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<document id="C6E85F7758F96729AA0C09E90F182F1B" ID-DOI="10.11646/zootaxa.5343.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8334181" ID-ZooBank="BB852CA9-6C6E-4016-9972-AAF2C378E41F" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1694429699348" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2023" docId="03A487CEFFFDFFDF85A551906823F861" docLanguage="en" docName="zootaxa.5343.3.2.pdf" docOrigin="Zootaxa 5343 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Securiops) primasia Kluge 2023, comb. n." docType="treatment" docVersion="1" lastPageNumber="270" masterDocId="FF9DFFB6FFE8FFC4853251076B7DFFDC" masterDocTitle="Redescription of the subgenus Securiops Jacobus, McCafferty &amp; Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)" masterLastPageNumber="272" masterPageNumber="243" pageNumber="264" updateTime="1694453124034" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods id="145B0BD6AC015EA8711E551D8BDD536D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="41A581B0D4148780AA1107F8477B437A">Redescription of the subgenus Securiops Jacobus, McCafferty &amp; Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="C39E34FF4AD6CFB4567AC360E35EFD6E">Kluge, Nikita J.</mods:namePart>
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<mods:title id="D5E31C318FD65BE5C05F8AB871AA03DD">Zootaxa</mods:title>
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<mods:part id="1597B3C9A0FDEAF6C84F165F7A3AA10E">
<mods:date id="9F288F7E709387B4AA5C3393E9E6F7E1">2023</mods:date>
<mods:detail id="C11B12D7AE0656CD2036C75CB42C8C9B" type="pubDate">
<mods:number id="89122DFCE3F9A8D893885B0BECBC78AC">2023-09-08</mods:number>
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<mods:number id="718A2F1EBE744662CB108393F07E8F48">5343</mods:number>
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<mods:number id="5754D54DCDFA8BD501394AD5773ACCD8">3</mods:number>
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<mods:start id="2672E145951856F2A429497F4878E97E">243</mods:start>
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<mods:url id="60D3009EC252A33652034D56D2CAB32D">http://dx.doi.org/10.11646/zootaxa.5343.3.2</mods:url>
</mods:location>
<mods:classification id="CE490380D394A0DD812125662770D608">journal article</mods:classification>
<mods:identifier id="791EA997B2597A208340EDA6782AE6F8" type="DOI">10.11646/zootaxa.5343.3.2</mods:identifier>
<mods:identifier id="26D7D9752A7291CDAFDCFCD10CFE2590" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CFEDF372ADAED07393F4DD27E48D8122" type="Zenodo-Dep">8334181</mods:identifier>
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</mods:mods>
<treatment id="03A487CEFFFDFFDF85A551906823F861" ID-DOI="http://doi.org/10.5281/zenodo.8334203" ID-Zenodo-Dep="8334203" LSID="urn:lsid:plazi:treatment:03A487CEFFFDFFDF85A551906823F861" httpUri="http://treatment.plazi.org/id/03A487CEFFFDFFDF85A551906823F861" lastPageId="27" lastPageNumber="270" pageId="21" pageNumber="264">
<subSubSection id="C3176553FFFDFFD185A5519068C5FF6D" box="[151,952,151,178]" pageId="21" pageNumber="264" type="nomenclature">
<paragraph id="8BB236D8FFFDFFD185A5519068C5FF6D" blockId="21.[151,952,151,213]" box="[151,952,151,178]" pageId="21" pageNumber="264">
<heading id="D0FA81B4FFFDFFD185A5519068C5FF6D" bold="true" box="[151,952,151,178]" fontSize="11" level="1" pageId="21" pageNumber="264" reason="1">
<emphasis id="B979EACAFFFDFFD185A5519068C5FF6D" bold="true" box="[151,952,151,178]" pageId="21" pageNumber="264">
<taxonomicName id="4C0D4D5BFFFDFFD185A55190683BFF6D" authority="(Kaltenbach et al. 2023)" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[151,838,151,178]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="264" phylum="Arthropoda" rank="species" species="primasia" status="comb. n." subGenus="Securiops">
<emphasis id="B979EACAFFFDFFD185A551906A6EFF6E" bold="true" box="[151,275,151,178]" italics="true" pageId="21" pageNumber="264">Procloeon</emphasis>
(
<emphasis id="B979EACAFFFDFFD1841651906AE6FF6D" bold="true" box="[292,411,151,177]" italics="true" pageId="21" pageNumber="264">Securiops</emphasis>
)
<emphasis id="B979EACAFFFDFFD1849E5190696BFF6D" bold="true" box="[428,534,151,177]" italics="true" pageId="21" pageNumber="264">primasia</emphasis>
(
<bibRefCitation id="EF9C4B29FFFDFFD1871751906843FF6D" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[549,830,151,178]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD187F25190698EFF6E" bold="true" box="[704,755,151,178]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24A57B1FFFDFFD1867F519068C5FF6D" box="[845,952,151,177]" pageId="21" pageNumber="264" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3176553FFFDFFD185A551BC6AEAFF09" box="[151,407,187,213]" pageId="21" pageNumber="264" type="description">
<paragraph id="8BB236D8FFFDFFD185A551BC6AEAFF09" blockId="21.[151,952,151,213]" box="[151,407,187,213]" pageId="21" pageNumber="264">
(
<figureCitation id="13362A5DFFFDFFD185AD51BC6A5DFF09" box="[159,288,187,213]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="21" pageNumber="264">Figs 18–24</figureCitation>
,
<figureCitation id="13362A5DFFFDFFD1841E51BC6AF2FF09" box="[300,399,187,213]" captionStart-0="FIGURES 106–115" captionStart-1="FIGURES 116–119" captionStart-2="FIGURES 120–125" captionStart-3="FIGURES 126–136" captionStart-4="FIGURES 137–142" captionStart-5="FIGURES 143–145" captionStartId-0="22.[151,264,1854,1878]" captionStartId-1="23.[151,264,1838,1862]" captionStartId-2="24.[151,264,1756,1780]" captionStartId-3="25.[151,264,1840,1864]" captionStartId-4="26.[151,264,1653,1677]" captionStartId-5="27.[151,264,1056,1080]" captionTargetBox-0="[151,1436,200,1831]" captionTargetBox-1="[151,1436,181,1814]" captionTargetBox-2="[151,1435,181,1738]" captionTargetBox-3="[151,1435,181,1816]" captionTargetBox-4="[151,1436,181,1629]" captionTargetBox-5="[151,1436,181,1032]" captionTargetId-0="figure-18@22.[151,1436,200,1831]" captionTargetId-1="figure-16@23.[151,1436,181,1814]" captionTargetId-2="figure-18@24.[151,1435,181,1738]" captionTargetId-3="figure-16@25.[151,1435,181,1816]" captionTargetId-4="figure-18@26.[151,1436,181,1629]" captionTargetId-5="figure-16@27.[151,1436,181,1032]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionTargetPageId-4="26" captionTargetPageId-5="27" captionText-0="FIGURES 106–115. Procloeon (Securiops) primasia, larval exuviae. 