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<document id="3D83998DC1B06A182A146403E402487A" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFD1E52250C6835B6738F855" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) nigroaeneus Cameron 1939" docType="treatment" docVersion="12" lastPageNumber="133" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="133" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="C2B6E9CA599F53B5BD9899CA8C24BD1D">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="8DEA29F5F4D19C8C993C433A20E7A827">Assing, Volker</mods:namePart>
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<mods:title id="534C44384DC500F3E1429F410825F642">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="A45628DC52D7C9056C16D231F741CF9E">2017</mods:date>
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<mods:number id="38C2332376C87E1A71FC6FAE8FC72CF5">2017-06-30</mods:number>
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<mods:number id="0B14A6049A45A54582B4724AD973F2BD">67</mods:number>
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<mods:identifier id="304EF2028F483C5055302FA19E3AB80B" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
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<treatment id="03A787BAFFD1E52250C6835B6738F855" ID-DOI="http://doi.org/10.5281/zenodo.5888411" ID-GBIF-Taxon="192498165" ID-Zenodo-Dep="5888411" LSID="urn:lsid:plazi:treatment:03A787BAFFD1E52250C6835B6738F855" httpUri="http://treatment.plazi.org/id/03A787BAFFD1E52250C6835B6738F855" lastPageNumber="133" pageId="16" pageNumber="133">
<subSubSection id="C3146527FFD1E52250C6835B619FFBDA" box="[227,660,1107,1134]" pageId="16" pageNumber="133" type="nomenclature">
<paragraph id="8BB136ACFFD1E52250C6835B619FFBDA" blockId="16.[227,660,1107,1167]" box="[227,660,1107,1134]" pageId="16" pageNumber="133">
<heading id="D0F981C0FFD1E52250C6835B619FFBDA" box="[227,660,1107,1134]" centered="true" fontSize="12" level="2" pageId="16" pageNumber="133" reason="2">
<taxonomicName id="4C0E4D2FFFD1E52250C6835B619FFBDA" ID-CoL="7GHYR" authority="CAMERON, 1939" authorityName="Cameron" authorityYear="1939" box="[227,660,1107,1134]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="nigroaeneus" subGenus="Zyras">
<emphasis id="B97AEABEFFD1E52250C6835B6212FBDA" box="[227,281,1107,1134]" italics="true" pageId="16" pageNumber="133">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD1E5225102835B6256FBDA" box="[295,349,1107,1134]" italics="true" pageId="16" pageNumber="133">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD1E522514F835B62E2FBDA" box="[362,489,1107,1134]" italics="true" pageId="16" pageNumber="133">nigroaeneus</emphasis>
CAMERON, 1939
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD1E52250C6837362EAFB3B" box="[227,481,1147,1167]" pageId="16" pageNumber="133" type="description">
<paragraph id="8BB136ACFFD1E52250C6837362EAFB3B" blockId="16.[227,660,1107,1167]" box="[227,481,1147,1167]" pageId="16" pageNumber="133">
(
<figureCitation id="13352A29FFD1E52250C28373622BFB3B" box="[231,288,1147,1167]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="16" pageNumber="133">Figs 22</figureCitation>
,
<figureCitation id="13352A29FFD1E522510283736237FB3B" box="[295,316,1147,1167]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="16" pageNumber="133">50</figureCitation>
,
<figureCitation id="13352A29FFD1E522516683736253FB3B" box="[323,344,1147,1167]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="16" pageNumber="133">81</figureCitation>
,
<figureCitation id="13352A29FFD1E522517A837362A8FB3B" box="[351,419,1147,1167]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="16" pageNumber="133">140–141</figureCitation>
,
<figureCitation id="13352A29FFD1E522518F837362D7FB3B" box="[426,476,1147,1167]" captionStart="Map 2" captionStartId="64.[189,234,1456,1477]" captionTargetBox="[189,1436,891,1446]" captionTargetId="figure-96@64.[189,1436,891,1446]" captionTargetPageId="64" captionText="Map 2: Revised distributions of Zyras pindarae (black circles) and Z. nigroaeneus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742395" httpUri="https://zenodo.org/record/5742395/files/figure.png" pageId="16" pageNumber="133">Map 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD1E522509883A46196FB75" box="[189,669,1195,1217]" pageId="16" pageNumber="133" type="reference_group">
