|
||
<taxonomicName id="4C1594CE2949FFB6FE1DF950EE13F903" authorityName="Motschulsky" authorityYear="1844" box="[449,573,1764,1790]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cribricollis">
|
||
<emphasis id="B961335F2949FFB6FE1DF950EE13F903" box="[449,573,1764,1790]" italics="true" pageId="9" pageNumber="10">cribricollis</emphasis>
|
||
</taxonomicName>
|
||
” and “
|
||
<taxonomicName id="4C1594CE2949FFB6FD50F950EF26F903" authorityName="Chaudoir" authorityYear="1882" box="[652,776,1764,1790]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F2949FFB6FD50F950EF26F903" box="[652,776,1764,1790]" italics="true" pageId="9" pageNumber="10">coelestinus</emphasis>
|
||
</taxonomicName>
|
||
” groups), or apical third (
|
||
<taxonomicName id="4C1594CE2949FFB6FBF7F951E8F3F903" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1067,1245,1764,1791]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ampliusculus">
|
||
<emphasis id="B961335F2949FFB6FBF7F951E812F903" box="[1067,1084,1765,1790]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FB95F950E8F3F903" box="[1097,1245,1764,1790]" italics="true" pageId="9" pageNumber="10">ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE2949FFB6FB37F951E9A2F903" authorityName="Jedlicka. Consists" authorityYear="1931" box="[1259,1420,1764,1791]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F2949FFB6FB37F951E8D2F903" box="[1259,1276,1765,1790]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FAD5F950E9A2F903" box="[1289,1420,1764,1790]" italics="true" pageId="9" pageNumber="10">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
), or near middle (
|
||
<taxonomicName id="4C1594CE2949FFB6FE97F8BDEDC6F8DF" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[331,488,1800,1827]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hunanensis">
|
||
<emphasis id="B961335F2949FFB6FE97F8BDED72F8DF" box="[331,348,1801,1826]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FEB6F8BCEDC6F8DF" box="[362,488,1800,1826]" italics="true" pageId="9" pageNumber="10">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE2949FFB6FE2AF8BDEEE5F8DF" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[502,715,1800,1827]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leigongshanicus">
|
||
<emphasis id="B961335F2949FFB6FE2AF8BDEE29F8DF" box="[502,519,1801,1826]" italics="true" pageId="9" pageNumber="10">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2949FFB6FDC9F8BCEEE5F8DF" box="[533,715,1800,1826]" italics="true" pageId="9" pageNumber="10">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
) of spermatheca; common oviduct with villous canal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FBCC62949FFB5FF1BF898EF01FF04" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="8BAAEF4D2949FFB6FF1BF898EDC9F807" blockId="9.[151,1437,467,2043]" pageId="9" pageNumber="10">
|
||
<emphasis id="B961335F2949FFB6FF1BF898EE1AF8BB" bold="true" box="[199,564,1836,1862]" pageId="9" pageNumber="10">Monophyly and relationships</emphasis>
|
||
. Species of
|
||
<taxonomicName id="4C1594CE2949FFB6FD0EF898EF76F8BB" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[722,856,1836,1862]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2949FFB6FD0EF898EF76F8BB" box="[722,856,1836,1862]" italics="true" pageId="9" pageNumber="10">Pseudoodes</emphasis>
|
||
</taxonomicName>
|
||
share three synapomorphies suggesting that the genus, as presently conceived, form a clade: (1) mentum tooth truncate to emarginate; (2) abdominal sternite 3 without ambulatory setae; and (3) internal sac of median lobe with elongate, helical sclerite. The genus is rather heterogeneous as its four species groups display significant morphological differences with each other. The great divergence seems to be a result of long-term evolution, colonization of new geographical areas and settlement in new (different) microhabitats.
