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<document id="D2DB7E97103118C165CCB48D8DD70708" ID-CLB-Dataset="3667" ID-DOI="10.11646/megataxa.2.1.1" ID-GBIF-Dataset="f7793976-e4e9-4562-a9cd-eda910d77fa0" ID-ISSN="2703-3090" ID-Zenodo-Dep="5597237" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614072591502" checkinUser="plazi" docAuthor="Miglio, Laura Tavares, Pérez-Miles, Fernando &amp; Bonaldo, Alexandre B." docDate="2020" docId="0975136AFF43CE8CFCD5F87CD9A93B71" docLanguage="en" docName="megataxa.2.1.1.pdf.imf" docOrigin="Megataxa 2 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Actinopus palmar Rios-Tamayo &amp; Goloboff 2018" docType="treatment" docVersion="11" lastPageNumber="210" masterDocId="F54C6B12FF8DCE5DFFFFFFD1DB533C2E" masterDocTitle="Taxonomic Revision of the Spider Genus Actinopus Perty, 1833 (Araneae, Mygalomorphae, Actinopodidae)" masterLastPageNumber="256" masterPageNumber="1" pageNumber="207" updateTime="1698930906725" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="9D35B2B73915478FDAA6F68A680B5BAD">Taxonomic Revision of the Spider Genus Actinopus Perty, 1833 (Araneae, Mygalomorphae, Actinopodidae)</mods:title>
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<mods:namePart id="8AAEBC001A913483C53FD0A15E7B6B1C">Miglio, Laura Tavares</mods:namePart>
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<mods:affiliation id="B3106834895ED108A336955C8306DA4F">Museu Paraense Emílio Goeldi, Coordenação de Zoologia, Laboratório de Aracnologia, Av. Perimetral, 1901, Terra Firme, Cep: 66077 - 830, Belém, Pará, Brazil. https: // orcid. org / 0000 - 0002 - 9101 - 8657,</mods:affiliation>
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<mods:namePart id="0401955CE374434D1558E212ECD2AAF0">Pérez-Miles, Fernando</mods:namePart>
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<mods:affiliation id="C8C57E76588035C61F8A9ACCB08EA5EA">Facultad de Ciencias, Sección Entomología, Iguá 4225, 11400, Montevideo, Uruguay. ® myga @ fcien. edu. uy, https: // orcid. org / 0000 - 0002 - 4367 - 6276</mods:affiliation>
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<mods:namePart id="371DE511A7AF086E97754E041E22EA67">Bonaldo, Alexandre B.</mods:namePart>
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<mods:affiliation id="ADE602B7CBDA497C28F9502FB3FDF88B">Museu Paraense Emílio Goeldi, Coordenação de Zoologia, Laboratório de Aracnologia, Av. Perimetral, 1901, Terra Firme, Cep: 66077 - 830, Belém, Pará, Brazil. https: // orcid. org / 0000 - 0002 - 9101 - 8657,</mods:affiliation>
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<mods:title id="9F11DFDF7D374CC33F9AD2AF515DC7D0">Megataxa</mods:title>
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<taxonomicName id="46DCD9FFFF43CE93FCD5F87CDEDD3BE9" ID-CoL="64PK9" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[810,1422,1964,1992]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="206" pageNumber="207" phylum="Arthropoda" rank="species" species="palmar">
<emphasis id="B3A87E6EFF43CE93FCD5F87CDEDD3BE9" bold="true" box="[810,1422,1964,1992]" pageId="206" pageNumber="207">
<emphasis id="B3A87E6EFF43CE93FCD5F87CDF523BE6" bold="true" box="[810,1025,1965,1992]" italics="true" pageId="206" pageNumber="207">Actinopus palmar</emphasis>
<bibRefCitation id="E54DDF8DFF43CE93FBF6F87DDEDD3BE9" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[1033,1422,1964,1991]" pageId="206" pageNumber="207" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</emphasis>
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<figureCitation id="19E7BEF9FF43CE93FCD5F803D8953BC3" box="[810,966,2002,2029]" captionStart-0="FIGURE 187" captionStart-1="FIGURE 188" captionStart-2="FIGURE 189" captionStartId-0="207.[136,235,1155,1179]" captionStartId-1="208.[136,235,1155,1179]" captionStartId-2="209.[136,235,1502,1526]" captionTargetBox-0="[180,1407,205,1115]" captionTargetBox-1="[180,1407,206,1116]" captionTargetBox-2="[159,1425,209,1451]" captionTargetId-0="figure-374@207.[151,1436,189,1132]" captionTargetId-1="figure-320@208.[151,1436,189,1132]" captionTargetId-2="figure-137@209.[151,1435,189,1478]" captionTargetPageId-0="207" captionTargetPageId-1="208" captionTargetPageId-2="209" captionText-0="FIGURE 187. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male:A–D MACN-Ar 19826:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral." captionText-1="FIGURE 188. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: 815–818 MACN-Ar 19826:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." captionText-2="FIGURE 189. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: A–E MACN 19826: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PAc- Prolateral Accessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrows: Serrated area. E. Rastellum, dorsal; F. Rastellum, ventral. Scales lines: A–C: 1 mm; D–E: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5597643" figureDoi-1="http://doi.org/10.5281/zenodo.5597646" figureDoi-2="http://doi.org/10.5281/zenodo.5597648" httpUri-0="https://zenodo.org/record/5597643/files/figure.png" httpUri-1="https://zenodo.org/record/5597646/files/figure.png" httpUri-2="https://zenodo.org/record/5597648/files/figure.png" pageId="206" pageNumber="207">Figs 187–189</figureCitation>
,
<figureCitation id="19E7BEF9FF43CE93FC2EF802DF753BC3" box="[977,1062,2003,2029]" captionStart="MAP 11" captionStartId="196.[136,192,1121,1145]" captionTargetBox="[157,1428,195,1089]" captionTargetId="figure-346@196.[151,1435,189,1097]" captionTargetPageId="196" captionText="MAP 11. Distribution map of Actinopus robustus O. Pickard-Cambridge, 1892, Actinopus guajara sp. nov., Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, Actinopus lomalinda sp. nov., Actinopus concinnus sp. nov. and Actinopus cucutaensis (Mello- Leit„o, 1941)." figureDoi="http://doi.org/10.5281/zenodo.5655450" httpUri="https://zenodo.org/record/5655450/files/figure.png" pageId="206" pageNumber="207">Map 11</figureCitation>
</paragraph>
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<paragraph id="8163A27CFF43CE92FCD5F7C7DBA7399A" blockId="206.[810,1451,2070,2092]" lastBlockId="207.[189,778,1342,1460]" lastPageId="207" lastPageNumber="208" pageId="206" pageNumber="207">
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<treatmentCitation id="007D846DFF43CE93FCD5F7C7DE6E3405" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[810,1341,2070,2092]" httpUri="http://treatment.plazi.org/id/C22D980AFFC6F35A730CF9FDD56DFA50" page="81" pageId="206" pageNumber="207" year="2018">
<taxonomicName id="46DCD9FFFF43CE93FCD5F7C7DE6E3405" ID-CoL="64PK9" authority="Rios-Tamayo &amp; Goloboff, 2018: 81" authorityName="Rios-Tamayo &amp; Goloboff" authorityPageNumber="81" authorityYear="2018" box="[810,1341,2070,2092]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="206" pageNumber="207" phylum="Arthropoda" rank="species" species="palmar">
<emphasis id="B3A87E6EFF43CE93FCD5F7C7D8853405" box="[810,982,2070,2091]" italics="true" pageId="206" pageNumber="207">Actinopus palmar</emphasis>
<bibRefCitation id="E54DDF8DFF43CE93FC21F7C7DE6E3405" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[990,1341,2070,2092]" pageId="206" pageNumber="207" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018: 81</bibRefCitation>
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, fig. 26 A–G, 45 (
<typeStatus id="5E671CDEFF42CE92FF02FAEEDA00397B" box="[253,339,1343,1365]" pageId="207" pageNumber="208" type="holotype">holotype</typeStatus>
♂, [
<geoCoordinate id="E4E8C4BBFF42CE92FE85FAEEDA82397A" box="[378,465,1343,1365]" degrees="31" direction="south" minutes="51" orientation="latitude" pageId="207" pageNumber="208" precision="925" value="-31.85">31º 51’ S</geoCoordinate>
<geoCoordinate id="E4E8C4BBFF42CE92FE29FAEED96B397B" box="[470,568,1343,1365]" degrees="58" direction="west" minutes="19" orientation="longitude" pageId="207" pageNumber="208" precision="925" value="-58.316666">58º 19’ W</geoCoordinate>
], Parque Nacional “El Palmar”,
<collectingRegion id="43186C9EFF42CE92FEE8FAB1DA2D395B" box="[279,382,1375,1397]" country="Argentina" name="Entre Rios" pageId="207" pageNumber="208">Entre Ríos</collectingRegion>
,
<date id="F56284BCFF42CE92FE78FA8ED95D395B" box="[391,526,1375,1397]" pageId="207" pageNumber="208" value="1980-02-12" valueMax="1980-02-16" valueMin="1980-02-12">12–16.ii.1980</date>
, P. Grosso and P. Goloboff leg., MACN-Ar 198206, examined);
<bibRefCitation id="E54DDF8DFF42CE92FDD2FAAEDBA3399A" author="World Spider Catalog" pageId="207" pageNumber="208" refId="ref140996" refString="World Spider Catalog (2020) World Spider Catalog. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 22 July 2020)" type="url" year="2020">World Spider Catalog, 2020</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
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<caption id="D5A3F2F4FF42CE92FF77FB52D9933891" ID-DOI="http://doi.org/10.5281/zenodo.5597643" ID-Zenodo-Dep="5597643" httpUri="https://zenodo.org/record/5597643/files/figure.png" pageId="207" pageNumber="208" startId="207.[136,235,1155,1179]" targetBox="[180,1407,205,1115]" targetPageId="207">
<paragraph id="8163A27CFF42CE92FF77FB52D9933891" blockId="207.[136,1452,1155,1215]" pageId="207" pageNumber="208">
<emphasis id="B3A87E6EFF42CE92FF77FB52DA4F38B5" bold="true" box="[136,284,1155,1179]" pageId="207" pageNumber="208">FIGURE 187.</emphasis>
<taxonomicName id="46DCD9FFFF42CE92FEDEFB55D84F38B5" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[289,796,1155,1179]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="palmar">
<emphasis id="B3A87E6EFF42CE92FEDEFB55DA8738B5" box="[289,468,1155,1179]" italics="true" pageId="207" pageNumber="208">Actinopus palmar</emphasis>
<bibRefCitation id="E54DDF8DFF42CE92FE26FB52D84F38B5" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[473,796,1155,1179]" pageId="207" pageNumber="208" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</taxonomicName>
, male: A–D MACN-Ar 19826: A. Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral.
</paragraph>
</caption>
<subSubSection id="C9C6F1F7FF42CE92FF77FA34D85B3B06" pageId="207" pageNumber="208" type="diagnosis">
<paragraph id="8163A27CFF42CE92FF77FA34D85B3B06" blockId="207.[136,778,1509,2092]" pageId="207" pageNumber="208">
<emphasis id="B3A87E6EFF42CE92FF77FA34DA5139D1" bold="true" box="[136,258,1509,1535]" pageId="207" pageNumber="208">Diagnosis.</emphasis>
Males of
<taxonomicName id="46DCD9FFFF42CE92FE87FA37D91239D1" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[376,577,1509,1536]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="palmar">
<emphasis id="B3A87E6EFF42CE92FE87FA37D91239D1" box="[376,577,1509,1536]" italics="true" pageId="207" pageNumber="208">Actinopus palmar</emphasis>
</taxonomicName>
differ from those of all other species of the group
<taxonomicName id="46DCD9FFFF42CE92FDD4F9DAD9FD3A0A" authorityName="Mello-Leitao" authorityYear="1941" box="[555,686,1547,1572]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="cucutaensis">
<emphasis id="B3A87E6EFF42CE92FDD4F9DAD9FD3A0A" box="[555,686,1547,1572]" italics="true" pageId="207" pageNumber="208">cucutaensis</emphasis>
</taxonomicName>
by the narrow ATP base, copulatory bulb with three prolateral keels in prolaterodorsal view and ATP with a big serrated area (
<figureCitation id="19E7BEF9FF42CE92FF38F9ABDA753ABA" box="[199,294,1658,1684]" captionStart="FIGURE 189" captionStartId="209.[136,235,1502,1526]" captionTargetBox="[159,1425,209,1451]" captionTargetId="figure-137@209.[151,1435,189,1478]" captionTargetPageId="209" captionText="FIGURE 189. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: A–E MACN 19826: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PAc- Prolateral Accessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrows: Serrated area. E. Rastellum, dorsal; F. Rastellum, ventral. Scales lines: A–C: 1 mm; D–E: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5597648" httpUri="https://zenodo.org/record/5597648/files/figure.png" pageId="207" pageNumber="208">Fig. 189</figureCitation>
A–C). Additionally, they differ from those of all other species of
<taxonomicName id="46DCD9FFFF42CE92FE60F94ED95F3A97" authorityName="Perty" authorityYear="1833" box="[415,524,1695,1721]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B3A87E6EFF42CE92FE60F94ED95F3A97" box="[415,524,1695,1721]" italics="true" pageId="207" pageNumber="208">Actinopus</emphasis>
</taxonomicName>
, except
<taxonomicName id="46DCD9FFFF42CE92FD88F94ED8503A97" baseAuthorityName="Keyserling" baseAuthorityYear="1891" box="[631,771,1695,1721]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="B3A87E6EFF42CE92FD88F94ED8503A97" box="[631,771,1695,1721]" italics="true" pageId="207" pageNumber="208">A. crassipes</emphasis>
</taxonomicName>
,
<taxonomicName id="46DCD9FFFF42CE92FF77F915DA333AF0" authorityName="Mello-Leitao" authorityYear="1923" box="[136,352,1732,1758]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="255" phylum="Arthropoda" rank="species" species="dubiomaculatus">
A.
<emphasis id="B3A87E6EFF42CE92FF51F915DA333AF0" box="[174,352,1732,1758]" italics="true" pageId="207" pageNumber="208">dubiomaculatus</emphasis>
</taxonomicName>
<emphasis id="B3A87E6EFF42CE92FE9FF915DBA73B2D" italics="true" pageId="207" pageNumber="208">
,
<taxonomicName id="46DCD9FFFF42CE92FE90F914DABA3AF0" authorityName="Mello-Leitao" authorityYear="1920" box="[367,489,1732,1759]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pusillus">A. pusillus</taxonomicName>
,
<taxonomicName id="46DCD9FFFF42CE92FE07F914D9053AF0" box="[504,598,1733,1759]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="emas">A. emas</taxonomicName>
,
<taxonomicName id="46DCD9FFFF42CE92FD9AF914D9B03AF0" box="[613,739,1732,1759]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="bocaina">A. bocaina</taxonomicName>
,
<taxonomicName id="46DCD9FFFF42CE92FD0DF914DBA73B2D" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="laventana">A. laventana</taxonomicName>
</emphasis>
and
<taxonomicName id="46DCD9FFFF42CE92FEC4F93BD9783B2D" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[315,555,1769,1796]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="gerschiapelliarum">
<emphasis id="B3A87E6EFF42CE92FEC4F93BD9783B2D" box="[315,555,1769,1796]" italics="true" pageId="207" pageNumber="208">A. gerschiapelliarum</emphasis>
</taxonomicName>
, by the distal leg articles being paler than proximal leg articles and the palp.
</paragraph>
</subSubSection>
<subSubSection id="C9C6F1F7FF42CE8DFF42F8E5DF3A3426" lastPageId="208" lastPageNumber="209" pageId="207" pageNumber="208" type="description">
<paragraph id="8163A27CFF42CE8DFF42F8E5D9EA3BB7" blockId="207.[136,778,1509,2092]" lastBlockId="208.[136,778,1327,1945]" lastPageId="208" lastPageNumber="209" pageId="207" pageNumber="208">
MALE (MACN 19826): Total length 8.62; Carapace, long 4.3; wide 3.6. Carapace anterior part tapering. Eyes. Anterior row slightly procurved, posterior row recurved (
<figureCitation id="19E7BEF9FF42CE92FF6FF872DA5F3B93" box="[144,268,1955,1981]" captionStart="FIGURE 187" captionStartId="207.[136,235,1155,1179]" captionTargetBox="[180,1407,205,1115]" captionTargetId="figure-374@207.[151,1436,189,1132]" captionTargetPageId="207" captionText="FIGURE 187. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male:A–D MACN-Ar 19826:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597643" httpUri="https://zenodo.org/record/5597643/files/figure.png" pageId="207" pageNumber="208">Fig. 187 B</figureCitation>
). One bristle between AME-clypeus and also ALE–PLE. Sternum with eight sigilla near center and two distal sigilla very fused to each other (
<figureCitation id="19E7BEF9FF42CE92FD83F83CD9AE3429" box="[636,765,2029,2055]" captionStart="FIGURE 187" captionStartId="207.[136,235,1155,1179]" captionTargetBox="[180,1407,205,1115]" captionTargetId="figure-374@207.[151,1436,189,1132]" captionTargetPageId="207" captionText="FIGURE 187. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male:A–D MACN-Ar 19826:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597643" httpUri="https://zenodo.org/record/5597643/files/figure.png" pageId="207" pageNumber="208">Fig. 187 C</figureCitation>
). Rastellum protuberant, inverted V-shaped, hirsute, with many spines on apex (
<figureCitation id="19E7BEF9FF42CE92FBC8FAEFDFC93977" box="[1079,1178,1342,1369]" captionStart="FIGURE 189" captionStartId="209.[136,235,1502,1526]" captionTargetBox="[159,1425,209,1451]" captionTargetId="figure-137@209.[151,1435,189,1478]" captionTargetPageId="209" captionText="FIGURE 189. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: A–E MACN 19826: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PAc- Prolateral Accessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrows: Serrated area. E. Rastellum, dorsal; F. Rastellum, ventral. Scales lines: A–C: 1 mm; D–E: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5597648" httpUri="https://zenodo.org/record/5597648/files/figure.png" pageId="207" pageNumber="208">Fig. 189</figureCitation>
D–E). Chelicerae with denticles along Prolateral row (
<figureCitation id="19E7BEF9FF42CE92FB6CFAB2DE453953" box="[1171,1302,1379,1405]" captionStart="FIGURE 187" captionStartId="207.[136,235,1155,1179]" captionTargetBox="[180,1407,205,1115]" captionTargetId="figure-374@207.[151,1436,189,1132]" captionTargetPageId="207" captionText="FIGURE 187. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male:A–D MACN-Ar 19826:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597643" httpUri="https://zenodo.org/record/5597643/files/figure.png" pageId="207" pageNumber="208">Fig. 187 D</figureCitation>
). Patella and tibia III with distal crown of well-developed thorns (
<figureCitation id="19E7BEF9FF42CE92FA81FA56D80739EB" captionStart="FIGURE 188" captionStartId="208.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1116]" captionTargetId="figure-320@208.[151,1436,189,1132]" captionTargetPageId="208" captionText="FIGURE 188. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: 815–818 MACN-Ar 19826:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597646" httpUri="https://zenodo.org/record/5597646/files/figure.png" pageId="207" pageNumber="208">Fig. 188</figureCitation>
A–B).Patellae III and IV with spines on prolaterodorsal face (
<figureCitation id="19E7BEF9FF42CE92FC95FA1EDF5C39C7" box="[874,1039,1487,1513]" captionStart="FIGURE 188" captionStartId="208.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1116]" captionTargetId="figure-320@208.[151,1436,189,1132]" captionTargetPageId="208" captionText="FIGURE 188. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: 815–818 MACN-Ar 19826:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597646" httpUri="https://zenodo.org/record/5597646/files/figure.png" pageId="207" pageNumber="208">Fig. 188 A, C</figureCitation>
). Patella IV without spines dorsally (
<figureCitation id="19E7BEF9FF42CE92FCCDFA22D8E03A20" box="[818,947,1523,1550]" captionStart="FIGURE 188" captionStartId="208.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1116]" captionTargetId="figure-320@208.[151,1436,189,1132]" captionTargetPageId="208" captionText="FIGURE 188. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: 815–818 MACN-Ar 19826:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597646" httpUri="https://zenodo.org/record/5597646/files/figure.png" pageId="207" pageNumber="208">Fig. 188 D</figureCitation>
). Ventral pseudoscopulae occupying 30% of tarsi I, 50% of II and 100% of III and IV. Pseudoscopula of tarsi I and II with setae spaced, forming diffuse group of bristles; On tarsi III and IV, compact. Carapace and Chelicerae reddish brown, sternum yellowish brown, coxae, trochantera, femora and patellae reddish brown, tibiae, metatarsi and tarsi yellowish brown, abdomen pale gray (
<figureCitation id="19E7BEF9FF42CE92FC95F920D8B63B25" box="[874,997,1777,1803]" captionStart="FIGURE 187" captionStartId="207.[136,235,1155,1179]" captionTargetBox="[180,1407,205,1115]" captionTargetId="figure-374@207.[151,1436,189,1132]" captionTargetPageId="207" captionText="FIGURE 187. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male:A–D MACN-Ar 19826:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597643" httpUri="https://zenodo.org/record/5597643/files/figure.png" pageId="207" pageNumber="208">Fig. 187 A</figureCitation>
).
<emphasis id="B3A87E6EFF42CE92FC07F920DF973B24" bold="true" box="[1016,1220,1776,1803]" pageId="207" pageNumber="208">Eyes: Diameters:</emphasis>
PME 0.15 PLE 0.18 ALE 0.28, AME 0.21; MOQ: Length 0.68, front width 1.78, back width 1.5;
<emphasis id="B3A87E6EFF42CE92FBD4F8E8DF8D3B7D" bold="true" box="[1067,1246,1849,1875]" pageId="207" pageNumber="208">Interdistances:</emphasis>
PME–PME 0.90, PLE–PME 0.12, AME–AME 0.09, ALE–AME 0.65, ALE–PLE 0.34, AME–PME 0.25.
<emphasis id="B3A87E6EFF42CE92FB37F850DE353BB2" bold="true" box="[1224,1382,1921,1948]" pageId="207" pageNumber="208">Ocular area:</emphasis>
OAL 2.53, OAW 2.62, and IF 1.4.
<emphasis id="B3A87E6EFF42CE92FB96F877DFE23BEE" bold="true" box="[1129,1201,1958,1984]" pageId="207" pageNumber="208">Body:</emphasis>
Clypeus: 0.15; Fovea: 1.3; Labium: long 0.85; wide 0.75; Chelicerae: long 2.43; wide 1.18; Sternum: long 2.8; wide 2.1. Abdomen: long 4.56; wide 3.5.
<emphasis id="B3A87E6EFF42CE92FC23F7C3DF943402" bold="true" box="[988,1223,2066,2092]" pageId="207" pageNumber="208">Leg measurements:</emphasis>
I: Fe 3.62/ Pa 1.62/ Ti 2.12/ Me 3.25/ Ta 1.75/ total 12.37. II: 4/ 1.75/ 2.5/ 2.87/ 1.87/ 13. III: 3/ 2.37/ 1.75/ 3/ 2.125/ 12.25. IV: 4.12/ 1.37/ 3.25/ 3.12/ 2.37/ 14.25. Formula 4123.
<emphasis id="B3A87E6EFF5DCE8DFD72FAA8D85939BD" bold="true" box="[653,778,1401,1427]" pageId="208" pageNumber="209">Spination:</emphasis>
I—Fe v0, d0, p0, r0; Pa v0-0-1, d0, p0, r0; ti v5-6-8, d0, p0, r0; Me v5-14-11, d0, p1-0-2, r0; ta v6-10-9, d0, p0-0- 2, r1-3-4. II—Fe v0, d0, p0, r0; Pa v0-0-2, d0, p0, r0; ti v4-4-6, d0, p0-0-1, r4-8-6; Me v5-5-8, d0, p0-0-2, r2-2-4; ta v7-11-13, d0, p0-0-1, r3-3-7. III—Fe v0, d1-4-3, p0, r0; Pa v0-0-2, d5-3-14, p2-0-3, r0-0-2; ti v0-0-2, d1-0-9, p0- 0-3, r0-0-2; Me v0-0-3, d1-3-0, p0-2-1, r0-1-2; ta v0, d0, p1-2-7, r0-2-5; IV—Fe v0, d0, p0, r0; Pa v0, d16-16-8, p0-1-1, r0; ti v0-0-2, d0, p0-0-1, r0; Me v0-0-2, d0, p0-2- 3, r0; ta v0, d0, p2-6-7, r0-1-2.
<emphasis id="B3A87E6EFF5DCE8DFE11F93AD97E3B2B" bold="true" box="[494,557,1771,1797]" pageId="208" pageNumber="209">Palp:</emphasis>
PA well-developed embolus with three keels (Pa, PI and PS). ATP almost 1/2 size of embolus. BTA developed; serrated area large, from beside embolus base to middle of embolus; embolus inserted at 80°; embolar base thin (
<figureCitation id="19E7BEF9FF5DCE8DFDF0F8AFD93D3BB7" box="[527,622,1918,1945]" captionStart="FIGURE 189" captionStartId="209.[136,235,1502,1526]" captionTargetBox="[159,1425,209,1451]" captionTargetId="figure-137@209.[151,1435,189,1478]" captionTargetPageId="209" captionText="FIGURE 189. Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, male: A–E MACN 19826: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PAc- Prolateral Accessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrows: Serrated area. E. Rastellum, dorsal; F. Rastellum, ventral. Scales lines: A–C: 1 mm; D–E: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5597648" httpUri="https://zenodo.org/record/5597648/files/figure.png" pageId="208" pageNumber="209">Fig. 189</figureCitation>
A–C).
</paragraph>
<caption id="D5A3F2F4FF5DCE8DFF77FB52D9C73891" ID-DOI="http://doi.org/10.5281/zenodo.5597646" ID-Zenodo-Dep="5597646" httpUri="https://zenodo.org/record/5597646/files/figure.png" pageId="208" pageNumber="209" startId="208.[136,235,1155,1179]" targetBox="[180,1407,206,1116]" targetPageId="208">
<paragraph id="8163A27CFF5DCE8DFF77FB52D9C73891" blockId="208.[136,1452,1155,1215]" pageId="208" pageNumber="209">
<emphasis id="B3A87E6EFF5DCE8DFF77FB52DA4E38B5" bold="true" box="[136,285,1155,1179]" pageId="208" pageNumber="209">FIGURE 188.</emphasis>
<taxonomicName id="46DCD9FFFF5DCE8DFEDDFB55D84C38B5" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[290,799,1155,1179]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="208" pageNumber="209" phylum="Arthropoda" rank="species" species="palmar">
<emphasis id="B3A87E6EFF5DCE8DFEDDFB55DA8638B5" box="[290,469,1155,1179]" italics="true" pageId="208" pageNumber="209">Actinopus palmar</emphasis>
<bibRefCitation id="E54DDF8DFF5DCE8DFE24FB52D84C38B5" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[475,799,1155,1179]" pageId="208" pageNumber="209" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</taxonomicName>
, male: 815–818 MACN-Ar 19826: A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal.
</paragraph>
</caption>
<paragraph id="8163A27CFF5DCE8DFCD5FAFED89C3967" blockId="208.[810,975,1327,1353]" box="[810,975,1327,1353]" pageId="208" pageNumber="209">
<emphasis id="B3A87E6EFF5DCE8DFCD5FAFED89C3967" bold="true" box="[810,975,1327,1353]" pageId="208" pageNumber="209">VARIATION</emphasis>
</paragraph>
<paragraph id="8163A27CFF5DCE8DFCD5FAABDF3A3426" blockId="208.[810,1452,1401,2093]" pageId="208" pageNumber="209">
<materialsCitation id="31B4A821FF5DCE8DFCD5FAABDF353426" ID-GBIF-Occurrence="3399910652" collectionCode="MALES, PME, PLE, ALE, PME-PME, AME-AME, ALE-AME, AME-PME, II, III, IV" location="Ocular area" pageId="208" pageNumber="209" specimenCount="1">
<collectionCode id="E7CD3AB9FF5DCE8DFCD5FAABD8DA39BA" box="[810,905,1402,1428]" pageId="208" pageNumber="209">MALES</collectionCode>
(n=2):
<emphasis id="B3A87E6EFF5DCE8DFC1FFAABDFFC39BD" bold="true" box="[992,1199,1401,1428]" pageId="208" pageNumber="209">Eyes: Diameters:</emphasis>
<collectionCode id="E7CD3AB9FF5DCE8DFB48FAABDFA239BA" box="[1207,1265,1402,1428]" pageId="208" pageNumber="209">PME</collectionCode>
0.12–0.15,
<collectionCode id="E7CD3AB9FF5DCE8DFA85FAABDEFF39BA" box="[1402,1452,1402,1428]" pageId="208" pageNumber="209">PLE</collectionCode>
0.15–0.18,
<collectionCode id="E7CD3AB9FF5DCE8DFC4CFA4ED8BA3997" box="[947,1001,1439,1465]" pageId="208" pageNumber="209">ALE</collectionCode>
0.28–0.31, AME 0.12–0.21; MOQ: Length 0.62–0.68, front width 1.68–1.78, back width 1.5– 1.56; I
<emphasis id="B3A87E6EFF5DCE8DFC8AFA39DF4E3A2C" bold="true" box="[885,1053,1512,1538]" pageId="208" pageNumber="209">nterdistances:</emphasis>
<collectionCode id="E7CD3AB9FF5DCE8DFBD8FA38DFFB3A2D" box="[1063,1192,1513,1539]" pageId="208" pageNumber="209">PME–PME</collectionCode>
0.84–0.90, PLE–PME 0.09–0.12,
<collectionCode id="E7CD3AB9FF5DCE8DFC5BF9DFDF7D3A06" box="[932,1070,1550,1576]" pageId="208" pageNumber="209">AME–AME</collectionCode>
0.09–0.15,
<collectionCode id="E7CD3AB9FF5DCE8DFB52F9DFDE633A06" box="[1197,1328,1550,1576]" pageId="208" pageNumber="209">ALE–AME</collectionCode>
0.59–0.65, ALE–PLE 0.28–0.34,
<collectionCode id="E7CD3AB9FF5DCE8DFBC7F9E3DFED3A62" box="[1080,1214,1586,1612]" pageId="208" pageNumber="209">AME–PME</collectionCode>
0.25–0.28.
<emphasis id="B3A87E6EFF5DCE8DFAA7F9E3D8343A5F" bold="true" pageId="208" pageNumber="209">
<location id="8403F4A7FF5DCE8DFAA7F9E3D8313A5F" LSID="urn:lsid:plazi:treatment:0975136AFF43CE8CFCD5F87CD9A93B71:8403F4A7FF5DCE8DFAA7F9E3D8313A5F" name="Ocular area" pageId="208" pageNumber="209">Ocular area</location>
:
</emphasis>
OAL 2.5–2.53, OAW 2.12–2.62, and IF 1.12–1.4.
<emphasis id="B3A87E6EFF5DCE8DFCD5F9ADD8223AB8" bold="true" box="[810,881,1660,1686]" pageId="208" pageNumber="209">Body:</emphasis>
Total length: 6.75–8.62; Carapace: long 3.62–4.3; wide 3.25–3.6; Clypeus: 0.12–0.15; Fovea: 1.30–1.40; Labium: long 0.84–0.85; wide 0.75–0.76; Chelicerae: long 1.56–2.43; wide 1.03–1.18; Sternum: long 2.24–2.8; wide 2–2.1. Abdomen: long 3.62–4.56; wide 2.75–3.5.
<emphasis id="B3A87E6EFF5DCE8DFCD5F8E4DF403B61" bold="true" box="[810,1043,1845,1871]" pageId="208" pageNumber="209">Leg measurements:</emphasis>
I: Fe 3.62–4/ Pa 1.37–1.62/ Ti 2.12– 2.37/ Me 2.87–3.25/ Ta 1.75–1.87/ total 12.37–12.5.
<collectionCode id="E7CD3AB9FF5DCE8DFA6EF88BDEF43B5A" box="[1425,1447,1882,1908]" pageId="208" pageNumber="209">II</collectionCode>
: 3.75–4/ 1.12–1.75/ 2.12–2.5/ 2.87/ 1.75–1.87/ 11.62–13.
<collectionCode id="E7CD3AB9FF5DCE8DFCD5F875D8193B90" box="[810,842,1956,1982]" pageId="208" pageNumber="209">III</collectionCode>
: 2.75–3/ 1.5–2.37/ 1.25–1.75/ 2.5–3/ 2–2.12/ 10–12.25.
<collectionCode id="E7CD3AB9FF5DCE8DFCD5F818D8193BCD" box="[810,842,1993,2019]" pageId="208" pageNumber="209">IV</collectionCode>
: 3.62–4.12/ 1.37–1.62/ 2.75–3.25/ 3–3.12/ 2.25–2.37/ 13.25–14.25. Formula 4123
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9C6F1F7FF5DCE8CFCA0F7C3D9993B14" lastPageId="209" lastPageNumber="210" pageId="208" pageNumber="209" type="materials_examined">
<paragraph id="8163A27CFF5DCE8CFCA0F7C3D9993B14" blockId="208.[810,1452,1401,2093]" lastBlockId="209.[136,778,1750,1887]" lastPageId="209" lastPageNumber="210" pageId="208" pageNumber="209">
<emphasis id="B3A87E6EFF5DCE8DFCA0F7C3DF013402" bold="true" box="[863,1106,2066,2092]" pageId="208" pageNumber="209">Material examined.</emphasis>
<materialsCitation id="31B4A821FF5DCE8CFB97F7C3DB843B15" ID-GBIF-Occurrence="3399910799" collectingDate="1980-02-12" collectingDateMax="1995-03" collectingDateMin="1980-02-12" collectionCode="MACN" collectorName="Grosson &amp; P. A. Goloboff" country="Argentina" lastPageId="209" lastPageNumber="210" latitude="-58.3225" location="Parque Nacional" longLatPrecision="17" longitude="-31.853056" pageId="208" pageNumber="209" specimenCode="MACN 19826, MACN 19839" specimenCount="1" stateProvince="Entre Rios">
<collectingCountry id="F9CBE2ECFF5DCE8DFB97F7C3DE593402" box="[1128,1290,2066,2092]" name="Argentina" pageId="208" pageNumber="209">ARGENTINA</collectingCountry>
:
<collectingRegion id="43186C9EFF5DCE8DFADCF7C2DEF43402" box="[1315,1447,2066,2093]" country="Argentina" name="Entre Rios" pageId="208" pageNumber="209">Entre Ríos</collectingRegion>
,
<location id="8403F4A7FF5CCE8CFF77F906DA6A3ADF" LSID="urn:lsid:plazi:treatment:0975136AFF43CE8CFCD5F87CD9A93B71:8403F4A7FF5CCE8CFF77F906DA6A3ADF" box="[136,313,1751,1777]" country="Argentina" latitude="-58.3225" longLatPrecision="17" longitude="-31.853056" name="Parque Nacional" pageId="209" pageNumber="210" stateProvince="Entre Rios">Parque Nacional</location>
“El Palmar”, [
<geoCoordinate id="E4E8C4BBFF5CCE8CFE2EF907D90C3ADF" box="[465,607,1750,1777]" degrees="58" direction="south" minutes="19" orientation="latitude" pageId="209" pageNumber="210" precision="15" seconds="21" value="-58.3225">58° 19’ 21” S</geoCoordinate>
<geoCoordinate id="E4E8C4BBFF5CCE8CFD9BF906D9AD3ADF" box="[612,766,1750,1777]" degrees="31" direction="west" minutes="51" orientation="longitude" pageId="209" pageNumber="210" precision="15" seconds="11" value="-31.853056">31° 51’ 11” W</geoCoordinate>
], 1♂,
<date id="F56284BCFF5CCE8CFF46F92ADA1C3B3B" box="[185,335,1787,1813]" pageId="209" pageNumber="210" value="1980-02-12" valueMax="1980-02-16" valueMin="1980-02-12">
<collectingDate id="E5267D54FF5CCE8CFF46F92ADA1C3B3B" box="[185,335,1787,1813]" pageId="209" pageNumber="210" value="1980-02-12" valueMax="1980-02-16" valueMin="1980-02-12">12–16.ii.1980</collectingDate>
</date>
,
<collectorName id="2C29C7AAFF5CCE8CFEA4F92ADAE73B3B" box="[347,436,1787,1813]" pageId="209" pageNumber="210">Grosson</collectorName>
&amp;
<collectorName id="2C29C7AAFF5CCE8CFE25F92DD92B3B3B" box="[474,632,1787,1814]" pageId="209" pageNumber="210">P. A. Goloboff</collectorName>
leg. (
<specimenCode id="D17A0A07FF5CCE8CFD49F92ADB9C3B15" collectionCode="MACN" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="209" pageNumber="210" type="Museum">MACN 19826</specimenCode>
)
</materialsCitation>
<materialsCitation id="31B4A821FF5CCE8CFF28F8F1D9953B14" ID-GBIF-Occurrence="3399910645" box="[215,710,1824,1851]" collectingDate="1980-02-12" collectingDateMax="1995-03" collectingDateMin="1980-02-12" collectionCode="MACN" collectorName="C. J. Grismado" country="Argentina" latitude="-58.3225" location="Parque Nacional" longLatPrecision="17" longitude="-31.853056" pageId="209" pageNumber="210" specimenCode="MACN 19826, MACN 19839" specimenCount="1" stateProvince="Entre Rios">
;
<date id="F56284BCFF5CCE8CFF1DF8F1DA653B15" box="[226,310,1824,1851]" pageId="209" pageNumber="210" value="1995-03">
<collectingDate id="E5267D54FF5CCE8CFF1DF8F1DA653B15" box="[226,310,1824,1851]" pageId="209" pageNumber="210" value="1995-03">iii.1995</collectingDate>
</date>
,
<collectorName id="2C29C7AAFF5CCE8CFEBFF8F1DAB23B14" box="[320,481,1824,1851]" pageId="209" pageNumber="210">C. J. Grismado</collectorName>
leg. (
<specimenCode id="D17A0A07FF5CCE8CFDE3F8F1D9EE3B14" box="[540,701,1824,1850]" collectionCode="MACN" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="209" pageNumber="210" type="Museum">MACN 19839</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="D5A3F2F4FF5CCE8CFF77FA0FD9B43A4C" ID-DOI="http://doi.org/10.5281/zenodo.5597648" ID-Zenodo-Dep="5597648" httpUri="https://zenodo.org/record/5597648/files/figure.png" pageId="209" pageNumber="210" startId="209.[136,235,1502,1526]" targetBox="[159,1425,209,1451]" targetPageId="209">
<paragraph id="8163A27CFF5CCE8CFF77FA0FD9B43A4C" blockId="209.[136,1452,1502,1634]" pageId="209" pageNumber="210">
<emphasis id="B3A87E6EFF5CCE8CFF77FA0FDA7339D8" bold="true" box="[136,288,1502,1526]" pageId="209" pageNumber="210">FIGURE 189.</emphasis>
<taxonomicName id="46DCD9FFFF5CCE8CFED5FA0ED86E39D8" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[298,829,1502,1526]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="palmar">
<emphasis id="B3A87E6EFF5CCE8CFED5FA0EDAB139D8" box="[298,482,1502,1526]" italics="true" pageId="209" pageNumber="210">Actinopus palmar</emphasis>
<bibRefCitation id="E54DDF8DFF5CCE8CFE13FA0FD86E39D8" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[492,829,1502,1526]" pageId="209" pageNumber="210" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</taxonomicName>
, male: A–E MACN 19826: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral.
<emphasis id="B3A87E6EFF5CCE8CFCFAF9D3D87E3A34" bold="true" box="[773,813,1538,1562]" pageId="209" pageNumber="210">PA-</emphasis>
Paraembolic apophysis;
<emphasis id="B3A87E6EFF5CCE8CFBD4F9D3DF0E3A34" bold="true" box="[1067,1117,1538,1562]" pageId="209" pageNumber="210">BTA</emphasis>
-Basal Tegular Apophysis;
<emphasis id="B3A87E6EFF5CCE8CFA87F9D3DEF03A34" bold="true" box="[1400,1443,1538,1562]" pageId="209" pageNumber="210">PAc</emphasis>
- Prolateral Accessory keel;
<emphasis id="B3A87E6EFF5CCE8CFE67F9F7DAE23A10" bold="true" box="[408,433,1574,1598]" pageId="209" pageNumber="210">PI</emphasis>
-Prolateral Inferior keel;
<emphasis id="B3A87E6EFF5CCE8CFD51F9F7D99F3A10" bold="true" box="[686,716,1574,1598]" pageId="209" pageNumber="210">PS</emphasis>
-Prolateral Superior keel; Arrows: Serrated area. E. Rastellum, dorsal; F. Rastellum, ventral. Scales lines: A–C: 1 mm; D–E: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C9C6F1F7FF5CCE8CFF42F895D9A93B71" box="[189,762,1860,1887]" pageId="209" pageNumber="210" type="distribution">
<paragraph id="8163A27CFF5CCE8CFF42F895D9A93B71" blockId="209.[136,778,1750,1887]" box="[189,762,1860,1887]" pageId="209" pageNumber="210">
<emphasis id="B3A87E6EFF5CCE8CFF42F895DA043B70" bold="true" box="[189,343,1860,1886]" pageId="209" pageNumber="210">Distribution.</emphasis>
<collectingCountry id="F9CBE2ECFF5CCE8CFEA3F894D9503B71" box="[348,515,1861,1887]" name="Argentina" pageId="209" pageNumber="210">ARGENTINA</collectingCountry>
.
<collectingRegion id="43186C9EFF5CCE8CFDF2F894D9D73B71" box="[525,644,1861,1887]" country="Argentina" name="Entre Rios" pageId="209" pageNumber="210">Entre Ríos</collectingRegion>
(
<figureCitation id="19E7BEF9FF5CCE8CFD6CF894D9BF3B71" box="[659,748,1861,1887]" captionStart="MAP 11" captionStartId="196.[136,192,1121,1145]" captionTargetBox="[157,1428,195,1089]" captionTargetId="figure-346@196.[151,1435,189,1097]" captionTargetPageId="196" captionText="MAP 11. Distribution map of Actinopus robustus O. Pickard-Cambridge, 1892, Actinopus guajara sp. nov., Actinopus palmar Ríos-Tamayo &amp; Goloboff, 2018, Actinopus lomalinda sp. nov., Actinopus concinnus sp. nov. and Actinopus cucutaensis (Mello- Leit„o, 1941)." figureDoi="http://doi.org/10.5281/zenodo.5655450" httpUri="https://zenodo.org/record/5655450/files/figure.png" pageId="209" pageNumber="210">Map 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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