106–108, dark individual (female); 109–115, light individual (male); 106, 115, abdominal terga and sterna; 107, 110, half of pronotum and mesonotum; 108, 111, left thoracic pleura and half of metanotum; 109, part of head; 112–114, fore, middle and hind legs." captionText-1="FIGURES 116–119. Procloeon (Securiops) primasia, exuviae of labium. 116, labium (ventral setae of glossae directed proximally); 117, 2nd and 3rd segments of labial palp; 118–119, glossae in ventral and dorsal views (ventral setae directed proximally). Abbreviations: pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." captionText-2="FIGURES 120–125. Procloeon (Securiops) primasia, larval exuviae. 120, base of tibia of fore leg, with focus on dorsal side; 121, the same with focus on ventral side; 122, abdominal tergum V; 123, abdominal sternum VIII; 124, cercus and paracercus; 125, portion of cercus." captionText-3="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." captionText-4="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." captionText-5="FIGURES 143–145. 143–144, eggs of Procloeon (Securiops) primasia. 145, male imaginal genitalia of Cheleocloeon excisum (Barnard 1932), reared from larva in Eerste river 9.II.2019 by N. Kluge and L. Sheyko [the same locality as for P. (S.) falcatum]. Abbreviation: mp, micropyle." figureDoi-0="http://doi.org/10.5281/zenodo.8334231" figureDoi-1="http://doi.org/10.5281/zenodo.8334235" figureDoi-2="http://doi.org/10.5281/zenodo.8334239" figureDoi-3="http://doi.org/10.5281/zenodo.8334245" figureDoi-4="http://doi.org/10.5281/zenodo.8334247" figureDoi-5="http://doi.org/10.5281/zenodo.8334251" httpUri-0="https://zenodo.org/record/8334231/files/figure.png" httpUri-1="https://zenodo.org/record/8334235/files/figure.png" httpUri-2="https://zenodo.org/record/8334239/files/figure.png" httpUri-3="https://zenodo.org/record/8334245/files/figure.png" httpUri-4="https://zenodo.org/record/8334247/files/figure.png" httpUri-5="https://zenodo.org/record/8334251/files/figure.png" pageId="21" pageNumber="264">106–144</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3176553FFFDFFD185A55007688AFEC4" box="[151,1015,256,280]" pageId="21" pageNumber="264" type="reference_group">
<paragraph id="8BB236D8FFFDFFD185A55007688AFEC4" blockId="21.[151,1015,256,280]" box="[151,1015,256,280]" pageId="21" pageNumber="264">
<taxonomicName id="4C0D4D5BFFFDFFD185A550076AA5FEC4" authority="Kaltenbach" authorityName="Kaltenbach" box="[151,472,256,280]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="264" phylum="Arthropoda" rank="species" species="primasia">
<emphasis id="B979EACAFFFDFFD185A550076A26FECB" box="[151,347,256,279]" italics="true" pageId="21" pageNumber="264">Securiops primasia</emphasis>
Kaltenbach
</taxonomicName>
, Phlai-ngam, Suttinun &amp; Gattolliat 2023: 129 (larva).
</paragraph>
</subSubSection>
<subSubSection id="C3176553FFFDFFD185A550426F29FE10" pageId="21" pageNumber="264" type="materials_examined">
<paragraph id="8BB236D8FFFDFFD185A550426F29FE10" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185A550426AFDFE83" bold="true" box="[151,384,325,351]" pageId="21" pageNumber="264">Material examined.</emphasis>
<materialsCitation id="3B653C85FFFDFFD184B550416AD0FE74" collectingDate="2015-01-30" collectingDateMax="2015-02-01" collectingDateMin="2015-01-30" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" county="river Taphoen" location="Lat Ya" municipality="river Kwai-Yai" pageId="21" pageNumber="264" specimenCount="1" specimenCount-female="1" stateProvince="Kanchanaburi">
<collectingCountry id="F31A7648FFFDFFD184B550416961FEBC" box="[391,540,326,352]" name="Thailand" pageId="21" pageNumber="264">THAILAND</collectingCountry>
:
<collectingRegion id="49C9F83AFFFDFFD1871450416852FEBC" box="[550,815,326,352]" country="Thailand" name="Kanchanaburi" pageId="21" pageNumber="264">Kanchanaburi province</collectingRegion>
,
<collectingCounty id="62D34E54FFFDFFD1860B504168A9FEBC" box="[825,980,326,352]" pageId="21" pageNumber="264">river Taphoen</collectingCounty>
near falling into
<collectingMunicipality id="6BD6ACA2FFFDFFD181A650416E42FEBC" box="[1172,1343,326,352]" pageId="21" pageNumber="264">river Kwai-Yai</collectingMunicipality>
, Lad-Ya (=
<location id="8ED26003FFFDFFD1858B506D6A75FE58" LSID="urn:lsid:plazi:treatment:03A487CEFFFDFFDF85A551906823F861:8ED26003FFFDFFD1858B506D6A75FE58" box="[185,264,362,388]" country="Thailand" county="river Taphoen" municipality="river Kwai-Yai" name="Lat Ya" pageId="21" pageNumber="264" stateProvince="Kanchanaburi">Lat Ya</location>
), resort «Island Resort River Kwai»,
<date id="FFB31018FFFDFFD18781506D682BFE59" box="[691,854,362,389]" pageId="21" pageNumber="264" value="2015-01-30" valueMax="2015-02-01" valueMin="2015-01-30">
<collectingDate id="EFF7E9F0FFFDFFD18781506D682BFE59" box="[691,854,362,389]" pageId="21" pageNumber="264" value="2015-01-30" valueMax="2015-02-01" valueMin="2015-01-30">30.I–1.II.2015</collectingDate>
</date>
, coll.
<collectorName id="26F8530EFFFDFFD186AF506D6F78FE58" box="[925,1029,362,388]" pageId="21" pageNumber="264">N. Kluge</collectorName>
&amp;
<collectorName id="26F8530EFFFDFFD1811E506D6FD8FE58" box="[1068,1189,362,388]" pageId="21" pageNumber="264">L. Sheyko</collectorName>
: 5 L-S-IJ, 1 L-SJ, 1 S-IJ, 3 L-S-I
<specimenCount id="9D0BFD51FFFDFFD1840850896A35FE74" box="[314,328,398,424]" pageId="21" pageNumber="264" type="female">♀</specimenCount>
, 1 L-S
<specimenCount id="9D0BFD51FFFDFFD184AC50896AD0FE74" box="[414,429,398,424]" pageId="21" pageNumber="264" type="female">♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B653C85FFFDFFD1848950896F9DFE74" box="[443,1248,398,425]" collectingDate="2015-02-01" collectingDateMax="2015-02-04" collectingDateMin="2015-02-01" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" location="Erawan Falls" pageId="21" pageNumber="264" specimenCount="1" specimenCount-female="1" stateProvince="Kanchanaburi province">
<location id="8ED26003FFFDFFD184895089692FFE74" LSID="urn:lsid:plazi:treatment:03A487CEFFFDFFDF85A551906823F861:8ED26003FFFDFFD184895089692FFE74" box="[443,594,398,424]" country="Thailand" name="Erawan Falls" pageId="21" pageNumber="264" stateProvince="Kanchanaburi province">Erawan Falls</location>
,
<date id="FFB31018FFFDFFD1876D5089699EFE75" box="[607,739,398,425]" pageId="21" pageNumber="264" value="2015-02-01" valueMax="2015-02-04" valueMin="2015-02-01">
<collectingDate id="EFF7E9F0FFFDFFD1876D5089699EFE75" box="[607,739,398,425]" pageId="21" pageNumber="264" value="2015-02-01" valueMax="2015-02-04" valueMin="2015-02-01">1–4.II.2015</collectingDate>
</date>
, coll.
