211 lines
33 KiB
XML
211 lines
33 KiB
XML
<document id="1F3887E4B342F785C35E8414F0A1032F" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE4E517509887FB60F2FC15" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) ambulans Assing 2017, spec. nov." docType="treatment" docUuid="600E1CB3-AA42-431F-992D-88C3B2569849" docUuidSource="ZooBank" docVersion="11" lastPageNumber="154" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="154" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="FDE6C29516ED88ACDD65DB321EA91A2E">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="31857B47BBB2F41E300B511480D25F9D">Assing, Volker</mods:namePart>
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<mods:title id="81A5F48C3DF86B63B89AC866FB9C6EB6">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="CDB2E9BCC18BFABE48684EB125B5300D">
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<mods:date id="4204A053FB2176A733057A1A290B5703">2017</mods:date>
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<mods:number id="66A5A94C457CF2C3FBBAB9FA41A1F8A1">2017-06-30</mods:number>
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<mods:number id="9B364FB94E372A83B10FDBA821B058C0">67</mods:number>
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<treatment id="03A787BAFFE4E517509887FB60F2FC15" ID-DOI="http://doi.org/10.5281/zenodo.5888485" ID-GBIF-Taxon="192498179" ID-Zenodo-Dep="5888485" LSID="urn:lsid:zoobank.org:act:600E1CB3-AA42-431F-992D-88C3B2569849" httpUri="http://treatment.plazi.org/id/03A787BAFFE4E517509887FB60F2FC15" lastPageNumber="154" pageId="37" pageNumber="154">
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<subSubSection id="C3146527FFE4E517509887FB61D9FE9B" pageId="37" pageNumber="154" type="nomenclature">
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<paragraph id="8BB136ACFFE4E517509887FB6112FEBA" blockId="37.[189,722,243,335]" box="[189,537,243,270]" pageId="37" pageNumber="154">
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<taxonomicName id="4C0E4D2FFFE4E517509887FB62A1FEBA" ID-CoL="7GHVN" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[189,426,243,270]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="ambulans" status="sp. nov." subGenus="Zyras">
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<emphasis id="B97AEABEFFE4E517509887FB63F8FEBA" box="[189,243,243,270]" italics="true" pageId="37" pageNumber="154">Zyras</emphasis>
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(
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<emphasis id="B97AEABEFFE4E517512487FB623CFEBA" box="[257,311,243,270]" italics="true" pageId="37" pageNumber="154">Zyras</emphasis>
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)
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<emphasis id="B97AEABEFFE4E517516087FB62A1FEBA" box="[325,426,243,270]" italics="true" pageId="37" pageNumber="154">ambulans</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957C5FFE4E517519587FB6112FEBA" box="[432,537,243,270]" pageId="37" pageNumber="154" rank="species">spec. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BB136ACFFE4E5175098861361D9FE9B" blockId="37.[189,722,243,335]" box="[189,722,283,303]" pageId="37" pageNumber="154">
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<uri id="FF9F3AAEFFE4E5175098861361D9FE9B" box="[189,722,283,303]" pageId="37" pageNumber="154">
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urn:lsid:zoobank.org:pub:
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<uuid id="FFA80C79FFE4E5175157861361D9FE9B" box="[370,722,283,303]" pageId="37" pageNumber="154">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFE4E5175098863362C6FEFB" box="[189,461,315,335]" pageId="37" pageNumber="154" type="description">
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<paragraph id="8BB136ACFFE4E5175098863362C6FEFB" blockId="37.[189,722,243,335]" box="[189,461,315,335]" pageId="37" pageNumber="154">
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(
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<figureCitation id="13352A29FFE4E51750E4863363F1FEFB" box="[193,250,315,335]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="37" pageNumber="154">Figs 34</figureCitation>
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,
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<figureCitation id="13352A29FFE4E51751248633621DFEFB" box="[257,278,315,335]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">70</figureCitation>
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,
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<figureCitation id="13352A29FFE4E517513886336230FEFB" box="[285,315,315,335]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="37" pageNumber="154">107</figureCitation>
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,
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<figureCitation id="13352A29FFE4E517516786336283FEFB" box="[322,392,315,335]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="37" pageNumber="154">212–215</figureCitation>
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,
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<figureCitation id="13352A29FFE4E51751AB863362C2FEFB" box="[398,457,315,335]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="37" pageNumber="154">Map 10</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE4E51750B2866162B0FE55" pageId="37" pageNumber="154" type="materials_examined">
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<paragraph id="8BB136ACFFE4E51750B2866162B0FE55" blockId="37.[151,756,361,481]" pageId="37" pageNumber="154">
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<emphasis id="B97AEABEFFE4E51750B28661623CFE35" bold="true" box="[151,311,361,385]" pageId="37" pageNumber="154">Type material</emphasis>
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||
:
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||
<materialsCitation id="3B663CF1FFE4E5175168866162B0FE55" ID-GBIF-Occurrence="3450162325" collectingDate="1996-01-01" collectingDateMax="1996-01-04" collectingDateMin="1996-01-01" collectorName="Schulz & Vock" country="Thailand" determinerName="V. Assing" elevation="300" location="Thale Ban N. P." pageId="37" pageNumber="154" specimenCount="1" stateProvince="Satun Prov." typeStatus="holotype">
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||
<typeStatus id="54B5880EFFE4E5175168866162BBFE35" box="[333,432,361,385]" pageId="37" pageNumber="154" type="holotype">Holotype</typeStatus>
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||
<specimenCount id="9D08FD25FFE4E51751E5866262D0FE35" box="[448,475,362,385]" pageId="37" pageNumber="154">♂:</specimenCount>
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“
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||
<collectingCountry id="F319763CFFE4E51751D186616153FE35" box="[500,600,361,385]" name="Thailand" pageId="37" pageNumber="154">Thailand</collectingCountry>
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,
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<collectingRegion id="49CAF84EFFE4E5175249866161FAFE35" box="[620,753,361,385]" country="Thailand" name="Satun" pageId="37" pageNumber="154">Satun Prov.</collectingRegion>
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,
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<location id="8ED16077FFE4E51750B286816230FE15" LSID="urn:lsid:plazi:treatment:03A787BAFFE4E517509887FB60F2FC15:8ED16077FFE4E51750B286816230FE15" box="[151,315,393,417]" country="Thailand" name="Thale Ban N. P." pageId="37" pageNumber="154" stateProvince="Satun Prov.">Thale Ban N.P.</location>
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,
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<quantity id="4CF69B49FFE4E517516286816281FE15" box="[327,394,393,417]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="154" unit="km" value="20.0">20 km</quantity>
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E Satun
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</locationDeviation>
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,
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<quantity id="4CF69B49FFE4E51751D686816167FE15" box="[499,620,393,417]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="37" pageNumber="154" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
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<elevation id="0023D19FFFE4E51751D686816167FE15" box="[499,620,393,417]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="37" pageNumber="154" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</elevation>
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</quantity>
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,
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<date id="FFB0106CFFE4E517525D868161FBFE15" box="[632,752,393,417]" pageId="37" pageNumber="154" value="1996-01-01" valueMax="1996-01-04" valueMin="1996-01-01">
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<collectingDate id="EFF4E984FFE4E517525D868161FBFE15" box="[632,752,393,417]" pageId="37" pageNumber="154" value="1996-01-01" valueMax="1996-01-04" valueMin="1996-01-01">1.–4.I.1996</collectingDate>
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</date>
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||
, leg.
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||
<collectorName id="26FB537AFFE4E51750E686A16202FE75" box="[195,265,425,449]" pageId="37" pageNumber="154">Schulz</collectorName>
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&
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||
<collectorName id="26FB537AFFE4E517510F86A1626BFE75" box="[298,352,425,449]" pageId="37" pageNumber="154">Vock</collectorName>
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||
/ Holotypus ♂
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||
<taxonomicName id="4C0E4D2FFFE4E517522D86A261A4FE75" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[520,687,425,449]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="ambulans" status="sp. nov.">
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<emphasis id="B97AEABEFFE4E517522D86A261A4FE75" box="[520,687,425,449]" italics="true" pageId="37" pageNumber="154">Zyras ambulans</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A24957C5FFE4E517529386A261E4FE75" box="[694,751,426,449]" pageId="37" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<determinerName id="7E6DDA68FFE4E51750E086C2622CFE55" box="[197,295,457,481]" pageId="37" pageNumber="154">V. Assing</determinerName>
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||
2016” (cAss).
|
||
</materialsCitation>
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE4E51750B2850162F6FDD5" pageId="37" pageNumber="154" type="etymology">
|
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<paragraph id="8BB136ACFFE4E51750B2850162F6FDD5" blockId="37.[151,756,521,609]" pageId="37" pageNumber="154">
|
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<emphasis id="B97AEABEFFE4E51750B285016205FD95" bold="true" box="[151,270,521,545]" pageId="37" pageNumber="154">Etymology</emphasis>
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: The specific epithet is the present participle of the Latin verb ambulare (to walk) and alludes to absence of functional hind wings.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE4E51750B285816710FD15" pageId="37" pageNumber="154" type="description">
|
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<paragraph id="8BB136ACFFE4E51750B28581611BFC75" blockId="37.[151,757,649,2081]" pageId="37" pageNumber="154">
|
||
<emphasis id="B97AEABEFFE4E51750B285816212FD15" bold="true" box="[151,281,649,673]" pageId="37" pageNumber="154">Description</emphasis>
|
||
: Body length
|
||
<quantity id="4CF69B49FFE4E517519A85816118FD15" box="[447,531,649,673]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="37" pageNumber="154" unit="mm" value="6.2">6.2 mm</quantity>
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||
; length of forebody
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<quantity id="4CF69B49FFE4E51750B285A163E0FD75" box="[151,235,681,705]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="37" pageNumber="154" unit="mm" value="2.5">2.5 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="13352A29FFE4E51751A185A162D8FD75" box="[388,467,681,705]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="37" pageNumber="154">Figs 34</figureCitation>
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||
,
|
||
<figureCitation id="13352A29FFE4E51751C185A1610BFD75" box="[484,512,681,705]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">70</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE4E517523585A16131FD75" box="[528,570,681,705]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="37" pageNumber="154">107</figureCitation>
|
||
): head blackishbrown; pronotum and elytra dark-brown; abdomen with segments III–V reddish, tergite VI dark-brown with reddish margins, VII and VIII dark-brown with the anterior and posterior margins narrowly paler; legs yellowish with the apices of the meso- and metafemora brown; antennae dark-brown with antennomeres I–II reddish and XI yellow; maxillary palpi dark-yellowish with the terminal palpomere pale-yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E51750B284C16202FA15" blockId="37.[151,757,649,2081]" pageId="37" pageNumber="154">
|
||
Head (
|
||
<figureCitation id="13352A29FFE4E51750C584C16222FC55" box="[224,297,969,993]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">Fig. 70</figureCitation>
|
||
) moderately transverse, middle extensively impunctate; punctation in lateral dorsal portions sparse and fine. Eyes large and bulging, much longer than postocular region in dorsal view. Antenna (
|
||
<figureCitation id="13352A29FFE4E517526083216185FBF5" box="[581,654,1065,1089]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="37" pageNumber="154">Fig. 34</figureCitation>
|
||
)
|
||
<quantity id="4CF69B49FFE4E5175281832161FFFBF5" box="[676,756,1065,1089]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="37" pageNumber="154" unit="mm" value="2.3">2.3 mm</quantity>
|
||
long; antennomeres IV–V oblong, VI weakly oblong, VII as long as broad or weakly oblong, VIII–IX very weakly transverse, X less than 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X. Pronotum (
|
||
<figureCitation id="13352A29FFE4E517513983C1626CFB55" box="[284,359,1225,1249]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">Fig. 70</figureCitation>
|
||
) weakly transverse, approximately 1.1 times as broad as long and 1.2 times as broad as head, broadest anteriorly, distinctly tapering posteriad; lateral margins straight in posterior two-thirds (dorsal view); punctation shallow and sparse; midline broadly impunctate; lateral margins anteriorly with two very long, stout, dark setae.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E51750B282A16248F9D5" blockId="37.[151,757,649,2081]" pageId="37" pageNumber="154">
|
||
Elytra (
|
||
<figureCitation id="13352A29FFE4E51750CF82A16232FA75" box="[234,313,1449,1473]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">Fig. 70</figureCitation>
|
||
) short, approximately 0.7 times as long as pronotum; punctation moderately dense, moderately coarse, and regularly distributed. Hind wings reduced. Metatarsomere I longer than the combined length of II and III, but shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E51750B2816162CAF795" blockId="37.[151,757,649,2081]" pageId="37" pageNumber="154">
|
||
Abdomen (
|
||
<figureCitation id="13352A29FFE4E517513081616267F935" box="[277,364,1641,1665]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="37" pageNumber="154">Fig. 107</figureCitation>
|
||
) broader than elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V without non-setiferous punctures; tergites III–VI with a lateral puncture bearing a long black seta on either side and with a transverse row of approximately 10 setiferous punctures at or near posterior margins, at least the lateral punctures of these rows bearing long black setae; tergite VII anteriorly with a cluster of rather sparse shallow non-setiferous punctures and posteriorly with sparse setiferous punctures, posterior margin with narrow palisade fringe; tergite VIII with sparse setiferous punctures only in posterior portion, posterior margin smoothly convex, without median excision or other modifications.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E517533F87E267CBFE35" blockId="37.[794,1399,233,385]" pageId="37" pageNumber="154">
|
||
♂: sternite VIII with convex posterior margin; median lobe of aedeagus
|
||
<quantity id="4CF69B49FFE4E51753D78601675AFE95" box="[1010,1105,265,289]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="154" unit="mm" value="0.65">0.65 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="13352A29FFE4E517533F862160A3FEF5" box="[794,936,297,321]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="37" pageNumber="154">Figs 212–213</figureCitation>
|
||
; paramere (
|
||
<figureCitation id="13352A29FFE4E517541D86216784FEF5" box="[1080,1167,297,321]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="37" pageNumber="154">Fig. 214</figureCitation>
|
||
) much longer than median lobe (
|
||
<quantity id="4CF69B49FFE4E517539786416702FED5" box="[946,1033,329,353]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="154" unit="mm" value="0.9">0.9 mm</quantity>
|
||
), with long, flattened, and somewhat club-shaped apical lobe (
|
||
<figureCitation id="13352A29FFE4E517547F866167BEFE35" box="[1114,1205,361,385]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="37" pageNumber="154">Fig. 215</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E517533F86A16710FD15" blockId="37.[794,1399,425,673]" pageId="37" pageNumber="154">
|
||
<emphasis id="B97AEABEFFE4E517533F86A160F8FE75" bold="true" box="[794,1011,425,449]" pageId="37" pageNumber="154">Comparative notes</emphasis>
|
||
:
|
||
<taxonomicName id="4C0E4D2FFFE4E517542286A267BCFE75" authorityName="Assing" authorityYear="2017" box="[1031,1207,425,449]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="ambulans">
|
||
<emphasis id="B97AEABEFFE4E517542286A267BCFE75" box="[1031,1207,425,449]" italics="true" pageId="37" pageNumber="154">Zyras ambulans</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all the species previously known from
|
||
<collectingCountry id="F319763CFFE4E517553386C1667DFE55" box="[1302,1398,457,481]" name="Thailand" pageId="37" pageNumber="154">Thailand</collectingCountry>
|
||
and Peninsular
|
||
<collectingCountry id="F319763CFFE4E51753F586E16725FDB5" box="[976,1070,489,513]" name="Malaysia" pageId="37" pageNumber="154">Malaysia</collectingCountry>
|
||
by the reduced hind wings and the morphology of the aedeagus (shape of median lobe; modifications of paramere). It is additionally characterized by the coloration (body, legs, and antennae), a slender pronotum, short elytra, and the punctation pattern of the abdomen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE4E517533F85C160F2FC15" pageId="37" pageNumber="154" type="distribution">
|
||
<paragraph id="8BB136ACFFE4E517533F85C160F2FC15" blockId="37.[794,1399,713,929]" pageId="37" pageNumber="154">
|
||
<emphasis id="B97AEABEFFE4E517533F85C16787FD55" bold="true" box="[794,1164,713,737]" pageId="37" pageNumber="154">Distribution and natural history</emphasis>
|
||
: Thale Ban National Park is situated in the extreme south of
|
||
<collectingCountry id="F319763CFFE4E51754D085E1665CFCB5" box="[1269,1367,745,769]" name="Thailand" pageId="37" pageNumber="154">Thailand</collectingCountry>
|
||
at approximately
|
||
<geoCoordinate id="EE3A506BFFE4E51753E684016704FC95" box="[963,1039,777,801]" degrees="6" direction="north" minutes="43" orientation="latitude" pageId="37" pageNumber="154" precision="925" value="6.7166667">6°43'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3A506BFFE4E517540484016789FC95" box="[1057,1154,777,801]" degrees="100" direction="east" minutes="10" orientation="longitude" pageId="37" pageNumber="154" precision="925" value="100.166664">100°10'E</geoCoordinate>
|
||
, close to the border with
|
||
<collectingCountry id="F319763CFFE4E5175372842160BCFCF5" box="[855,951,809,833]" name="Malaysia" pageId="37" pageNumber="154">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="13352A29FFE4E51753E984216729FCF5" box="[972,1058,809,833]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="37" pageNumber="154">Map 10</figureCitation>
|
||
). The
|
||
<typeStatus id="54B5880EFFE4E5175451842167D8FCF5" box="[1140,1235,809,833]" pageId="37" pageNumber="154" type="holotype">holotype</typeStatus>
|
||
was collected at an altitude between 200 and
|
||
<specimenCount id="9D08FD25FFE4E5175451844167B5FCD5" box="[1140,1214,841,865]" pageId="37" pageNumber="154" type="generic">
|
||
<quantity id="4CF69B49FFE4E5175451844167B1FCD5" box="[1140,1210,841,865]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="154" unit="m" value="400.0">400 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Since one of the collectors is a myrmecologist, it may have been found associated with ants.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |