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<document id="9674D41267B29E3727F662A1CF6C1DCF" ID-CLB-Dataset="9504" ID-DOI="10.1080/00222933.2020.1746426" ID-GBIF-Dataset="a33c8231-6ff0-4198-ac1e-3e52fe251db3" ID-ISSN="1464-5262" ID-Zenodo-Dep="4290471" ID-ZooBank="61D1A59D-3702-480A-B146-73067C29CD82" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1606311042563" checkinUser="carolina" docAuthor="Noort, Simon van &amp; Copeland, Robert S." docDate="2020" docId="14566979FFDB3A6A84A2FCF2FDB76702" docLanguage="en" docName="JNATHIST.54.9-12.703-722.pdf" docOrigin="Journal of Natural History 54 (9)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Tanaostigma lasallei van Noort 2020, sp. nov" docType="treatment" docVersion="9" lastPageNumber="712" masterDocId="E86F1101FFDC3A608530FF99FFFD6500" masterDocTitle="First record of the genus Tanaostigma (Hymenopteraı Chalcidoideaı Tanaostigmatidae) from the Afrotropical region with description of three new species" masterLastPageNumber="722" masterPageNumber="703" pageNumber="709" updateTime="1698846559214" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5D99A58E77D7D92C0D54FC5B47A5CA97">First record of the genus Tanaostigma (Hymenopteraı Chalcidoideaı Tanaostigmatidae) from the Afrotropical region with description of three new species</mods:title>
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<mods:namePart id="A0275242565848794EB8918D5ABE4DB3">Noort, Simon van</mods:namePart>
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<mods:namePart id="E379D8409196739A0A52674E42AE0573">Copeland, Robert S.</mods:namePart>
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<mods:title id="98F87F20AD038CBBFE2978AEF848B1D4">Journal of Natural History</mods:title>
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<mods:date id="D98568D6888E9C0DCD4B51300C4EEB9C">2020</mods:date>
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<mods:number id="BEA1D37A170F971AE619AADE7476F56C">2020-09-23</mods:number>
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<mods:number id="8879559EB87FAB5939DD5BDEF24D0A97">54</mods:number>
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<mods:classification id="6F05E101F1FBD7C97462EEF9748F6264">journal article</mods:classification>
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<treatment id="14566979FFDB3A6A84A2FCF2FDB76702" ID-DOI="http://doi.org/10.5281/zenodo.4328725" ID-GBIF-Taxon="170529782" ID-Zenodo-Dep="4328725" LSID="urn:lsid:plazi:treatment:14566979FFDB3A6A84A2FCF2FDB76702" httpUri="http://treatment.plazi.org/id/14566979FFDB3A6A84A2FCF2FDB76702" lastPageId="10" lastPageNumber="712" pageId="7" pageNumber="709">
<subSubSection id="D4E58BE4FFDB3A6784A2FCF2FC6F6685" box="[402,914,875,901]" pageId="7" pageNumber="709" type="nomenclature">
<paragraph id="9C40D86FFFDB3A6784A2FCF2FC6F6685" blockId="7.[402,914,875,935]" box="[402,914,875,901]" pageId="7" pageNumber="709">
<heading id="C7086F03FFDB3A6784A2FCF2FC6F6685" box="[402,914,875,901]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="709" reason="2">
<emphasis id="AE8B047DFFDB3A6784A2FCF2FC6F6685" bold="true" box="[402,914,875,901]" pageId="7" pageNumber="709">
<taxonomicName id="5BFFA3ECFFDB3A6784A2FCF2FCD46685" authority="van Noort" authorityName="van Noort" authorityYear="2020" box="[402,809,875,901]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="709" phylum="Arthropoda" rank="species" species="lasallei" status="sp. nov.">
<emphasis id="AE8B047DFFDB3A6784A2FCF2FD616685" bold="true" box="[402,668,875,901]" italics="true" pageId="7" pageNumber="709">Tanaostigma lasallei</emphasis>
van Noort
</taxonomicName>
<taxonomicNameLabel id="B5B8B906FFDB3A678601FCF2FC6F6685" box="[817,914,875,901]" pageId="7" pageNumber="709" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D4E58BE4FFDB3A67873CFC14FCEA66A7" box="[524,791,909,935]" pageId="7" pageNumber="709" type="description">
<paragraph id="9C40D86FFFDB3A67873CFC14FCEA66A7" blockId="7.[402,914,875,935]" box="[524,791,909,935]" pageId="7" pageNumber="709">
(
<figureCitation id="04C4C4EAFFDB3A678724FC14FD4666A7" box="[532,699,909,935]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="7" pageNumber="709">
Figures 1(a
<emphasis id="AE8B047DFFDB3A6787ACFC14FD5666A7" box="[668,683,909,935]" italics="true" pageId="7" pageNumber="709">–</emphasis>
f)
</figureCitation>
,
<figureCitation id="04C4C4EAFFDB3A6787FAFC14FCFB66A7" box="[714,774,909,935]" captionStart="Figure 2" captionStartId="8.[160,225,1287,1310]" captionTargetBox="[164,1152,143,1254]" captionTargetId="figure-114@8.[163,1153,142,1255]" captionTargetPageId="8" captionText="Figure 2. Tanaostigma lasallei Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="7" pageNumber="709">
2(a
<emphasis id="AE8B047DFFDB3A6787DFFC14FD0366A7" box="[751,766,909,935]" italics="true" pageId="7" pageNumber="709">–</emphasis>
f
</figureCitation>
))
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFDB3A678590FC5AFEDA6143" pageId="7" pageNumber="709" type="materials_examined">
<paragraph id="9C40D86FFFDB3A678590FC5AFEDA6143" blockId="7.[160,1156,963,1091]" pageId="7" pageNumber="709">
<materialsCitation id="2C97D232FFDB3A678590FC5AFEDA6143" ID-GBIF-Occurrence="2985539304" collectingDate="2010-10-18" collectingDateMax="2010-10-20" collectingDateMin="2010-10-18" collectingMethod="Malaise trap" collectionCode="SAMC" collectorName="S. van Noort" country="South Africa" elevation="50" latitude="-27.3303" location="Mabibi campsite" longLatPrecision="1" longitude="32.746532" pageId="7" pageNumber="709" specimenCode="SAMC 2019" specimenCount="1" specimenCount-female="1" stateProvince="Kwazulu-Natal" typeStatus="holotype">
<emphasis id="AE8B047DFFDB3A678590FC5AFEFC66DB" box="[160,257,963,987]" italics="true" pageId="7" pageNumber="709">
<typeStatus id="434466CDFFDB3A678590FC5AFEFC66DB" box="[160,257,963,987]" pageId="7" pageNumber="709" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="8AF913E6FFDB3A678439FC5AFEE066DB" box="[265,285,963,987]" pageId="7" pageNumber="709" type="female">♀</specimenCount>
(deposited in
<collectionCode id="FAEE40AAFFDB3A6784F1FC5AFDF566DB" box="[449,520,963,987]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="7" pageNumber="709" type="Museum">SAMC</collectionCode>
), point mounted:
<collectingCountry id="E4E898FFFFDB3A6787E1FC5AFC8366DB" box="[721,894,963,987]" name="South Africa" pageId="7" pageNumber="709">SOUTH AFRICA</collectingCountry>
,
<collectingRegion id="5E3B168DFFDB3A6786B8FC5AFBD666DB" box="[904,1067,963,987]" country="South Africa" name="KwaZulu-Natal" pageId="7" pageNumber="709">Kwazulu-Natal</collectingRegion>
,
<location id="99208EB4FFDB3A678106FC5AFEF566FE" LSID="urn:lsid:plazi:treatment:14566979FFDB3A6A84A2FCF2FDB76702:99208EB4FFDB3A678106FC5AFEF566FE" country="South Africa" latitude="-27.3303" longLatPrecision="1" longitude="32.746532" name="Mabibi campsite" pageId="7" pageNumber="709" stateProvince="Kwazulu-Natal">Mabibi campsite</location>
,
<quantity id="5B07758AFFDB3A678422FC7FFEB066FE" box="[274,333,998,1022]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="709" unit="m" value="50.0">
<elevation id="17D23F5CFFDB3A678422FC7FFEB066FE" box="[274,333,998,1022]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="709" unit="m" value="50.0">50 m</elevation>
</quantity>
,
<geoCoordinate id="F9CBBEA8FFDB3A678468FC7FFE2566FE" box="[344,472,998,1022]" degrees="27" direction="south" minutes="19.818" orientation="latitude" pageId="7" pageNumber="709" precision="1" value="-27.3303">
27°19.818
<emphasis id="AE8B047DFFDB3A6784F5FC7FFE3666FE" bold="true" box="[453,459,998,1022]" italics="true" pageId="7" pageNumber="709">′</emphasis>
S
</geoCoordinate>
<geoCoordinate id="F9CBBEA8FFDB3A6784D0FC7FFD9E66FE" box="[480,611,998,1022]" degrees="32" direction="east" minutes="44.792" orientation="longitude" pageId="7" pageNumber="709" precision="1" value="32.746532">
32°44.792
<emphasis id="AE8B047DFFDB3A67877DFC7FFDAE66FE" bold="true" box="[589,595,998,1022]" italics="true" pageId="7" pageNumber="709">′</emphasis>
E
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,
<date id="E841FEAFFFDB3A67875DFC7FFCA966FE" box="[621,852,998,1022]" pageId="7" pageNumber="709" value="2010-10-18" valueMax="2010-10-20" valueMin="2010-10-18">
<collectingDate id="F8050747FFDB3A67875DFC7FFCA966FE" box="[621,852,998,1022]" pageId="7" pageNumber="709" value="2010-10-18" valueMax="2010-10-20" valueMin="2010-10-18">
18
<emphasis id="AE8B047DFFDB3A6787B9FC7FFD6B66FE" box="[649,662,998,1022]" italics="true" pageId="7" pageNumber="709">–</emphasis>
20 October 2010
</collectingDate>
</date>
,
<collectorName id="310ABDB9FFDB3A67866EFC7FFC1666FE" box="[862,1003,998,1022]" pageId="7" pageNumber="709">S. van Noort</collectorName>
,
<collectingMethod id="45BEA078FFDB3A6786C5FC7FFB7D66FE" box="[1013,1152,998,1022]" pageId="7" pageNumber="709">Malaise trap</collectingMethod>
, SA10-MAB-M01, Maputaland Coastal Belt, SAM-HYM-P082674, imaged WaspWeb, LAS 4.9, SAMC 2019.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFDB3A698590FBE5FE9966E7" lastPageId="9" lastPageNumber="711" pageId="7" pageNumber="709" type="description">
<paragraph id="9C40D86FFFDB3A678590FBE5FE636195" blockId="7.[160,1156,1148,1727]" box="[160,414,1148,1173]" pageId="7" pageNumber="709">
<emphasis id="AE8B047DFFDB3A678590FBE5FE636195" bold="true" box="[160,414,1148,1173]" italics="true" pageId="7" pageNumber="709">Description of female</emphasis>
</paragraph>
<paragraph id="9C40D86FFFDB3A688590FB07FE306312" blockId="7.[160,1156,1148,1727]" lastBlockId="8.[160,1155,1426,1727]" lastPageId="8" lastPageNumber="710" pageId="7" pageNumber="709">
<emphasis id="AE8B047DFFDB3A678590FB07FEA761B7" bold="true" box="[160,346,1182,1207]" italics="true" pageId="7" pageNumber="709">Size and colour.</emphasis>
Length
<quantity id="5B07758AFFDB3A6784F1FB06FDEA61B7" box="[449,535,1183,1207]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="709" unit="mm" value="3.3">3.3 mm</quantity>
. Head and body black with extensive areas of the head, mesosoma, and metasoma covered with white squamiferous setae, absent on mesopleura, antero-medial portion of mesoscutum, anterior half of scutellum, medial dorsal and anterior ventral areas of metasoma. Scape and pedicel orange-brown, anelli and first funicle segment dark brown grading into black for rest of flagellar segments except for last two club segments, which are orange-yellow. Distal two-thirds of hind tibiae orange. All tarsi yellow-orange, as is distal third of meso-tibia. Finer, white setae present on legs.
<emphasis id="AE8B047DFFDB3A6785F0FA0BFF0460AA" box="[192,249,1426,1450]" italics="true" pageId="7" pageNumber="709">Head</emphasis>
(
<figureCitation id="04C4C4EAFFDB3A678438FA0BFE7A60AA" box="[264,391,1426,1450]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="7" pageNumber="709">Figure 1(e))</figureCitation>
1.3 times wider than high. Lateral ocellus closer to eye margin than to median ocellus (OOL:LOL: 0.37), situated directly adjacent to sharp occipital carinal crest, which curves inwards towards median ocellus. Scrobal impression shallow, glabrate. Inter-antennal projection small. Genal sulcus complete. Head sculpture elongate reticulate on face grading to imbricate on frons, with evenly dispersed white squamiferous setae present on face and frons, grading into normal white setae on vertex, setae largely absent on posterior of head including genae posterior of genal sulcus. Mandible tridentate (
<figureCitation id="04C4C4EAFFDB3A678439F91CFE80639D" box="[265,381,1669,1693]" captionStart="Figure 2" captionStartId="8.[160,225,1287,1310]" captionTargetBox="[164,1152,143,1254]" captionTargetId="figure-114@8.[163,1153,142,1255]" captionTargetPageId="8" captionText="Figure 2. Tanaostigma lasallei Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="7" pageNumber="709">Figure 2(a</figureCitation>
)). Antenna (
<figureCitation id="04C4C4EAFFDB3A67873FF91CFD5C639D" box="[527,673,1669,1693]" captionStart="Figure 2" captionStartId="8.[160,225,1287,1310]" captionTargetBox="[164,1152,143,1254]" captionTargetId="figure-114@8.[163,1153,142,1255]" captionTargetPageId="8" captionText="Figure 2. Tanaostigma lasallei Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="7" pageNumber="709">Figures 2(a,b</figureCitation>
)) with scape 4.7 times longer than wide, with flattened ventral expansion present on distal half. Pedicel 1.5 times longer than wide. Two anelli, first about half as long as, and slightly narrower than second. First funicular segment longer than wide, F2-F4 as wide as long, F5-F6 wider than long, not laterally compressed. Club 1.4 times longer than wide, proximally as broad as preceding funicle segment, narrowing to tip.
</paragraph>
<caption id="C88088E7FFD43A688590FA9EFB78603B" ID-DOI="http://doi.org/10.5281/zenodo.4290475" ID-Zenodo-Dep="4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="8" pageNumber="710" startId="8.[160,225,1287,1310]" targetBox="[164,1152,143,1254]" targetPageId="8">
<paragraph id="9C40D86FFFD43A688590FA9EFB78603B" blockId="8.[160,1157,1287,1339]" pageId="8" pageNumber="710">
<emphasis id="AE8B047DFFD43A688590FA9EFF04601E" bold="true" box="[160,249,1287,1310]" pageId="8" pageNumber="710">Figure 2.</emphasis>
<taxonomicName id="5BFFA3ECFFD43A6885CFFA9EFE3F601E" authorityName="van Noort" authorityYear="2020" box="[255,450,1287,1310]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="710" phylum="Arthropoda" rank="species" species="lasallei">
<emphasis id="AE8B047DFFD43A6885CFFA9EFE3F601E" box="[255,450,1287,1310]" italics="true" pageId="8" pageNumber="710">Tanaostigma lasallei</emphasis>
</taxonomicName>
Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view.
</paragraph>
</caption>
<paragraph id="9C40D86FFFD43A6985F0F984FDB767D1" blockId="8.[160,1155,1426,1727]" lastBlockId="9.[160,1156,142,929]" lastPageId="9" lastPageNumber="711" pageId="8" pageNumber="710">
<emphasis id="AE8B047DFFD43A6885F0F984FEC96335" box="[192,308,1565,1589]" italics="true" pageId="8" pageNumber="710">Mesosoma</emphasis>
dorsally minutely reticulate, punctiferous where setae present, covered with white squamiferous setae, except for mesoscutum anterior of notauli and axillae which have small, fine white setae. Notauli complete but indistinct posteriorly, meeting before trans-scutellar articulation. Scutellum (
<figureCitation id="04C4C4EAFFD43A68877BF91CFD31639D" box="[587,716,1669,1693]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="8" pageNumber="710">Figure 1(d))</figureCitation>
without lateral glabrate area, devoid of white setae in anterior third, replaced by small, fine brown setae. Propodeum (
<figureCitation id="04C4C4EAFFD43A688133F93EFB7E63BF" box="[1027,1155,1703,1727]" captionStart="Figure 2" captionStartId="8.[160,225,1287,1310]" captionTargetBox="[164,1152,143,1254]" captionTargetId="figure-114@8.[163,1153,142,1255]" captionTargetPageId="8" captionText="Figure 2. Tanaostigma lasallei Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="8" pageNumber="710">Figure 2(d))</figureCitation>
laterally glabrate to medially reticulate with complete medial carina; metanotal trough with two large metanotal fovea laterad of central smaller fovea, with narrow elongate fovea anteriorly situated, extending medially beyond central fovea. Mesopleuron (
<figureCitation id="04C4C4EAFFD53A698597FF6FFED7640E" box="[167,298,246,270]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="9" pageNumber="711">Figure 1(c))</figureCitation>
dorsally reticulate, ventrally elongate reticulate, medially finely reticulate, without setae. Sternopleural suture fused with mesopleural suture. Metapleuron covered with a dense cluster of long white setae (
<figureCitation id="04C4C4EAFFD53A698757FEA5FD1A6454" box="[615,743,316,340]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="9" pageNumber="711">Figure 1(c))</figureCitation>
; which extend to the proximal dorsal surface of the hind coxa. Leg segments laterally flattened; all tibiae with sharp dorsal margin. Hind femora with a distal ventral angulate extension, curving inwards to end of femur; hind tibiae with a proximal dorsal carina. Wings hyaline, except for fore wing (
<figureCitation id="04C4C4EAFFD53A698597FE5FFED864DE" box="[167,293,454,478]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="9" pageNumber="711">Figure 1(f))</figureCitation>
which has a medial infuscate patch extending from marginal and stigmal veins halfway towards apical and ventral margins in an axe shape; larger, modified, dark setae present below marginal vein grading outwards into normal setae; small triangular patch of larger more widely dispersed setae present directly below submarginal vein, adjacent to glabrous area. Venation dark brown, submarginal vein lighter. Stigmal vein strong, distinctly curved, longer than post-marginal vein. Fore wing marginal fringe present apically, extending ventrally to anal angle. Costal cell: marginal vein: stigmal vein: postmarginal vein = 50:32:15:12.
</paragraph>
<paragraph id="9C40D86FFFD53A6985F0FD45FD5666A1" blockId="9.[160,1156,142,929]" pageId="9" pageNumber="711">
<emphasis id="AE8B047DFFD53A6985F0FD45FECF67F4" box="[192,306,732,756]" italics="true" pageId="9" pageNumber="711">Metasoma</emphasis>
(
<figureCitation id="04C4C4EAFFD53A698473FD45FE4267F4" box="[323,447,732,756]" captionStart="Figure 2" captionStartId="8.[160,225,1287,1310]" captionTargetBox="[164,1152,143,1254]" captionTargetId="figure-114@8.[163,1153,142,1255]" captionTargetPageId="8" captionText="Figure 2. Tanaostigma lasallei Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="9" pageNumber="711">Figure 2(f))</figureCitation>
with elongate reticulate surface sculpture; T2 with transverse rows of scattered setae, T3-T5 without setae medially; T6 with a row of setae anteriorly, T7 with setae laterally. Posterior margin of T2 with shallow medial invagination; faint medial line present; posterior margin of T3-T7 straight, without medial incision, or medial line. Ovipositor short. Each pygostyle with four setae, two extremely elongate, half as long as metasoma, the remaining pair half this length.
</paragraph>
<paragraph id="9C40D86FFFD53A698590FC57FE9966E7" blockId="9.[160,356,974,999]" box="[160,356,974,999]" pageId="9" pageNumber="711">
<emphasis id="AE8B047DFFD53A698590FC57FF1C66E7" bold="true" box="[160,225,974,999]" italics="true" pageId="9" pageNumber="711">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFD53A698590FB8AFE306334" pageId="9" pageNumber="711" type="diagnosis">
<paragraph id="9C40D86FFFD53A698590FB8AFE306334" blockId="9.[160,1156,1043,1588]" pageId="9" pageNumber="711">
<emphasis id="AE8B047DFFD53A698590FB8AFEE7612C" bold="true" box="[160,282,1043,1068]" italics="true" pageId="9" pageNumber="711">Diagnosis.</emphasis>
<taxonomicName id="5BFFA3ECFFD53A69841AFB8DFDF2612C" authorityName="van Noort" authorityYear="2020" box="[298,527,1044,1068]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="species" species="lasallei">
<emphasis id="AE8B047DFFD53A69841AFB8DFDF2612C" box="[298,527,1044,1068]" italics="true" pageId="9" pageNumber="711">Tanaostigma lasallei</emphasis>
</taxonomicName>
is distinguished from other
<taxonomicName id="5BFFA3ECFFD53A698665FB8DFC1B612C" authorityName="Howard" authorityYear="1890" box="[853,998,1044,1068]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="genus">
<emphasis id="AE8B047DFFD53A698665FB8DFC1B612C" box="[853,998,1044,1068]" italics="true" pageId="9" pageNumber="711">Tanaostigma</emphasis>
</taxonomicName>
by having an infuscate area on the fore wing (
<figureCitation id="04C4C4EAFFD53A698721FBAEFD72614F" box="[529,655,1079,1103]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="9" pageNumber="711">Figure 1(f))</figureCitation>
. The modified setae are restricted to a small patch below the marginal vein and do not contribute to the colour pattern on the wing. Presence of modified setae is shared with the Neotropical species
<taxonomicName id="5BFFA3ECFFD53A69813BFBE5FB836194" box="[1035,1150,1148,1172]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="AE8B047DFFD53A69813BFBE5FB836194" box="[1035,1150,1148,1172]" italics="true" pageId="9" pageNumber="711">T. stanleyi</emphasis>
</taxonomicName>
, which however has a patterned appearance as a result, and the other two African species where the setae form an obvious dark transverse band. The stigmal vein is strongly curved (
<figureCitation id="04C4C4EAFFD53A69845AFB7DFE1361FC" box="[362,494,1252,1276]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="9" pageNumber="711">Figure 1(f))</figureCitation>
. Scutellum lacks lateral glabrate area (
<figureCitation id="04C4C4EAFFD53A69868EFB7DFBB461FC" box="[958,1097,1252,1276]" captionStart="Figure 1" captionStartId="4.[160,225,1287,1310]" captionTargetBox="[164,1152,144,1254]" captionTargetId="figure-95@4.[163,1153,142,1255]" captionTargetPageId="4" captionText="Figure 1. Tanaostigma lasallei Holotype female. (a) Habitus, lateral view (b) Habitus, dorsal view (c) Head and mesosoma, lateral view (d) Head and mesosoma, dorsal view (e) Head, anterior view (f) Wings (inset: data labels)." figureDoi="http://doi.org/10.5281/zenodo.4290473" httpUri="https://zenodo.org/record/4290473/files/figure.png" pageId="9" pageNumber="711">Figure 1(d))</figureCitation>
; and posterior margin of T2-T4 without medial incision (
<figureCitation id="04C4C4EAFFD53A698624FA9EFC67601F" box="[788,922,1287,1311]" captionStart="Figure 2" captionStartId="8.[160,225,1287,1310]" captionTargetBox="[164,1152,143,1254]" captionTargetId="figure-114@8.[163,1153,142,1255]" captionTargetPageId="8" captionText="Figure 2. Tanaostigma lasallei Holotype female. (a) Clypeus and mandibles (b) Antenna (c) Pedicel and anelli (d) Scutellum, metanotum and propodeum (e) Trans-scutellar suture (f) Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4290475" httpUri="https://zenodo.org/record/4290475/files/figure.png" pageId="9" pageNumber="711">Figure 2(f))</figureCitation>
. All three of these character states are shared with the other two African species, which have a very different wing pattern, and with the Neotropical
<taxonomicName id="5BFFA3ECFFD53A6987D1FAD5FCAF6064" box="[737,850,1356,1380]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="AE8B047DFFD53A6987D1FAD5FCAF6064" box="[737,850,1356,1380]" italics="true" pageId="9" pageNumber="711">T. stanleyi</emphasis>
</taxonomicName>
, which, in contrast to the black African species, is strikingly coloured with pink, orange and white areas on the pleural regions and face. The remaining species of
<taxonomicName id="5BFFA3ECFFD53A6987D6FA0BFC8A60AA" authorityName="Howard" authorityYear="1890" box="[742,887,1426,1450]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="genus">
<emphasis id="AE8B047DFFD53A6987D6FA0BFC8A60AA" box="[742,887,1426,1450]" italics="true" pageId="9" pageNumber="711">Tanaostigma</emphasis>
</taxonomicName>
have a straight stigmal vein, glabrate areas on the scutellum and medial incisions on the metasomal tergites. Notauli of
<taxonomicName id="5BFFA3ECFFD53A698410FA4EFE7760EF" authorityName="van Noort" authorityYear="2020" box="[288,394,1495,1519]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="species" species="lasallei">
<emphasis id="AE8B047DFFD53A698410FA4EFE7760EF" box="[288,394,1495,1519]" italics="true" pageId="9" pageNumber="711">T. lasallei</emphasis>
</taxonomicName>
are complete, but indistinct posteriorly, whereas the other two African species have incomplete notauli and the New World and Oriental species have distinct, complete notauli.
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFD53A6A8590F9F8FB8665EC" lastPageId="10" lastPageNumber="712" pageId="9" pageNumber="711" type="discussion">
<paragraph id="9C40D86FFFD53A6A8590F9F8FB8665EC" blockId="9.[160,1156,1633,1727]" lastBlockId="10.[160,1155,142,236]" lastPageId="10" lastPageNumber="712" pageId="9" pageNumber="711">
<emphasis id="AE8B047DFFD53A698590F9F8FEF3637A" bold="true" box="[160,270,1633,1658]" italics="true" pageId="9" pageNumber="711">Affinities.</emphasis>
This species shares antennal morphology with the other two African species, and this is most similar to the Neotropical species,
<taxonomicName id="5BFFA3ECFFD53A6987D3F91CFC98639D" baseAuthorityName="Crawford" baseAuthorityYear="1911" box="[739,869,1669,1693]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="711" phylum="Arthropoda" rank="species" species="slossonae">
<emphasis id="AE8B047DFFD53A6987D3F91CFC98639D" box="[739,869,1669,1693]" italics="true" pageId="9" pageNumber="711">T. slossonae</emphasis>
</taxonomicName>
, which also has funicular segments that are not flattened, but which are subequal in length. The fore wing has only a small patch of weakly modified setae below the marginal vein, and does not have the extensive modified setae on the fore wing of the other two African species, but this shared character state suggests a possible relationship with the Neotropical species,
<taxonomicName id="5BFFA3ECFFD63A6A8138FF4DFB8865EC" box="[1032,1141,212,236]" class="Insecta" family="Tanaostigmatidae" genus="Tanaostigma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="712" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="AE8B047DFFD63A6A8138FF4DFB8865EC" box="[1032,1141,212,236]" italics="true" pageId="10" pageNumber="712">T. stanleyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFD63A6A8590FE81FC026431" box="[160,1023,280,305]" pageId="10" pageNumber="712" type="etymology">
<paragraph id="9C40D86FFFD63A6A8590FE81FC026431" blockId="10.[160,1023,280,305]" box="[160,1023,280,305]" pageId="10" pageNumber="712">
<emphasis id="AE8B047DFFD63A6A8590FE81FEDE6431" bold="true" box="[160,291,280,305]" italics="true" pageId="10" pageNumber="712">Etymology.</emphasis>
The species is named in memory of John LaSalle. Genitive case.
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFD63A6A8590FEC7FC486476" box="[160,949,350,375]" pageId="10" pageNumber="712" type="distribution">
<paragraph id="9C40D86FFFD63A6A8590FEC7FC486476" blockId="10.[160,949,350,375]" box="[160,949,350,375]" pageId="10" pageNumber="712">
<emphasis id="AE8B047DFFD63A6A8590FEC7FECE6477" bold="true" box="[160,307,350,375]" italics="true" pageId="10" pageNumber="712">Distribution.</emphasis>
<collectingCountry id="E4E898FFFFD63A6A8473FEC7FE326476" box="[323,463,350,374]" name="South Africa" pageId="10" pageNumber="712">South Africa</collectingCountry>
. Only known from
<collectingRegion id="5E3B168DFFD63A6A8795FEC7FC4C6476" box="[677,945,350,374]" country="South Africa" name="KwaZulu-Natal" pageId="10" pageNumber="712">Kwazulu-Natal Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFD63A6A8590FE3AFE7F64BC" box="[160,386,419,444]" pageId="10" pageNumber="712" type="biology_ecology">
<paragraph id="9C40D86FFFD63A6A8590FE3AFE7F64BC" blockId="10.[160,386,419,444]" box="[160,386,419,444]" pageId="10" pageNumber="712">
<emphasis id="AE8B047DFFD63A6A8590FE3AFF0364BC" bold="true" box="[160,254,419,444]" italics="true" pageId="10" pageNumber="712">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4E58BE4FFD63A6A8590FE70FDB76702" box="[160,586,489,514]" pageId="10" pageNumber="712" type="biology_ecology">
<paragraph id="9C40D86FFFD63A6A8590FE70FDB76702" blockId="10.[160,586,489,514]" box="[160,586,489,514]" pageId="10" pageNumber="712">
<emphasis id="AE8B047DFFD63A6A8590FE70FEFD6702" bold="true" box="[160,256,489,514]" italics="true" pageId="10" pageNumber="712">Habitat.</emphasis>
Coastal bushveld/grassland.
</paragraph>
</subSubSection>
</treatment>
</document>
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