<collectorName id="26F8530EFFFDFFD18618508968EFFE74" box="[810,914,398,424]" pageId="21" pageNumber="264">N. Kluge</collectorName>
&amp;
<collectorName id="26F8530EFFFDFFD1868B50896F4FFE74" box="[953,1074,398,424]" pageId="21" pageNumber="264">L. Sheyko</collectorName>
: 2 L/S
<specimenCount id="9D0BFD51FFFDFFD181B650896FE9FE74" box="[1156,1172,398,424]" pageId="21" pageNumber="264" type="female">♀</specimenCount>
, 1 L
<specimenCount id="9D0BFD51FFFDFFD181FD50896F9DFE74" box="[1231,1248,398,424]" pageId="21" pageNumber="264" type="female">♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B653C85FFFDFFD181DC50896F2DFE10" collectingDate="2015-02-09" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" county="Province" location="Pai" municipality="river Pai" pageId="21" pageNumber="264" specimenCount="1" stateProvince="Kanchanaburi province">
Mae-Hong-Son
<collectingCounty id="62D34E54FFFDFFD185A550B56B80FE10" box="[151,253,434,460]" pageId="21" pageNumber="264">Province</collectingCounty>
,
<collectingMunicipality id="6BD6ACA2FFFDFFD1843B50B56A14FE10" box="[265,361,434,460]" pageId="21" pageNumber="264">river Pai</collectingMunicipality>
upstream of
<location id="8ED26003FFFDFFD184C950B5695CFE10" LSID="urn:lsid:plazi:treatment:03A487CEFFFDFFDF85A551906823F861:8ED26003FFFDFFD184C950B5695CFE10" box="[507,545,434,460]" country="Thailand" county="Province" municipality="river Pai" name="Pai" pageId="21" pageNumber="264" stateProvince="Kanchanaburi province">Pai</location>
,
<date id="FFB31018FFFDFFD1871F50B569EBFE11" box="[557,662,434,461]" pageId="21" pageNumber="264" value="2015-02-09">
<collectingDate id="EFF7E9F0FFFDFFD1871F50B569EBFE11" box="[557,662,434,461]" pageId="21" pageNumber="264" value="2015-02-09">9.II.2015</collectingDate>
</date>
, coll.
<collectorName id="26F8530EFFFDFFD187E850B5683DFE10" box="[730,832,434,460]" pageId="21" pageNumber="264">N. Kluge</collectorName>
&amp;
<collectorName id="26F8530EFFFDFFD1865650B568A6FE10" box="[868,987,434,460]" pageId="21" pageNumber="264">L. Sheyko</collectorName>
: 1 L-S-IJ
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176553FFFDFFDF85F550D26823F861" lastPageId="27" lastPageNumber="270" pageId="21" pageNumber="264" type="description">
<paragraph id="8BB236D8FFFDFFD185F550D2681EFE33" blockId="21.[151,1437,325,2008]" box="[199,867,469,496]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F550D2681EFE33" bold="true" box="[199,867,469,496]" pageId="21" pageNumber="264">Additions and corrections to larval and egg descriptions</emphasis>
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F550FE694CFD15" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F550FE6AC7FDC8" bold="true" box="[199,442,505,532]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F550FE6ACEFDC8" bold="true" box="[199,435,505,532]" italics="true" pageId="21" pageNumber="264">Cuticular coloration</emphasis>
.
</emphasis>
Larvae of the both sexes have two sharply different forms of cuticular coloration, in one of which abdominal terga have very contrasting dark brown median part and light lateral parts (
<figureCitation id="13362A5DFFFDFFD181D653196E35FDE4" box="[1252,1352,542,568]" captionStart="FIGURES 106–115" captionStartId="22.[151,264,1854,1878]" captionTargetBox="[151,1436,200,1831]" captionTargetId="figure-18@22.[151,1436,200,1831]" captionTargetPageId="22" captionText="FIGURES 106–115. Procloeon (Securiops) primasia, larval exuviae. 106–108, dark individual (female); 109–115, light individual (male); 106, 115, abdominal terga and sterna; 107, 110, half of pronotum and mesonotum; 108, 111, left thoracic pleura and half of metanotum; 109, part of head; 112–114, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.8334231" httpUri="https://zenodo.org/record/8334231/files/figure.png" pageId="21" pageNumber="264">Fig. 106</figureCitation>
), while another form has abdominal terga mostly light, with small brown markings (
<figureCitation id="13362A5DFFFDFFD1813953456F12FD81" box="[1035,1135,578,605]" captionStart="FIGURES 106–115" captionStartId="22.[151,264,1854,1878]" captionTargetBox="[151,1436,200,1831]" captionTargetId="figure-18@22.[151,1436,200,1831]" captionTargetPageId="22" captionText="FIGURES 106–115. Procloeon (Securiops) primasia, larval exuviae. 106–108, dark individual (female); 109–115, light individual (male); 106, 115, abdominal terga and sterna; 107, 110, half of pronotum and mesonotum; 108, 111, left thoracic pleura and half of metanotum; 109, part of head; 112–114, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.8334231" httpUri="https://zenodo.org/record/8334231/files/figure.png" pageId="21" pageNumber="264">Fig. 115</figureCitation>
).
<bibRefCitation id="EF9C4B29FFFDFFD181B653456EE1FD80" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[1156,1436,578,604]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1803F53446E3CFD80" box="[1293,1345,578,604]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. (2023)
</bibRefCitation>
wrongly assumed that the first form is the normal one, and the second form has ́colour not yet fully developed» (
<bibRefCitation id="EF9C4B29FFFDFFD185AD538D6AD4FD78" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[159,425,650,676]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1841B538C6A20FD78" box="[297,349,650,676]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: fig. 1). Actually, cuticle of the both forms is finally developed, that is testified by shed exuviae shown on the
<figureCitation id="13362A5DFFFDFFD184A053A96950FD15" box="[402,557,686,713]" captionStart="FIGURES 106–115" captionStartId="22.[151,264,1854,1878]" captionTargetBox="[151,1436,200,1831]" captionTargetId="figure-18@22.[151,1436,200,1831]" captionTargetPageId="22" captionText="FIGURES 106–115. Procloeon (Securiops) primasia, larval exuviae. 106–108, dark individual (female); 109–115, light individual (male); 106, 115, abdominal terga and sterna; 107, 110, half of pronotum and mesonotum; 108, 111, left thoracic pleura and half of metanotum; 109, part of head; 112–114, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.8334231" httpUri="https://zenodo.org/record/8334231/files/figure.png" pageId="21" pageNumber="264">Figs 106–115</figureCitation>
.
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F553D66A75FC84" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F553D66A61FD30" bold="true" box="[199,284,721,748]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F553D66A68FD30" bold="true" box="[199,277,721,748]" italics="true" pageId="21" pageNumber="264">Glossa</emphasis>
.
</emphasis>
As well as in other species of
<taxonomicName id="4C0D4D5BFFFDFFD1875553D569A8FD30" authorityName=": Kluge" authorityYear="2020" box="[615,725,722,748]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B979EACAFFFDFFD1875553D569A8FD30" box="[615,725,722,748]" italics="true" pageId="21" pageNumber="264">Securiops</emphasis>
</taxonomicName>
, glossa has an arched, subapical, transverse row of setae located on its dorsal side [erroneously reported as ́ventroapically» and in ́ventral view» (
<bibRefCitation id="EF9C4B29FFFDFFD1810653F16E4BFCCC" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[1076,1334,758,784]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1818E53F06F90FCCC" box="[1212,1261,758,784]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: fig. 4d)] and a row of long, straight setae near base on its ventral side (not reported in the original description) (
<figureCitation id="13362A5DFFFDFFD18000521D6B81FC84" captionStart="FIGURES 116–119" captionStartId="23.[151,264,1838,1862]" captionTargetBox="[151,1436,181,1814]" captionTargetId="figure-16@23.[151,1436,181,1814]" captionTargetPageId="23" captionText="FIGURES 116–119. Procloeon (Securiops) primasia, exuviae of labium. 116, labium (ventral setae of glossae directed proximally); 117, 2nd and 3rd segments of labial palp; 118–119, glossae in ventral and dorsal views (ventral setae directed proximally). Abbreviations: pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." figureDoi="http://doi.org/10.5281/zenodo.8334235" httpUri="https://zenodo.org/record/8334235/files/figure.png" pageId="21" pageNumber="264">Figs 116, 118–119</figureCitation>
).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F55266684EFC7C" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F552666A33FCA0" bold="true" box="[199,334,865,892]" pageId="21" pageNumber="264">
<taxonomicName id="4C0D4D5BFFFDFFD185F552666A3AFCA0" authorityName="Hampson" authorityYear="1906" box="[199,327,865,892]" class="Insecta" family="Pyralidae" genus="Paraglossa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B979EACAFFFDFFD185F552666A3AFCA0" bold="true" box="[199,327,865,892]" italics="true" pageId="21" pageNumber="264">Paraglossa</emphasis>
</taxonomicName>
.
</emphasis>
In contrast to other species, inner margins of paraglossae are sharply angulate, in apical part concave and diverging (
<figureCitation id="13362A5DFFFDFFD1847252816AD8FC7C" box="[320,421,902,928]" captionStart="FIGURES 116–119" captionStartId="23.[151,264,1838,1862]" captionTargetBox="[151,1436,181,1814]" captionTargetId="figure-16@23.[151,1436,181,1814]" captionTargetPageId="23" captionText="FIGURES 116–119. Procloeon (Securiops) primasia, exuviae of labium. 116, labium (ventral setae of glossae directed proximally); 117, 2nd and 3rd segments of labial palp; 118–119, glossae in ventral and dorsal views (ventral setae directed proximally). Abbreviations: pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." figureDoi="http://doi.org/10.5281/zenodo.8334235" httpUri="https://zenodo.org/record/8334235/files/figure.png" pageId="21" pageNumber="264">Figs 116</figureCitation>
;
<bibRefCitation id="EF9C4B29FFFDFFD18483528169CFFC7C" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[433,690,902,928]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1870552806915FC7C" box="[567,616,902,928]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: figs 4a, e).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F552AE6AE8FB88" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F552AE6A2DFC18" bold="true" box="[199,336,937,964]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F552AE6A34FC18" bold="true" box="[199,329,937,964]" italics="true" pageId="21" pageNumber="264">Labial palp</emphasis>
.
</emphasis>
Setal field located on apical-dorsal side, proximally is bordered by sparse, regular row of especially large setae in high sockets (in contrast to the African species) (
<figureCitation id="13362A5DFFFDFFD1865E52C968ADFC34" box="[876,976,974,1000]" captionStart="FIGURES 116–119" captionStartId="23.[151,264,1838,1862]" captionTargetBox="[151,1436,181,1814]" captionTargetId="figure-16@23.[151,1436,181,1814]" captionTargetPageId="23" captionText="FIGURES 116–119. Procloeon (Securiops) primasia, exuviae of labium. 116, labium (ventral setae of glossae directed proximally); 117, 2nd and 3rd segments of labial palp; 118–119, glossae in ventral and dorsal views (ventral setae directed proximally). Abbreviations: pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." figureDoi="http://doi.org/10.5281/zenodo.8334235" httpUri="https://zenodo.org/record/8334235/files/figure.png" pageId="21" pageNumber="264">Fig. 117</figureCitation>
).
<bibRefCitation id="EF9C4B29FFFDFFD186D752C96F81FC34" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[997,1276,974,1000]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1815C52C86FDFFC34" box="[1134,1186,974,1000]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. (2023)
</bibRefCitation>
described the labial palp as ́2-segmented», taking the 2nd and 3dr segments fore one; however, the border separating these segments and the muscle going from the 2nd segment to the 3rd one are retained, as in other
<taxonomicName id="4C0D4D5BFFFDFFD181AC55116E71FBEC" authorityName=": Kluge" authorityYear="2020" box="[1182,1292,1046,1072]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B979EACAFFFDFFD181AC55116E71FBEC" box="[1182,1292,1046,1072]" italics="true" pageId="21" pageNumber="264">Securiops</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9C4B29FFFDFFD1802955116A6FFB88" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD185A5553C6BB5FB88" box="[151,200,1082,1108]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: figs 4a–b).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F5555A68B2FB1D" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F5555A6A6BFBA4" bold="true" box="[199,278,1117,1144]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F5555A6A72FBA4" bold="true" box="[199,271,1117,1144]" italics="true" pageId="21" pageNumber="264">Tibiae</emphasis>
.
</emphasis>
The proximal end of patella-tibial suture is bent distally and connected under acute angle with row of long thin hairs (as in other
<taxonomicName id="4C0D4D5BFFFDFFD184F35585696AFB40" authorityName="Leach" authorityYear="1815" box="[449,535,1154,1180]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1, see above) (
<figureCitation id="13362A5DFFFDFFD187E055856849FB40" box="[722,820,1154,1180]" captionStart="FIGURES 120–125" captionStartId="24.[151,264,1756,1780]" captionTargetBox="[151,1435,181,1738]" captionTargetId="figure-18@24.[151,1435,181,1738]" captionTargetPageId="24" captionText="FIGURES 120–125. Procloeon (Securiops) primasia, larval exuviae. 120, base of tibia of fore leg, with focus on dorsal side; 121, the same with focus on ventral side; 122, abdominal tergum V; 123, abdominal sternum VIII; 124, cercus and paracercus; 125, portion of cercus." figureDoi="http://doi.org/10.5281/zenodo.8334239" httpUri="https://zenodo.org/record/8334239/files/figure.png" pageId="21" pageNumber="264">Fig. 120</figureCitation>
). In the original description, the bent portion of patella-tibial suture is not shown on the drawing (
<bibRefCitation id="EF9C4B29FFFDFFD1875C55A1680CFB1C" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[622,881,1190,1216]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD187C455A06855FB1C" box="[758,808,1190,1216]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: fig. 5j).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F555CE69B1FA8C" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F555CE6972FB38" bold="true" box="[199,527,1225,1252]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F555CE6975FB38" bold="true" box="[199,520,1225,1252]" italics="true" pageId="21" pageNumber="264">Lateral spines on abdomen</emphasis>
.
</emphasis>
Lateral spines are located on abdominal segments VIII–IX, varying from 2 to 10 larger or smaller spines on one side (not constantly seven and five spines which are reported in the original description); previous segments normally lack lateral spines; small posterolateral spines are present on abdominal segments III–
<collectionCode id="ED1CAE1DFFFDFFD1840354316A25FA8C" box="[305,344,1334,1360]" pageId="21" pageNumber="264">VII</collectionCode>
(
<bibRefCitation id="EF9C4B29FFFDFFD1845454316914FA8C" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[358,617,1334,1360]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD184DC54306962FA8C" box="[494,543,1334,1360]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: fig. 6a).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F5545E6976FA3C" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F5545E6964FAA8" bold="true" box="[199,537,1369,1396]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F5545E696EFAA8" bold="true" box="[199,531,1369,1396]" italics="true" pageId="21" pageNumber="264">Abdominal terga and sterna</emphasis>
.
