371 lines
51 KiB
XML
371 lines
51 KiB
XML
<document id="6B5362DB14F441B5E422C9D5DE03C725" ID-DOI="10.11646/zootaxa.5343.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8334181" ID-ZooBank="BB852CA9-6C6E-4016-9972-AAF2C378E41F" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1694429699348" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2023" docId="03A487CEFFE9FFC785A554E469AFFB35" docLanguage="en" docName="zootaxa.5343.3.2.pdf" docOrigin="Zootaxa 5343 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cloeonini sensu Kluge 2016" docType="treatment" docVersion="3" lastPageNumber="246" masterDocId="FF9DFFB6FFE8FFC4853251076B7DFFDC" masterDocTitle="Redescription of the subgenus Securiops Jacobus, McCafferty & Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)" masterLastPageNumber="272" masterPageNumber="243" pageNumber="244" updateTime="1694453124034" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="77E3A62336589FFEEA595B7CBA9E58F0">Redescription of the subgenus Securiops Jacobus, McCafferty & Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<mods:name id="39992EC2094BD4C7C85E9F31AD78767F" type="personal">
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<mods:namePart id="F050EB3DB1E8E2F60D892DE4C79370B7">Kluge, Nikita J.</mods:namePart>
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<mods:typeOfResource id="199132C9F044B0F0AD56DF59A8FD21E5">text</mods:typeOfResource>
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<mods:title id="4654351B8FDFDF176701E5FDCFAD11B0">Zootaxa</mods:title>
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<mods:part id="C5E1A98A72FE42EA26EE509FA396BA02">
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<mods:date id="3F36647533DFB66F9F4AC641F56CA7B7">2023</mods:date>
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<mods:number id="94C28241E446DD7E29FE3CDC72AA4273">2023-09-08</mods:number>
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<mods:number id="A35DE6183BFB430E1CCDDD5C5373ECDB">5343</mods:number>
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<mods:number id="FCC2E9BA1232EFCAF25FE02E1F260A92">3</mods:number>
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<mods:start id="CC40136AED12CFC3247C30DDAB53B5FA">243</mods:start>
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<mods:url id="2B7D28D4956135504DF111CF536FDDDC">http://dx.doi.org/10.11646/zootaxa.5343.3.2</mods:url>
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<mods:classification id="0F0DBAAC9407F3797DC850EC1620D119">journal article</mods:classification>
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<mods:identifier id="E7767D1DD26504646DB4E29FFAC38AB6" type="DOI">10.11646/zootaxa.5343.3.2</mods:identifier>
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<mods:identifier id="4909DC94D2EE79377325BEF1E9ED70CF" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03A487CEFFE9FFC785A554E469AFFB35" LSID="urn:lsid:plazi:treatment:03A487CEFFE9FFC785A554E469AFFB35" httpUri="http://treatment.plazi.org/id/03A487CEFFE9FFC785A554E469AFFB35" lastPageId="3" lastPageNumber="246" pageId="1" pageNumber="244">
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<subSubSection id="C3176553FFE9FFC585A554E46912FA21" box="[151,623,1507,1533]" pageId="1" pageNumber="244" type="nomenclature">
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<paragraph id="8BB236D8FFE9FFC585A554E46912FA21" blockId="1.[151,623,1507,1533]" box="[151,623,1507,1533]" pageId="1" pageNumber="244">
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<heading id="D0FA81B4FFE9FFC585A554E46912FA21" bold="true" box="[151,623,1507,1533]" fontSize="11" level="1" pageId="1" pageNumber="244" reason="1">
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<emphasis id="B979EACAFFE9FFC585A554E46912FA21" bold="true" box="[151,623,1507,1533]" pageId="1" pageNumber="244">
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Characters of
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<taxonomicName id="4C0D4D5BFFE9FFC5847054E46ACAFA21" authorityName="sensu Kluge" authorityYear="2016" box="[322,439,1507,1533]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="244" phylum="Arthropoda" rank="tribe" tribe="Cloeonini">Cloeonini</taxonomicName>
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, or Cloeon/fg1:
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3176553FFE9FFC785A5572269AFFB35" lastPageId="3" lastPageNumber="246" pageId="1" pageNumber="244" type="description">
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<paragraph id="8BB236D8FFE9FFC585A557226A2DF9B8" blockId="1.[151,1437,1573,2032]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585A557226A65F99C" bold="true" box="[151,280,1573,1600]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585A557226A6DF99C" bold="true" box="[151,272,1573,1600]" italics="true" pageId="1" pageNumber="244">Mandibles</emphasis>
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.
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</emphasis>
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Both larval mandibles have dense setae between prostheca and mola (
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<figureCitation id="13362A5DFFE9FFC5811857216FD1F99D" box="[1066,1196,1574,1601]" captionStart="FIGURES 31–39" captionStartId="5.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-16@5.[151,1436,181,1817]" captionTargetPageId="5" captionText="FIGURES 31–39. Procloeon (Securiops) falcatum, larval mouthparts. 31, apex of labrum, ventral view; 32, labrum, dorsal view; 33, apex of labrum, enlarged; 34–35, left and right mandibles; 36, maxilla; 37–38, apex of maxilla with focus on ventral and dorsal sides; 39, hypopharynx with superlinguae.Abbreviations: can, maxillary canines; ds1, ds2, ds3, 1st–3rd dentisetae," figureDoi="http://doi.org/10.5281/zenodo.8334189" httpUri="https://zenodo.org/record/8334189/files/figure.png" pageId="1" pageNumber="244">Figs 34–35</figureCitation>
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,
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<figureCitation id="13362A5DFFE9FFC5818557216E7FF99C" box="[1207,1282,1574,1600]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="1" pageNumber="244">98–99</figureCitation>
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). The same in some other taxa.
