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<document id="5FA30F6DCAA9E81364BECAD7A466DEE2" ID-DOI="10.5281/zenodo.12586111" ID-ISSN="2064-2474" ID-Zenodo-Dep="12586111" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="felipe" checkinTime="1719595482395" checkinUser="felipe" docAuthor="Papp, L., Merz, B. &amp; Földvári, M." docDate="2006" docId="03A487CEFFB3F53FE803FB0DD0EBFB30" docLanguage="en" docName="ActaZoolAcadSciH.52.2.97-269.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 52 (2), No. 2" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Heteropterna oroszi L. PAPP 2006, sp. n." docType="treatment" docVersion="3" lastPageNumber="132" masterDocId="FF9DFFB6FF92F51CEA41FF99D277FF96" masterDocTitle="DIPTERA OF THAILAND A summary of the families and genera with references to the species representations" masterLastPageNumber="269" masterPageNumber="97" pageNumber="130" updateTime="1719712550900" updateUser="admin" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="DA4851C79C8F0B500C551E389C9A8DAB">DIPTERA OF THAILAND A summary of the families and genera with references to the species representations</mods:title>
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<mods:namePart id="22DF310E85359D2E8B1AB218DE6577D2">Papp, L.</mods:namePart>
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<mods:namePart id="2597AD64C13D183793A18085366F26EA">Merz, B.</mods:namePart>
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<mods:namePart id="60F94B45EF42522EB3B1B6C6C629B4DA">Földvári, M.</mods:namePart>
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<mods:title id="FE76BC4634B7165CA10AD59E6CE4EF7C">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="25918AE52C6E5447571B6D993DBCE0E8">2006</mods:date>
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<mods:number id="C6CA2C36A15D1C7BCA766E93A8E78CAF">2006-07-31</mods:number>
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<mods:identifier id="0692E4F9DFB6A0992C1F4309DF3C083B" type="ISSN">2064-2474</mods:identifier>
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<treatment id="03A487CEFFB3F53FE803FB0DD0EBFB30" ID-DOI="http://doi.org/10.5281/zenodo.12586179" ID-Zenodo-Dep="12586179" LSID="urn:lsid:plazi:treatment:03A487CEFFB3F53FE803FB0DD0EBFB30" httpUri="http://treatment.plazi.org/id/03A487CEFFB3F53FE803FB0DD0EBFB30" lastPageId="35" lastPageNumber="132" pageId="33" pageNumber="130">
<subSubSection id="C3176553FFB3F53DE803FB0DD187FB38" box="[578,1008,1172,1198]" pageId="33" pageNumber="130" type="nomenclature">
<paragraph id="8BB236D8FFB3F53DE803FB0DD187FB38" blockId="33.[578,1008,1172,1198]" box="[578,1008,1172,1198]" pageId="33" pageNumber="130">
<heading id="D0FA81B4FFB3F53DE803FB0DD187FB38" box="[578,1008,1172,1198]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="130" reason="6">
<taxonomicName id="4C0D4D5BFFB3F53DE803FB0DD1D4FB3B" authority="L. PAPP" authorityName="L. PAPP" authorityYear="2006" box="[578,931,1172,1198]" class="Insecta" family="Keroplatidae" genus="Heteropterna" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="130" phylum="Arthropoda" rank="species" species="oroszi" status="sp. nov.">
<emphasis id="B979EACAFFB3F53DE803FB0DD149FB38" bold="true" box="[578,830,1172,1198]" pageId="33" pageNumber="130">Heteropterna oroszi</emphasis>
L.
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</taxonomicName>
,
<taxonomicNameLabel id="A24A57B1FFB3F53DE9F3FB0DD187FB38" box="[946,1008,1172,1198]" pageId="33" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3176553FFB3F53FE88FFB2ED0EBFB30" lastPageId="35" lastPageNumber="132" pageId="33" pageNumber="130" type="description">
<paragraph id="8BB236D8FFB3F53DE88FFB2ED112FB47" blockId="33.[718,869,1207,1233]" box="[718,869,1207,1233]" pageId="33" pageNumber="130">
<heading id="D0FA81B4FFB3F53DE88FFB2ED112FB47" box="[718,869,1207,1233]" centered="true" fontSize="11" level="3" pageId="33" pageNumber="130" reason="3">
(
<figureCitation id="13362A5DFFB3F53DE897FB2ED12AFB47" box="[726,861,1207,1233]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="33" pageNumber="130">Figs 22–25</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BB236D8FFB3F53DEB3FFB62D106FADF" blockId="33.[317,1270,1275,1353]" pageId="33" pageNumber="130">
<materialsCitation id="3B653C85FFB3F53DEB3FFB62D106FADF" collectingDate="2004-11-14" collectionCode="HNHM, PAPP, FOLDVARI" country="Thailand" location="Khao Chong Botanic Garden" pageId="33" pageNumber="130" specimenCount="1" stateProvince="Trang" typeStatus="holotype">
<typeStatus id="54B6887AFFB3F53DEB3FFB62D3A0FA86" box="[382,471,1275,1296]" pageId="33" pageNumber="130" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED1CAE1DFFB3F53DE859FB62D012FA86" box="[536,613,1275,1296]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="33" pageNumber="130">HNHM</collectionCode>
, left hind leg lost, abdomen with genitalia in a plastic microvial with glycerol):
<collectingCountry id="F31A7648FFB3F53DEBE0FA81D056FABB" box="[417,545,1304,1325]" name="Thailand" pageId="33" pageNumber="130">THAILAND</collectingCountry>
:
<collectingRegion id="49C9F83AFFB3F53DE86BFA81D0D6FABB" box="[554,673,1304,1325]" country="Thailand" name="Trang" pageId="33" pageNumber="130">Trang Prov.</collectingRegion>
,
<location id="8ED26003FFB3F53DE8EBFA81D1B1FABB" LSID="urn:lsid:plazi:treatment:03A487CEFFB3F53FE803FB0DD0EBFB30:8ED26003FFB3F53DE8EBFA81D1B1FABB" box="[682,966,1304,1325]" country="Thailand" name="Khao Chong Botanic Garden" pageId="33" pageNumber="130" stateProvince="Trang">Khao Chong Botanic Garden</location>
, along the stream below waterfall,
<date id="FFB31018FFB3F53DEB2AFAADD384FADF" box="[363,499,1332,1353]" pageId="33" pageNumber="130" value="2004-11-14">
<collectingDate id="EFF7E9F0FFB3F53DEB2AFAADD384FADF" box="[363,499,1332,1353]" pageId="33" pageNumber="130" value="2004-11-14">Nov 14, 2004</collectingDate>
</date>
, No. 30, leg. L.
<smallCapsWord id="8D54A004FFB3F53DE8D7FAADD0B2FADE" baselines="1348,1348" box="[662,709,1332,1353]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Papp" pageId="33" pageNumber="130">
<collectionCode id="ED1CAE1DFFB3F53DE8D7FAADD0B2FADE" box="[662,709,1332,1353]" pageId="33" pageNumber="130">PAPP</collectionCode>
</smallCapsWord>
&amp; M.
<smallCapsWord id="8D54A004FFB3F53DE94BFAADD11CFADE" baselines="1348,1348" box="[778,875,1332,1353]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Foldvari" pageId="33" pageNumber="130">
<collectionCode id="ED1CAE1DFFB3F53DE94BFAADD11CFADE" box="[778,875,1332,1353]" pageId="33" pageNumber="130">FÖLDVÁRI</collectionCode>
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.
</materialsCitation>
</paragraph>
<paragraph id="8BB236D8FFB3F53DEB3FFAF7D60BFA15" blockId="33.[317,1271,1390,1785]" box="[382,1148,1390,1411]" pageId="33" pageNumber="130">Measurements in mm: body length 4.46, wing length 3.37, wing breadth 1.38.</paragraph>
<paragraph id="8BB236D8FFB3F53DEB3FFA12D643F9A6" blockId="33.[317,1271,1390,1785]" pageId="33" pageNumber="130">
Head dark, covered with grey microtomentum, only ocelli are shiny. Palpi minute and black like proboscis. Two large lateral ocelli and a minute anterior one. Face linear at middle. Eyes covered by short whitish, very dense cilia. Basal 11 flagellomeres dark brown, only anterior lateral part of flagellomeres 2–6 with diffuse yellowish stripe. Flagellomeres 12–13 yellow, apical flagellomere bicolorous (
<figureCitation id="13362A5DFFB3F53DEBEEFA67D382F985" box="[431,501,1534,1555]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="33" pageNumber="130">Fig. 25</figureCitation>
) and slightly bifid apically (best seen subdorsally): medial part yellow, rest dark brown. Flagellomeres much flattened, medial ones 4 times broader than long.
</paragraph>
<paragraph id="8BB236D8FFB3F53DEB3FF9A1D6C4F96F" blockId="33.[317,1271,1390,1785]" pageId="33" pageNumber="130">Scutum dark brown but anterior half yellow with a thin sagittal brown line and a pair of rectangular brown stripe, which embrace (with the dark lateral and caudal colour) a pair of quadrate yellow spots. Supraalar area with long dense black setae. Postpronotum, antepronotum and proepisternum dark brown, bulging, latter two sclerites with long black setae. Prosternum with some lateral setae. Anepisternum dark brown, katepisternum (preepisternum), anepimeron and laterotergite less dark brown, metepisternum, metepimeron and metanotum dark brown again. Scutellum with medium long apical setae, no discal setae. Subscutellar membraneous area forms a regular triangle.</paragraph>
<caption id="DF726650FFB0F53EEB7CF950D682F96D" ID-DOI="http://doi.org/10.5281/zenodo.12586123" ID-Zenodo-Dep="12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="34" pageNumber="131" startId="34.[317,360,1737,1758]" targetBox="[322,1267,478,1678]" targetPageId="34" targetType="figure">
<paragraph id="8BB236D8FFB0F53EEB7CF950D682F96D" blockId="34.[317,1270,1737,1816]" pageId="34" pageNumber="131">
<emphasis id="B979EACAFFB0F53EEB7CF950D3DDF948" bold="true" box="[317,426,1737,1758]" pageId="34" pageNumber="131">Figs 22–25</emphasis>
.
<taxonomicName id="4C0D4D5BFFB0F53EEBF7F953D00FF949" authorityName="Papp &amp; Merz &amp; Földvári" authorityYear="2006" box="[438,632,1738,1759]" class="Insecta" family="Keroplatidae" genus="Heteropterna" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="131" phylum="Arthropoda" rank="species" species="oroszi" status="sp. nov.">
<emphasis id="B979EACAFFB0F53EEBF7F953D00FF949" box="[438,632,1738,1759]" italics="true" pageId="34" pageNumber="131">Heteropterna oroszi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A57B1FFB0F53EE83FF953D0C4F949" box="[638,691,1738,1759]" pageId="34" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner
</paragraph>
</caption>
<paragraph id="8BB236D8FFB0F53EEB92F89AD617F88E" blockId="34.[317,1270,1737,1816]" box="[467,1120,1795,1816]" pageId="34" pageNumber="131">
(lateral) view. Scales:
<quantity id="4CF59B3DFFB0F53EE8EBF89AD084F88E" box="[682,755,1795,1816]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="131" unit="mm" value="0.2">0.2 mm</quantity>
for
<figureCitation id="13362A5DFFB0F53EE95AF89AD1FEF88E" box="[795,905,1795,1816]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="34" pageNumber="131">Figs 22–24</figureCitation>
,
<quantity id="4CF59B3DFFB0F53EE9D3F89AD184F88E" box="[914,1011,1795,1816]" metricMagnitude="-4" metricUnit="m" metricValue="1.09" pageId="34" pageNumber="131" unit="mm" value="0.109">0.109 mm</quantity>
for
<figureCitation id="13362A5DFFB0F53EEE5DF89AD617F88E" box="[1052,1120,1795,1816]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="34" pageNumber="131">Fig. 25</figureCitation>
</paragraph>
<paragraph id="8BB236D8FFB1F53FEB3FFE2DD6B0FDB6" blockId="35.[317,1270,436,976]" pageId="35" pageNumber="132">
Coxae all blackish brown, bases of femora dark. Legs mainly light brown. Coxae, particularly cx1 and cx2 with dense long black setae. Tibial spurs: 1 (brown): 2 (white): 2 (white). Hind tibia thickened in its apical half (almost bat-shaped) and darkened. Hind basitarsus very long (
<quantity id="4CF59B3DFFB1F53FEED0FE77D69CFD95" box="[1169,1259,494,515]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="35" pageNumber="132" unit="mm" value="1.13">1.13 mm</quantity>
), thick, flattened laterally, width
<quantity id="4CF59B3DFFB1F53FE831FD92D0B0FDB6" box="[624,711,523,544]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="132" unit="mm" value="0.12">0.12 mm</quantity>
at middle. Claws short thick with ventral dentation.
</paragraph>
<paragraph id="8BB236D8FFB1F53FEB3FFDB1D136FD7F" blockId="35.[317,1270,436,976]" pageId="35" pageNumber="132">
Wing’s basic colour light brown and covered with dense microtrichia. Veins thick, dark brown, particularly so for radial veins. A large dark brown spot in costal, subcosta and r1 cells to the apex of Sc, down to M1–2 fork base; another quadrate dark spot from 2/5 of Sc-R1 costal section to the apex of R4 into r5 cell, where it is confluent with the apical (somewhat less dark) spot. R1 setose in its whole length and R5 setose from base of M1–2 to apex, Sc and other veins bare.
<collectingRegion id="49C9F83AFFB1F53FEE34FD02D6DBFD26" box="[1141,1196,667,688]" country="Tanzania" name="Pwani" pageId="35" pageNumber="132">Costa</collectingRegion>
continued on 2/5 of R5-M1 costal section (
<quantity id="4CF59B3DFFB1F53FE8EFFD21D160FD5B" box="[686,791,696,717]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" pageId="35" pageNumber="132" unit="mm" value="0.215">0.215 mm</quantity>
). M2 does not reach wing margin. Cu1 strongly downcurving, Cu2 distinct distally to Cu1 curvature.
</paragraph>
<paragraph id="8BB236D8FFB1F53FEB3FFD68D699FC46" blockId="35.[317,1270,436,976]" pageId="35" pageNumber="132">
First abdominal sclerites dark brown, tergites 2- yellow with brown caudal margin. Sternites 2- with a pair of lateral and 2 medial brown spots, which are confluent on S3 and S4, S5 to S7 with broad brown caudal margin, T8 and S8 dark brown. Tergite 9 (
<figureCitation id="13362A5DFFB1F53FE9E4FCB2D19BFCD6" box="[933,1004,811,832]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="35" pageNumber="132">Fig. 22</figureCitation>
) rather long, with a pair of digitiform lateral caudal processes, proctiger with almost sharp lateral projections, cerci normal with medium-long thin setae. Gonocoxites (
<figureCitation id="13362A5DFFB1F53FE8F4FCFDD08BFCEF" box="[693,764,868,889]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="35" pageNumber="132">Fig. 23</figureCitation>
) with a medium-deep but broad caudal incision and with a pair of medial processes, which bear short thick black setae. Gonostylus (
<figureCitation id="13362A5DFFB1F53FEE7EFC18D6F2FC00" box="[1087,1157,897,918]" captionStart="Figs 22–25" captionStartId="34.[317,360,1737,1758]" captionTargetBox="[322,1267,478,1678]" captionTargetId="figure-22@34.[317,1269,476,1684]" captionTargetPageId="34" captionText="Figs 22–25. Heteropterna oroszi sp. n., holotype male. 22 = tergite 9 with cerci and proctiger, 23 = gonocoxite and gonostylus, ventral view, 24 = gonostylus, lateral view, 25 = apex of antenna, inner" figureDoi="http://doi.org/10.5281/zenodo.12586123" httpUri="https://zenodo.org/record/12586123/files/figure.png" pageId="35" pageNumber="132">Fig. 24</figureCitation>
) long, apex quadrate in lateral view, basal process rather large, with a black apical and another black subapical teeth. No long setae on basal process, otherwise gonostlus covered by almost evenly thick setae.
</paragraph>
<paragraph id="8BB236D8FFB1F53FEB3FFC6ED19CFBA3" blockId="35.[317,1269,1015,1077]" pageId="35" pageNumber="132">
This species runs to
<taxonomicName id="4C0D4D5BFFB1F53FE826FC61D1C6FB87" authority="MATILE, 1990" authorityName="MATILE" authorityYear="1990" box="[615,945,1015,1042]" class="Insecta" family="Keroplatidae" genus="Heteropterna" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="132" phylum="Arthropoda" rank="species" species="fenestralis">
<emphasis id="B979EACAFFB1F53FE826FC61D174FB84" box="[615,771,1016,1042]" italics="true" pageId="35" pageNumber="132">H. fenestralis</emphasis>
<bibRefCitation id="EF9C4B29FFB1F53FE948FC6ED1C6FB87" author="MATILE, L." box="[777,945,1015,1041]" pageId="35" pageNumber="132" pagination="1 - 682" refId="ref85528" refString="MATILE, L. (1990) Recherches sur la systematique et l'evolution des Keroplatidae (Diptera, Mycetophiloidea). Memoires du Museum National d'Histoire Naturelle Paris, Zoologie 148: 1 - 682." type="journal article" year="1990">
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,
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</taxonomicName>
in
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s key, but wings are much different: subcostal “windows” are much longer.
</paragraph>
<paragraph id="8BB236D8FFB1F53FEB3FFBC1D0EBFB30" blockId="35.[317,1270,1112,1190]" pageId="35" pageNumber="132">
I name this new species to the honour of Mr.
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(Dept. of Zoology, HNHM), who was our partner during the collection trip, and who collected so many valuable dipterous specimens for the
<taxonomicName id="4C0D4D5BFFB1F53FEBC4FB08D3BAFB30" box="[389,461,1169,1190]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="132" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
Collection formerly.
</paragraph>
</subSubSection>
</treatment>
</document>
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