</emphasis>
Scales are entirely absent; all abdominal terga with numerous small, pointed, brown protuberances; all abdominal sterna with numerous colorless protuberances; besides this, abdominal terga and sterna with ring-like sensilla and fine setae, which are located sparsely and irregularly, not forming regular rows or constant pairs (
<figureCitation id="13362A5DFFFDFFD1846D54C16A82FA3C" box="[351,511,1478,1504]" captionStart="FIGURES 120–125" captionStartId="24.[151,264,1756,1780]" captionTargetBox="[151,1435,181,1738]" captionTargetId="figure-18@24.[151,1435,181,1738]" captionTargetPageId="24" captionText="FIGURES 120–125. Procloeon (Securiops) primasia, larval exuviae. 120, base of tibia of fore leg, with focus on dorsal side; 121, the same with focus on ventral side; 122, abdominal tergum V; 123, abdominal sternum VIII; 124, cercus and paracercus; 125, portion of cercus." figureDoi="http://doi.org/10.5281/zenodo.8334239" httpUri="https://zenodo.org/record/8334239/files/figure.png" pageId="21" pageNumber="264">Figs 122–123</figureCitation>
).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F554EE683EF900" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F554EE6A5AF9D8" bold="true" box="[199,295,1513,1540]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F554EE6A5DF9D8" bold="true" box="[199,288,1513,1540]" italics="true" pageId="21" pageNumber="264">Tergalii</emphasis>
.
</emphasis>
Second lamella (see characteristics of
<taxonomicName id="4C0D4D5BFFFDFFD187E754ED6854F9D8" authorityName="Leach" authorityYear="1815" box="[725,809,1514,1540]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1 above) is present on tergalii of I–
<collectionCode id="ED1CAE1DFFFDFFD181F454ED6F9EF9D8" box="[1222,1251,1514,1540]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="21" pageNumber="264">VI</collectionCode>
pairs and absent on tergalii of the
<collectionCode id="ED1CAE1DFFFDFFD1846557096A03F9F4" box="[343,382,1550,1576]" pageId="21" pageNumber="264">VII</collectionCode>
pair (
<figureCitation id="13362A5DFFFDFFD184F25709693EF9F4" box="[448,579,1550,1576]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="21" pageNumber="264">Figs 18–24</figureCitation>
). In the original description, the second lamella was erroneously reported on tergalius
<collectionCode id="ED1CAE1DFFFDFFD185CF57356A59F990" box="[253,292,1586,1612]" pageId="21" pageNumber="264">VII</collectionCode>
(
<bibRefCitation id="EF9C4B29FFFDFFD184035735691BF990" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[305,614,1586,1612]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1848A57346A95F990" box="[440,488,1586,1612]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023: 136
</bibRefCitation>
) because on the drawing (
<bibRefCitation id="EF9C4B29FFFDFFD186B457356FFAF990" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[902,1159,1586,1612]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1813C57346F43F990" box="[1038,1086,1586,1612]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: fig. 7b) the tergalius
<collectionCode id="ED1CAE1DFFFDFFD1804757356EE1F990" box="[1397,1436,1586,1612]" pageId="21" pageNumber="264">VII</collectionCode>
was overturned, its sharply convex costal margin was taken for the anal margin, and an occasional fold along costal margin was taken for a gap separating two lamellae. Actually, the costal margin is bordered with the costal rib and cannot be expanded into second lamella; the second lamella, characteristic for
<taxonomicName id="4C0D4D5BFFFDFFD1813557996F20F964" authorityName="Leach" authorityYear="1815" box="[1031,1117,1694,1720]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1, if present, represents an outgrowth of the anal margin, which is not bordered by a rib.
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F557E16F36F8F8" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
Tergalii of
<taxonomicName id="4C0D4D5BFFFDFFD1840D57E06A90F8DC" box="[319,493,1766,1792]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="252" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFFDFFD1840D57E06A2DF8DC" box="[319,336,1767,1792]" italics="true" pageId="21" pageNumber="264">P</emphasis>
. (
<emphasis id="B979EACAFFFDFFD1845457E16A09F8DC" box="[358,372,1766,1792]" italics="true" pageId="21" pageNumber="264">S</emphasis>
.)
<emphasis id="B979EACAFFFDFFD184BB57E06A90F8DC" box="[393,493,1767,1792]" italics="true" pageId="21" pageNumber="264">primasia</emphasis>
</taxonomicName>
have characteristic species-specific shape resembling linden leaf: with apex stretched and pointed, and with costal margin more convex than in other species (
<figureCitation id="13362A5DFFFDFFD1868E560D6F42F8F8" box="[956,1087,1802,1828]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="21" pageNumber="264">Figs 18–24</figureCitation>
).
</paragraph>
<paragraph id="8BB236D8FFFDFFD185F556296FE9F804" blockId="21.[151,1437,325,2008]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F556296A71F894" bold="true" box="[199,268,1838,1864]" pageId="21" pageNumber="264">
<emphasis id="B979EACAFFFDFFD185F556296A78F894" bold="true" box="[199,261,1838,1864]" italics="true" pageId="21" pageNumber="264">Cerci</emphasis>
.
</emphasis>
Cerci of
<taxonomicName id="4C0D4D5BFFFDFFD1844256286963F894" box="[368,542,1838,1864]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="252" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFFDFFD1844256286AFCF894" box="[368,385,1839,1864]" italics="true" pageId="21" pageNumber="264">P</emphasis>
. (
<emphasis id="B979EACAFFFDFFD184A556296AD8F894" box="[407,421,1838,1864]" italics="true" pageId="21" pageNumber="264">S</emphasis>
.)
<emphasis id="B979EACAFFFDFFD1848856286963F894" box="[442,542,1839,1864]" italics="true" pageId="21" pageNumber="264">primasia</emphasis>
</taxonomicName>
have unique structure: the enlarged spines on lateral side of cercus (characteristic for
<taxonomicName id="4C0D4D5BFFFDFFD185F356556A44F8B0" authorityName="sensu Kluge" authorityYear="2016" box="[193,313,1874,1900]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="264" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
/g1) are sharply differentiated: in proximal portion of the cercus all spines are stout, conic and not longer than next cercomere; in distal portion of the cercus all spines are delicate, widened from base and greatly enlarged, being as long as 2–3 next cercomeres (
<figureCitation id="13362A5DFFFDFFD187E0569D680CF869" box="[722,881,1946,1973]" captionStart="FIGURES 120–125" captionStartId="24.[151,264,1756,1780]" captionTargetBox="[151,1435,181,1738]" captionTargetId="figure-18@24.[151,1435,181,1738]" captionTargetPageId="24" captionText="FIGURES 120–125. Procloeon (Securiops) primasia, larval exuviae. 120, base of tibia of fore leg, with focus on dorsal side; 121, the same with focus on ventral side; 122, abdominal tergum V; 123, abdominal sternum VIII; 124, cercus and paracercus; 125, portion of cercus." figureDoi="http://doi.org/10.5281/zenodo.8334239" httpUri="https://zenodo.org/record/8334239/files/figure.png" pageId="21" pageNumber="264">Fig. 124–125</figureCitation>
). Original description was based on larvae with broken cerci (
<bibRefCitation id="EF9C4B29FFFDFFD1840356B96949F804" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[305,564,1982,2008]" pageId="21" pageNumber="264" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFFDFFD1848B56B86A97F804" box="[441,490,1982,2008]" italics="true" pageId="21" pageNumber="264">et al</emphasis>
. 2023
</bibRefCitation>
: fig. 1), and this important character was not reported.
</paragraph>
<caption id="DF726650FFFEFFD285A556396880F843" ID-DOI="http://doi.org/10.5281/zenodo.8334231" ID-Zenodo-Dep="8334231" httpUri="https://zenodo.org/record/8334231/files/figure.png" pageId="22" pageNumber="265" startId="22.[151,264,1854,1878]" targetBox="[151,1436,200,1831]" targetPageId="22" targetType="figure">
<paragraph id="8BB236D8FFFEFFD285A556396880F843" blockId="22.[151,1437,1854,1951]" pageId="22" pageNumber="265">
<emphasis id="B979EACAFFFEFFD285A556396A0FF88B" bold="true" box="[151,370,1854,1879]" pageId="22" pageNumber="265">FIGURES 106–115.</emphasis>
<taxonomicName id="4C0D4D5BFFFEFFD2844F563869B1F88A" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[381,716,1855,1879]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="265" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFFEFFD2844F56386A9BF88B" box="[381,486,1855,1879]" italics="true" pageId="22" pageNumber="265">Procloeon</emphasis>
(
<emphasis id="B979EACAFFFEFFD284C856386923F88A" box="[506,606,1855,1878]" italics="true" pageId="22" pageNumber="265">Securiops</emphasis>
)
<emphasis id="B979EACAFFFEFFD28740563869B1F88A" box="[626,716,1855,1878]" italics="true" pageId="22" pageNumber="265">primasia</emphasis>
</taxonomicName>
, larval exuviae. 106–108, dark individual (female); 109–115, light individual (male); 106, 115, abdominal terga and sterna; 107, 110, half of pronotum and mesonotum; 108, 111, left thoracic pleura and half of metanotum; 109, part of head; 112–114, fore, middle and hind legs.
</paragraph>
</caption>
<caption id="DF726650FFFFFFD385A556296E1FF852" ID-DOI="http://doi.org/10.5281/zenodo.8334235" ID-Zenodo-Dep="8334235" httpUri="https://zenodo.org/record/8334235/files/figure.png" pageId="23" pageNumber="266" startId="23.[151,264,1838,1862]" targetBox="[151,1436,181,1814]" targetPageId="23" targetType="figure">
<paragraph id="8BB236D8FFFFFFD385A556296E1FF852" blockId="23.[151,1437,1838,1934]" pageId="23" pageNumber="266">
<emphasis id="B979EACAFFFFFFD385A556296A12F89A" bold="true" box="[151,367,1838,1862]" pageId="23" pageNumber="266">FIGURES 116–119.</emphasis>
<taxonomicName id="4C0D4D5BFFFFFFD3844E562969B6F89A" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[380,715,1838,1862]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="266" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFFFFFD3844E56296A98F89A" box="[380,485,1838,1862]" italics="true" pageId="23" pageNumber="266">Procloeon</emphasis>
(
<emphasis id="B979EACAFFFFFFD384CA56286921F89A" box="[504,604,1839,1862]" italics="true" pageId="23" pageNumber="266">Securiops</emphasis>
)
<emphasis id="B979EACAFFFFFFD38743562869B6F89A" box="[625,715,1839,1862]" italics="true" pageId="23" pageNumber="266">primasia</emphasis>
</taxonomicName>
, exuviae of labium. 116, labium (ventral setae of glossae directed proximally); 117, 2nd and 3rd segments of labial palp; 118–119, glossae in ventral and dorsal views (ventral setae directed proximally). Abbreviations: pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa.
</paragraph>
</caption>
<caption id="DF726650FFF0FFDC85A557DB6A00F8E0" ID-DOI="http://doi.org/10.5281/zenodo.8334239" ID-Zenodo-Dep="8334239" httpUri="https://zenodo.org/record/8334239/files/figure.png" pageId="24" pageNumber="267" startId="24.[151,264,1756,1780]" targetBox="[151,1435,181,1738]" targetPageId="24" targetType="figure">
<paragraph id="8BB236D8FFF0FFDC85A557DB6A00F8E0" blockId="24.[151,1437,1756,1853]" pageId="24" pageNumber="267">
<emphasis id="B979EACAFFF0FFDC85A557DB6A13F929" bold="true" box="[151,366,1756,1781]" pageId="24" pageNumber="267">FIGURES 120–125.</emphasis>
<taxonomicName id="4C0D4D5BFFF0FFDC844757DA69C7F928" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[373,698,1757,1781]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="267" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFF0FFDC844757DA6AA3F929" box="[373,478,1757,1781]" italics="true" pageId="24" pageNumber="267">Procloeon</emphasis>
(
<emphasis id="B979EACAFFF0FFDC84DF57DA692CF928" box="[493,593,1757,1780]" italics="true" pageId="24" pageNumber="267">Securiops</emphasis>
)
<emphasis id="B979EACAFFF0FFDC875257DA69C7F928" box="[608,698,1757,1780]" italics="true" pageId="24" pageNumber="267">primasia</emphasis>
</taxonomicName>
, larval exuviae. 120, base of tibia of fore leg, with focus on dorsal side; 121, the same with focus on ventral side; 122, abdominal tergum V; 123, abdominal sternum VIII; 124, cercus and paracercus; 125, portion of cercus.
</paragraph>
</caption>
<caption id="DF726650FFF1FFDD85A556376955F868" ID-DOI="http://doi.org/10.5281/zenodo.8334245" ID-Zenodo-Dep="8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="25" pageNumber="268" startId="25.[151,264,1840,1864]" targetBox="[151,1435,181,1816]" targetPageId="25" targetType="figure">
<paragraph id="8BB236D8FFF1FFDD85A556376955F868" blockId="25.[151,1436,1840,1972]" pageId="25" pageNumber="268">
<emphasis id="B979EACAFFF1FFDD85A556376A13F894" bold="true" box="[151,366,1840,1864]" pageId="25" pageNumber="268">FIGURES 126–136.</emphasis>
<taxonomicName id="4C0D4D5BFFF1FFDD8446563769C4F894" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[372,697,1840,1864]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="268" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFF1FFDD844656376AA0F894" box="[372,477,1840,1864]" italics="true" pageId="25" pageNumber="268">Procloeon</emphasis>
(
<emphasis id="B979EACAFFF1FFDD84DE5637692DF89B" box="[492,592,1840,1863]" italics="true" pageId="25" pageNumber="268">Securiops</emphasis>
)
<emphasis id="B979EACAFFF1FFDD876D563669C4F894" box="[607,697,1841,1864]" italics="true" pageId="25" pageNumber="268">primasia</emphasis>
</taxonomicName>
. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite.
</paragraph>
</caption>
<caption id="DF726650FFF2FFDE85A5577268D9F909" ID-DOI="http://doi.org/10.5281/zenodo.8334247" ID-Zenodo-Dep="8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="26" pageNumber="269" startId="26.[151,264,1653,1677]" targetBox="[151,1436,181,1629]" targetPageId="26" targetType="figure">
<paragraph id="8BB236D8FFF2FFDE85A5577268D9F909" blockId="26.[151,1436,1653,1749]" pageId="26" pageNumber="269">
<emphasis id="B979EACAFFF2FFDE85A557726A11F951" bold="true" box="[151,364,1653,1677]" pageId="26" pageNumber="269">FIGURES 137–142.</emphasis>
<taxonomicName id="4C0D4D5BFFF2FFDE8443577269CFF951" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[369,690,1653,1677]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFF2FFDE844357726AA7F951" box="[369,474,1653,1677]" italics="true" pageId="26" pageNumber="269">Procloeon</emphasis>
(
<emphasis id="B979EACAFFF2FFDE84D557716936F951" box="[487,587,1654,1677]" italics="true" pageId="26" pageNumber="269">Securiops</emphasis>
)
<emphasis id="B979EACAFFF2FFDE876A577169CFF951" box="[600,690,1654,1677]" italics="true" pageId="26" pageNumber="269">primasia</emphasis>
</taxonomicName>
. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia.
</paragraph>
</caption>
<paragraph id="8BB236D8FFF2FFDE85F556036B87F82A" blockId="26.[151,1436,1795,2038]" pageId="26" pageNumber="269">
<emphasis id="B979EACAFFF2FFDE85F556036B81F8C2" bold="true" box="[199,252,1796,1822]" italics="true" pageId="26" pageNumber="269">Egg.</emphasis>
Elongate-oval; chorion with even convex relief in a form of net with cells elongated along egg axis; micropyle is very small, round (
<figureCitation id="13362A5DFFF2FFDE84CF562069E1F89E" box="[509,668,1831,1858]" captionStart="FIGURES 143–145" captionStartId="27.[151,264,1056,1080]" captionTargetBox="[151,1436,181,1032]" captionTargetId="figure-16@27.[151,1436,181,1032]" captionTargetPageId="27" captionText="FIGURES 143–145. 143–144, eggs of Procloeon (Securiops) primasia. 145, male imaginal genitalia of Cheleocloeon excisum (Barnard 1932), reared from larva in Eerste river 9.II.2019 by N. Kluge and L. Sheyko [the same locality as for P. (S.) falcatum]. Abbreviation: mp, micropyle." figureDoi="http://doi.org/10.5281/zenodo.8334251" httpUri="https://zenodo.org/record/8334251/files/figure.png" pageId="26" pageNumber="269">Figs 143–144</figureCitation>
).
<bibRefCitation id="EF9C4B29FFF2FFDE879D562068C2F89E" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[687,959,1831,1858]" pageId="26" pageNumber="269" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFF2FFDE8607562F681AF89D" box="[821,871,1831,1857]" italics="true" pageId="26" pageNumber="269">et al</emphasis>
. (2023)
</bibRefCitation>
erroneously described eggs of this species as having ́four longitudinal rows of wide, sub-rectangular structural elements» (
<bibRefCitation id="EF9C4B29FFF2FFDE811A564C6E18F8B9" author="Kaltenbach, T. &amp; Suttinun, Ch. &amp; Gattolliat, J. - L." box="[1064,1381,1867,1894]" pageId="26" pageNumber="269" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. &amp; Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty &amp; Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
Kaltenbach
<emphasis id="B979EACAFFF2FFDE8183564B6F9EF8B9" box="[1201,1251,1867,1893]" italics="true" pageId="26" pageNumber="269">et al</emphasis>
. 2023: 136
</bibRefCitation>
, figs 7c–d). Eggs described by them, were not developed; their ́sub-rectangular structural elements» are tissues of ovarium, but not elements of the egg chorion. The same egg structure is visible on scanning electron photographs in all cases when non-developed eggs are examined (e.g., see
<bibRefCitation id="EF9C4B29FFF2FFDE860256B06FCEF80E" author="Kopelke, J. - P. &amp; Muller-Liebenau, I." box="[816,1203,1975,2002]" pageId="26" pageNumber="269" pagination="7 - 25" refId="ref13151" refString="Kopelke, J. - P. &amp; Muller-Liebenau, I. (1982) Eistrukturen bei Ephemeroptera und deren Bedeutung fur die Aufstellung von Artengruppen am Beispiel der europaischen Arten der Gattung Baetis Leach, 1815, (Insecta: Baetidae). Teil I: alpinus -, lutheri -, pavidus - und lapponicus - Gruppe. Gewasser und Abwasser, 68 / 69, 7 - 25. https: // doi. org / 10.1002 / mmnd. 4800280102" type="journal article" year="1982">Kopelke &amp; Müller-Liebenau 1982</bibRefCitation>
: fig. 6: egg of
<taxonomicName id="4C0D4D5BFFF2FFDE806456BF6B94F829" baseAuthorityName="Pictet" baseAuthorityYear="1843" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B979EACAFFF2FFDE806456BF6B94F829" italics="true" pageId="26" pageNumber="269">Baetis alpinus</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF726650FFF3FFDF85A555276AB8FB5C" ID-DOI="http://doi.org/10.5281/zenodo.8334251" ID-Zenodo-Dep="8334251" httpUri="https://zenodo.org/record/8334251/files/figure.png" pageId="27" pageNumber="270" startId="27.[151,264,1056,1080]" targetBox="[151,1436,181,1032]" targetPageId="27" targetType="figure">
<paragraph id="8BB236D8FFF3FFDF85A555276AB8FB5C" blockId="27.[151,1437,1056,1152]" pageId="27" pageNumber="270">
<emphasis id="B979EACAFFF3FFDF85A555276A10FBE4" bold="true" box="[151,365,1056,1080]" pageId="27" pageNumber="270">FIGURES 143–145.</emphasis>
143–144, eggs of
<taxonomicName id="4C0D4D5BFFF3FFDF871A55276811FBE4" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Kaltenbach et al." baseAuthorityYear="2023" box="[552,876,1056,1080]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="270" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
<emphasis id="B979EACAFFF3FFDF871A552769ECFBE4" box="[552,657,1056,1080]" italics="true" pageId="27" pageNumber="270">Procloeon</emphasis>
(
<emphasis id="B979EACAFFF3FFDF87AD5527687EFBEB" box="[671,771,1056,1079]" italics="true" pageId="27" pageNumber="270">Securiops</emphasis>
)
<emphasis id="B979EACAFFF3FFDF862055266811FBE4" box="[786,876,1057,1080]" italics="true" pageId="27" pageNumber="270">primasia</emphasis>
</taxonomicName>
. 145, male imaginal genitalia of
<taxonomicName id="4C0D4D5BFFF3FFDF818B55276A49FB80" authority="(Barnard 1932)" baseAuthorityName="Barnard" baseAuthorityYear="1932" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="270" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="B979EACAFFF3FFDF818B55276EE0FBE4" box="[1209,1437,1056,1080]" italics="true" pageId="27" pageNumber="270">Cheleocloeon excisum</emphasis>
(
<bibRefCitation id="EF9C4B29FFF3FFDF85AC55436A50FB80" author="Barnard, K. H." box="[158,301,1092,1116]" pageId="27" pageNumber="270" pagination="201 - 259" refId="ref11827" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
)
</taxonomicName>
, reared from larva in Eerste river 9.II.2019 by N. Kluge and L. Sheyko [the same locality as for
<taxonomicName id="4C0D4D5BFFF3FFDF81C255426EF3FB80" authorityName="Crass" authorityYear="1947" box="[1264,1422,1092,1116]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="252" phylum="Arthropoda" rank="species" species="falcatum" subGenus="Securiops">
<emphasis id="B979EACAFFF3FFDF81C255426F82FB80" box="[1264,1279,1093,1116]" italics="true" pageId="27" pageNumber="270">P</emphasis>
. (
<emphasis id="B979EACAFFF3FFDF802655436E5CFB87" box="[1300,1313,1092,1115]" italics="true" pageId="27" pageNumber="270">S</emphasis>
.)
<emphasis id="B979EACAFFF3FFDF800755436EF3FB80" box="[1333,1422,1092,1116]" italics="true" pageId="27" pageNumber="270">falcatum</emphasis>
</taxonomicName>
]. Abbreviation: mp, micropyle.
</paragraph>
</caption>
<paragraph id="8BB236D8FFF3FFDF85A555D56A8BFB30" blockId="27.[151,502,1234,1260]" box="[151,502,1234,1260]" pageId="27" pageNumber="270">
<heading id="D0FA81B4FFF3FFDF85A555D56A8BFB30" bold="true" box="[151,502,1234,1260]" fontSize="11" level="1" pageId="27" pageNumber="270" reason="1">
<emphasis id="B979EACAFFF3FFDF85A555D56A8BFB30" bold="true" box="[151,502,1234,1260]" pageId="27" pageNumber="270">Descriptions of winged stages</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236D8FFF3FFDF85A5541D6FBFFA34" blockId="27.[151,1437,1306,1981]" pageId="27" pageNumber="270">
<emphasis id="B979EACAFFF3FFDF85A5541D6A6EFAE9" bold="true" box="[151,275,1306,1333]" italics="true" pageId="27" pageNumber="270">Subimago.</emphasis>
CUTICULAR COLORATION. Head nearly colorless, antennal flagellum brownish. Pronotum colorless. Mesonotum mostly colorless, with contrasting brown grooves bordering anterolateral scutal costa, contrasting brown prealar bridge and contrasting brown lateral sclerite of parascutellum (
<figureCitation id="13362A5DFFF3FFDF810354656FE4FAA0" box="[1073,1177,1378,1404]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Fig. 129</figureCitation>
). Thoracic pleura and sterna mostly colorless, with contrasting brown area of pleural suture, contrasting brown postsubalar sclerite with lateropostnotal crest and brown pleura of metathorax (
<figureCitation id="13362A5DFFF3FFDF863054AD681BFA18" box="[770,870,1450,1476]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Fig. 130</figureCitation>
). Wing cuticle colorless, microtrichia ochre. Legs colorless. Abdominal terga and sterna colorless. Gonostyli colorless (
<figureCitation id="13362A5DFFF3FFDF86AC54C96F7DFA34" box="[926,1024,1486,1512]" captionStart="FIGURES 137–142" captionStartId="26.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-18@26.[151,1436,181,1629]" captionTargetPageId="26" captionText="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." figureDoi="http://doi.org/10.5281/zenodo.8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="27" pageNumber="270">Fig. 142</figureCitation>
). Cerci colorless.
</paragraph>
<paragraph id="8BB236D8FFF3FFDF85F554F46E40F9D0" blockId="27.[151,1437,1306,1981]" box="[199,1341,1522,1549]" pageId="27" pageNumber="270">
TEXTURE. On legs of all pairs, all tarsal segments covered with pointed microlepides (as in
<figureCitation id="13362A5DFFF3FFDF81EF54F56E4CF9D0" box="[1245,1329,1522,1548]" captionStart="FIGURES 70–76" captionStartId="15.[151,264,1837,1861]" captionTargetBox="[152,1435,181,1814]" captionTargetId="figure-16@15.[151,1435,181,1814]" captionTargetPageId="15" captionText="FIGURES 70–76. Procloeon (Securiops) macafertiorum. 70, subimaginal exuviae: apex of tibia and base of tarsus of fore leg of male; 71–76, larval exuviae: 71, caudalii; 72, fragment of cercus; 73, fragment of fore tarsus; 74, abdominal tergum IV; 75, abdominal sternum V; 76, abdominal tergum X, paraproct and margin of tergum IX." figureDoi="http://doi.org/10.5281/zenodo.8334211" httpUri="https://zenodo.org/record/8334211/files/figure.png" pageId="27" pageNumber="270">Fig. 70</figureCitation>
).
</paragraph>
<paragraph id="8BB236D8FFF3FFDF85F557106AF3F8F0" blockId="27.[151,1437,1306,1981]" pageId="27" pageNumber="270">
<emphasis id="B979EACAFFF3FFDF85F557106A25F9ED" bold="true" box="[199,344,1558,1585]" italics="true" pageId="27" pageNumber="270">Imago, male</emphasis>
(
<figureCitation id="13362A5DFFF3FFDF845A5711697AF9EC" box="[360,519,1558,1584]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Figs 126–127</figureCitation>
). Head brown. Turbinate eyes dark brown. Thorax brown with membranes ochre. Wings colorless, with veins ochre. Pterostigma with several complete, non-branched, oblique crossveins. Legs ochre; either all legs without maculae, or fore femur with diffusive macula on outer side near base (as in
<figureCitation id="13362A5DFFF3FFDF804257596BB8F940" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Fig. 132</figureCitation>
). Middle and hind legs with tarsus longer than tibia, with one apical spine on initial 3rd tarsomere (actual 2nd tarsomere) (as in
<figureCitation id="13362A5DFFF3FFDF846957A16A84F91D" box="[347,505,1702,1729]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Figs 135–136</figureCitation>
). Abdomen mostly ochre; terga with diffusive brownish markings; posterior part of tergum
<collectionCode id="ED1CAE1DFFF3FFDF85D957CC6A6FF939" box="[235,274,1739,1765]" pageId="27" pageNumber="270">VII</collectionCode>
and terga VIII–IX brownish. Cerci uniformly ochre. Gonostyli ochre, long and slender (
<figureCitation id="13362A5DFFF3FFDF81D157CD6E39F938" box="[1251,1348,1738,1764]" captionStart="FIGURES 137–142" captionStartId="26.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-18@26.[151,1436,181,1629]" captionTargetPageId="26" captionText="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." figureDoi="http://doi.org/10.5281/zenodo.8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="27" pageNumber="270">Fig. 137</figureCitation>
). Penial bridge forming projection semicircular in ventral view (
<figureCitation id="13362A5DFFF3FFDF863757E9681BF8D4" box="[773,870,1774,1800]" captionStart="FIGURES 137–142" captionStartId="26.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-18@26.[151,1436,181,1629]" captionTargetPageId="26" captionText="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." figureDoi="http://doi.org/10.5281/zenodo.8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="27" pageNumber="270">Fig. 137</figureCitation>
), apically with projection bent dorsally under right angle (
<figureCitation id="13362A5DFFF3FFDF85D156156AFFF8F0" box="[227,386,1810,1836]" captionStart="FIGURES 137–142" captionStartId="26.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-18@26.[151,1436,181,1629]" captionTargetPageId="26" captionText="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." figureDoi="http://doi.org/10.5281/zenodo.8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="27" pageNumber="270">Figs 138–141</figureCitation>
).
</paragraph>
<paragraph id="8BB236D8FFF3FFDF85F556306E5AF844" blockId="27.[151,1437,1306,1981]" pageId="27" pageNumber="270">
<emphasis id="B979EACAFFF3FFDF85F556306A0DF88D" bold="true" box="[199,368,1846,1873]" italics="true" pageId="27" pageNumber="270">Imago, female</emphasis>
(
<figureCitation id="13362A5DFFF3FFDF84B356316A98F88C" box="[385,485,1846,1872]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Fig. 128</figureCitation>
). Head, thorax and abdomen ochre with brown markings. Fore tarsus 5-segmented, with small apical spine on 3rd segment (
<figureCitation id="13362A5DFFF3FFDF875B565D6878F8A8" box="[617,773,1882,1908]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Fig. 132–133</figureCitation>
); middle and hind tarsi 4-segmented, with apical spine on initial 3rd segment (actual 2nd segment) (
<figureCitation id="13362A5DFFF3FFDF87585679687BF845" box="[618,774,1918,1945]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="27" pageNumber="270">Fig. 135–136</figureCitation>
). Coloration of wings, legs, and cerci as in male.
</paragraph>
<paragraph id="8BB236D8FFF3FFDF85F556A56823F861" blockId="27.[151,1437,1306,1981]" box="[199,862,1954,1981]" pageId="27" pageNumber="270">
<emphasis id="B979EACAFFF3FFDF85F556A56A33F860" bold="true" box="[199,334,1954,1980]" pageId="27" pageNumber="270">Dimension.</emphasis>
Fore wing length of male and female
<quantity id="4CF59B3DFFF3FFDF87CB56A56824F861" box="[761,857,1954,1981]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="27" pageNumber="270" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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