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</paragraph>
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<paragraph id="8BB236D8FFE9FFC585F5576A6E03F928" blockId="1.[151,1437,1573,2032]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F5576A6A2EF954" bold="true" box="[199,339,1645,1672]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F5576A6A31F954" bold="true" box="[199,332,1645,1672]" italics="true" pageId="1" pageNumber="244">Labial palp</emphasis>
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.
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</emphasis>
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The 3rd segment of labial palp is truncate, and the 2nd segment lacks apical-median projection, so that these two segments form an integral figure widened from base toward apex. At the same time, the border between these segments is retained, and the muscle going from the 2nd segment to the base of the 3rd segment is retained (
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<figureCitation id="13362A5DFFE9FFC5843357DD6A28F928" box="[257,341,1754,1780]" captionStart="FIGURES 40–46" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-16@7.[151,1436,181,1817]" captionTargetPageId="7" captionText="FIGURES 40–46. Procloeon (Securiops) falcatum, labium. 40, integral labium; 41–46, parts of exuviae: 41, glossae and paraglossae, ventral view; 42, half of labium of another individual (ventral setae of glossae directed distally); 43, the same individual, with ventral setae of glossae directed proximally; 44, distal segments of labial palp; 45–46, enlarged glossae in ventral and dorsal view (ventral setae directed distally). Abbreviations: m2-3, muscle going from 2nd palpomere to base of 3rd palpomere; pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." figureDoi="http://doi.org/10.5281/zenodo.8334193" httpUri="https://zenodo.org/record/8334193/files/figure.png" pageId="1" pageNumber="244">Fig. 40</figureCitation>
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). An autapomorphy of
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<taxonomicName id="4C0D4D5BFFE9FFC5876557DD69B8F928" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[599,709,1754,1780]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="244" phylum="Arthropoda" rank="genus">
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<emphasis id="B979EACAFFE9FFC5876557DD69B8F928" box="[599,709,1754,1780]" italics="true" pageId="1" pageNumber="244">Securiops</emphasis>
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</taxonomicName>
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is the great widening of this triangular formation (see below).
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</paragraph>
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<paragraph id="8BB236D8FFE9FFC585F557FA6894F8E0" blockId="1.[151,1437,1573,2032]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F557FA697FF8C4" bold="true" box="[199,514,1789,1816]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F557FA6A86F8C4" bold="true" box="[199,507,1789,1816]" italics="true" pageId="1" pageNumber="244">Larval patella-tibial suture</emphasis>
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.
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</emphasis>
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The proximal end of patella-tibial suture is bent distally and connected under acute angle with a row of long thin hairs (autapomorphy of
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<taxonomicName id="4C0D4D5BFFE9FFC587C35625683BF8E0" authorityName="Leach" authorityYear="1815" box="[753,838,1826,1852]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
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/fg1) (
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<figureCitation id="13362A5DFFE9FFC586BA562568A1F8E0" box="[904,988,1826,1852]" captionStart="FIGURES 47–52" captionStartId="8.[151,264,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-18@8.[151,1435,181,1646]" captionTargetPageId="8" captionText="FIGURES 47–52. Procloeon (Securiops) falcatum, larva. 47, articulation of femur and tibia of fore leg, dorsal view; 48, the same, compressed to show junction of patella-tibial suture with row of fine hairs; 49, fragment of tarsus; 50, claw; 51, abdominal tergum X and margin of tergum IX; 52, abdominal tergum X and paraprocts." figureDoi="http://doi.org/10.5281/zenodo.8334199" httpUri="https://zenodo.org/record/8334199/files/figure.png" pageId="1" pageNumber="244">Fig. 48</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB236D8FFE9FFC585F556426A16F875" blockId="1.[151,1437,1573,2032]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F556426A2BF8BC" bold="true" box="[199,342,1861,1888]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F556426A32F8BC" bold="true" box="[199,335,1861,1888]" italics="true" pageId="1" pageNumber="244">Larval claw</emphasis>
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.
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</emphasis>
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In all
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<taxonomicName id="4C0D4D5BFFE9FFC584AE56416A8DF8BC" authorityName="Leach" authorityYear="1815" box="[412,496,1862,1888]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
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/fg1 larval claw is symmetric, slightly curved, either with 2 similar rows of denticles, or without denticles (plesiomorphy within Liberevenata). In all known species of
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<taxonomicName id="4C0D4D5BFFE9FFC5811C566D6FE1F858" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[1070,1180,1898,1924]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="244" phylum="Arthropoda" rank="genus">
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<emphasis id="B979EACAFFE9FFC5811C566D6FE1F858" box="[1070,1180,1898,1924]" italics="true" pageId="1" pageNumber="244">Securiops</emphasis>
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</taxonomicName>
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claw is long and lacks denticles (
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<figureCitation id="13362A5DFFE9FFC5843856896A23F875" box="[266,350,1934,1961]" captionStart="FIGURES 47–52" captionStartId="8.[151,264,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-18@8.[151,1435,181,1646]" captionTargetPageId="8" captionText="FIGURES 47–52. Procloeon (Securiops) falcatum, larva. 47, articulation of femur and tibia of fore leg, dorsal view; 48, the same, compressed to show junction of patella-tibial suture with row of fine hairs; 49, fragment of tarsus; 50, claw; 51, abdominal tergum X and margin of tergum IX; 52, abdominal tergum X and paraprocts." figureDoi="http://doi.org/10.5281/zenodo.8334199" httpUri="https://zenodo.org/record/8334199/files/figure.png" pageId="1" pageNumber="244">Fig. 50</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB236D8FFE9FFC585F556B66F72F82C" blockId="1.[151,1437,1573,2032]" pageId="1" pageNumber="244">
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<emphasis id="B979EACAFFE9FFC585F556B66931F810" bold="true" box="[199,588,1969,1996]" pageId="1" pageNumber="244">
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||
<emphasis id="B979EACAFFE9FFC585F556B66938F810" bold="true" box="[199,581,1969,1996]" italics="true" pageId="1" pageNumber="244">Lateral spines on larval abdomen</emphasis>
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.
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</emphasis>
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Lateral margins of abdominal segment IX and some previous segments bear spines (autapomorphy of
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<taxonomicName id="4C0D4D5BFFE9FFC5848756D16977F82C" authorityName="Leach" authorityYear="1815" box="[437,522,2006,2032]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
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||
/fg1) (
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||
<figureCitation id="13362A5DFFE9FFC5877956D169E2F82C" box="[587,671,2006,2032]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="1" pageNumber="244">Fig. 17</figureCitation>
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||
;
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||
<bibRefCitation id="EF9C4B29FFE9FFC5879856D168D1F82C" author="Kaltenbach, T. & Suttinun, Ch. & Gattolliat, J. - L." box="[682,940,2006,2032]" pageId="1" pageNumber="244" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. & Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty & Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
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||
Kaltenbach
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||
<emphasis id="B979EACAFFE9FFC5860356D0681FF82C" box="[817,866,2006,2032]" italics="true" pageId="1" pageNumber="244">et al</emphasis>
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||
. 2023
|
||
</bibRefCitation>
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||
: fig. 6a).
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||
</paragraph>
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||
<caption id="DF726650FFEAFFC685A5578F6E4BF934" ID-DOI="http://doi.org/10.5281/zenodo.8334183" ID-Zenodo-Dep="8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="2" pageNumber="245" startId="2.[151,264,1672,1696]" targetBox="[155,1427,188,1633]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BB236D8FFEAFFC685A5578F6E4BF934" blockId="2.[151,1437,1672,1769]" pageId="2" pageNumber="245">
|
||
<emphasis id="B979EACAFFEAFFC685A5578F6A3BF97C" bold="true" box="[151,326,1672,1696]" pageId="2" pageNumber="245">FIGURES 1–24.</emphasis>
|
||
Right tergalii of
|
||
<taxonomicName id="4C0D4D5BFFEAFFC684C3578F6928F943" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[497,597,1672,1695]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACAFFEAFFC684C3578F6928F943" box="[497,597,1672,1695]" italics="true" pageId="2" pageNumber="245">Securiops</emphasis>
|
||
</taxonomicName>
|
||
with dorsal lamella turned out. 1–7,
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||
<taxonomicName id="4C0D4D5BFFEAFFC686F6578F6E78F97C" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Crass" baseAuthorityYear="1947" box="[964,1285,1672,1696]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="falcatum" subGenus="Securiops">
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||
<emphasis id="B979EACAFFEAFFC686F6578F6F50F97C" box="[964,1069,1672,1696]" italics="true" pageId="2" pageNumber="245">Procloeon</emphasis>
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||
(
|
||
<emphasis id="B979EACAFFEAFFC68108578F6FE3F943" box="[1082,1182,1672,1695]" italics="true" pageId="2" pageNumber="245">Securiops</emphasis>
|
||
)
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||
<emphasis id="B979EACAFFEAFFC6819E578F6E78F97C" box="[1196,1285,1672,1696]" italics="true" pageId="2" pageNumber="245">falcatum</emphasis>
|
||
</taxonomicName>
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||
, tergalii I–VII; 8–17, two individuals of
|
||
