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<document id="1F68CA0E2C9525EA5C9D8172DF694B7B" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE2E5165098811B6283FDD5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) brevilobatus Assing 2017, spec. nov." docType="treatment" docUuid="6C6FB4E9-8A7B-40FB-B1C9-938C01CCD505" docUuidSource="ZooBank" docVersion="11" lastPageNumber="153" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="152" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="D7BD4CD22F23192C80FE34767A0DD6FF">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
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<mods:role id="C335EBB55B68BBB14B8AEB5FAD09B5CD">
<mods:roleTerm id="D93DA5E654CE938B795E8BC2A3105D1D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="162FE8624CC6D9B77D40CC69E6DA419E">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="E4A4B51AF1E5673D6587F4533BCA74DC">text</mods:typeOfResource>
<mods:relatedItem id="ADB586E87ADB5E0BBB0A66F8D147CD24" type="host">
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<mods:title id="5B28F94F2E700DAB5237ED49F2E6BF4B">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="E076A093FDFF89D1AFAAD02D4B9A1C4F">
<mods:date id="D51563ADE45FEF9FA10834DE13991F37">2017</mods:date>
<mods:detail id="65B84BA88AFC19BFE31E4D45C2F31DE0" type="pubDate">
<mods:number id="D7394A8D6E399942FCA1F27FF71257EF">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="734664B6EC7F73E4BAF53937818B8777" type="volume">
<mods:number id="82A7A8A6C9AF6403355F3ED7F65288C9">67</mods:number>
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<mods:detail id="5B5AEB694C048EF23F0E2D55DD826EAA" type="issue">
<mods:number id="4F0C28EFE7C1D1961A91B09C412FF77E">1</mods:number>
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<mods:classification id="42252FC031C16BC97C4D319754A8E5C3">journal article</mods:classification>
<mods:identifier id="D92CEA6603ED5FDD135DE12B33ECB830" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="88BF0885519999587F25C21AA126EF69" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="A140736E61134636918473FEA1B8FB04" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="9AA86A5C489C492DCE3816076C36EB2E" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="9848D741AFA9A337E520AACA63B9C6FB" type="Zenodo-Dep">5742363</mods:identifier>
<mods:identifier id="41BB7DCB51DF5D3BDC0DC4205D6367C4" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03A787BAFFE2E5165098811B6283FDD5" ID-DOI="http://doi.org/10.5281/zenodo.5888481" ID-GBIF-Taxon="192498181" ID-Zenodo-Dep="5888481" LSID="urn:lsid:zoobank.org:act:6C6FB4E9-8A7B-40FB-B1C9-938C01CCD505" httpUri="http://treatment.plazi.org/id/03A787BAFFE2E5165098811B6283FDD5" lastPageId="36" lastPageNumber="153" pageId="35" pageNumber="152">
<subSubSection id="C3146527FFE2E5115098811B61C3F9FB" pageId="35" pageNumber="152" type="nomenclature">
<paragraph id="8BB136ACFFE2E5115098811B6139F99A" blockId="35.[189,712,1555,1647]" box="[189,562,1555,1582]" pageId="35" pageNumber="152">
<heading id="D0F981C0FFE2E5115098811B6139F99A" box="[189,562,1555,1582]" fontSize="12" level="2" pageId="35" pageNumber="152" reason="2">
<taxonomicName id="4C0E4D2FFFE2E5115098811B62C8F99A" ID-CoL="5DDB8" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[189,451,1555,1582]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="brevilobatus" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFE2E5115098811B63F8F99A" box="[189,243,1555,1582]" italics="true" pageId="35" pageNumber="152">Zyras</emphasis>
(
<emphasis id="B97AEABEFFE2E5115124811B623CF99A" box="[257,311,1555,1582]" italics="true" pageId="35" pageNumber="152">Zyras</emphasis>
)
<emphasis id="B97AEABEFFE2E5115160811B62C8F99A" box="[325,451,1555,1582]" italics="true" pageId="35" pageNumber="152">brevilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFE2E51151EC811B6139F99A" box="[457,562,1555,1582]" pageId="35" pageNumber="152" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BB136ACFFE2E5115098813361C3F9FB" blockId="35.[189,712,1555,1647]" box="[189,712,1595,1615]" pageId="35" pageNumber="152">
<uri id="FF9F3AAEFFE2E5115098813361C3F9FB" box="[189,712,1595,1615]" pageId="35" pageNumber="152">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C79FFE2E511514A813361C3F9FB" box="[367,712,1595,1615]" pageId="35" pageNumber="152">6C6FB4E9-8A7B-40FB-B1C9-938C01CCD505</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE2E5115098815362C6F9DB" box="[189,461,1627,1647]" pageId="35" pageNumber="152" type="description">
<paragraph id="8BB136ACFFE2E5115098815362C6F9DB" blockId="35.[189,712,1555,1647]" box="[189,461,1627,1647]" pageId="35" pageNumber="152">
(
<figureCitation id="13352A29FFE2E51150E4815363F1F9DB" box="[193,250,1627,1647]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="35" pageNumber="152">Figs 32</figureCitation>
,
<figureCitation id="13352A29FFE2E51151248153621DF9DB" box="[257,278,1627,1647]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="35" pageNumber="152">68</figureCitation>
,
<figureCitation id="13352A29FFE2E511513881536230F9DB" box="[285,315,1627,1647]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="35" pageNumber="152">102</figureCitation>
,
<figureCitation id="13352A29FFE2E511516781536283F9DB" box="[322,392,1627,1647]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="35" pageNumber="152">203–205</figureCitation>
,
<figureCitation id="13352A29FFE2E51151AB815362C2F9DB" box="[398,457,1627,1647]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="35" pageNumber="152">Map 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE2E51150B2818162AAF835" pageId="35" pageNumber="152" type="materials_examined">
<paragraph id="8BB136ACFFE2E51150B2818162AAF835" blockId="35.[151,757,1673,1921]" pageId="35" pageNumber="152">
<emphasis id="B97AEABEFFE2E51150B28181623FF915" bold="true" box="[151,308,1673,1697]" pageId="35" pageNumber="152">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFE2E5115162818161A5F8F5" ID-GBIF-Occurrence="3450162345" collectingDate="1988-04-27" collectingDateMax="1988-05-06" collectingDateMin="1988-04-27" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.5" location="Thung Yai Wildlife Sanctuary" longLatPrecision="1284" longitude="98.8" municipality="Umphang District" pageId="35" pageNumber="152" specimenCount="1" stateProvince="Tak Province" typeStatus="holotype">
<typeStatus id="54B5880EFFE2E5115162818162A1F915" box="[327,426,1673,1697]" pageId="35" pageNumber="152" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFE2E5115192818262C7F915" box="[439,460,1674,1697]" pageId="35" pageNumber="152">♂</specimenCount>
: “W.
<collectingCountry id="F319763CFFE2E51152218181619BF915" box="[516,656,1673,1697]" name="Thailand" pageId="35" pageNumber="152">THAILAND</collectingCountry>
:
<quantity id="4CF69B49FFE2E51152BA818161E7F915" box="[671,748,1673,1697]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="152" unit="m" value="300.0">
<elevation id="0023D19FFFE2E51152BA818161E7F915" box="[671,748,1673,1697]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="152" unit="m" value="300.0">300 m</elevation>
</quantity>
.,
<location id="8ED16077FFE2E51150B281A162D7F975" LSID="urn:lsid:plazi:treatment:03A787BAFFE2E5165098811B6283FDD5:8ED16077FFE2E51150B281A162D7F975" box="[151,476,1705,1729]" country="Thailand" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Umphang District" name="Thung Yai Wildlife Sanctuary" pageId="35" pageNumber="152" stateProvince="Tak Province">Thung Yai Wildlife Sanctuary</location>
.
<geoCoordinate id="EE3A506BFFE2E51151C281A16137F975" box="[487,572,1705,1729]" degrees="15" direction="north" minutes="30" orientation="latitude" pageId="35" pageNumber="152" precision="925" value="15.5">15°30'N</geoCoordinate>
–
<geoCoordinate id="EE3A506BFFE2E511527981A161A4F975" box="[604,687,1705,1729]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="35" pageNumber="152" precision="925" value="98.8">98°48'E</geoCoordinate>
. /
<collectingRegion id="49CAF84EFFE2E51152E881A163FCF955" country="Thailand" name="Tak" pageId="35" pageNumber="152">Tak Province</collectingRegion>
,
<collectingMunicipality id="6BD5ACD6FFE2E511512281C162DAF955" box="[263,465,1737,1761]" pageId="35" pageNumber="152">Umphang District</collectingMunicipality>
,
<location id="8ED16077FFE2E51151C581C2615FF955" LSID="urn:lsid:plazi:treatment:03A787BAFFE2E5165098811B6283FDD5:8ED16077FFE2E51151C581C2615FF955" box="[480,596,1737,1761]" country="Thailand" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Umphang District" name="Mae Chan" pageId="35" pageNumber="152" stateProvince="Tak Province">Mae Chan</location>
–
<location id="8ED16077FFE2E511525C81C26205F8B5" LSID="urn:lsid:plazi:treatment:03A787BAFFE2E5165098811B6283FDD5:8ED16077FFE2E511525C81C26205F8B5" country="Thailand" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Umphang District" name="Mae Klong confluence" pageId="35" pageNumber="152" stateProvince="Tak Province">Mae Klong confluence</location>
.
<date id="FFB0106CFFE2E511510681E162CAF8B5" box="[291,449,1769,1793]" pageId="35" pageNumber="152" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">
<collectingDate id="EFF4E984FFE2E511510681E162CAF8B5" box="[291,449,1769,1793]" pageId="35" pageNumber="152" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">27.iv.–6.v.1988</collectingDate>
</date>
. / Mixed riverside forest.
<collectorName id="26FB537AFFE2E51150B280026249F895" box="[151,322,1801,1825]" pageId="35" pageNumber="152">M.J.D. Brendell.</collectorName>
B.M. 1988-183 / Holotypus ♂
<taxonomicName id="4C0E4D2FFFE2E511529F8002621DF8F5" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="brevilobatus" status="sp. nov.">
<emphasis id="B97AEABEFFE2E511529F8002621DF8F5" italics="true" pageId="35" pageNumber="152">Zyras brevilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFE2E511510480226253F8F5" box="[289,344,1834,1857]" pageId="35" pageNumber="152" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="7E6DDA68FFE2E51151B1802262FDF8F5" box="[404,502,1833,1857]" pageId="35" pageNumber="152">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFE2E5115275802161ADF8F5" box="[592,678,1833,1857]" pageId="35" pageNumber="152">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFE2E511529E80226297F835" ID-GBIF-Occurrence="3450162317" collectingDate="1988-04-27" collectingDateMax="1988-05-06" collectingDateMin="1988-04-27" collectionCode="BMNH" collectorName="M. J. D. Brendell." country="Thailand" determinerName="V. Assing" elevation="300" latitude="15.5" location="Mae Chan" longLatPrecision="1284" longitude="98.8" municipality="Umphang District" pageId="35" pageNumber="152" specimenCount="1" stateProvince="Tak" typeStatus="paratype">
<typeStatus id="54B5880EFFE2E511529E802263D9F8D6" pageId="35" pageNumber="152" type="paratype">Paratypes</typeStatus>
: 2 ♂♂ [both without aedeagus!]: same data as holotype (
<collectionCode id="ED1FAE69FFE2E51151278061625DF835" box="[258,342,1897,1921]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="35" pageNumber="152">BMNH</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE2E51150B280A161C9F855" pageId="35" pageNumber="152" type="etymology">
<paragraph id="8BB136ACFFE2E51150B280A161C9F855" blockId="35.[151,756,1961,2017]" pageId="35" pageNumber="152">
<emphasis id="B97AEABEFFE2E51150B280A16205F875" bold="true" box="[151,270,1961,1985]" pageId="35" pageNumber="152">Etymology</emphasis>
: The specific epithet (adjective) alludes to the conspicuously short ventral process of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE2E516533F87E162B5FE35" lastPageId="36" lastPageNumber="153" pageId="35" pageNumber="152" type="description">
<paragraph id="8BB136ACFFE2E511533F87E1608DFDF5" blockId="35.[794,1399,233,1985]" pageId="35" pageNumber="152">
<emphasis id="B97AEABEFFE2E511533F87E16094FEB5" bold="true" box="[794,927,233,257]" pageId="35" pageNumber="152">Description</emphasis>
: Body length 6.0–
<quantity id="4CF69B49FFE2E511545587E167CEFEB5" box="[1136,1221,233,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="35" pageNumber="152" unit="mm" value="6.3">6.3 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFE2E511537E860160D4FE95" box="[859,991,265,289]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="35" pageNumber="152" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFE2E5115453860167CCFE95" box="[1142,1223,265,289]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="35" pageNumber="152">Figs 32</figureCitation>
,
<figureCitation id="13352A29FFE2E51154F3860167F9FE95" box="[1238,1266,265,289]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="35" pageNumber="152">68</figureCitation>
,
<figureCitation id="13352A29FFE2E511552486016627FE95" box="[1281,1324,265,289]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="35" pageNumber="152">102</figureCitation>
): head black; pronotum and elytra dark-brown; abdomen with segments II–V pale-reddish, segment VI reddish or brown with reddish margins, segment VII bicoloured with the anterior fourth pale-reddish and the posterior three-fourths blackish-brown, and segments VIII–X blackish-brown; legs pale-yellowish; antennae blackish-brown with antennomeres I–II dark-yellowish and the apex of XI slightly paler brown; maxillary palpi yellowish.
</paragraph>
<paragraph id="8BB136ACFFE2E511533F85416748FC35" blockId="35.[794,1399,233,1985]" pageId="35" pageNumber="152">
Head (
<figureCitation id="13352A29FFE2E5115340854160A4FDD5" box="[869,943,585,609]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="35" pageNumber="152">Fig. 68</figureCitation>
) distinctly transverse, middle and posterior portion extensively impunctate; punctation in lateral and anterior portions sparse and moderately coarse. Eyes large and bulging, approximately twice as long as postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFE2E51154BA85C167E2FD55" box="[1183,1257,713,737]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="35" pageNumber="152">Fig. 32</figureCitation>
) 1.8–2.0 mm long; antennomeres IV–V very weakly oblong, VI approximately as broad as long, VII–IX weakly transverse, X less than 1.5 times as broad as long, and XI longer than the combined length of IX and X, but shorter than the combined length of VIII–X.
</paragraph>
<paragraph id="8BB136ACFFE2E511533F84826789FB35" blockId="35.[794,1399,233,1985]" pageId="35" pageNumber="152">
Pronotum (
<figureCitation id="13352A29FFE2E51153B7848160D7FC15" box="[914,988,905,929]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="35" pageNumber="152">Fig. 68</figureCitation>
) strongly transverse, nearly 1.3 times as broad as long and 1.2 times as broad as head, broadest in anterior half, distinctly tapering posteriad; lateral margins straight in posterior two-thirds (dorsal view); punctation coarse, moderately dense, and slightly irregularly distributed, antero-lateral portions of disc with an impunctate patch on either side; midline narrowly impunctate; lateral margins with four long dark setae.
</paragraph>
<paragraph id="8BB136ACFFE2E511533F83816725FA95" blockId="35.[794,1399,233,1985]" pageId="35" pageNumber="152">
Elytra (
<figureCitation id="13352A29FFE2E5115342838160BAFB15" box="[871,945,1161,1185]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="35" pageNumber="152">Fig. 68</figureCitation>
) approximately 0.9 times as long as pronotum; punctation coarse, moderately dense anteriorly and very sparse posteriorly. Hind wings fully developed. Metatarsomere I as long as, or slightly shorter than the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFE2E511533F8221660DF8F5" blockId="35.[794,1399,233,1985]" pageId="35" pageNumber="152">
Abdomen (
<figureCitation id="13352A29FFE2E51153B0822160E7FAF5" box="[917,1004,1321,1345]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="35" pageNumber="152">Fig. 102</figureCitation>
) narrower than elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a row of moderately coarse non-setiferous punctures; tergite III with a lateral setiferous puncture on either side and with four setiferous punctures at posterior margin; tergites IV–V with a lateral setiferous puncture on either side and with six setiferous punctures at posterior margin; tergite VI with a somewhat irregular transverse band of rather sparse and fine non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, and with six setiferous punctures at posterior margin; tergite VII with sparse and fine non-setiferous punctation in anterior portion and with two transverse rows of setiferous punctures in posterior portion, posterior margin with palisade fringe; tergite VIII with long black setae near posterior margin, the latter with distinct concavity in the middle.
</paragraph>
<paragraph id="8BB136ACFFE2E511533F80426606F875" blockId="35.[794,1399,233,1985]" pageId="35" pageNumber="152">
♂: sternite VIII with convex posterior margin; median lobe of aedeagus (
<figureCitation id="13352A29FFE2E51153C48061677AF835" box="[993,1137,1897,1921]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="35" pageNumber="152">Figs 203–204</figureCitation>
)
<quantity id="4CF69B49FFE2E51154A7806167DFF835" box="[1154,1236,1897,1921]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="152" unit="mm" value="0.8">0.8 mm</quantity>
long and with conspicuously short ventral process; paramere (
<figureCitation id="13352A29FFE2E511553280816664F815" box="[1303,1391,1929,1953]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="35" pageNumber="152">Fig. 205</figureCitation>
)
<quantity id="4CF69B49FFE2E511533F80A16060F875" box="[794,875,1961,1985]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="35" pageNumber="152" unit="mm" value="0.9">0.9 mm</quantity>
long, with moderately long apical lobe.
</paragraph>
<paragraph id="8BB136ACFFE2E516533F80E162B5FE35" blockId="35.[794,1398,2025,2081]" lastBlockId="36.[189,794,233,385]" lastPageId="36" lastPageNumber="153" pageId="35" pageNumber="152">
<emphasis id="B97AEABEFFE2E511533F80E160F1F7B5" bold="true" box="[794,1018,2025,2049]" pageId="35" pageNumber="152">Comparative notes</emphasis>
: This species is characterized particularly by the conspicuously short ventral process of the aedeagus. It additionally differs from other species recorded from
<collectingCountry id="F319763CFFE5E5165151860162DFFE95" box="[372,468,265,289]" name="Thailand" pageId="36" pageNumber="153">Thailand</collectingCountry>
by the strongly transverse pronotum, by the coloration of the abdomen in combination with completely dark elytra, and by the punctation pattern of the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE5E516509886A16283FDD5" pageId="36" pageNumber="153" type="distribution">
<paragraph id="8BB136ACFFE5E516509886A16283FDD5" blockId="36.[189,794,425,609]" pageId="36" pageNumber="153">
<emphasis id="B97AEABEFFE5E516509886A16124FE75" bold="true" box="[189,559,425,449]" pageId="36" pageNumber="153">Distribution and natural history</emphasis>
: The
<typeStatus id="54B5880EFFE5E516525386A261AFFE76" box="[630,676,426,450]" pageId="36" pageNumber="153">type</typeStatus>
locality is situated in West
<collectingCountry id="F319763CFFE5E516514886C162C6FE55" box="[365,461,457,481]" name="Thailand" pageId="36" pageNumber="153">Thailand</collectingCountry>
(
<figureCitation id="13352A29FFE5E51651FE86C1613BFE55" box="[475,560,457,481]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="36" pageNumber="153">Map 10</figureCitation>
). The specimens were collected in a mixed forest near a river at an altitude of
<quantity id="4CF69B49FFE5E51650988501620AFD95" box="[189,257,521,545]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="36" pageNumber="153" unit="m" value="300.0">300 m</quantity>
, most likely with Malaise or flight interception traps, together with numerous specimens of
<taxonomicName id="4C0E4D2FFFE5E51652B985226012FDF5" authorityName="CAMERON" authorityYear="1939" box="[668,793,554,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B97AEABEFFE5E51652B985226012FDF5" box="[668,793,554,577]" italics="true" pageId="36" pageNumber="153">Z. proximus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E4D2FFFE5E51650CE85426289FDD5" authorityName="Assing" authorityYear="2017" box="[235,386,586,609]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="novinversus">
<emphasis id="B97AEABEFFE5E51650CE85426289FDD5" box="[235,386,586,609]" italics="true" pageId="36" pageNumber="153">Z. novinversus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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