|
||
</paragraph>
|
||
<paragraph id="8BAAEF4D294AFFB5FF1BFF23EF01FF04" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="11">
|
||
The species of
|
||
<taxonomicName id="4C1594CE294AFFB5FEB0FF23EDDCFF4C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[364,498,151,177]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F294AFFB5FEB0FF23EDDCFF4C" box="[364,498,151,177]" italics="true" pageId="10" pageNumber="11">Pseudoodes</emphasis>
|
||
</taxonomicName>
|
||
share one synapomorphy with those of
|
||
<taxonomicName id="4C1594CE294AFFB5FC75FF23E81EFF4C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[937,1072,151,177]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F294AFFB5FC75FF23E81EFF4C" box="[937,1072,151,177]" italics="true" pageId="10" pageNumber="11">Sundaoodes</emphasis>
|
||
</taxonomicName>
|
||
—the spermatheca differentiated into long and thinner seminal canal and short and wider receptaculum (
|
||
<figureCitation id="132EF3C8294AFFB5FC60FF0FE839FF28" box="[956,1047,187,213]" captionStart="FIGURE 2" captionStartId="14.[151,250,1894,1918]" captionTargetBox="[257,1283,227,1844]" captionTargetId="figure-20@14.[219,1367,181,1870]" captionTargetPageId="14" captionText="FIGURE 2. Pseudoodes cribristernis Bates, 1892 (A–B: female specimen, China, Yunnan Prov., Fugong County, Guquan Village; C: male specimen, same locality; D–E: female specimen, Thailand, Chiang Dao). A: mentum; B: prosternal process; C: right protarsomeres; D: left gonocoxite, ventral view; E: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–C, E); = 0.2 mm (Fig. D)." figureDoi="http://doi.org/10.5281/zenodo.4407076" httpUri="https://zenodo.org/record/4407076/files/figure.png" pageId="10" pageNumber="11">Figs 2E</figureCitation>
|
||
,
|
||
<figureCitation id="132EF3C8294AFFB5FBF8FF0FE866FF28" box="[1060,1096,187,213]" captionStart="FIGURE 7" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[261,1303,218,1851]" captionTargetId="figure-20@22.[214,1372,181,1884]" captionTargetPageId="22" captionText="FIGURE 7. Pseudoodes coelestinus Chaudoir, 1882 (A: male specimen, Malaysia, Mulu National Park; B: male specimen, Malaysia, Kapit; C–D: female specimen, Malaysia, Mulu National Park).A: median lobe of aedeagus, left lateral view; B: same, dorsal view; C: left gonocoxite, ventral view; D: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–B, D); = 0.2 mm (Fig. C)." figureDoi="http://doi.org/10.5281/zenodo.4407086" httpUri="https://zenodo.org/record/4407086/files/figure.png" pageId="10" pageNumber="11">7D</figureCitation>
|
||
,
|
||
<figureCitation id="132EF3C8294AFFB5FB89FF0FE850FF28" box="[1109,1150,187,213]" captionStart="FIGURE 14" captionStartId="39.[151,250,1891,1915]" captionTargetBox="[250,1329,208,1852]" captionTargetId="figure-17@39.[221,1367,181,1866]" captionTargetPageId="39" captionText="FIGURE 14. Sundaoodes hainanensis sp. n. (A–H:holotype; I–J: female paratype).A: habitus and labels; B: head and pronotum; C: mentum; D: prosternal process; E: left metepisternum and metacoxa; F: median lobe of aedeagus, left lateral view; G: same, ventral view; H: same, dorsal view; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Fig. A); = 1 mm (Figs B, E); = 0.5 mm (Figs C–D, F–H, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407100" httpUri="https://zenodo.org/record/4407100/files/figure.png" pageId="10" pageNumber="11">14J</figureCitation>
|
||
,
|
||
<figureCitation id="132EF3C8294AFFB5FB57FF0EE8F6FF28" box="[1163,1240,186,213]" captionStart="FIGURE 15" captionStartId="41.[151,250,1858,1882]" captionTargetBox="[258,1316,199,1824]" captionTargetId="figure-17@41.[232,1356,181,1834]" captionTargetPageId="41" captionText="FIGURE 15. Sundaoodes kalimantanensis sp. n. (A, C–D, G–H: holotype; B, E–F, I–K: paratype). A: habitus and labels; B: habitus; C: head and pronotum; D: mentum (white spots show locations of submental setiferous punctures); E: prosternal process; F: left metepisternum and metacoxa; G: median lobe of aedeagus, left lateral view; H: same, anterior half, dorsal view; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, dorsal view; K: same, ventral view. Scale lines = 2 mm (Figs A–B); = 1 mm (Figs C, F); = 0.5 mm (Fig. D–E, G–H, J–K); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407104" httpUri="https://zenodo.org/record/4407104/files/figure.png" pageId="10" pageNumber="11">15J, K</figureCitation>
|
||
). The two genera might be adelphotaxa, but evidence to support this is weak.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FBCC6294AFFB5FF1BFEB7ED88FD54" pageId="10" pageNumber="11" type="etymology">
|
||
<subSubSection id="C30FBCC6294AFFB5FF1BFEB7E88FFEBC" pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph id="8BAAEF4D294AFFB5FF1BFEB7E88FFEBC" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="11">
|
||
<emphasis id="B961335F294AFFB5FF1BFEB7ED66FEE0" bold="true" box="[199,328,259,285]" pageId="10" pageNumber="11">Etymology</emphasis>
|
||
. A compound name which derives from the Greek stem ψευδής [pseudḗs] (false, fake) and the generic name
|
||
<taxonomicName id="4C1594CE294AFFB5FEE8FE93ED52FEBC" box="[308,380,295,321]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F294AFFB5FEE8FE93ED52FEBC" box="[308,380,295,321]" italics="true" pageId="10" pageNumber="11">Oodes</emphasis>
|
||
</taxonomicName>
|
||
also of Greek origin (see Bousquet 2012: 955). Its gender is masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FBCC6294AFFB5FF1BFEFEEF18FE08" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="8BAAEF4D294AFFB5FF1BFEFEEF18FE08" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="11">
|
||
<emphasis id="B961335F294AFFB5FF1BFEFEEE9FFE99" bold="true" box="[199,689,330,357]" pageId="10" pageNumber="11">Geographical distribution and diversity</emphasis>
|
||
. The genus includes ten species:
|
||
<taxonomicName id="4C1594CE294AFFB5FB95FEFFE95DFE98" authority="(Bates, 1873)" authorityName="Guéorguiev & Liang" authorityYear="2020" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1097,1395,331,357]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="B961335F294AFFB5FB95FEFFE874FE99" box="[1097,1114,331,356]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FBB1FEFFE8E9FE99" box="[1133,1223,331,356]" italics="true" pageId="10" pageNumber="11">vicarius</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8492BC294AFFB5FB00FEFFE944FE98" author="Bates, H. W." box="[1244,1386,331,357]" pageId="10" pageNumber="11" pagination="219 - 322" refId="ref59849" refString="Bates, H. W. (1873) On the geodephagous Coleoptera of Japan. The Transactions of the Entomological Society of London for the year 1873, 1873, 219 - 322. https: // doi. org / 10.1111 / j. 1365 - 2311.1873. tb 00643. x" type="journal article" year="1873">Bates, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FA58FEFFEDC8FE74" authority="(Chaudoir, 1882)" baseAuthorityName="Chaudoir" baseAuthorityYear="1882" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="coelestinus">
|
||
<emphasis id="B961335F294AFFB5FA58FEFFE9BBFE99" box="[1412,1429,331,356]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FF4BFEDBED3DFE74" box="[151,275,367,393]" italics="true" pageId="10" pageNumber="11">coelestinus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8492BC294AFFB5FEFBFEDBEDF0FE74" author="Chaudoir, M. de" box="[295,478,367,393]" pageId="10" pageNumber="11" pagination="317 - 378" refId="ref60500" refString="Chaudoir, M. de (1882) Monographie des oodides 1 re partie. Annales de la Societe Entomologique de France, Series 6, 2, 317 - 378." type="journal article" year="1882">Chaudoir, 1882</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FE2AFEDBEF51FE74" authority="(Chaudoir, 1882)" baseAuthorityName="Chaudoir" baseAuthorityYear="1882" box="[502,895,367,393]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="subcoriaceus">
|
||