<taxonomicName id="4C0D4D5BFFEAFFC684AD57AA69FCF918" box="[415,641,1708,1732]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="macafertiorum" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEAFFC684AD57AA6AD3F918" box="[415,430,1709,1732]" italics="true" pageId="2" pageNumber="245">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEAFFC684F457AB6AAEF91F" box="[454,467,1708,1731]" italics="true" pageId="2" pageNumber="245">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEAFFC684D857AB69FCF918" box="[490,641,1708,1732]" italics="true" pageId="2" pageNumber="245">macafertiorum</emphasis>
|
||
</taxonomicName>
|
||
: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24,
|
||
<taxonomicName id="4C0D4D5BFFEAFFC686CE57D66FE1F934" box="[1020,1180,1744,1768]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEAFFC686CE57D66F76F934" box="[1020,1035,1745,1768]" italics="true" pageId="2" pageNumber="245">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEAFFC6811257D76F50F93B" box="[1056,1069,1744,1767]" italics="true" pageId="2" pageNumber="245">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEAFFC6817057D66FE1F934" box="[1090,1180,1745,1768]" italics="true" pageId="2" pageNumber="245">primasia</emphasis>
|
||
</taxonomicName>
|
||
, tergalii I–VII.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB236D8FFEAFFC685F55611689AF838" blockId="2.[151,1436,1814,2021]" pageId="2" pageNumber="245">
|
||
<emphasis id="B979EACAFFEAFFC685F556116A89F8EC" bold="true" box="[199,500,1814,1841]" pageId="2" pageNumber="245">
|
||
<emphasis id="B979EACAFFEAFFC685F556116A93F8ED" bold="true" box="[199,494,1814,1841]" italics="true" pageId="2" pageNumber="245">Scales on larval abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
Abdominal terga and some other body parts of larva bear translucent scales in wide, opened, semicircular sockets. The same in some other baetid taxa, in contrast to angulate and operculated sockets, which are probably initial for Turbanoculata (
|
||
<bibRefCitation id="EF9C4B29FFEAFFC687A356596867F8A4" author="Kluge, N. J." box="[657,794,1886,1912]" pageId="2" pageNumber="245" refId="ref12481" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
|
||
). In most species of
|
||
<taxonomicName id="4C0D4D5BFFEAFFC686C956596F32F8A4" authorityName="Leach" authorityYear="1815" box="[1019,1103,1886,1912]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg1 such scales densely cover most part of surface of abdominal terga and sterna. Among
|
||
<taxonomicName id="4C0D4D5BFFEAFFC68618568468E5F841" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[810,920,1923,1949]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACAFFEAFFC68618568468E5F841" box="[810,920,1923,1949]" italics="true" pageId="2" pageNumber="245">Securiops</emphasis>
|
||
</taxonomicName>
|
||
, such scales are well-developed on abdominal terga III–X and sterna IV–IX in
|
||
<taxonomicName id="4C0D4D5BFFEAFFC684CC56A069D0F81C" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Crass" baseAuthorityYear="1947" box="[510,685,1958,1985]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="falcatum" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEAFFC684CC56A06972F81C" box="[510,527,1959,1984]" italics="true" pageId="2" pageNumber="245">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEAFFC6871456A06949F81D" box="[550,564,1959,1985]" italics="true" pageId="2" pageNumber="245">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEAFFC6877956A169D0F81C" box="[587,685,1958,1984]" italics="true" pageId="2" pageNumber="245">falcatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13362A5DFFEAFFC6878956A16843F81D" box="[699,830,1958,1985]" captionStart="FIGURES 53–58" captionStartId="10.[151,264,1932,1956]" captionTargetBox="[151,1436,274,1908]" captionTargetId="figure-63@10.[151,1436,273,1908]" captionTargetPageId="10" captionText="FIGURES 53–58. Procloeon (Securiops) falcatum, exuviae of larval abdomen. 53–56, terga II, III, IV and IX; 57, sternum IV; 58, constant bifurcate seta on sublateral part of sternum VIII." figureDoi="http://doi.org/10.5281/zenodo.8334201" httpUri="https://zenodo.org/record/8334201/files/figure.png" pageId="2" pageNumber="245">Figs 55–56</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C0D4D5BFFEAFFC6865C56A06F5BF81C" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[878,1062,1958,1985]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="mutadens" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEAFFC6865C56A06802F81C" box="[878,895,1959,1984]" italics="true" pageId="2" pageNumber="245">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEAFFC686A456A068D9F81D" box="[918,932,1959,1985]" italics="true" pageId="2" pageNumber="245">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEAFFC6868956A16F5BF81C" box="[955,1062,1958,1984]" italics="true" pageId="2" pageNumber="245">mutadens</emphasis>
|
||
</taxonomicName>
|
||
such scales are sparse (
|
||
<figureCitation id="13362A5DFFEAFFC6801F56A16EF2F81C" box="[1325,1423,1958,1984]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="2" pageNumber="245">Fig. 103</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C0D4D5BFFEAFFC6858656CC6AD5F838" box="[180,424,1994,2021]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="macafertiorum" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEAFFC6858656CC6BB8F838" box="[180,197,1995,2020]" italics="true" pageId="2" pageNumber="245">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEAFFC685EE56CC6B97F839" box="[220,234,1995,2021]" italics="true" pageId="2" pageNumber="245">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEAFFC6843056CD6AD5F838" box="[258,424,1994,2020]" italics="true" pageId="2" pageNumber="245">macafertiorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D5BFFEAFFC684ED56CC69EDF838" box="[479,656,1994,2021]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="252" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEAFFC684ED56CC6A8DF838" box="[479,496,1995,2020]" italics="true" pageId="2" pageNumber="245">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEAFFC6873556CC6968F839" box="[519,533,1995,2021]" italics="true" pageId="2" pageNumber="245">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEAFFC6871E56CC69EDF838" box="[556,656,1995,2020]" italics="true" pageId="2" pageNumber="245">primasia</emphasis>
|
||
</taxonomicName>
|
||
the scales are completely lost.