<paragraph id="8BB136ACFFD1E522509883A46196FB75" blockId="16.[189,669,1195,1217]" box="[189,669,1195,1217]" pageId="16" pageNumber="133">
<treatmentCitationGroup id="AB1E1182FFD1E522509883A46196FB75" box="[189,669,1195,1217]" pageId="16" pageNumber="133">
<treatmentCitation id="0AAF10BDFFD1E522509883A46180FB75" author="CAMERON, M." box="[189,651,1195,1217]" page="543" pageId="16" pageNumber="133" year="1939">
<taxonomicName id="4C0E4D2FFFD1E522509883A46180FB75" ID-CoL="7GHYR" authority="CAMERON, 1939 a: 543" authorityName="Cameron" authorityPageNumber="543" authorityYear="1939" box="[189,651,1195,1217]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="nigroaeneus" subGenus="Zyras">
<emphasis id="B97AEABEFFD1E522509883A463FAFB75" box="[189,241,1196,1217]" italics="true" pageId="16" pageNumber="133">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD1E52250DA83A46238FB75" box="[255,307,1196,1217]" italics="true" pageId="16" pageNumber="133">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD1E522516483A462B9FB75" box="[321,434,1196,1217]" italics="true" pageId="16" pageNumber="133">nigroaeneus</emphasis>
<bibRefCitation id="EF9F4B5DFFD1E522519283A36180FB75" author="CAMERON, M." box="[439,651,1195,1217]" pageId="16" pageNumber="133" refId="ref57482" refString="CAMERON, M. 1939 a: Coleoptera: Staphylinidae. Volume IV Parts I-II. - Sewell, R. B. S. (ed): The Fauna of British India, including Ceylon and Burma. - London: 691 pp." type="book" year="1939">
CAMERON, 1939a:
<specimenCount id="9D08FD25FFD1E522524D83A36180FB75" box="[616,651,1195,1217]" pageId="16" pageNumber="133" type="generic">543</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD1E522509883E16266FA15" pageId="16" pageNumber="133" type="materials_examined">
<paragraph id="8BB136ACFFD1E522509883E16266FA15" blockId="16.[189,794,1257,1441]" pageId="16" pageNumber="133">
<emphasis id="B97AEABEFFD1E522509883E162FBFAB5" bold="true" box="[189,496,1257,1281]" pageId="16" pageNumber="133">Type material examined</emphasis>
:
<materialsCitation id="3B663CF1FFD1E522522A83E16262FA15" ID-GBIF-Occurrence="3450162393" collectingDate="1921-09-14" collectionCode="BMNH" collectorName="Dr. Cameron. &amp; V. Assing" location="Simla Hills" pageId="16" pageNumber="133" specimenCount="1" stateProvince="Chin" typeStatus="syntype">
<typeStatus id="54B5880EFFD1E522522A83E1616EFAB5" box="[527,613,1257,1281]" pageId="16" pageNumber="133" type="syntype">Syntype</typeStatus>
<specimenCount id="9D08FD25FFD1E522525A83E2619FFAB5" box="[639,660,1258,1281]" pageId="16" pageNumber="133" type="generic">♂</specimenCount>
[aedeagus missing]: “Narkanda 9230',
<location id="8ED16077FFD1E522522382016183FA95" LSID="urn:lsid:plazi:treatment:03A787BAFFD1E52250C6835B6738F855:8ED16077FFD1E522522382016183FA95" box="[518,648,1289,1313]" name="Simla Hills" pageId="16" pageNumber="133" stateProvince="Chin">Simla Hills</location>
/ Fungus /
<collectorName id="26FB537AFFD1E522509882226258FAF5" box="[189,339,1321,1345]" pageId="16" pageNumber="133">Dr. Cameron.</collectorName>
<date id="FFB0106CFFD1E5225147822162D7FAF5" box="[354,476,1321,1345]" pageId="16" pageNumber="133" value="1921-09-14">
<collectingDate id="EFF4E984FFD1E5225147822162D7FAF5" box="[354,476,1321,1345]" pageId="16" pageNumber="133" value="1921-09-14">14-IX.1921</collectingDate>
</date>
/
<taxonomicName id="4C0E4D2FFFD1E52252208222619EFAF5" box="[517,661,1322,1345]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="nigraeneus">
<emphasis id="B97AEABEFFD1E52252208222619EFAF5" box="[517,661,1322,1345]" italics="true" pageId="16" pageNumber="133">Z. nigraeneus</emphasis>
</taxonomicName>
[sic] Cam Type / M. Cameron. Bequest. B.M. 1955-147 / Holotype / Syntypus
<taxonomicName id="4C0E4D2FFFD1E52251188262617DFA35" authority="Cameron" authorityName="Cameron" authorityYear="1939" box="[317,630,1385,1409]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B97AEABEFFD1E522511882626108FA35" box="[317,515,1386,1409]" italics="true" pageId="16" pageNumber="133">Zyras nigroaeneus</emphasis>
Cameron
</taxonomicName>
, rev.
<collectorName id="26FB537AFFD1E522529082626011FA35" box="[693,794,1385,1409]" pageId="16" pageNumber="133">V. Assing</collectorName>
2016” (
<collectionCode id="ED1FAE69FFD1E522512F8281626AFA15" box="[266,353,1417,1441]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="16" pageNumber="133">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD1E522509882C161CFF975" pageId="16" pageNumber="133" type="discussion">
<paragraph id="8BB136ACFFD1E522509882C1629DF9D5" blockId="16.[189,794,1481,1633]" pageId="16" pageNumber="133">
<emphasis id="B97AEABEFFD1E522509882C16227FA55" bold="true" box="[189,300,1481,1505]" pageId="16" pageNumber="133">Comment</emphasis>
: The original description is based on an unspecified number of
<typeStatus id="54B5880EFFD1E52251EA82E26120F9B6" box="[463,555,1514,1538]" pageId="16" pageNumber="133" type="syntype">syntypes</typeStatus>
from “Simla Hills: Narkanda, alt.
<quantity id="4CF69B49FFD1E5225144810162CCF995" box="[353,455,1545,1569]" metricMagnitude="3" metricUnit="m" metricValue="2.813304" pageId="16" pageNumber="133" unit="ft" value="9230.0">9230 feet</quantity>
” (
<bibRefCitation id="EF9F4B5DFFD1E52251C681016195F995" author="CAMERON, M." box="[483,670,1545,1569]" pageId="16" pageNumber="133" refId="ref57482" refString="CAMERON, M. 1939 a: Coleoptera: Staphylinidae. Volume IV Parts I-II. - Sewell, R. B. S. (ed): The Fauna of British India, including Ceylon and Burma. - London: 691 pp." type="book" year="1939">CAMERON 1939a</bibRefCitation>
). Only
<specimenCount id="9D08FD25FFD1E52252D68102621AF9F6" pageId="16" pageNumber="133" type="generic" typeStatus="syntype">one syntype</specimenCount>
, a male without aedeagus, was located in the Cameron collection.
</paragraph>
<paragraph id="8BB136ACFFD1E5225098818161CFF975" blockId="16.[189,794,1673,1729]" pageId="16" pageNumber="133">
<emphasis id="B97AEABEFFD1E522509881816108F915" bold="true" box="[189,515,1673,1697]" pageId="16" pageNumber="133">Additional material examined</emphasis>
:
<emphasis id="B97AEABEFFD1E52252288181614CF915" bold="true" box="[525,583,1673,1697]" pageId="16" pageNumber="133">
<collectingCountry id="F319763CFFD1E52252288181614CF915" box="[525,583,1673,1697]" name="India" pageId="16" pageNumber="133">India</collectingCountry>
</emphasis>
: 1 ♀,
<collectingRegion id="49CAF84EFFD1E52252A48182601DF915" box="[641,790,1673,1697]" country="India" name="Uttar Pradesh" pageId="16" pageNumber="133">Uttar Pradesh</collectingRegion>
, Nainital Div., Kumaon, leg. Champion (BMNH).
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD1E522509881E160A0F895" pageId="16" pageNumber="133" type="description">
<paragraph id="8BB136ACFFD1E522509881E1622CF855" blockId="16.[189,794,1769,2081]" pageId="16" pageNumber="133">
<emphasis id="B97AEABEFFD1E522509881E1625DF8B5" bold="true" box="[189,342,1769,1793]" pageId="16" pageNumber="133">Redescription</emphasis>
: Species of slender habitus. Body length 5.1–6.0 mm; length of forebody
<quantity id="4CF69B49FFD1E522523C80016191F895" box="[537,666,1801,1825]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="16" pageNumber="133" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFD1E52250E18021621FF8F5" box="[196,276,1833,1857]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="16" pageNumber="133">Figs 22</figureCitation>
,
<figureCitation id="13352A29FFD1E522513B80216231F8F5" box="[286,314,1833,1857]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="16" pageNumber="133">50</figureCitation>
,
<figureCitation id="13352A29FFD1E52251618021626BF8F5" box="[324,352,1833,1857]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="16" pageNumber="133">81</figureCitation>
): whole body black, forebody with a faint blueish hue; legs bicoloured: femora blackish-brown, tibiae and tarsi pale-yellowish; antennae blackish-brown with antennomeres I–II and the base of III yellowish; maxillary palpi brown, with the terminal palpomere yellowish.
</paragraph>
<paragraph id="8BB136ACFFD1E522509880E16777FE75" blockId="16.[189,794,1769,2081]" lastBlockId="16.[831,1437,233,1633]" pageId="16" pageNumber="133">
Head (
<figureCitation id="13352A29FFD1E522512F80E1625EF7B5" box="[266,341,2025,2049]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="16" pageNumber="133">Fig. 50</figureCitation>
) moderately transverse, median portion extensively impunctate; punctures in lateral portions sparse and coarse. Eyes longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFD1E522547486016790FE95" box="[1105,1179,265,289]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="16" pageNumber="133">Fig. 22</figureCitation>
) approximately
<quantity id="4CF69B49FFD1E522556E86016696FE95" box="[1355,1437,265,289]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="133" unit="mm" value="1.8">1.8 mm</quantity>
long and moderately slender; antennomeres IV–V approximately as long as broad, VI–X increasingly transverse and of gradually increasing width, X approximately 1.5 times as broad as long, and XI barely as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFD1E522531A86C26611FD15" blockId="16.[831,1437,233,1633]" pageId="16" pageNumber="133">
Pronotum (
<figureCitation id="13352A29FFD1E522539886C16703FE55" box="[957,1032,457,481]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="16" pageNumber="133">Fig. 50</figureCitation>
) approximately 1.1 times as broad as long and approximately 1.25 times as broad as head, not distinctly tapering posteriad, broadest approximately in the middle; lateral margins weakly convex in dorsal view; punctation coarse, sparse, and very irregularly distributed, with rather large impunctate areas on either side of midline; midline broadly impunctate.
</paragraph>
<paragraph id="8BB136ACFFD1E522531A85A167B9FCD5" blockId="16.[831,1437,233,1633]" pageId="16" pageNumber="133">
Elytra (
<figureCitation id="13352A29FFD1E52253B185A160D4FD75" box="[916,991,681,705]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="16" pageNumber="133">Fig. 50</figureCitation>
) 0.86–0.93 times as long as pronotum; punctation moderately coarse, defined, rather sparse, and regularly distributed; interstices on average approximately three times as broad as diameter of punctures. Hind wings fully developed. Metatarsomere I shorter than the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFD1E522531A846160B2FA55" blockId="16.[831,1437,233,1633]" pageId="16" pageNumber="133">
Abdomen (
<figureCitation id="13352A29FFD1E52253928461670AFC35" box="[951,1025,873,897]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="16" pageNumber="133">Fig. 81</figureCitation>
) narrower than elytra, gradually tapering from base to apex, with sharp paratergites; anterior impressions on tergites III–V rather shallow; anterior impressions of tergites III–V each with a transverse row of sparse and fine non-setiferous punctures; tergites III–IV with a lateral setiferous puncture on either side and with four setiferous punctures at posterior margin; tergite V with a lateral setiferous puncture on either side and with six setiferous punctures at posterior margin; tergite VI with a transverse row of non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, with a median pair of setiferous punctures in posterior portion, and with six setiferous punctures at posterior margin; tergite VII with a transverse band of sparse and fine non-setiferous punctures anteriorly and with scattered setiferous punctures in posterior portion, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFD1E522552082616657FA35" box="[1285,1372,1385,1409]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="16" pageNumber="133">Fig. 140</figureCitation>
) with sparse setiferous punctures only in posterior portion, posterior margin convex and with distinct concavity in the middle.
</paragraph>
<paragraph id="8BB136ACFFD1E522531A82E2666BF995" blockId="16.[831,1437,233,1633]" pageId="16" pageNumber="133">
♂: sternite VIII (
<figureCitation id="13352A29FFD1E52253D682E16740F9B5" box="[1011,1099,1513,1537]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="16" pageNumber="133">Fig. 141</figureCitation>
) much longer than tergite VIII, posteriorly strongly convex; aedeagus not available.
</paragraph>
<paragraph id="8BB136ACFFD1E5225365812260B2F9D5" blockId="16.[831,1437,233,1633]" pageId="16" pageNumber="133">♀: posterior margin of sternite VIII shallowly concave in the middle.</paragraph>
<paragraph id="8BB136ACFFD1E5225365818160A0F895" blockId="16.[832,1436,1673,1825]" pageId="16" pageNumber="133">
<emphasis id="B97AEABEFFD1E522536581816711F915" bold="true" box="[832,1050,1673,1697]" pageId="16" pageNumber="133">Comparative notes</emphasis>
: Among its geographically close congeners,
<taxonomicName id="4C0E4D2FFFD1E522539A81A26753F975" authorityName="Cameron" authorityYear="1939" box="[959,1112,1706,1729]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B97AEABEFFD1E522539A81A26753F975" box="[959,1112,1706,1729]" italics="true" pageId="16" pageNumber="133">Z. nigroaeneus</emphasis>
</taxonomicName>
is characterized particularly by its slender habitus, the conspicuous coloration of the body and the legs, and by the punctation pattern of the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD1E522536580416738F855" pageId="16" pageNumber="133" type="distribution">
<paragraph id="8BB136ACFFD1E522536580416738F855" blockId="16.[832,1436,1865,2017]" pageId="16" pageNumber="133">
<emphasis id="B97AEABEFFD1E5225365804167C6F8D5" bold="true" box="[832,1229,1865,1889]" pageId="16" pageNumber="133">Distribution and natural history</emphasis>
: This species is currently known only from its type locality in
<collectingRegion id="49CAF84EFFD1E52255118061609DF815" country="India" name="Himachal Pradesh" pageId="16" pageNumber="133">Himachal Pradesh</collectingRegion>
and from one locality in
<collectingRegion id="49CAF84EFFD1E522549080826645F815" box="[1205,1358,1929,1953]" country="India" name="Uttar Pradesh" pageId="16" pageNumber="133">Uttar Pradesh</collectingRegion>
, North
<collectingCountry id="F319763CFFD1E522536580A16073F875" box="[832,888,1961,1985]" name="India" pageId="16" pageNumber="133">India</collectingCountry>
(
<figureCitation id="13352A29FFD1E52253A080A160C7F875" box="[901,972,1961,1985]" captionStart="Map 2" captionStartId="64.[189,234,1456,1477]" captionTargetBox="[189,1436,891,1446]" captionTargetId="figure-96@64.[189,1436,891,1446]" captionTargetPageId="64" captionText="Map 2: Revised distributions of Zyras pindarae (black circles) and Z. nigroaeneus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742395" httpUri="https://zenodo.org/record/5742395/files/figure.png" pageId="16" pageNumber="133">Map 2</figureCitation>
). The
<typeStatus id="54B5880EFFD1E522542B80A2676BF876" box="[1038,1120,1962,1986]" pageId="16" pageNumber="133" type="syntype">syntype</typeStatus>
was collected at an altitude of approximately
<quantity id="4CF69B49FFD1E52253FB80C16724F855" box="[990,1071,1993,2017]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="133" unit="m" value="2800.0">2800 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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