<emphasis id="B961335F294AFFB5FE2AFEDBEE29FE75" box="[502,519,367,392]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FDC5FEDBEE83FE74" box="[537,685,367,393]" italics="true" pageId="10" pageNumber="11">subcoriaceus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8492BC294AFFB5FD1CFEDBEF59FE74" author="Chaudoir, M. de" box="[704,887,367,393]" pageId="10" pageNumber="11" pagination="317 - 378" refId="ref60500" refString="Chaudoir, M. de (1882) Monographie des oodides 1 re partie. Annales de la Societe Entomologique de France, Series 6, 2, 317 - 378." type="journal article" year="1882">Chaudoir, 1882</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FC53FEDBE8C8FE74" authority="(Bates, 1892)" baseAuthorityName="Bates" baseAuthorityYear="1892" box="[911,1254,367,393]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cribristernis">
|
||
<emphasis id="B961335F294AFFB5FC53FEDBEF8EFE75" box="[911,928,367,392]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FC6EFEDBE812FE74" box="[946,1084,367,393]" italics="true" pageId="10" pageNumber="11">cribristernis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8492BC294AFFB5FB8CFEDBE8F0FE74" author="Bates, H. W." box="[1104,1246,367,393]" pageId="10" pageNumber="11" pagination="267 - 428" refId="ref59903" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, 32, 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FB2AFEDBED63FE50" authority="(Jedlicka, 1931)" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F294AFFB5FB2AFEDBE929FE75" box="[1270,1287,367,392]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FAC6FEDBE9B3FE74" box="[1306,1437,367,393]" italics="true" pageId="10" pageNumber="11">rambouseki</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8492BC294AFFB5FF43FE27ED6AFE50" author="Jedlicka, A." box="[159,324,403,429]" pageId="10" pageNumber="11" pagination="21 - 30" refId="ref62314" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China-Szetschuan. Casopis Ceskoslovenske Spolecnosti Entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička, 1931</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FE84FE27EE24FE50" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[344,522,403,429]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ampliusculus" status="sp. nov.">
|
||
<emphasis id="B961335F294AFFB5FE84FE27ED47FE51" box="[344,361,403,428]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FEAAFE27EE24FE50" box="[374,522,403,429]" italics="true" pageId="10" pageNumber="11">ampliusculus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961335F294AFFB5FDCCFE27EE60FE50" bold="true" box="[528,590,403,429]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A2528E24294AFFB5FDCCFE27EE60FE50" box="[528,590,403,429]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FD87FE27EF3EFE50" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[603,784,403,429]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="emeishanicus" status="sp. nov.">
|
||
<emphasis id="B961335F294AFFB5FD87FE27EE42FE51" box="[603,620,403,428]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FDA5FE27EF3EFE50" box="[633,784,403,429]" italics="true" pageId="10" pageNumber="11">emeishanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961335F294AFFB5FCCAFE27EF7AFE50" bold="true" box="[790,852,403,429]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A2528E24294AFFB5FCCAFE27EF7AFE50" box="[790,852,403,429]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FCBDFE27EFD3FE50" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[865,1021,403,429]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. nov.">
|
||
<emphasis id="B961335F294AFFB5FCBDFE27EF5CFE51" box="[865,882,403,428]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FCA3FE27EFD3FE50" box="[895,1021,403,429]" italics="true" pageId="10" pageNumber="11">hunanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961335F294AFFB5FBDFFE27E86CFE50" bold="true" box="[1027,1090,403,429]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A2528E24294AFFB5FBDFFE27E86CFE50" box="[1027,1090,403,429]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C1594CE294AFFB5FB92FE27E90DFE50" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1102,1315,403,429]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="leigongshanicus" status="sp. nov.">
|
||
<emphasis id="B961335F294AFFB5FB92FE27E871FE51" box="[1102,1119,403,428]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FBB1FE27E90DFE50" box="[1133,1315,403,429]" italics="true" pageId="10" pageNumber="11">leigongshanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961335F294AFFB5FAF5FE27E949FE50" bold="true" box="[1321,1383,403,429]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A2528E24294AFFB5FAF5FE27E949FE50" box="[1321,1383,403,429]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C1594CE294AFFB5FF4BFE03ED16FE2C" authority="Guéorguiev & Liang, 2020" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[151,312,439,465]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tianlinensis" status="sp. nov.">
|
||
<emphasis id="B961335F294AFFB5FF4BFE03EC86FE2D" box="[151,168,439,464]" italics="true" pageId="10" pageNumber="11">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FF6BFE03ED16FE2C" box="[183,312,439,465]" italics="true" pageId="10" pageNumber="11">tianlinensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961335F294AFFB5FEE3FE03ED51FE2C" bold="true" box="[319,383,439,465]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A2528E24294AFFB5FEE3FE03ED51FE2C" box="[319,383,439,465]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The species occur in East and Southeast Asia, ranging from
|
||
<collectingCountry id="F302AFDD294AFFB5FBF7FE03E845FE2C" box="[1067,1131,439,465]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
|
||
in the north through South
|
||
<collectingCountry id="F302AFDD294AFFB5FF4BFE6FECF4FE08" box="[151,218,475,501]" name="China" pageId="10" pageNumber="11">China</collectingCountry>
|
||
to Indochina and the Malay Archipelago in the south.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FBCC6294AFFB5FF1BFE4BED88FD54" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="8BAAEF4D294AFFB5FF1BFE4BED88FD54" blockId="10.[151,1437,151,681]" pageId="10" pageNumber="11">
|
||
<emphasis id="B961335F294AFFB5FF1BFE4BED24FDE4" bold="true" box="[199,266,511,537]" pageId="10" pageNumber="11">Notes</emphasis>
|
||
. The number of submental setae, prosternal process narrowly rounded apically and the structure of spermatheca (
|
||
<bibRefCitation id="EF8492BC294AFFB5FEEFFD97EE7AFDC0" author="Bousquet, Y." box="[307,596,547,573]" pageId="10" pageNumber="11" pagination="443 - 537" refId="ref60231" refString="Bousquet, Y. (1996) Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae). The Canadian Entomologist, 128, 443 - 537. https: // doi. org / 10.4039 / Ent 128443 - 3" type="journal article" year="1996">Bousquet 1996: 471–472</bibRefCitation>
|
||
, 532, fig. 131) suggest that the North American species
|
||
<taxonomicName id="4C1594CE294AFFB5FB0BFD97ED02FD9C" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amaroides">
|
||
<emphasis id="B961335F294AFFB5FB0BFD97E9B2FDC0" box="[1239,1436,547,573]" italics="true" pageId="10" pageNumber="11">Oodes amaroides</emphasis>
|
||
Dejean, 1831
|
||
</taxonomicName>
|
||
may also belong to this genus. However, this species has prosternal process bordered apically, a state opposite to that in the taxa of
|
||
<taxonomicName id="4C1594CE294AFFB5FE2BFDDFEE53FD78" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[503,637,619,645]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F294AFFB5FE2BFDDFEE53FD78" box="[503,637,619,645]" italics="true" pageId="10" pageNumber="11">Pseudoodes</emphasis>
|
||
</taxonomicName>
|
||
. We have not studied any specimens of four Nearctic
|
||
<taxonomicName id="4C1594CE294AFFB5FB2FFDDFE915FD78" box="[1267,1339,619,645]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F294AFFB5FB2FFDDFE915FD78" box="[1267,1339,619,645]" italics="true" pageId="10" pageNumber="11">Oodes</emphasis>
|
||
</taxonomicName>
|
||
species, including
|
||
<taxonomicName id="4C1594CE294AFFB5FED4FD3BEDB1FD54" authorityName="Dejean" authorityYear="1831" box="[264,415,655,681]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amaroides">
|
||
<emphasis id="B961335F294AFFB5FED4FD3BED32FD54" box="[264,284,655,681]" italics="true" pageId="10" pageNumber="11">O</emphasis>
|
||
.
|
||
<emphasis id="B961335F294AFFB5FEF6FD3BEDB1FD54" box="[298,415,655,681]" italics="true" pageId="10" pageNumber="11">amaroides</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |