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<document id="E9388A431341BC21305625EC25BCA5F3" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFD2E526509887FB617CFED5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) truncatus Assing 2017, spec. nov." docType="treatment" docUuid="D18CCF00-B59C-4826-9750-E0991FFD871E" docUuidSource="ZooBank" docVersion="12" lastPageNumber="137" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="136" updateTime="1699471407760" updateUser="ExternalLinkService">
<mods:mods id="028B7A7D748FF8D865EEF4ED4C5C29FE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6AE76678CFDA50225BE9709F0E76C42F">
<mods:title id="7DC3DE647F1AB7CCBAD7CF8A0D92FAF8">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
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<mods:role id="3755C6256D7A781DCFF7E612FEA4DAD4">
<mods:roleTerm id="EAA1E07148D22D7DA0815E5B7960A7D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2FF7241550391848B6D108F8E6C63D75">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="914C593D525F8DBA83160109BA1C367F">text</mods:typeOfResource>
<mods:relatedItem id="D0BDBBFC0F3D729F9010C41F286CE659" type="host">
<mods:titleInfo id="7AA407D05A69A3CB61CF055260D973DB">
<mods:title id="54A3DD44180A502289A1A0C3090CE9F6">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="04960D817C482486B841DEE8D08F7B34">
<mods:date id="E8BB38F05D470D55D9B4EC739A580C58">2017</mods:date>
<mods:detail id="408AF74097FF5C77906B88C3AFF399B2" type="pubDate">
<mods:number id="7402707080845600B79C721CADFC31E3">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="7D69E7B3168B80FDEEE4041C5DDA1024" type="volume">
<mods:number id="37D03A463F499CEC939AD22E7CECD757">67</mods:number>
</mods:detail>
<mods:detail id="9AC2BDA195F0C7E3F67AC4F185A00E73" type="issue">
<mods:number id="B9FA397E7C67348D0E8366CDEB23C602">1</mods:number>
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<mods:start id="A988E4F87E8B2DAA7256E12390FB7923">117</mods:start>
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AF1E1B8E3D1460E8ACA80CB5F2F8E986">journal article</mods:classification>
<mods:identifier id="E140EE711B66227A038DE1E436A43F04" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="59FD4207D437485F0BFE65D9911DB454" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="7B7F385EDBBF5B5037904BF7311BCF0E" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="9AB007DC79578E0D4109703EB6CCD213" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="8A220DD84203BC62C9132CF4F36CD7EA" type="Zenodo-Dep">5742363</mods:identifier>
<mods:identifier id="7C572FDC88B2744E64C6FA81341C52DE" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03A787BAFFD2E526509887FB617CFED5" ID-DOI="http://doi.org/10.5281/zenodo.5888413" ID-GBIF-Taxon="192498166" ID-Zenodo-Dep="5888413" LSID="urn:lsid:zoobank.org:act:D18CCF00-B59C-4826-9750-E0991FFD871E" httpUri="http://treatment.plazi.org/id/03A787BAFFD2E526509887FB617CFED5" lastPageId="20" lastPageNumber="137" pageId="19" pageNumber="136">
<subSubSection id="C3146527FFD2E521509887FB61CAFE9B" pageId="19" pageNumber="136" type="nomenclature">
<paragraph id="8BB136ACFFD2E521509887FB611DFEBA" blockId="19.[189,705,243,335]" box="[189,534,243,270]" pageId="19" pageNumber="136">
<heading id="D0F981C0FFD2E521509887FB611DFEBA" box="[189,534,243,270]" fontSize="12" level="2" pageId="19" pageNumber="136" reason="2">
<taxonomicName id="4C0E4D2FFFD2E521509887FB62ACFEBA" ID-CoL="5DDGC" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[189,423,243,270]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="truncatus" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFD2E521509887FB63F8FEBA" box="[189,243,243,270]" italics="true" pageId="19" pageNumber="136">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD2E521512487FB623CFEBA" box="[257,311,243,270]" italics="true" pageId="19" pageNumber="136">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD2E521516087FB62ACFEBA" box="[325,423,243,270]" italics="true" pageId="19" pageNumber="136">truncatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFD2E521518887FB611DFEBA" box="[429,534,243,270]" pageId="19" pageNumber="136" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BB136ACFFD2E5215098861361CAFE9B" blockId="19.[189,705,243,335]" box="[189,705,283,303]" pageId="19" pageNumber="136">
<uri id="FF9F3AAEFFD2E5215098861361CAFE9B" box="[189,705,283,303]" pageId="19" pageNumber="136">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C79FFD2E5215148861361CAFE9B" box="[365,705,283,303]" pageId="19" pageNumber="136">D18CCF00-B59C-4826-9750-E0991FFD871E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD2E5215098863362B0FEFB" box="[189,443,315,335]" pageId="19" pageNumber="136" type="description">
<paragraph id="8BB136ACFFD2E5215098863362B0FEFB" blockId="19.[189,705,243,335]" box="[189,443,315,335]" pageId="19" pageNumber="136">
(
<figureCitation id="13352A29FFD2E52150E4863363F1FEFB" box="[193,250,315,335]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="19" pageNumber="136">Figs 11</figureCitation>
,
<figureCitation id="13352A29FFD2E52151248633621DFEFB" box="[257,278,315,335]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="19" pageNumber="136">52</figureCitation>
,
<figureCitation id="13352A29FFD2E521513886336239FEFB" box="[285,306,315,335]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="19" pageNumber="136">91</figureCitation>
,
<figureCitation id="13352A29FFD2E521511C86336276FEFB" box="[313,381,315,335]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="19" pageNumber="136">150–155</figureCitation>
,
<figureCitation id="13352A29FFD2E52151A0863362BCFEFB" box="[389,439,315,335]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="19" pageNumber="136">Map 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD2E52150B28661618BFD35" pageId="19" pageNumber="136" type="materials_examined">
<paragraph id="8BB136ACFFD2E52150B28661618BFD35" blockId="19.[151,757,361,641]" pageId="19" pageNumber="136">
<emphasis id="B97AEABEFFD2E52150B286616227FE35" bold="true" box="[151,300,361,385]" pageId="19" pageNumber="136">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFD2E52151128661629AFE55" ID-GBIF-Occurrence="3450162368" collectingDate="2004-05-25" collectorName="A. Kleeberg" country="Nepal" county="Himalaya" determinerName="V. Assing" elevation="2500" location="Region Parbat" municipality="Region Parbat" pageId="19" pageNumber="136" specimenCount="1" stateProvince="Dhawalagiri" typeStatus="holotype">
<typeStatus id="54B5880EFFD2E521511286616291FE35" box="[311,410,361,385]" pageId="19" pageNumber="136" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFD2E52151BB866262B8FE35" box="[414,435,362,385]" pageId="19" pageNumber="136" type="generic">♂</specimenCount>
: “
<collectingCountry id="F319763CFFD2E52151E386616113FE35" box="[454,536,361,385]" name="Nepal" pageId="19" pageNumber="136">NEPAL</collectingCountry>
<collectingCounty id="62D04E20FFD2E52152398661618EFE35" box="[540,645,361,385]" pageId="19" pageNumber="136">Himalaya</collectingCounty>
,
<collectingRegion id="49CAF84EFFD2E52152A8866163B5FE15" country="Nepal" name="Dhawalagiri" pageId="19" pageNumber="136">Dhawalagiri</collectingRegion>
, 2004,
<location id="8ED16077FFD2E5215135868162A0FE15" LSID="urn:lsid:plazi:treatment:03A787BAFFD2E526509887FB617CFED5:8ED16077FFD2E5215135868162A0FE15" box="[272,427,393,417]" country="Nepal" county="Himalaya" municipality="Region Parbat" name="Region Parbat" pageId="19" pageNumber="136" stateProvince="Dhawalagiri">Region Parbat</location>
/
<location id="8ED16077FFD2E52151ED86816105FE15" LSID="urn:lsid:plazi:treatment:03A787BAFFD2E526509887FB617CFED5:8ED16077FFD2E52151ED86816105FE15" box="[456,526,393,417]" country="Nepal" county="Himalaya" municipality="Region Parbat" name="Chitre" pageId="19" pageNumber="136" stateProvince="Dhawalagiri">Chitre</location>
,
<quantity id="4CF69B49FFD2E521523986816166FE15" box="[540,621,393,417]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="136" unit="m" value="2500.0">
<elevation id="0023D19FFFD2E521523986816166FE15" box="[540,621,393,417]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="136" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="FFB0106CFFD2E521525E868161FBFE15" box="[635,752,393,417]" pageId="19" pageNumber="136" value="2004-05-25">
<collectingDate id="EFF4E984FFD2E521525E868161FBFE15" box="[635,752,393,417]" pageId="19" pageNumber="136" value="2004-05-25">25.05.2004</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFD2E52150E486A16234FE75" box="[193,319,425,449]" pageId="19" pageNumber="136">A. Kleeberg</collectorName>
/ Holotypus ♂
<taxonomicName id="4C0E4D2FFFD2E52151C486A2618FFE75" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[481,644,426,449]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="truncatus" status="sp. nov.">
<emphasis id="B97AEABEFFD2E52151C486A2618FFE75" box="[481,644,426,449]" italics="true" pageId="19" pageNumber="136">Zyras truncatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFD2E52152AC86A261C9FE75" box="[649,706,426,449]" pageId="19" pageNumber="136" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E6DDA68FFD2E52150B286C263F2FE55" box="[151,249,457,481]" pageId="19" pageNumber="136">V. Assing</determinerName>
2016” (cAss)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFD2E52151BB86C26177FD35" ID-GBIF-Occurrence="3450162449" collectingDate="2004-05-24" collectorName="A. Kleeberg" country="Nepal" county="Himalaya" elevation="2400" location="Chitre" municipality="Region Parbat" pageId="19" pageNumber="136" specimenCount="1" stateProvince="Dhawalagiri" typeStatus="paratype">
<typeStatus id="54B5880EFFD2E52151BB86C26103FE56" box="[414,520,458,482]" pageId="19" pageNumber="136" type="paratype">Paratypes</typeStatus>
: 1 ♂: “
<collectingCountry id="F319763CFFD2E521527786C161AFFE55" box="[594,676,457,481]" name="Nepal" pageId="19" pageNumber="136">NEPAL</collectingCountry>
<collectingCounty id="62D04E20FFD2E521528B86C163C8FDB5" pageId="19" pageNumber="136">Himalaya</collectingCounty>
,
<collectingRegion id="49CAF84EFFD2E52150F186E16253FDB5" box="[212,344,489,513]" country="Nepal" name="Dhawalagiri" pageId="19" pageNumber="136">Dhawalagiri</collectingRegion>
, 2004,
<collectingMunicipality id="6BD5ACD6FFD2E521518A86E16147FDB5" box="[431,588,489,513]" pageId="19" pageNumber="136">Region Parbat</collectingMunicipality>
/ near
<location id="8ED16077FFD2E521528F86E161FBFDB5" LSID="urn:lsid:plazi:treatment:03A787BAFFD2E526509887FB617CFED5:8ED16077FFD2E521528F86E161FBFDB5" box="[682,752,489,513]" country="Nepal" county="Himalaya" municipality="Region Parbat" name="Chitre" pageId="19" pageNumber="136" stateProvince="Dhawalagiri">Chitre</location>
,
<location id="8ED16077FFD2E52150B28501626EFD95" LSID="urn:lsid:plazi:treatment:03A787BAFFD2E526509887FB617CFED5:8ED16077FFD2E52150B28501626EFD95" box="[151,357,521,545]" country="Nepal" county="Himalaya" municipality="Region Parbat" name="Ghar Khola valley" pageId="19" pageNumber="136" stateProvince="Dhawalagiri">Ghar Khola valley</location>
, ~
<quantity id="4CF69B49FFD2E52151A7850162DDFD95" box="[386,470,521,545]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="136" unit="m" value="2400.0">
<elevation id="0023D19FFFD2E52151A7850162DDFD95" box="[386,470,521,545]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="136" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<date id="FFB0106CFFD2E52151C385016150FD95" box="[486,603,521,545]" pageId="19" pageNumber="136" value="2004-05-24">
<collectingDate id="EFF4E984FFD2E52151C385016150FD95" box="[486,603,521,545]" pageId="19" pageNumber="136" value="2004-05-24">24.05.2004</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFD2E52152B9850163CCFDF5" pageId="19" pageNumber="136">A. Kleeberg</collectorName>
” (cKle); 1 ♀: “
<collectingCountry id="F319763CFFD2E5215145852162B9FDF5" box="[352,434,553,577]" name="Nepal" pageId="19" pageNumber="136">NEPAL</collectingCountry>
Himalaya,
<collectingRegion id="49CAF84EFFD2E5215209852161BBFDF5" box="[556,688,553,577]" country="Nepal" name="Dhawalagiri" pageId="19" pageNumber="136">Dhawalagiri</collectingRegion>
, 2004, Region Parbat / near Chitre,
<quantity id="4CF69B49FFD2E52151EF85416146FDD5" box="[458,589,585,609]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="19" pageNumber="136" unit="m" value="2.5" valueMax="2.6" valueMin="2.4">
<elevation id="0023D19FFFD2E52151EF85416146FDD5" box="[458,589,585,609]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="19" pageNumber="136" unit="m" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.600 m</elevation>
</quantity>
[sic], 27.05.200 [sic], leave [sic] litter, leg.
<collectorName id="26FB537AFFD2E521518C85616123FD35" box="[425,552,617,641]" pageId="19" pageNumber="136">A. Kleeberg</collectorName>
” (cKle)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD2E52150B285A1615EFD55" pageId="19" pageNumber="136" type="etymology">
<paragraph id="8BB136ACFFD2E52150B285A1615EFD55" blockId="19.[151,756,681,737]" pageId="19" pageNumber="136">
<emphasis id="B97AEABEFFD2E52150B285A16205FD75" bold="true" box="[151,270,681,705]" pageId="19" pageNumber="136">Etymology</emphasis>
: The specific epithet (Latin, adjective) alludes to the apically truncate male sternite VIII.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD2E52150B284016746F7B5" pageId="19" pageNumber="136" type="description">
<paragraph id="8BB136ACFFD2E52150B2840161BAFB35" blockId="19.[151,756,777,2081]" pageId="19" pageNumber="136">
<emphasis id="B97AEABEFFD2E52150B284016212FC95" bold="true" box="[151,281,777,801]" pageId="19" pageNumber="136">Description</emphasis>
: Body length
<quantity id="4CF69B49FFD2E521519A8401614BFC95" box="[447,576,777,801]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.3" metricValueMin="5.6" pageId="19" pageNumber="136" unit="mm" value="5.949999999999999" valueMax="6.3" valueMin="5.6">5.6–6.3 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFD2E52150F984216256FCF5" box="[220,349,809,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="19" pageNumber="136" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFD2E52151DC84216143FCF5" box="[505,584,809,833]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="19" pageNumber="136">Figs 11</figureCitation>
,
<figureCitation id="13352A29FFD2E521527E8421617CFCF5" box="[603,631,809,833]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="19" pageNumber="136">52</figureCitation>
,
<figureCitation id="13352A29FFD2E52152AC842161ADFCF5" box="[649,678,809,833]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="19" pageNumber="136">91</figureCitation>
): head black; pronotum blackish-brown to black; elytra blackish with the humeral portion reddish-yellow and the suture narrowly dark-reddish; abdomen with segments II–V brown, segments VI–VII blackish (except for the partly brown margins), and segments VIII–X dark-brown to blackish-brown; legs dark-yellowish to yellowish-brown; antennae dark-brown with antennomeres I–II and the base of III yellowish-brown and the apical 1–2 antennomeres more or less distinctly pale-brown; maxillary palpi dark-yellowish with palpomere IV paler yellowish.
</paragraph>
<paragraph id="8BB136ACFFD2E52150B28381628DFA35" blockId="19.[151,756,777,2081]" pageId="19" pageNumber="136">
Head (
<figureCitation id="13352A29FFD2E52150FA83816221FB15" box="[223,298,1161,1185]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="19" pageNumber="136">Fig. 52</figureCitation>
) rather weakly transverse, median portion extensively impunctate; punctures in lateral portions very sparse and moderately coarse. Eyes slightly longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFD2E521528783E161E0FAB5" box="[674,747,1257,1281]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="19" pageNumber="136">Fig. 11</figureCitation>
) approximately
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long and very slender; antennomeres IV–IX oblong, X approximately as broad as long, and XI approximately as long as the combined length of IX and X, or nearly so.
</paragraph>
<paragraph id="8BB136ACFFD2E52150B2828262B7F955" blockId="19.[151,756,777,2081]" pageId="19" pageNumber="136">
Pronotum (
<figureCitation id="13352A29FFD2E521513A82816261FA15" box="[287,362,1417,1441]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="19" pageNumber="136">Fig. 52</figureCitation>
) slender, approximately 1.1 times as broad as long and approximately 1.2 times as broad as head, broadest in anterior half, moderately tapering posteriad; lateral margins sinuate in posterior half in dorsal view; punctation rather coarse and very irregularly distributed, arranged in a cluster behind the middle of lateral surface; antero-lateral and postero-lateral portion of lateral surface impunctate or nearly so; midline broadly impunctate; lateral margins each with four long and erect black setae; pubescence of disc pale, thin, rather short, and more or less depressed.
</paragraph>
<paragraph id="8BB136ACFFD2E52150B281E161BBF855" blockId="19.[151,756,777,2081]" pageId="19" pageNumber="136">
Elytra (
<figureCitation id="13352A29FFD2E52150D581E16231F8B5" box="[240,314,1769,1793]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="19" pageNumber="136">Fig. 52</figureCitation>
) approximately 0.85 times as long as pronotum; punctation moderately coarse and moderately irregularly distributed, rather dense anteriorly, gradually becoming sparser posteriorly, and sparse to very sparse near posterior margin; disc with moderately long, fine, and depressed pale pubescence. Hind wings fully developed. Metatarsomere I approximately as long as, or slightly shorter than the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFD2E52150B280E16766FD75" blockId="19.[151,756,777,2081]" lastBlockId="19.[794,1399,233,1025]" pageId="19" pageNumber="136">
Abdomen (
<figureCitation id="13352A29FFD2E521513080E1626AF7B5" box="[277,353,2025,2049]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="19" pageNumber="136">Fig. 91</figureCitation>
) slightly narrower than elytra, with rather deep anterior impressions on tergites III–V; tergite II impunctate; anterior impressions of tergites III–V and anterior portion of tergite VI each with a transverse row of coarse, but weakly delimited pits of non-setiferous punctures; tergite III with a lateral setiferous puncture on either side and with a transverse row of six setiferous punctures near posterior margin; tergites IV–VI with a lateral setiferous puncture on either side and with 8–10 setiferous punctures at or near posterior margins; tergite VII with scattered fine nonsetiferous punctures anteriorly and with two irregular transverse series of few setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFD2E5215304856160B1FD35" box="[801,954,617,641]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="19" pageNumber="136">Figs 152, 154</figureCitation>
) with moderately dense dark setae in posterior third, without sexual dimorphism, posterior margin with distinct concavity.
</paragraph>
<paragraph id="8BB136ACFFD2E521533F85C26610FC15" blockId="19.[794,1399,233,1025]" pageId="19" pageNumber="136">
♂: sternite VIII (
<figureCitation id="13352A29FFD2E52153EA85C1672CFD55" box="[975,1063,713,737]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="19" pageNumber="136">Fig. 153</figureCitation>
) nearly as long as broad, much longer than tergite VIII, strongly tapering posteriorly and with distinctly truncate posterior margin; median lobe of aedeagus (
<figureCitation id="13352A29FFD2E521538F84216730FCF5" box="[938,1083,809,833]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="19" pageNumber="136">Figs 150–151</figureCitation>
)
<quantity id="4CF69B49FFD2E521546E842167EFFCF5" box="[1099,1252,809,833]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="19" pageNumber="136" unit="mm" value="0.725" valueMax="0.75" valueMin="0.7">0.70–0.75 mm</quantity>
long; ventral process slender, subapically very slender, and apically very acute in lateral view; paramere approximately
<quantity id="4CF69B49FFD2E521533F84816073FC15" box="[794,888,905,929]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="19" pageNumber="136" unit="mm" value="0.85">0.85 mm</quantity>
long, apical lobe very long and slender.
</paragraph>
<paragraph id="8BB136ACFFD2E521533F84A260A4FBB5" blockId="19.[794,1399,233,1025]" pageId="19" pageNumber="136">
♀: sternite VIII (
<figureCitation id="13352A29FFD2E52153EB84A1672EFC75" box="[974,1061,937,961]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="19" pageNumber="136">Fig. 155</figureCitation>
) much shorter and more transverse than in male, posterior margin indistinctly concave in the middle.
</paragraph>
<paragraph id="8BB136ACFFD2E521533F83216602F875" blockId="19.[794,1399,1065,2049]" pageId="19" pageNumber="136">
<emphasis id="B97AEABEFFD2E521533F832160FBFBF5" bold="true" box="[794,1008,1065,1089]" pageId="19" pageNumber="136">Comparative notes</emphasis>
: Based on several derived character conditions, particularly the slender body, long and slender antennae, a conspicuously irregular distribution of the pronotal punctation, irregularly punctate elytra, the punctation pattern of the pronotum, a posteriorly strongly tapering male sternite VIII with a distinctly truncate posterior margin, an apically very slender ventral process of the aedeagus, and a conspicuously long apical lobe of the paramere,
<taxonomicName id="4C0E4D2FFFD2E521546B822267C5FAF5" authorityName="Assing" authorityYear="2017" box="[1102,1230,1322,1345]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B97AEABEFFD2E521546B822267C5FAF5" box="[1102,1230,1322,1345]" italics="true" pageId="19" pageNumber="136">Z. truncatus</emphasis>
</taxonomicName>
is undoubtedly very closely related to
<taxonomicName id="4C0E4D2FFFD2E521542A8242667DFAD5" authority="SCHEERPELTZ, 1965" authorityName="SCHEERPELTZ" authorityYear="1965" box="[1039,1398,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B97AEABEFFD2E521542A82426790FAD5" box="[1039,1179,1353,1377]" italics="true" pageId="19" pageNumber="136">Z. glabricollis</emphasis>
<bibRefCitation id="EF9F4B5DFFD2E52154868241667DFAD5" author="SCHEERPELTZ, O." box="[1187,1398,1353,1377]" pageId="19" pageNumber="136" pagination="93 - 371" refId="ref59167" refString="SCHEERPELTZ, O. 1965: Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). - Arkiv for Zoologie 17 (2): 93 - 371." type="journal article" year="1965">SCHEERPELTZ, 1965</bibRefCitation>
</taxonomicName>
from Northeast
<collectingCountry id="F319763CFFD2E52153EF82626739FA36" box="[970,1074,1386,1410]" name="Myanmar" pageId="19" pageNumber="136">Myanmar</collectingCountry>
and related species (
<taxonomicName id="4C0E4D2FFFD2E521553C826260DAFA15" authority="ASSING, 2016" authorityName="ASSING" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="extensus">
<emphasis id="B97AEABEFFD2E521553C82626031FA15" italics="true" pageId="19" pageNumber="136">Z. extensus</emphasis>
ASSING, 2016
</taxonomicName>
and
<taxonomicName id="4C0E4D2FFFD2E521542C828267F7FA15" authority="ASSING, 2016" authorityName="ASSING" authorityYear="2016" box="[1033,1276,1417,1441]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="B97AEABEFFD2E521542C82826769FA15" box="[1033,1122,1418,1441]" italics="true" pageId="19" pageNumber="136">Z. rectus</emphasis>
ASSING, 2016
</taxonomicName>
, both from Northwest
<collectingRegion id="49CAF84EFFD2E52153B682A260E2FA75" box="[915,1001,1450,1473]" country="China" name="Yunnan" pageId="19" pageNumber="136">Yunnan</collectingRegion>
,
<collectingCountry id="F319763CFFD2E52153D382A16737FA75" box="[1014,1084,1449,1473]" name="China" pageId="19" pageNumber="136">China</collectingCountry>
). It is distinguished from all these species by a more slender pronotum with (more) distinctly sinuate lateral margins in posterior half and additionally as follows: from
<taxonomicName id="4C0E4D2FFFD2E521537F812260EDF9F5" authorityName="SCHEERPELTZ" authorityYear="1965" box="[858,998,1577,1601]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B97AEABEFFD2E521537F812260EDF9F5" box="[858,998,1577,1601]" italics="true" pageId="19" pageNumber="136">Z. glabricollis</emphasis>
</taxonomicName>
by more extensively blackish elytra and by a longer median lobe of the aedeagus with a less strongly curved ventral process in lateral view; from
<taxonomicName id="4C0E4D2FFFD2E5215371818260C3F915" authorityName="ASSING" authorityYear="2016" box="[852,968,1674,1697]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="extensus">
<emphasis id="B97AEABEFFD2E5215371818260C3F915" box="[852,968,1674,1697]" italics="true" pageId="19" pageNumber="136">Z. extensus</emphasis>
</taxonomicName>
by denser punctation of the more extensively blackish elytra and by a shorter median lobe of the aedeagus with an apically shorter and less slender ventral process of the aedeagus; from
<taxonomicName id="4C0E4D2FFFD2E5215345800260B2F895" authorityName="ASSING" authorityYear="2016" box="[864,953,1802,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="B97AEABEFFD2E5215345800260B2F895" box="[864,953,1802,1825]" italics="true" pageId="19" pageNumber="136">Z. rectus</emphasis>
</taxonomicName>
by more slender antennae (
<taxonomicName id="4C0E4D2FFFD2E52155328002667BF895" authorityName="ASSING" authorityYear="2016" box="[1303,1392,1802,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="B97AEABEFFD2E52155328002667BF895" box="[1303,1392,1802,1825]" italics="true" pageId="19" pageNumber="136">Z. rectus</emphasis>
</taxonomicName>
: antennomere IX not oblong), more densely and less irregularly distributed punctation of the elytra, a posteriorly distinctly truncate male sternite VIII, and an aedeagus with an apically more slender ventral process (lateral view) and with a less prominent crista apicalis.
</paragraph>
<paragraph id="8BB136ACFFD2E521533F80C26746F7B5" blockId="19.[794,1399,1065,2049]" pageId="19" pageNumber="136">
For illustrations of
<taxonomicName id="4C0E4D2FFFD2E521543380C267A9F855" authorityName="SCHEERPELTZ" authorityYear="1965" box="[1046,1186,1993,2017]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B97AEABEFFD2E521543380C267A9F855" box="[1046,1186,1993,2017]" italics="true" pageId="19" pageNumber="136">Z. glabricollis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0E4D2FFFD2E521549B80C26639F855" authorityName="ASSING" authorityYear="2016" box="[1214,1330,1994,2017]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="extensus">
<emphasis id="B97AEABEFFD2E521549B80C26639F855" box="[1214,1330,1994,2017]" italics="true" pageId="19" pageNumber="136">Z. extensus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C0E4D2FFFD2E521533F80E26078F7B5" authorityName="ASSING" authorityYear="2016" box="[794,883,2026,2049]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="B97AEABEFFD2E521533F80E26078F7B5" box="[794,883,2026,2049]" italics="true" pageId="19" pageNumber="136">Z. rectus</emphasis>
</taxonomicName>
see
<bibRefCitation id="EF9F4B5DFFD2E521538580E16742F7B5" author="ASSING, V." box="[928,1097,2025,2049]" pageId="19" pageNumber="136" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD5E526509887E1617CFED5" pageId="20" pageNumber="137" type="distribution">
<paragraph id="8BB136ACFFD5E526509887E1617CFED5" blockId="20.[189,794,233,353]" pageId="20" pageNumber="137">
<emphasis id="B97AEABEFFD5E526509887E16120FEB5" bold="true" box="[189,555,233,257]" pageId="20" pageNumber="137">Distribution and natural history</emphasis>
: The specimens were collected in three close localities near Chitre in the
<collectingRegion id="49CAF84EFFD5E52650988621623BFEF5" box="[189,304,297,321]" country="Nepal" name="Dhawalagiri" pageId="20" pageNumber="137">Dhaulagiri</collectingRegion>
range, Central
<collectingCountry id="F319763CFFD5E52651CC86216123FEF5" box="[489,552,297,321]" name="Nepal" pageId="20" pageNumber="137">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFD5E526521886216188FEF5" box="[573,643,297,321]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="20" pageNumber="137">Map 3</figureCitation>
), at altitudes between 2400 and approximately
<quantity id="4CF69B49FFD5E526520486416179FED5" box="[545,626,329,353]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="137" unit="m" value="2500.0">2500 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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