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F5519168BDFDC5" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F551916A5AFF6D" bold="true" box="[199,295,150,177]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F551916A5DFF6D" bold="true" box="[199,288,150,177]" italics="true" pageId="3" pageNumber="246">Tergalii</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tergalii have the costal rib, but lack the anal rib and can bear a second lobe, which represents an analproximal expansion curved dorsally (presence of such lobe is an autapomorphy of
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7811751BC6F01FF09" authorityName="Leach" authorityYear="1815" box="[1061,1148,187,213]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg1). In various species of
|
||
<taxonomicName id="4C0D4D5BFFEBFFC785A551D86B90FF25" authorityName="Leach" authorityYear="1815" box="[151,237,223,249]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg1, such second lobe is either present on all tergalii I–VII, or is absent on tergalius VII or some other tergalii of posterior pairs, or is absent on all tergalii. Tergalius VII usually lacks the second lobe and often has a peculiar form sharply different from other tergalii: its anal margin (lacking rib) is straight, and the costal margin (armed with the costal rib) is sharply convex (
|
||
<figureCitation id="13362A5DFFEBFFC78737504C69DCFEB9" box="[517,673,331,357]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="3" pageNumber="246">Figs 7, 17, 24</figureCitation>
|
||
); being the last one, this tergalius is directed posteriorly and adjacent by its anal margin to the lateral margins of abdominal segments VIII–IX (
|
||
<figureCitation id="13362A5DFFEBFFC786D250686F48FE55" box="[992,1077,367,393]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="3" pageNumber="246">Fig. 17</figureCitation>
|
||
). Such shape of tergalius VII is especially characteristic for
|
||
<taxonomicName id="4C0D4D5BFFEBFFC784E85094692EFE71" authorityName="sensu Kluge" authorityYear="2016" box="[474,595,403,429]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1, but can be found in some other taxa, sometimes varying individually (e.g.,
|
||
<bibRefCitation id="EF9C4B29FFEBFFC785E450B06A90FE0D" author="Kluge, N. J. & Novikova, E. A." box="[214,493,439,465]" pageId="3" pageNumber="246" pagination="29 - 54" refId="ref13007" refString="Kluge, N. J. & Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54]" type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
|
||
: figs 7.7–8). Because of this difference between tergalius VII and other tergalii, some authors confuse its costal and anal margins and draw it in overturned position (
|
||
<bibRefCitation id="EF9C4B29FFEBFFC786CC50DC6FF1FE29" author="Gillies, M. T." box="[1022,1164,475,501]" pageId="3" pageNumber="246" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies 1988</bibRefCitation>
|
||
: fig. 19;
|
||
<bibRefCitation id="EF9C4B29FFEBFFC781DE50DC6EE5FE29" author="Gattolliat, J. - L." box="[1260,1432,475,501]" pageId="3" pageNumber="246" pagination="1 - 18" refId="ref11982" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003</bibRefCitation>
|
||
: figs 19, 23;
|
||
<bibRefCitation id="EF9C4B29FFEBFFC7842F50F86A85FDC5" author="Jacobus, L. M. & McCafferty, W. P. & Gattolliat, J. - L." box="[285,504,511,537]" pageId="3" pageNumber="246" pagination="129 - 140" refId="ref12262" refString="Jacobus, L. M., McCafferty, W. P. & Gattolliat, J. - L. (2006) Taxonomy of Afrotropical Securiops, new genus, and Cloeodes Traver (Ephemeroptera: Baetidae). African Entomology, 14 (1), 129 - 140." type="journal article" year="2006">
|
||
Jacobus
|
||
<emphasis id="B979EACAFFEBFFC7844F50F86AD3FDC5" box="[381,430,511,537]" italics="true" pageId="3" pageNumber="246">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
: fig. 19;
|
||
<bibRefCitation id="EF9C4B29FFEBFFC7876850F86821FDC5" author="Kaltenbach, T. & Suttinun, Ch. & Gattolliat, J. - L." box="[602,860,511,537]" pageId="3" pageNumber="246" pagination="127 - 143" refId="ref12315" refString="Kaltenbach, T., Phlai-ngam, S., Suttinun, Ch. & Gattolliat, J. - L. (2023) First report of the Afrotropical genus Securiops Jacobus, McCafferty & Gattolliat (Ephemeroptera, Baetidae) from Southeast Asia, with description of a new species. ZooKeys, 1157, 127 - 143. https: // doi. org / 10.3897 / zookeys. 1157.99642" type="journal article" year="2023">
|
||
Kaltenbach
|
||
<emphasis id="B979EACAFFEBFFC787D350F8686FFDC5" box="[737,786,511,537]" italics="true" pageId="3" pageNumber="246">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
: fig. 7b).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F553256E23FDE1" blockId="3.[151,1437,150,1257]" box="[199,1374,546,573]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553256ADBFDE1" bold="true" box="[199,422,546,573]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553256AE2FDE1" bold="true" box="[199,415,546,573]" italics="true" pageId="3" pageNumber="246">Mobility of tergalii</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tergalii are able to make intensive rhythmic respiratory movements (plesiomorphy).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F5534168A5FD59" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553416A02FDBD" bold="true" box="[199,383,582,609]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553416A0AFDBD" bold="true" box="[199,375,582,609]" italics="true" pageId="3" pageNumber="246">Larval caudalii</emphasis>
|
||
.
|
||
</emphasis>
|
||
Cerci and paracercus have dark rings on every 4th joining; segment preceding this joint bears a row of elongate denticles on its posterior margin (
|
||
<figureCitation id="13362A5DFFEBFFC787EA536C6827FD59" box="[728,858,619,645]" captionStart="FIGURES 70–76" captionStartId="15.[151,264,1837,1861]" captionTargetBox="[152,1435,181,1814]" captionTargetId="figure-16@15.[151,1435,181,1814]" captionTargetPageId="15" captionText="FIGURES 70–76. Procloeon (Securiops) macafertiorum. 70, subimaginal exuviae: apex of tibia and base of tarsus of fore leg of male; 71–76, larval exuviae: 71, caudalii; 72, fragment of cercus; 73, fragment of fore tarsus; 74, abdominal tergum IV; 75, abdominal sternum V; 76, abdominal tergum X, paraproct and margin of tergum IX." figureDoi="http://doi.org/10.5281/zenodo.8334211" httpUri="https://zenodo.org/record/8334211/files/figure.png" pageId="3" pageNumber="246">Figs 71–72</figureCitation>
|
||
,
|
||
<figureCitation id="13362A5DFFEBFFC78657536D68B1FD59" box="[869,972,618,645]" captionStart="FIGURES 120–125" captionStartId="24.[151,264,1756,1780]" captionTargetBox="[151,1435,181,1738]" captionTargetId="figure-18@24.[151,1435,181,1738]" captionTargetPageId="24" captionText="FIGURES 120–125. Procloeon (Securiops) primasia, larval exuviae. 120, base of tibia of fore leg, with focus on dorsal side; 121, the same with focus on ventral side; 122, abdominal tergum V; 123, abdominal sternum VIII; 124, cercus and paracercus; 125, portion of cercus." figureDoi="http://doi.org/10.5281/zenodo.8334239" httpUri="https://zenodo.org/record/8334239/files/figure.png" pageId="3" pageNumber="246">124–125</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F553896948FD11" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553896ACFFD75" bold="true" box="[199,434,654,681]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553896AD6FD75" bold="true" box="[199,427,654,681]" italics="true" pageId="3" pageNumber="246">Subimaginal tarsus</emphasis>
|
||
.
|
||
</emphasis>
|
||
In subimago all tarsomeres on legs of all pairs are covered with pointed microlepides, without blunt microlepides (
|
||
<figureCitation id="13362A5DFFEBFFC784E653B46954FD11" box="[468,553,691,717]" captionStart="FIGURES 70–76" captionStartId="15.[151,264,1837,1861]" captionTargetBox="[152,1435,181,1814]" captionTargetId="figure-16@15.[151,1435,181,1814]" captionTargetPageId="15" captionText="FIGURES 70–76. Procloeon (Securiops) macafertiorum. 70, subimaginal exuviae: apex of tibia and base of tarsus of fore leg of male; 71–76, larval exuviae: 71, caudalii; 72, fragment of cercus; 73, fragment of fore tarsus; 74, abdominal tergum IV; 75, abdominal sternum V; 76, abdominal tergum X, paraproct and margin of tergum IX." figureDoi="http://doi.org/10.5281/zenodo.8334211" httpUri="https://zenodo.org/record/8334211/files/figure.png" pageId="3" pageNumber="246">Fig. 70</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F553D1691EFCC9" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553D16911FD2D" bold="true" box="[199,620,726,753]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F553D16918FD2D" bold="true" box="[199,613,726,753]" italics="true" pageId="3" pageNumber="246">Pleuron of subimaginal mesothorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Posterodorsal process of postsubalar sclerite is diminished, pointed, with concave antero-dorsal margin (
|
||
<figureCitation id="13362A5DFFEBFFC784C153FC6928FCC9" box="[499,597,763,789]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="3" pageNumber="246">Fig. 130</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F552186ACCFC81" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F552186A31FCE5" bold="true" box="[199,332,799,825]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F552186A38FCE5" bold="true" box="[199,325,799,825]" italics="true" pageId="3" pageNumber="246">Fore wing</emphasis>
|
||
.
|
||
</emphasis>
|
||
Fore wing has no more than one marginal intercalary in each space (plesiomorphy within Liberevenata) (
|
||
<figureCitation id="13362A5DFFEBFFC7847052446AD9FC81" box="[322,420,835,861]" captionStart="FIGURES 126–136" captionStartId="25.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1816]" captionTargetId="figure-16@25.[151,1435,181,1816]" captionTargetPageId="25" captionText="FIGURES 126–136. Procloeon (Securiops) primasia. 126–127, male imagines; 128, female imago; 129, subimaginal exuviae of half of mesonotum; 130, subimaginal exuviae of thoracic pleura; 131–133, fore leg of female imago; 134–136, its middle leg. Abbreviations: cx2, cx3, coxae of fore, middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334245" httpUri="https://zenodo.org/record/8334245/files/figure.png" pageId="3" pageNumber="246">Fig. 126</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F552616B9BFBCD" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F552616A35FC5D" bold="true" box="[199,328,870,897]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F552616A3CFC5D" bold="true" box="[199,321,870,897]" italics="true" pageId="3" pageNumber="246">Hind wing</emphasis>
|
||
.
|
||
</emphasis>
|
||
Hind wing, if present, is narrow, with no more than two longitudinal veins, always with prominent, narrow, hooked costal process (
|
||
<figureCitation id="13362A5DFFEBFFC784CB528C6930FC79" box="[505,589,907,933]" captionStart="FIGURES 77–87" captionStartId="16.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-18@16.[151,1436,181,1758]" captionTargetPageId="16" captionText="FIGURES 77–87. Procloeon (Securiops) macafertiorum. 77, 78, male imagines; 79, female imago; 80–87, parts of male imago and subimago reared from larva: 80–81, head and thorax; 82, abdomen; 83, hind wing; 84, subimaginal exuviae of thoracic pleura; 85, subimaginal exuviae of half of mesonotum; 86–87, fore and hind legs. Abbreviations: cx2, cx3, coxae of middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334215" httpUri="https://zenodo.org/record/8334215/files/figure.png" pageId="3" pageNumber="246">Fig. 83</figureCitation>
|
||
); in some species hind wings are completely absent, but no one species has hind wing lacking costal process. Among
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7874A52A8699BFC15" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[632,742,943,969]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACAFFEBFFC7874A52A8699BFC15" box="[632,742,943,969]" italics="true" pageId="3" pageNumber="246">Securiops</emphasis>
|
||
</taxonomicName>
|
||
, hind wings of such structure are present in
|
||
<taxonomicName id="4C0D4D5BFFEBFFC781DB52A86EE1FC15" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Crass" baseAuthorityYear="1947" box="[1257,1436,943,969]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="falcatum" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEBFFC781DB52A86F87FC14" box="[1257,1274,943,968]" italics="true" pageId="3" pageNumber="246">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEBFFC7802152A86E5CFC15" box="[1299,1313,943,969]" italics="true" pageId="3" pageNumber="246">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEBFFC7800852A86EE1FC15" box="[1338,1436,943,969]" italics="true" pageId="3" pageNumber="246">falcatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13362A5DFFEBFFC785AD52D46B8DFC31" box="[159,240,979,1005]" captionStart="FIGURES 59–65" captionStartId="11.[151,264,1814,1838]" captionTargetBox="[157,1430,561,1784]" captionTargetId="figure-179@11.[157,1430,561,1791]" captionTargetPageId="11" captionText="FIGURES 59–65. Procloeon (Securiops) falcatum. 59–60, parts of male subimago extracted from larva: 59, abdomen; 60, genitalia; 61–64, young male subimago: 61, abdomen; 62, genitalia; 63, fore femur; 64, hind wing (partly crumpled); 65, genitalia of male imago (after Crass 1947). Abbreviations: gs1, gs2, gs3, 1st–3rd segments of gonostylus; pb, median projection of penial bridge; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.8334205" httpUri="https://zenodo.org/record/8334205/files/figure.png" pageId="3" pageNumber="246">Fig. 64</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7841C52D4695CFC31" box="[302,545,979,1005]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="macafertiorum" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEBFFC7841C52D46A42FC30" box="[302,319,979,1004]" italics="true" pageId="3" pageNumber="246">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEBFFC7846452D46A19FC31" box="[342,356,979,1005]" italics="true" pageId="3" pageNumber="246">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEBFFC7844952D4695CFC31" box="[379,545,979,1005]" italics="true" pageId="3" pageNumber="246">macafertiorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13362A5DFFEBFFC7870352D469F9FC31" box="[561,644,979,1005]" captionStart="FIGURES 77–87" captionStartId="16.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-18@16.[151,1436,181,1758]" captionTargetPageId="16" captionText="FIGURES 77–87. Procloeon (Securiops) macafertiorum. 77, 78, male imagines; 79, female imago; 80–87, parts of male imago and subimago reared from larva: 80–81, head and thorax; 82, abdomen; 83, hind wing; 84, subimaginal exuviae of thoracic pleura; 85, subimaginal exuviae of half of mesonotum; 86–87, fore and hind legs. Abbreviations: cx2, cx3, coxae of middle and hind legs; lpnc, lateropostnotal crest; mtpl, pleurite of metathorax; pls, pleural suture; psa, postsubalar sclerite." figureDoi="http://doi.org/10.5281/zenodo.8334215" httpUri="https://zenodo.org/record/8334215/files/figure.png" pageId="3" pageNumber="246">Fig. 83</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7878752D46810FC31" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[693,877,979,1005]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="mutadens" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEBFFC7878752D469BBFC30" box="[693,710,979,1004]" italics="true" pageId="3" pageNumber="246">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEBFFC787EF52D46996FC31" box="[733,747,979,1005]" italics="true" pageId="3" pageNumber="246">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEBFFC7863052D46810FC31" box="[770,877,979,1005]" italics="true" pageId="3" pageNumber="246">mutadens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7864952D46F56FC30" box="[891,1067,979,1005]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="primasia" subGenus="Securiops">
|
||
<emphasis id="B979EACAFFEBFFC7864952D468F1FC30" box="[891,908,979,1004]" italics="true" pageId="3" pageNumber="246">P</emphasis>
|
||
. (
|
||
<emphasis id="B979EACAFFEBFFC7869152D468CCFC31" box="[931,945,979,1005]" italics="true" pageId="3" pageNumber="246">S</emphasis>
|
||
.)
|
||
<emphasis id="B979EACAFFEBFFC786F552D46F56FC30" box="[967,1067,979,1004]" italics="true" pageId="3" pageNumber="246">primasia</emphasis>
|
||
</taxonomicName>
|
||
and other species hind wings are absent.
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F5551D686EFB7D" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F5551D6A40FBE9" bold="true" box="[199,317,1050,1077]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F5551D6A48FBE9" bold="true" box="[199,309,1050,1077]" italics="true" pageId="3" pageNumber="246">Gonostyli</emphasis>
|
||
.
|
||
</emphasis>
|
||
The 2nd segment of gonostylus is thickened apically, and the 3rd segment is small and petiolate (autapomorphy of
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7845D55386AB9FB85" authorityName="Leach" authorityYear="1815" box="[367,452,1087,1113]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="271" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg1) (
|
||
<figureCitation id="13362A5DFFEBFFC7873A553869ECFB85" box="[520,657,1087,1113]" captionStart="FIGURES 88–91" captionStartId="17.[151,264,1895,1919]" captionTargetBox="[151,1436,417,1872]" captionTargetId="figure-134@17.[151,1436,417,1872]" captionTargetPageId="17" captionText="FIGURES 88–91. Procloeon (Securiops) macafertiorum. 88, genitalia of male imago reared from larva; 89, its subimaginal exuviae; 90, genitalia of another specimen, ventral view; 91, the same, with focus on penis." figureDoi="http://doi.org/10.5281/zenodo.8334217" httpUri="https://zenodo.org/record/8334217/files/figure.png" pageId="3" pageNumber="246">Figs 88, 90</figureCitation>
|
||
,
|
||
<figureCitation id="13362A5DFFEBFFC78792553869B3FB85" box="[672,718,1087,1113]" captionStart="FIGURES 137–142" captionStartId="26.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-18@26.[151,1436,181,1629]" captionTargetPageId="26" captionText="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." figureDoi="http://doi.org/10.5281/zenodo.8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="3" pageNumber="246">137</figureCitation>
|
||
). Being developed under larval cuticle, subimaginal gonostyli are bent in the
|
||
<taxonomicName id="4C0D4D5BFFEBFFC7847255646AE2FBA1" authorityName="Leach" authorityYear="1815" box="[320,415,1123,1149]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACAFFEBFFC7847255646AE2FBA1" box="[320,415,1123,1149]" italics="true" pageId="3" pageNumber="246">́Cloeon</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54B6887AFFEBFFC7849455646AA4FBA1" box="[422,473,1123,1149]" pageId="3" pageNumber="246">type</typeStatus>
|
||
» pose, that is their 2nd segments are directed laterally, and 3rd segments are directed medially (plesiomorphy within Turbanoculata) (
|
||
<figureCitation id="13362A5DFFEBFFC787805580687BFB7D" box="[690,774,1159,1185]" captionStart="FIGURES 59–65" captionStartId="11.[151,264,1814,1838]" captionTargetBox="[157,1430,561,1784]" captionTargetId="figure-179@11.[157,1430,561,1791]" captionTargetPageId="11" captionText="FIGURES 59–65. Procloeon (Securiops) falcatum. 59–60, parts of male subimago extracted from larva: 59, abdomen; 60, genitalia; 61–64, young male subimago: 61, abdomen; 62, genitalia; 63, fore femur; 64, hind wing (partly crumpled); 65, genitalia of male imago (after Crass 1947). Abbreviations: gs1, gs2, gs3, 1st–3rd segments of gonostylus; pb, median projection of penial bridge; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.8334205" httpUri="https://zenodo.org/record/8334205/files/figure.png" pageId="3" pageNumber="246">Fig. 60</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFEBFFC785F555AC69AFFB35" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F555AC6A73FB19" bold="true" box="[199,270,1195,1221]" pageId="3" pageNumber="246">
|
||
<emphasis id="B979EACAFFEBFFC785F555AC6A7AFB19" bold="true" box="[199,263,1195,1221]" italics="true" pageId="3" pageNumber="246">Penis</emphasis>
|
||
.
|
||
</emphasis>
|
||
In male imago gonovectes are small and incapable for protraction; penial bridge is sclerotized and projected between unistyligers (
|
||
<figureCitation id="13362A5DFFEBFFC784CE55C8692EFB35" box="[508,595,1231,1257]" captionStart="FIGURES 88–91" captionStartId="17.[151,264,1895,1919]" captionTargetBox="[151,1436,417,1872]" captionTargetId="figure-134@17.[151,1436,417,1872]" captionTargetPageId="17" captionText="FIGURES 88–91. Procloeon (Securiops) macafertiorum. 88, genitalia of male imago reared from larva; 89, its subimaginal exuviae; 90, genitalia of another specimen, ventral view; 91, the same, with focus on penis." figureDoi="http://doi.org/10.5281/zenodo.8334217" httpUri="https://zenodo.org/record/8334217/files/figure.png" pageId="3" pageNumber="246">Figs 91</figureCitation>
|
||
,
|
||
<figureCitation id="13362A5DFFEBFFC7876D55C869B8FB35" box="[607,709,1231,1257]" captionStart="FIGURES 137–142" captionStartId="26.[151,264,1653,1677]" captionTargetBox="[151,1436,181,1629]" captionTargetId="figure-18@26.[151,1436,181,1629]" captionTargetPageId="26" captionText="FIGURES 137–142. Procloeon (Securiops) primasia. 137–141, genitalia of male imagines: 137, ventral view; 138, lateral view, right gonostylus and styliger removed; 139, lateral view of penis; 140, apex of penis with focus on its dorsal margin; 141, the same penis with focus on ventral side; 142, subimaginal exuviae of genitalia." figureDoi="http://doi.org/10.5281/zenodo.8334247" httpUri="https://zenodo.org/record/8334247/files/figure.png" pageId="3" pageNumber="246">137–141</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |