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<document id="9AF91DE692464161914697AD36444D47" ID-DOI="10.11646/zootaxa.3670.4.10" ID-GBIF-Dataset="3bdb16f3-3beb-4596-86e3-efd169b4b8e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="217870" ID-ZooBank="DAEFA201-93E5-4109-8C11-AA7AD84FDC70" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460635206891" checkinUser="plazi" docAuthor="Pepato, Almir R. &amp; Silveira, Paulo Sergio Amorim Da" docDate="2013" docId="762787ECFF88FFFFD2D909F8FAFCCC1D" docLanguage="en" docName="zt03670p600.pdf" docOrigin="Zootaxa 3670 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halacarus omului Pepato &amp; Silveira, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="597" masterDocId="8A1EFF94FF89FFF9D24E0D71FF94CE42" masterDocTitle="Two new species of Halacarus (Acari, Prostigmata) from Brazil" masterLastPageNumber="600" masterPageNumber="591" pageNumber="592" updateTime="1698335175665" updateUser="plazi">
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<mods:title id="D098961F51E9F078DA3A646C7E8A0A01">Two new species of Halacarus (Acari, Prostigmata) from Brazil</mods:title>
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<mods:namePart id="9355E863C2E159DDC5414F7B1367906B">Pepato, Almir R.</mods:namePart>
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<mods:namePart id="016E8DA213AD50801FBB8985CF378F68">Silveira, Paulo Sergio Amorim Da</mods:namePart>
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<mods:date id="EF9307341FD9DD1495B28F2169579137">2013</mods:date>
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<treatment id="762787ECFF88FFFFD2D909F8FAFCCC1D" ID-DOI="http://doi.org/10.5281/zenodo.5620639" ID-GBIF-Taxon="119570297" ID-Zenodo-Dep="5620639" LSID="urn:lsid:plazi:treatment:762787ECFF88FFFFD2D909F8FAFCCC1D" httpUri="http://treatment.plazi.org/id/762787ECFF88FFFFD2D909F8FAFCCC1D" lastPageId="6" lastPageNumber="597" pageId="1" pageNumber="592">
<subSubSection id="B6946571FF88FFF8D2D909F8FE9CCA86" pageId="1" pageNumber="592" type="nomenclature">
<paragraph id="FE3136FAFF88FFF8D2D909F8FE45CAE0" blockId="1.[151,465,1160,1220]" box="[151,465,1160,1187]" pageId="1" pageNumber="592">
<heading id="A5798196FF88FFF8D2D909F8FE45CAE0" bold="true" box="[151,465,1160,1187]" fontSize="11" level="1" pageId="1" pageNumber="592" reason="1">
<emphasis id="CCFAEAE8FF88FFF8D2D909F8FE45CAE0" bold="true" box="[151,465,1160,1187]" pageId="1" pageNumber="592">
<taxonomicName id="398E4D79FF88FFF8D2D909F8FEFBCAE1" box="[151,367,1161,1187]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="omului" status="sp. nov.">
<emphasis id="CCFAEAE8FF88FFF8D2D909F8FEFBCAE1" bold="true" box="[151,367,1161,1187]" italics="true" pageId="1" pageNumber="592">Halacarus omului</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7C95793FF88FFF8D33809F9FE45CAE0" box="[374,465,1160,1186]" pageId="1" pageNumber="592" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FE3136FAFF88FFF8D2D909DDFE9CCA86" blockId="1.[151,465,1160,1220]" box="[151,264,1196,1220]" pageId="1" pageNumber="592">
(
<figureCitation id="66B52A7FFF88FFF8D2D009DDFF6BCA86" box="[158,255,1196,1220]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1921,1944]" captionStartId-1="4.[151,250,1914,1937]" captionTargetBox-0="[271,1316,195,1870]" captionTargetBox-1="[266,1320,193,1893]" captionTargetId-0="figure@3.[271,1316,195,1870]" captionTargetId-1="figure@4.[266,1320,193,1894]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." captionText-1="FIGURE 2. Halacarus omului sp. nov., Deutonymph: A. Idiosoma, dorsal. B. Idiosoma, ventral. C. Leg IV. D. Leg I. E. Leg III. F. Leg II. P rotonymph: G. Idiosoma, dorsal. H. Idiosoma, ventral. I. Leg III. J. Leg I. K. Leg IV. L. Leg II. Scale bars: 50 μm." httpUri-0="https://zenodo.org/record/217871/files/figure.png" httpUri-1="https://zenodo.org/record/217872/files/figure.png" pageId="1" pageNumber="592">Figs 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6946571FF88FFF8D2D90985FB29C99E" pageId="1" pageNumber="592" type="materials_examined">
<paragraph id="FE3136FAFF88FFF8D2D90985FE05CB6D" blockId="1.[151,1436,1268,2013]" pageId="1" pageNumber="592">
<emphasis id="CCFAEAE8FF88FFF8D2D90985FE9FCB4E" bold="true" box="[151,267,1268,1292]" pageId="1" pageNumber="592">
<typeStatus id="21358858FF88FFF8D2D90985FE92CB4E" box="[151,262,1268,1292]" pageId="1" pageNumber="592" type="holotype">Holotype</typeStatus>
:
</emphasis>
Female (UFMG-AC1200163), on
<taxonomicName id="398E4D79FF88FFF8D0C10984FBA3CB4E" authority="C. Agardh, 1820" authorityName="C. Agardh" authorityYear="1820" box="[655,1079,1268,1292]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="592" phylum="Heterokontophyta" rank="species" species="cymosum">
<emphasis id="CCFAEAE8FF88FFF8D0C10984FCE1CB4E" box="[655,885,1269,1292]" italics="true" pageId="1" pageNumber="592">Sargassum cymosum</emphasis>
C. Agardh, 1820
</taxonomicName>
at Lázaro Beach, Ubatuba,
<date id="8A30103AFF88FFF8D7200985FF44CB6D" pageId="1" pageNumber="592" value="1991-07">July 1991</date>
, coll. R. Curvelo.
</paragraph>
<paragraph id="FE3136FAFF88FFF8D2D9084DFB29C99E" blockId="1.[151,1436,1268,2013]" pageId="1" pageNumber="592">
<emphasis id="CCFAEAE8FF88FFF8D2D9084DFE81CB16" bold="true" box="[151,277,1340,1364]" pageId="1" pageNumber="592">
<typeStatus id="21358858FF88FFF8D2D9084DFE85CB16" box="[151,273,1340,1364]" pageId="1" pageNumber="592" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Seventeen females (UFMG-AC12000881200096, 12001101200116, 1200155), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D077082EFCB5CB35" box="[569,801,1375,1399]" pageId="1" pageNumber="592" value="2001-09-27">27 September 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D13D082EFC52CB35" box="[883,966,1375,1399]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; six females (
<collectionCode id="989FAE3FFF88FFF8D693082EFABACB35" box="[1245,1326,1375,1399]" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="1" pageNumber="592">UFMG</collectionCode>
1200144 1200147, 12001531200154), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D65808F5FB63CBDE" box="[1046,1271,1412,1436]" pageId="1" pageNumber="592" value="2001-09-28">28 September 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D70C08F5FA07CBDE" box="[1346,1427,1412,1436]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/ FAPESP; three females (UFMG-AC12001361200138) on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D76008D6FF44CBA6" pageId="1" pageNumber="592" value="2001-03-16">16 March 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D35D08BDFEF2CBA6" box="[275,358,1484,1508]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; two females (UFMG-AC1200162, 1200166), on
<taxonomicName id="398E4D79FF88FFF8D1B608BCFA08CBA6" authority="C. Agardh, 1820" authorityName="C. Agardh" authorityYear="1820" box="[1016,1436,1484,1508]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="592" phylum="Heterokontophyta" rank="species" species="cymosum">
<emphasis id="CCFAEAE8FF88FFF8D1B608BCFB49CBA6" box="[1016,1245,1485,1508]" italics="true" pageId="1" pageNumber="592">Sargassum cymosum</emphasis>
C. Agardh, 1820
</taxonomicName>
at Lázaro Beach, Ubatuba,
<date id="8A30103AFF88FFF8D384089EFDAEC845" box="[458,570,1519,1543]" pageId="1" pageNumber="592" value="1991-07">July 1991</date>
, coll. R. Curvelo; seventeen males (UFMG-AC12000971200103, 1200117 1200126), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D1790B65FB8CC86E" box="[823,1048,1556,1580]" pageId="1" pageNumber="592" value="2001-09-27">27 September 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D62A0B65FB23C86E" box="[1124,1207,1556,1580]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; six males (UFMG-AC12001301200135), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D6750B46FB67C80D" box="[1083,1267,1591,1615]" pageId="1" pageNumber="592" value="2001-03-16">16 March 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D70C0B46FA07C80D" box="[1346,1427,1591,1615]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/ FAPESP; nine males (UFMG-AC1200152, 12001601200161, 1200168, 12001701200171, 12001731200175), on
<taxonomicName id="398E4D79FF88FFF8D2D90BF1FDC5C8D5" authority="C. Agardh, 1820" authorityName="C. Agardh" authorityYear="1820" box="[151,593,1663,1688]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="592" phylum="Heterokontophyta" rank="species" species="cymosum">
<emphasis id="CCFAEAE8FF88FFF8D2D90BF1FE15C8DA" box="[151,385,1664,1688]" italics="true" pageId="1" pageNumber="592">Sargassum cymosum</emphasis>
C. Agardh, 1820
</taxonomicName>
at Lázaro Beach, Ubatuba,
<date id="8A30103AFF88FFF8D1D10B0EFB87C8D5" box="[927,1043,1663,1687]" pageId="1" pageNumber="592" value="1991-07">July 1991</date>
, coll. R. Curvelo; quiescent male (UFMG-AC1200109), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D1FB0BD5FB07C8FE" box="[949,1171,1700,1724]" pageId="1" pageNumber="592" value="2001-09-27">27 September 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D6950BD5FABAC8FE" box="[1243,1326,1700,1724]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; eight deutonymphs (UFMG-AC12001041200108, 12001271200129), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D3070B9DFDBBC946" box="[329,559,1772,1796]" pageId="1" pageNumber="592" value="2001-09-27">27 September 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D0300B9DFD45C946" box="[638,721,1772,1796]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; three deutonymphs (UFMG-AC1200143, 1200149 1200150), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D1520A61FC53C96A" box="[796,967,1808,1832]" pageId="1" pageNumber="592" value="2001-03-16">16 March 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D6450A61FBCAC96A" box="[1035,1118,1808,1832]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; four deutonymphs (UFMG-AC 1200151, 1200159, 1200167, 1200172), on
<taxonomicName id="398E4D79FF88FFF8D1510A44FB45C90E" authority="C. Agardh, 1820" authorityName="C. Agardh" authorityYear="1820" box="[799,1233,1844,1868]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="592" phylum="Heterokontophyta" rank="species" species="cymosum">
<emphasis id="CCFAEAE8FF88FFF8D1510A44FB93C90E" box="[799,1031,1845,1868]" italics="true" pageId="1" pageNumber="592">Sargassum cymosum</emphasis>
C. Agardh, 1820
</taxonomicName>
at Lázaro Beach, Ubatuba,
<date id="8A30103AFF88FFF8D3450A29FE15C932" box="[267,385,1880,1904]" pageId="1" pageNumber="592" value="1991-07">July 1991</date>
, coll. R. Curvelo; deutonymph (UFMG-AC1200148), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D30D0A0DFD8FC9D6" box="[323,539,1916,1940]" pageId="1" pageNumber="592" value="2001-09-28">28 September 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D0100A0DFD25C9D6" box="[606,689,1916,1940]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; quiescent deutonymph (UFMG-AC1200139), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D06E0AD1FD5EC9FA" box="[544,714,1952,1976]" pageId="1" pageNumber="592" value="2001-03-16">16 March 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D1450AD1FCCAC9FA" box="[779,862,1952,1976]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP; three protonymphs (UFMG-AC1200140 1200142), on algae at Massaguaçu Island, Caraguatatuba,
<date id="8A30103AFF88FFF8D15A0AB5FC2AC99E" box="[788,958,1988,2012]" pageId="1" pageNumber="592" value="2001-03-16">16 March 2001</date>
, coll.
<collectionCode id="989FAE3FFF88FFF8D1B10AB5FBC4C99E" box="[1023,1104,1988,2012]" pageId="1" pageNumber="592">BIOTA</collectionCode>
/FAPESP.
</paragraph>
</subSubSection>
<subSubSection id="B6946571FF8BFFFBD2890DE6FC9FC836" pageId="2" pageNumber="593" type="description">
<paragraph id="FE3136FAFF8BFFFBD2890DE6FBD4CF5E" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
<emphasis id="CCFAEAE8FF8BFFFBD2890DE6FE23CEF2" bold="true" box="[199,439,151,176]" pageId="2" pageNumber="593">Description. Female</emphasis>
: Idiosoma length 475538, 5565 μm long anterior spine included, width 260330. AD length 175188, width 7083, almost indiscernible, only discriminated from remaining idiosoma by its slightly reticulate ornamentation instead of faint maze-like marks or striations in the surrounding area. Anterior spine 0.27 0.41 of AD length. Ds-1 on AD, anterior to gp-1. Gp-1 at level of 0.550.62 of AD.
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890C56FE91CF8D" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
OC lacking, except for smooth corneae (
<figureCitation id="66B52A7FFF8BFFFBD0C00C56FD5ACF7D" box="[654,718,295,319]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
B). Ds-2 on membranous cuticle between AD and corneae. Gp- 2 placed laterally, at level of 0.320.38 of idiosoma. Pore canaliculi on striated cuticle, near lateral eyes. Gp-3 at level of 0.400.45 of idiosoma. PD absent, tegument of posterior idiosoma maze-like, as shown in
<figureCitation id="66B52A7FFF8BFFFBD7740C1EFAEACFC5" box="[1338,1406,367,391]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
D. Insertions of ds-5 and adanal setae adjacent to gp-4 and gp-5 (
<figureCitation id="66B52A7FFF8BFFFBD11B0CE5FC00CFEE" box="[853,916,404,428]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
C), respectively. Ds-5 at level of 0.690.74 of idiosoma.
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890CADFE27CC55" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">AE length 6580, width 245280, with three pairs of setae and without epimeral pores. PE with three ventral setae and one dorsal seta.</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890F55FE1ACC8E" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
One pair of pgs placed on membranous cuticle, at 2858 μm or at 0.661.19 relative to the distance between the anterior margins of GA and GO from anterior edge of GA. GA length 90113, width 125148. GO length 43 53, width 3548, at 4358 μm from anterior edge of GA. Two pairs of pgs on genitoanal plate, close
<collectingCountry id="8699766AFF8BFFFBD75B0F1CFACBCCC6" box="[1301,1375,620,644]" name="Togo" pageId="2" pageNumber="593">to GO</collectingCountry>
. Five pairs of sgs on genital sclerites, grouped 2:3. Epicuticle raised on GA, appearing as refrigent area indicated by a dotted line in
<figureCitation id="66B52A7FFF8BFFFBD37C0FC5FEE4CC8E" box="[306,368,692,716]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
D.
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890FA6FBF8CD56" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
Gnathosoma length 153172, width 7588. Rostrum length 92105, equal to 0.580.63 of gnathosoma length. Trito- and basirostral setae on rostrum, two small cuticular processes at its tip (
<figureCitation id="66B52A7FFF8BFFFBD6440F8DFBDDCD56" box="[1034,1097,764,788]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
F).
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890E50FD2ACA4D" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
Legs with integument faintly reticulated or without conspicuous ornamentation. Leg chaetotaxy as follows: Leg I, 1, 2, 10 (2S), 10 (2S), 13 (4S), 5 (2S); leg II, 1, 4, 8, 10 (1S, 1B), 12 (2S, 2B),7; leg III, 2, 2, 7, 7, 9, 8; leg IV, 1, 2, 5, 6, 9, 7. Tarsus I with three dorsal and two ventral setae, three pairs of eupathidia as parambulacral setae, and setiform famulus and solenidion. Tarsus II with three dorsal and four ventral setae, three pairs of eupathidia and a setiform solenidion. Tarsi III-IV with two pairs of ventral setae and a pair of parambulacral setae. Tarsi III and IV with four and three dorsal setae, respectively (
<figureCitation id="66B52A7FFF8BFFFBD0D60EA5FD42CDAE" box="[664,726,980,1004]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
K, H). Length of telofemora, genua and tibiae of leg I, 143160 μm, 130143 μm, and 120133 μm, respectively.
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD289096DFA9FCA76" blockId="2.[151,1437,150,1976]" box="[199,1291,1052,1077]" pageId="2" pageNumber="593">
Lateral claws not pectinate, with a narrow accessory process. Median claw bidentate (
<figureCitation id="66B52A7FFF8BFFFBD6C6096DFB52CA76" box="[1160,1222,1052,1076]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">
Fig.
<date id="8A30103AFF8BFFFBD6F2096DFB52CA76" box="[1212,1222,1052,1076]" pageId="2" pageNumber="593">1</date>
</figureCitation>
IM).
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890931FCC5CA3E" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
<emphasis id="CCFAEAE8FF8BFFFBD2890931FE99CA1A" bold="true" box="[199,269,1088,1112]" pageId="2" pageNumber="593">Male:</emphasis>
Idiosoma length 430470, width 260330. Similar to female in most features except for those in the genital area and the plumose parambulacral setae on tarsus IV.
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2D909F6FB70CB4E" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
GA length 91111, width 119138. A pair of pgs on membranous cuticle at 1224 μm from the anterior edge of GA. Genital opening length 4247, width 3341, at 3340 μm from the anterior margin of GA. 5882 perigenital setae surrounding GO, beside a pair of distinctly stouter and longer (outlying) setae. Five pairs of sgs on genital sclerites, grouped 2:3. Spermatophorotype extending 1724 μm beyond genital opening (
<figureCitation id="66B52A7FFF8BFFFBD6330985FB29CB4E" box="[1149,1213,1268,1292]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[271,1316,195,1870]" captionTargetId="figure@3.[271,1316,195,1870]" captionTargetPageId="3" captionText="FIGURE 1. Halacarus omului sp. nov., Female: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds- 5 and gp- 4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I. Male: G. GA. M. Tarsus IV. Scale bars: A, E I, M: 50 μm; B D: 10 μm; L: 25 μm." httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="2" pageNumber="593">Fig. 1</figureCitation>
G).
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2890869FCCBCBFD" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
<emphasis id="CCFAEAE8FF8BFFFBD2890869FEF8CB72" bold="true" box="[199,364,1304,1328]" pageId="2" pageNumber="593">Deutonymph:</emphasis>
Idiosoma length 320425. Integument of posterior dorsal portion of idiosoma striated instead of maze-like, as found in adults. Two pairs of pgs on striated cuticle near genital plate. A pair of shorter setae on plate close to primordial GO. Underneath the genital plate, two pairs of genital acetabula found (
<figureCitation id="66B52A7FFF8BFFFBD6DD082EFB4CCB35" box="[1171,1240,1375,1399]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1937]" captionTargetBox="[266,1320,193,1893]" captionTargetId="figure@4.[266,1320,193,1894]" captionTargetPageId="4" captionText="FIGURE 2. Halacarus omului sp. nov., Deutonymph: A. Idiosoma, dorsal. B. Idiosoma, ventral. C. Leg IV. D. Leg I. E. Leg III. F. Leg II. P rotonymph: G. Idiosoma, dorsal. H. Idiosoma, ventral. I. Leg III. J. Leg I. K. Leg IV. L. Leg II. Scale bars: 50 μm." httpUri="https://zenodo.org/record/217872/files/figure.png" pageId="2" pageNumber="593">Fig. 2</figureCitation>
B). Leg chaetotaxy: Leg I, 1, 2, 8 (2S),8 (2S), 11 (4S), 5 (2S); leg II, 1, 4, 6, 8 (1S), 10 (2S, 1B),7; leg III, 2, 2, 5, 6, 8, 6; leg IV, 1, 2, 3, 4, 78, 5. Ventral setae on tarsi I-IV: 2, 4, 2, 2.
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD28908BDFD5FC86E" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
<emphasis id="CCFAEAE8FF8BFFFBD28908BDFEF2CBA6" bold="true" box="[199,358,1484,1508]" pageId="2" pageNumber="593">Protonymph:</emphasis>
Idiosoma length 213320. Similar to deutonymphs regarding dorsal aspects of idiosoma. PE with two ventral setae and one dorsal seta. A single pair of pgs close to primordial GO. Underneath genital sclerite, a single pair of genital acetabula found (
<figureCitation id="66B52A7FFF8BFFFBD0170B65FD33C86E" box="[601,679,1556,1580]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1937]" captionTargetBox="[266,1320,193,1893]" captionTargetId="figure@4.[266,1320,193,1894]" captionTargetPageId="4" captionText="FIGURE 2. Halacarus omului sp. nov., Deutonymph: A. Idiosoma, dorsal. B. Idiosoma, ventral. C. Leg IV. D. Leg I. E. Leg III. F. Leg II. P rotonymph: G. Idiosoma, dorsal. H. Idiosoma, ventral. I. Leg III. J. Leg I. K. Leg IV. L. Leg II. Scale bars: 50 μm." httpUri="https://zenodo.org/record/217872/files/figure.png" pageId="2" pageNumber="593">Figs. 2</figureCitation>
H).
</paragraph>
<paragraph id="FE3136FAFF8BFFFBD2D90B48FC9FC836" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">Leg chaetotaxy: Leg I, 1, 2, 6 (2S), 6 (2S), 9 (4S), 5 (2S); leg II, 1, 3, 5, 6 (1S), 6 (1B), 5; leg III, 2, 2, 3, 3, 56, 5; leg IV, 0, 3, 3, 5, 3. Ventral setae on tarsi I-IV: 2, 2, 1, 0.</paragraph>
</subSubSection>
<subSubSection id="B6946571FF8BFFFBD2890BF1FB11C946" pageId="2" pageNumber="593" type="etymology">
<paragraph id="FE3136FAFF8BFFFBD2890BF1FB11C946" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
<emphasis id="CCFAEAE8FF8BFFFBD2890BF1FEDAC8DA" bold="true" box="[199,334,1664,1688]" pageId="2" pageNumber="593">Etymology.</emphasis>
The specific name refers to Omulu, an African god that is a member of the Candomble and the Umbanda pantheons, who is syncretically taken in Umbanda as Saint Lazarus. Hence, the name refers indirectly to the
<typeStatus id="21358858FF8BFFFBD28F0BB8FF65C8A3" box="[193,241,1737,1761]" pageId="2" pageNumber="593">type</typeStatus>
locality, Lázaro Beach. It refers also to the fact that both Saint Lazarus and Omulu are depicted with skin pustules in their representations, and
<taxonomicName id="398E4D79FF8BFFFBD0790B9DFD39C946" box="[567,685,1772,1796]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="CCFAEAE8FF8BFFFBD0790B9DFD39C946" box="[567,685,1772,1796]" italics="true" pageId="2" pageNumber="593">Halacarus</emphasis>
</taxonomicName>
species have well-developed gland pores.
</paragraph>
</subSubSection>
<subSubSection id="B6946571FF8BFFFFD2890A61FAFCCC1D" lastPageId="6" lastPageNumber="597" pageId="2" pageNumber="593" type="discussion">
<paragraph id="FE3136FAFF8BFFFBD2890A61FEDFC9FA" blockId="2.[151,1437,150,1976]" pageId="2" pageNumber="593">
<emphasis id="CCFAEAE8FF8BFFFBD2890A61FEAFC96A" bold="true" box="[199,315,1808,1832]" pageId="2" pageNumber="593">Remarks.</emphasis>
<taxonomicName id="398E4D79FF8BFFFBD30A0A61FD85C96A" box="[324,529,1808,1832]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="omului" status="sp. nov.">
<emphasis id="CCFAEAE8FF8BFFFBD30A0A61FD85C96A" box="[324,529,1808,1832]" italics="true" pageId="2" pageNumber="593">Halacarus omului</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7C95793FF8BFFFBD0570A61FDE1C96A" box="[537,629,1808,1832]" pageId="2" pageNumber="593" rank="species">
<emphasis id="CCFAEAE8FF8BFFFBD0570A61FDE1C96A" bold="true" box="[537,629,1808,1832]" pageId="2" pageNumber="593">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the
<taxonomicName id="398E4D79FF8BFFFBD1690A60FCE9C96A" box="[807,893,1809,1832]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8BFFFBD1690A60FCE9C96A" box="[807,893,1809,1832]" italics="true" pageId="2" pageNumber="593">actenos</emphasis>
</taxonomicName>
species group, defined by the lacking of PD in both sexes, reduction or absence of OC, long apodemes on AE and PE, GA with rather uniform cerotegumental cover and two pairs of pgs adjacent
<collectingCountry id="8699766AFF8BFFFBD07F0A28FDE3C932" box="[561,631,1880,1904]" name="Togo" pageId="2" pageNumber="593">to GO</collectingCountry>
in the female, one pair (rarely two pairs) of pgs anterior to GA, one or two bipectinate setae on tibia II, claw shaft without tines, lateral claws often with pectinate accessory process (
<bibRefCitation id="9A1F4B0BFF8BFFFBD2D00AD1FED4C9FA" author="Bartsch" box="[158,320,1952,1976]" pageId="2" pageNumber="593" refString="Bartsch, I. (2011) Halacarus socius (Acari: Halacaridae), description of the male and diagnoses of species of the Halacarus actenos group. Zootaxa, 2800, 18 - 40." type="journal article" year="2011">Bartsch, 2011</bibRefCitation>
).
</paragraph>
<caption id="AAF16672FF8AFFFAD2D90AF0FC67C991" httpUri="https://zenodo.org/record/217871/files/figure.png" pageId="3" pageNumber="594" targetBox="[271,1316,195,1870]" targetPageId="3">
<paragraph id="FE3136FAFF8AFFFAD2D90AF0FC67C991" blockId="3.[151,1436,1921,2005]" pageId="3" pageNumber="594">
<emphasis id="CCFAEAE8FF8AFFFAD2D90AF0FE81C9D5" bold="true" box="[151,277,1921,1944]" pageId="3" pageNumber="594">FIGURE 1.</emphasis>
<taxonomicName id="398E4D79FF8AFFFAD3500AF3FE4CC9DA" box="[286,472,1922,1944]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="omului" status="sp. nov.">
<emphasis id="CCFAEAE8FF8AFFFAD3500AF3FE4CC9DA" box="[286,472,1922,1944]" italics="true" pageId="3" pageNumber="594">Halacarus omului</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7C95793FF8AFFFAD3AF0AF3FDA0C9D5" box="[481,564,1922,1943]" pageId="3" pageNumber="594" rank="species">
<emphasis id="CCFAEAE8FF8AFFFAD3AF0AF3FDA0C9D5" bold="true" box="[481,564,1922,1943]" pageId="3" pageNumber="594">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="CCFAEAE8FF8AFFFAD00A0AF3FD07C9D5" bold="true" box="[580,659,1922,1943]" pageId="3" pageNumber="594">Female</emphasis>
: A. Idiosoma, dorsal. B. Cornea. C. Sclerite with ds-5 and gp-4. D. Body surface on posterior idiosoma. E. Idiosoma, ventral. F. Gnathosoma ventral. H. Leg IV. I. Leg I. J. Leg II. K. Leg III. L. Tarsus I.
<emphasis id="CCFAEAE8FF8AFFFAD2E20AB1FF77C997" bold="true" box="[172,227,1984,2005]" pageId="3" pageNumber="594">Male</emphasis>
: G. GA. M. Tarsus IV. Scale bars: A, EI, M: 50 μm; BD: 10 μm; L: 25 μm.
</paragraph>
</caption>
<caption id="AAF16672FF8DFFFDD2D90A0BFF2BC98E" httpUri="https://zenodo.org/record/217872/files/figure.png" pageId="4" pageNumber="595" targetBox="[266,1320,193,1893]" targetPageId="4">
<paragraph id="FE3136FAFF8DFFFDD2D90A0BFF2BC98E" blockId="4.[151,1436,1914,1996]" pageId="4" pageNumber="595">
<emphasis id="CCFAEAE8FF8DFFFDD2D90A0BFE87C9D2" bold="true" box="[151,275,1914,1937]" pageId="4" pageNumber="595">FIGURE 2.</emphasis>
<taxonomicName id="398E4D79FF8DFFFDD3540A0AFE45C9D3" box="[282,465,1915,1937]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="omului" status="sp. nov.">
<emphasis id="CCFAEAE8FF8DFFFDD3540A0AFE45C9D3" box="[282,465,1915,1937]" italics="true" pageId="4" pageNumber="595">Halacarus omului</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7C95793FF8DFFFDD3970A0AFDBDC9D2" box="[473,553,1915,1936]" pageId="4" pageNumber="595" rank="species">
<emphasis id="CCFAEAE8FF8DFFFDD3970A0AFDBDC9D2" bold="true" box="[473,553,1915,1936]" pageId="4" pageNumber="595">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="CCFAEAE8FF8DFFFDD0790A0AFD57C9D2" bold="true" box="[567,707,1915,1936]" pageId="4" pageNumber="595">Deutonymph</emphasis>
: A. Idiosoma, dorsal. B. Idiosoma, ventral. C. Leg IV. D. Leg I. E. Leg III. F. Leg II. P
<emphasis id="CCFAEAE8FF8DFFFDD37F0AEBFE3DC9ED" bold="true" box="[305,425,1946,1967]" pageId="4" pageNumber="595">rotonymph</emphasis>
: G. Idiosoma, dorsal. H. Idiosoma, ventral. I. Leg III. J. Leg I. K. Leg IV. L. Leg II. Scale bars: 50 μm.
</paragraph>
</caption>
<paragraph id="FE3136FAFF8CFFFCD2890DE6FEB1CF8D" blockId="5.[151,1436,151,572]" pageId="5" pageNumber="596">
Currently, this group consists of 21 species:
<taxonomicName id="398E4D79FF8CFFFCD0AD0DE8FBA7CEED" authority="Trouessart, 1889" authorityName="Trouessart" authorityYear="1889" box="[739,1075,151,176]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8CFFFCD0AD0DE8FCF4CEF2" box="[739,864,153,176]" italics="true" pageId="5" pageNumber="596">H. actenos</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD1200DE9FBA7CEED" author="Trouessart" box="[878,1075,151,175]" pageId="5" pageNumber="596" refString="Trouessart, E. (1889) Revue synoptique de la famille des Halacaridae. Bulletin scientifiques de la France et de la Belgique, Serie 3, 20, 225 - 251." type="journal article" year="1889">Trouessart, 1889</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD60A0DE8FAE4CEED" authority="Bartsch, 1987" authorityName="Bartsch" authorityYear="1987" box="[1092,1392,151,176]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="CCFAEAE8FF8CFFFCD60A0DE8FB2ACEF2" box="[1092,1214,153,176]" italics="true" pageId="5" pageNumber="596">H. griseus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD6850DE6FAE4CEED" author="Bartsch" box="[1227,1392,151,175]" pageId="5" pageNumber="596" refString="Bartsch, I. (1987) Halacarus griseus sp. n., eine Halacaride (Acari) aus dem Mittelmeer. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 9, 75 - 79." type="journal article" year="1987">Bartsch, 1987</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD7CF0DE8FE21CE96" authority="Police, 1909" authorityName="Police" authorityYear="1909" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="longiunguis">
<emphasis id="CCFAEAE8FF8CFFFCD7CF0DE8FE8FCE96" italics="true" pageId="5" pageNumber="596">H. longiunguis</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD36A0DCDFE21CE96" author="Police" box="[292,437,188,212]" pageId="5" pageNumber="596" refString="Police, G. (1909) Alcune nuove specie di Halacaridae del Golfo di Napoli. Archivio Zoologico, Napoli, 3, 409 - 443." type="journal article" year="1909">Police, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD38D0DCCFC8CCE96" authority="Bartsch, 1996" authorityName="Bartsch" authorityYear="1996" box="[451,792,188,212]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="prolongatus">
<emphasis id="CCFAEAE8FF8CFFFCD38D0DCCFDFACE96" box="[451,622,188,212]" italics="true" pageId="5" pageNumber="596">H. prolongatus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD0390DCDFC8CCE96" author="Bartsch" box="[631,792,188,212]" pageId="5" pageNumber="596" refString="Bartsch, I. (1996) Halacarid mites (Acari) from the Mid-Atlantic Ridge. New records. Cahiers de Biologie Marine, 37, 159 - 167." type="journal article" year="1996">Bartsch, 1996</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="398E4D79FF8CFFFCD1190DCCFB16CE96" authority="Bartsch, 2002" authorityName="Bartsch" authorityYear="2002" box="[855,1154,188,212]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="leptopus">
<emphasis id="CCFAEAE8FF8CFFFCD1190DCCFC4FCE96" box="[855,987,188,212]" italics="true" pageId="5" pageNumber="596">H. leptopus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD1AA0DCDFB16CE96" author="Bartsch" box="[996,1154,188,212]" pageId="5" pageNumber="596" refString="Bartsch, I. (2002) Halacarids from the Great Meteor Seamount (Northeastern Atlantic). Description of new species of the genera Agauopsis, Atelopsalis and Halacarus and redescription of H. spiniger Bartsch (Arachnida: Acari: Halacaridae). Mitteilungen aus dem Hamburgischen Zoologische Museum und Institut, 99, 29 - 45." type="journal article" year="2002">Bartsch, 2002</bibRefCitation>
</taxonomicName>
from the North Atlantic and the Mediterranean;
<taxonomicName id="398E4D79FF8CFFFCD3EB0D90FC94CEB5" authority="Kishida, 1927" authorityName="Kishida" authorityYear="1927" box="[421,768,223,248]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="spongiphilus">
<emphasis id="CCFAEAE8FF8CFFFCD3EB0D90FDC3CEBA" box="[421,599,224,248]" italics="true" pageId="5" pageNumber="596">H. spongiphilus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD02E0DAEFC94CEB5" author="Kishida" box="[608,768,223,247]" pageId="5" pageNumber="596" refString="Kishida, K. (1927) Ushio-dani, Ushio-dani-ka [Halacarus spongiphilus, Halacaridae]. In: Iizuka, et al. (Eds.), Nippon Doubutsu Zukan [Illustrated Encyclopedia of Japanese Animals]. Hokuryukan, Tokyo, pp. 978." type="book chapter" year="1927">Kishida, 1927</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD1430D90FBD8CEB5" authority="Krantz,1973" authorityName="Krantz" authorityYear="1973" box="[781,1100,223,248]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="magniporus">
<emphasis id="CCFAEAE8FF8CFFFCD1430D90FC22CEBA" box="[781,950,225,248]" italics="true" pageId="5" pageNumber="596">H. magniporus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD1F00DAEFBD8CEB5" author="Krantz" box="[958,1100,223,247]" pageId="5" pageNumber="596" refString="Krantz, G. W. (1973) Four new predatory species of Halacaridae (Acari: Prostigmata) from Oregon, with remarks on their distribution in the intertidal mussel habitats (Pelecypoda: Mytilidae). Annals of the Entomological Society of America, 66, 975 - 985." type="journal article" year="1973">Krantz,1973</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="398E4D79FF8CFFFCD6C70D90FA08CEB5" authority="Sokolov, 1952" authorityName="Sokolov" authorityYear="1952" box="[1161,1436,223,248]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="mollis">
<emphasis id="CCFAEAE8FF8CFFFCD6C70D90FB65CEBA" box="[1161,1265,224,248]" italics="true" pageId="5" pageNumber="596">H. mollis</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD6B70DAEFA08CEB5" author="Sokolov" box="[1273,1436,223,247]" pageId="5" pageNumber="596" refString="Sokolov, I. I. (1952) Vodjanye kles c i. II. Halacarae. Fauna SSSR, 5, Izdatelstvo Akademii NAUK SSSR, Moskva, Leningrad, 201 pp." type="book" year="1952">Sokolov, 1952</bibRefCitation>
</taxonomicName>
from the North Pacific;
<taxonomicName id="398E4D79FF8CFFFCD3EA0C74FD50CF5E" authority="Newell, 1984" authorityName="Newell" authorityYear="1984" box="[420,708,260,284]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="higginsi">
<emphasis id="CCFAEAE8FF8CFFFCD3EA0C74FDB5CF5E" box="[420,545,260,284]" italics="true" pageId="5" pageNumber="596">H. higginsi</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD0670C75FD50CF5E" author="Newell" box="[553,708,260,284]" pageId="5" pageNumber="596" refString="Newell, I. M. (1984) Biology of the Antartic Seas. XV Antarctic Halacaroidea. Antarctic Research Series, 40, 1 - 284." type="journal article" year="1984">Newell, 1984</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="398E4D79FF8CFFFCD0B10C74FB88CF5E" authority="Newell, 1984" authorityName="Newell" authorityYear="1984" box="[767,1052,260,284]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="perditus">
<emphasis id="CCFAEAE8FF8CFFFCD0B10C74FCE9CF5E" box="[767,893,260,284]" italics="true" pageId="5" pageNumber="596">H. perditus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD1CA0C75FB88CF5E" author="Newell" box="[900,1052,260,284]" pageId="5" pageNumber="596" refString="Newell, I. M. (1984) Biology of the Antartic Seas. XV Antarctic Halacaroidea. Antarctic Research Series, 40, 1 - 284." type="journal article" year="1984">Newell, 1984</bibRefCitation>
</taxonomicName>
from the Southeastern Pacific;
<taxonomicName id="398E4D79FF8CFFFCD7CF0C74FE18CF7D" authority="Bartsch, 1979" authorityName="Bartsch" authorityYear="1979" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="CCFAEAE8FF8CFFFCD7CF0C74FF77CF02" italics="true" pageId="5" pageNumber="596">H. nitidus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD2A20C56FE18CF7D" author="Bartsch" box="[236,396,295,319]" pageId="5" pageNumber="596" refString="Bartsch, I. (1979) Five new species of Halacaridae (Acari) from New Zealand. New Zealand Journal of Marine &amp; Freshwater Research, 13, 175 - 185. http: // dx. doi. org / 10.1080 / 00288330.1979.9515792" type="journal article" year="1979">Bartsch, 1979</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="398E4D79FF8CFFFCD3850C58FD4FCF7D" authority="Bartsch, 1992" authorityName="Bartsch" authorityYear="1992" box="[459,731,295,320]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="socius">
<emphasis id="CCFAEAE8FF8CFFFCD3850C58FDA1CF02" box="[459,565,297,320]" italics="true" pageId="5" pageNumber="596">H. socius</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD0700C56FD4FCF7D" author="Bartsch" box="[574,731,295,319]" pageId="5" pageNumber="596" refString="Bartsch, I. (1992) Halacariden von den Inseln Moorea und Bora Bora, Gesellschaftsinseln (Arachnida: Acari). Senckenbergiana Biologica, 72, 465 - 488." type="journal article" year="1992">Bartsch, 1992</bibRefCitation>
</taxonomicName>
from the Tropical Southern Pacific;
<taxonomicName id="398E4D79FF8CFFFCD6CC0C58FF46CF26" authority="Bartsch, 1993" authorityName="Bartsch" authorityYear="1993" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="discophorus">
<emphasis id="CCFAEAE8FF8CFFFCD6CC0C58FAA2CF02" box="[1154,1334,296,320]" italics="true" pageId="5" pageNumber="596">H. discophorus,</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD7710C56FF46CF26" author="Bartsch" pageId="5" pageNumber="596" refString="Bartsch, I. (1993) Halacarus (Halacaridae, Acari) from south-western Australia. In: Wells, F. E., Walker, D. I., Kirkman, H. &amp; Lethbridge, R. (Eds.), The marine fauna and flora of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 19 - 43." type="book chapter" year="1993">Bartsch, 1993</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD2AE0C3CFD84CF26" authority="Bartsch, 1993" authorityName="Bartsch" authorityYear="1993" box="[224,528,332,356]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="flavellus">
<emphasis id="CCFAEAE8FF8CFFFCD2AE0C3CFEF0CF26" box="[224,356,332,356]" italics="true" pageId="5" pageNumber="596">H. flavellus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD3210C3DFD84CF26" author="Bartsch" box="[367,528,332,356]" pageId="5" pageNumber="596" refString="Bartsch, I. (1993) Halacarus (Halacaridae, Acari) from south-western Australia. In: Wells, F. E., Walker, D. I., Kirkman, H. &amp; Lethbridge, R. (Eds.), The marine fauna and flora of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 19 - 43." type="book chapter" year="1993">Bartsch, 1993</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD0510C3CFCD8CF26" authority="Bartsch, 1993" authorityName="Bartsch" authorityYear="1993" box="[543,844,332,356]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="CCFAEAE8FF8CFFFCD0510C3CFD34CF26" box="[543,672,332,356]" italics="true" pageId="5" pageNumber="596">H. fuscatus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD0E40C3DFCD8CF26" author="Bartsch" box="[682,844,332,356]" pageId="5" pageNumber="596" refString="Bartsch, I. (1993) Halacarus (Halacaridae, Acari) from south-western Australia. In: Wells, F. E., Walker, D. I., Kirkman, H. &amp; Lethbridge, R. (Eds.), The marine fauna and flora of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 19 - 43." type="book chapter" year="1993">Bartsch, 1993</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="398E4D79FF8CFFFCD1C30C3CFB49CF26" authority="Bartsch, 1981" authorityName="Bartsch" authorityYear="1981" box="[909,1245,332,356]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="multispinus">
<emphasis id="CCFAEAE8FF8CFFFCD1C30C3CFBA0CF26" box="[909,1076,332,356]" italics="true" pageId="5" pageNumber="596">H. multispinus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD6700C3DFB49CF26" author="Bartsch" box="[1086,1245,332,356]" pageId="5" pageNumber="596" refString="Bartsch, I. (1981) Halacaridae (Acari) aus dem Kanal von Mocambique. Cahiers de Biologie marine, 22, 35 - 63." type="journal article" year="1981">Bartsch, 1981</bibRefCitation>
</taxonomicName>
from the Indian Ocean;
<taxonomicName id="398E4D79FF8CFFFCD2BE0C00FDBCCFC5" authority="Newell, 1984" authorityName="Newell" authorityYear="1984" box="[240,552,367,392]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="CCFAEAE8FF8CFFFCD2BE0C00FE15CFCA" box="[240,385,368,392]" italics="true" pageId="5" pageNumber="596">H. echinatus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD3C50C1EFDBCCFC5" author="Newell" box="[395,552,367,391]" pageId="5" pageNumber="596" refString="Newell, I. M. (1984) Biology of the Antartic Seas. XV Antarctic Halacaroidea. Antarctic Research Series, 40, 1 - 284." type="journal article" year="1984">Newell, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD0780C00FC02CFC5" authority="Newell, 1984" authorityName="Newell" authorityYear="1984" box="[566,918,367,392]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="falklandensis">
<emphasis id="CCFAEAE8FF8CFFFCD0780C00FD64CFCA" box="[566,752,368,392]" italics="true" pageId="5" pageNumber="596">H. falklandensis</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD0B40C1EFC02CFC5" author="Newell" box="[762,918,367,391]" pageId="5" pageNumber="596" refString="Newell, I. M. (1984) Biology of the Antartic Seas. XV Antarctic Halacaroidea. Antarctic Research Series, 40, 1 - 284." type="journal article" year="1984">Newell, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD1EB0C00FA9CCFC5" authority="Newell, 1984" authorityName="Newell" authorityYear="1984" box="[933,1288,367,392]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="hemispinosus">
<emphasis id="CCFAEAE8FF8CFFFCD1EB0C00FBF6CFCA" box="[933,1122,368,392]" italics="true" pageId="5" pageNumber="596">H. hemispinosus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD6220C1EFA9CCFC5" author="Newell" box="[1132,1288,367,391]" pageId="5" pageNumber="596" refString="Newell, I. M. (1984) Biology of the Antartic Seas. XV Antarctic Halacaroidea. Antarctic Research Series, 40, 1 - 284." type="journal article" year="1984">Newell, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD7590C00FEDACFEE" authority="Lohmann, 1907" authorityName="Lohmann" authorityYear="1907" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="CCFAEAE8FF8CFFFCD7590C00FA08CFCA" box="[1303,1436,368,392]" italics="true" pageId="5" pageNumber="596">H. robustus</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD2D90CE5FEDACFEE" author="Lohmann" box="[151,334,404,428]" pageId="5" pageNumber="596" refString="Lohmann, H. (1907) Die Meeresmilben der Deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition 1901 - 1903, 9. G. Fischer, Jena, pp. 361 - 413." type="journal article" year="1907">Lohmann, 1907</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="398E4D79FF8CFFFCD3C10CE4FD2FCFEE" authority="Lohmann, 1907" authorityName="Lohmann" authorityYear="1907" box="[399,699,404,428]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="werthi">
<emphasis id="CCFAEAE8FF8CFFFCD3C10CE4FE6FCFEE" box="[399,507,404,428]" italics="true" pageId="5" pageNumber="596">H. werthi</emphasis>
<bibRefCitation id="9A1F4B0BFF8CFFFCD0480CE5FD2FCFEE" author="Lohmann" box="[518,699,404,428]" pageId="5" pageNumber="596" refString="Lohmann, H. (1907) Die Meeresmilben der Deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition 1901 - 1903, 9. G. Fischer, Jena, pp. 361 - 413." type="journal article" year="1907">Lohmann, 1907</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="8699766AFF8CFFFCD17C0CE5FC09CFEE" box="[818,925,404,428]" name="Antarctica" pageId="5" pageNumber="596">Antarctic</collectingCountry>
, Subantarctic and the adjacent sea (
<bibRefCitation id="9A1F4B0BFF8CFFFCD7700CE5FF46CF8D" author="Bartsch" pageId="5" pageNumber="596" refString="Bartsch, I. (2002) Halacarids from the Great Meteor Seamount (Northeastern Atlantic). Description of new species of the genera Agauopsis, Atelopsalis and Halacarus and redescription of H. spiniger Bartsch (Arachnida: Acari: Halacaridae). Mitteilungen aus dem Hamburgischen Zoologische Museum und Institut, 99, 29 - 45." type="journal article" year="2002">Bartsch, 2002</bibRefCitation>
,
<bibRefCitation id="9A1F4B0BFF8CFFFCD2930CC6FE8DCF8D" author="Bartsch" box="[221,281,439,463]" pageId="5" pageNumber="596" refString="Bartsch, I. (2011) Halacarus socius (Acari: Halacaridae), description of the male and diagnoses of species of the Halacarus actenos group. Zootaxa, 2800, 18 - 40." type="journal article" year="2011">2011</bibRefCitation>
).
</paragraph>
<paragraph id="FE3136FAFF8CFFFCD2890CADFDC8CC7E" blockId="5.[151,1436,151,572]" pageId="5" pageNumber="596">
Among these species, only
<taxonomicName id="398E4D79FF8CFFFCD3B60CACFDFBCFB6" box="[504,623,477,500]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8CFFFCD3B60CACFDFBCFB6" box="[504,623,477,500]" italics="true" pageId="5" pageNumber="596">H. actenos</emphasis>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD0330CACFD6ACFB6" box="[637,766,476,500]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="leptopus">
<emphasis id="CCFAEAE8FF8CFFFCD0330CACFD6ACFB6" box="[637,766,476,500]" italics="true" pageId="5" pageNumber="596">H. leptopus</emphasis>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD1450CACFCE9CFB6" box="[779,893,477,500]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="CCFAEAE8FF8CFFFCD1450CACFCE9CFB6" box="[779,893,477,500]" italics="true" pageId="5" pageNumber="596">H. griseus</emphasis>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD1C50CACFBA6CFB6" box="[907,1074,477,500]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="magniporus">
<emphasis id="CCFAEAE8FF8CFFFCD1C50CACFBA6CFB6" box="[907,1074,477,500]" italics="true" pageId="5" pageNumber="596">H. magniporus</emphasis>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD60E0CACFB7CCFB6" box="[1088,1256,476,500]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="prolongatus">
<emphasis id="CCFAEAE8FF8CFFFCD60E0CACFB7CCFB6" box="[1088,1256,476,500]" italics="true" pageId="5" pageNumber="596">H. prolongatus</emphasis>
</taxonomicName>
,
<taxonomicName id="398E4D79FF8CFFFCD6B80CACFA08CFB6" box="[1270,1436,476,500]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="longiunguis">
<emphasis id="CCFAEAE8FF8CFFFCD6B80CACFA08CFB6" box="[1270,1436,476,500]" italics="true" pageId="5" pageNumber="596">H. longiunguis</emphasis>
</taxonomicName>
and
<taxonomicName id="398E4D79FF8CFFFCD2890F70FEE7CC5A" box="[199,371,512,536]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="discophorus">
<emphasis id="CCFAEAE8FF8CFFFCD2890F70FEE7CC5A" box="[199,371,512,536]" italics="true" pageId="5" pageNumber="596">H. discophorus</emphasis>
</taxonomicName>
share the presence of ds-5 and gp-4, and adanal setae and gp-5, adjacent on the same sclerites with
<taxonomicName id="398E4D79FF8CFFFCD2810F54FEABCC7E" box="[207,319,548,572]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="omului" status="sp. nov.">
<emphasis id="CCFAEAE8FF8CFFFCD2810F54FEABCC7E" box="[207,319,548,572]" italics="true" pageId="5" pageNumber="596">H. omului</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7C95793FF8CFFFCD3080F55FE34CC7E" box="[326,416,548,572]" pageId="5" pageNumber="596" rank="species">
<emphasis id="CCFAEAE8FF8CFFFCD3080F55FE34CC7E" bold="true" box="[326,416,548,572]" pageId="5" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<bibRefCitation id="9A1F4B0BFF8CFFFCD3E10F55FDC4CC7E" author="Bartsch" box="[431,592,548,572]" pageId="5" pageNumber="596" refString="Bartsch, I. (2011) Halacarus socius (Acari: Halacaridae), description of the male and diagnoses of species of the Halacarus actenos group. Zootaxa, 2800, 18 - 40." type="journal article" year="2011">Bartsch, 2011</bibRefCitation>
).
</paragraph>
<caption id="AAF16672FF8CFFFCD2D90F17FCBFCCBF" ID-Table-UUID="AAF16672FF8CFFFCD2D90F17FCBFCCBF" httpUri="http://table.plazi.org/id/AAF16672FF8CFFFCD2D90F17FCBFCCBF" pageId="5" pageNumber="596" targetBox="[159,1420,916,1228]" targetIsTable="true" targetPageId="5">
<paragraph id="FE3136FAFF8CFFFCD2D90F17FCBFCCBF" blockId="5.[151,1436,614,765]" pageId="5" pageNumber="596">
<emphasis id="CCFAEAE8FF8CFFFCD2D90F17FE9FCC3C" bold="true" box="[151,267,614,638]" pageId="5" pageNumber="596">TABLE 1.</emphasis>
Character state distributions in species of the
<taxonomicName id="398E4D79FF8CFFFCD15C0F17FC4ACC3F" box="[786,990,614,637]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8CFFFCD15C0F17FC4ACC3F" box="[786,990,614,637]" italics="true" pageId="5" pageNumber="596">Halacarus actenos</emphasis>
</taxonomicName>
group (cf. Bartsch, 2011) that share the coexistence of ds-5 with gp-4 and ds-6 with gp-5. Abbreviations used here are as follows: abs, absent; ant, anterior; adj, adjacent to; co, cornea, ocular plate reduced; level, at same level; l, long; integ, striated integument; ling, linguiform; m, medium-sized; ov+co, ovate ocular plate with cornea; pres, present; post, posterior; rem, removed; sh, short; slen, slender; spin, spiniform; trian, triangular;?, no data available.
</paragraph>
</caption>
<paragraph id="FE3136FAFF8CFFFCD3220E68FBC2CD6D" blockId="5.[364,1110,793,815]" box="[364,1110,793,815]" pageId="5" pageNumber="596">idiosoma, length (in µm) frontal spine dorsum</paragraph>
<paragraph id="FE3136FAFF8CFFFCD2D10E3DFA18CD20" blockId="5.[159,1420,844,897]" box="[159,1420,844,866]" pageId="5" pageNumber="596">Species female male size size in relation to shape of posterior position gp-1 on</paragraph>
<paragraph id="FE3136FAFF8CFFFCD17F0E1AFA9ECDC3" blockId="5.[159,1420,844,897]" box="[817,1290,875,897]" pageId="5" pageNumber="596">length of AD AD AD</paragraph>
<paragraph id="FE3136FAFF8CFFFCD2D10EE4FA92CA89" pageId="5" pageNumber="596">
<table id="8C8EC45AFF8C0006D2D10EE5FA18CA8E" box="[159,1420,916,1228]" gridcols="6" gridrows="8" pageId="5" pageNumber="596">
<tr id="40BE34B8FF8C0006D2D10EE5FA18CDE9" box="[159,1420,916,939]" gridrow="0" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10EE5FE8FCDE9" box="[159,283,916,939]" gridcol="0" gridrow="0" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10EE4FF72CDE9" box="[159,230,917,939]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="omului">
<emphasis id="CCFAEAE8FF8CFFFCD2D10EE4FF72CDE9" box="[159,230,917,939]" italics="true" pageId="5" pageNumber="596">omului</emphasis>
</taxonomicName>
</th>
<th id="036F5DC4FF8C0006D3220EE5FE45CDE9" box="[364,465,916,939]" gridcol="1" gridrow="0" pageId="5" pageNumber="596">475538</th>
<th id="036F5DC4FF8C0006D06A0EE5FD40CDE9" box="[548,724,916,939]" gridcol="2" gridrow="0" pageId="5" pageNumber="596">430470 m</th>
<th id="036F5DC4FF8C0006D17F0EE5FCE9CDE9" box="[817,893,916,939]" gridcol="3" gridrow="0" pageId="5" pageNumber="596">0.30.4</th>
<th id="036F5DC4FF8C0006D6440EE5FBADCDE9" box="[1034,1081,916,939]" gridcol="4" gridrow="0" pageId="5" pageNumber="596">ling</th>
<th id="036F5DC4FF8C0006D6A80EE5FA18CDE9" box="[1254,1420,916,939]" gridcol="5" gridrow="0" pageId="5" pageNumber="596">0.6</th>
</tr>
<tr id="40BE34B8FF8C0006D2D10ECFFA18CD96" box="[159,1420,958,980]" gridrow="1" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10ECFFE8FCD96" box="[159,283,958,980]" gridcol="0" gridrow="1" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10ECEFF78CD96" box="[159,236,959,980]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8CFFFCD2D10ECEFF78CD96" box="[159,236,959,980]" italics="true" pageId="5" pageNumber="596">actenos</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3220ECFFE45CD96" box="[364,465,958,980]" gridcol="1" gridrow="1" pageId="5" pageNumber="596">785879</td>
<td id="036F5DC4FF8C0006D06A0ECFFD40CD96" box="[548,724,958,980]" gridcol="2" gridrow="1" pageId="5" pageNumber="596">555670 m</td>
<td id="036F5DC4FF8C0006D17F0ECFFCE9CD96" box="[817,893,958,980]" gridcol="3" gridrow="1" pageId="5" pageNumber="596">0.4</td>
<td id="036F5DC4FF8C0006D6440ECFFBADCD96" box="[1034,1081,958,980]" gridcol="4" gridrow="1" pageId="5" pageNumber="596">spin</td>
<td id="036F5DC4FF8C0006D6A80ECFFA18CD96" box="[1254,1420,958,980]" gridcol="5" gridrow="1" pageId="5" pageNumber="596">0.6</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10E96FA18CDBF" box="[159,1420,999,1021]" gridrow="2" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10E96FE8FCDBF" box="[159,283,999,1021]" gridcol="0" gridrow="2" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10E96FE8FCDBF" box="[159,283,999,1021]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="discophorus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10E96FE8FCDBF" box="[159,283,999,1021]" italics="true" pageId="5" pageNumber="596">discophorus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3220E96FE45CDBF" box="[364,465,999,1021]" gridcol="1" gridrow="2" pageId="5" pageNumber="596">675</td>
<td id="036F5DC4FF8C0006D06A0E96FD40CDBF" box="[548,724,999,1021]" gridcol="2" gridrow="2" pageId="5" pageNumber="596">470641 m</td>
<td id="036F5DC4FF8C0006D17F0E96FCE9CDBF" box="[817,893,999,1021]" gridcol="3" gridrow="2" pageId="5" pageNumber="596">0.3</td>
<td id="036F5DC4FF8C0006D6440E96FBADCDBF" box="[1034,1081,999,1021]" gridcol="4" gridrow="2" pageId="5" pageNumber="596">trian</td>
<td id="036F5DC4FF8C0006D6A80E96FA18CDBF" box="[1254,1420,999,1021]" gridcol="5" gridrow="2" pageId="5" pageNumber="596">0.6</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10961FA18CA65" box="[159,1420,1040,1063]" gridrow="3" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10961FE8FCA65" box="[159,283,1040,1063]" gridcol="0" gridrow="3" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10963FF7CCA65" box="[159,232,1042,1063]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10963FF7CCA65" box="[159,232,1042,1063]" italics="true" pageId="5" pageNumber="596">griseus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3220961FE45CA65" box="[364,465,1040,1063]" gridcol="1" gridrow="3" pageId="5" pageNumber="596">9141025</td>
<td id="036F5DC4FF8C0006D06A0961FD40CA65" box="[548,724,1040,1063]" gridcol="2" gridrow="3" pageId="5" pageNumber="596">790884 l</td>
<td id="036F5DC4FF8C0006D17F0961FCE9CA65" box="[817,893,1040,1063]" gridcol="3" gridrow="3" pageId="5" pageNumber="596">0.4</td>
<td id="036F5DC4FF8C0006D6440961FBADCA65" box="[1034,1081,1040,1063]" gridcol="4" gridrow="3" pageId="5" pageNumber="596">spin</td>
<td id="036F5DC4FF8C0006D6A80961FA18CA65" box="[1254,1420,1040,1063]" gridcol="5" gridrow="3" pageId="5" pageNumber="596">0.60.7</td>
</tr>
<tr id="40BE34B8FF8C0006D2D1094BFA18CA12" box="[159,1420,1082,1104]" gridrow="4" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D1094BFE8FCA12" box="[159,283,1082,1104]" gridcol="0" gridrow="4" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D1094BFF61CA12" box="[159,245,1082,1104]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="leptopus">
<emphasis id="CCFAEAE8FF8CFFFCD2D1094BFF61CA12" box="[159,245,1082,1104]" italics="true" pageId="5" pageNumber="596">leptopus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D322094BFE45CA12" box="[364,465,1082,1104]" gridcol="1" gridrow="4" pageId="5" pageNumber="596">580653</td>
<td id="036F5DC4FF8C0006D06A094BFD40CA12" box="[548,724,1082,1104]" gridcol="2" gridrow="4" pageId="5" pageNumber="596">562 l</td>
<td id="036F5DC4FF8C0006D17F094BFCE9CA12" box="[817,893,1082,1104]" gridcol="3" gridrow="4" pageId="5" pageNumber="596">0.5</td>
<td id="036F5DC4FF8C0006D644094BFBADCA12" box="[1034,1081,1082,1104]" gridcol="4" gridrow="4" pageId="5" pageNumber="596">ling</td>
<td id="036F5DC4FF8C0006D6A8094BFA18CA12" box="[1254,1420,1082,1104]" gridcol="5" gridrow="4" pageId="5" pageNumber="596">0.7</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10912FA18CA3B" box="[159,1420,1123,1145]" gridrow="5" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10912FE8FCA3B" box="[159,283,1123,1145]" gridcol="0" gridrow="5" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10912FE83CA3B" box="[159,279,1123,1145]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="longiunguis">
<emphasis id="CCFAEAE8FF8CFFFCD2D10912FE83CA3B" box="[159,279,1123,1145]" italics="true" pageId="5" pageNumber="596">longiunguis</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3220912FE45CA3B" box="[364,465,1123,1145]" gridcol="1" gridrow="5" pageId="5" pageNumber="596">572720</td>
<td id="036F5DC4FF8C0006D06A0912FD40CA3B" box="[548,724,1123,1145]" gridcol="2" gridrow="5" pageId="5" pageNumber="596">515559 l</td>
<td id="036F5DC4FF8C0006D17F0912FCE9CA3B" box="[817,893,1123,1145]" gridcol="3" gridrow="5" pageId="5" pageNumber="596">0.5</td>
<td id="036F5DC4FF8C0006D6440912FBADCA3B" box="[1034,1081,1123,1145]" gridcol="4" gridrow="5" pageId="5" pageNumber="596">ling</td>
<td id="036F5DC4FF8C0006D6A80912FA18CA3B" box="[1254,1420,1123,1145]" gridcol="5" gridrow="5" pageId="5" pageNumber="596">0.7</td>
</tr>
<tr id="40BE34B8FF8C0006D2D109FDFA18CAE1" box="[159,1420,1164,1187]" gridrow="6" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D109FDFE8FCAE1" box="[159,283,1164,1187]" gridcol="0" gridrow="6" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D109FFFE8CCAE1" box="[159,280,1166,1187]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="magniporus">
<emphasis id="CCFAEAE8FF8CFFFCD2D109FFFE8CCAE1" box="[159,280,1166,1187]" italics="true" pageId="5" pageNumber="596">magniporus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D32209FDFE45CAE1" box="[364,465,1164,1187]" gridcol="1" gridrow="6" pageId="5" pageNumber="596">825</td>
<td id="036F5DC4FF8C0006D06A09FDFD40CAE1" box="[548,724,1164,1187]" gridcol="2" gridrow="6" pageId="5" pageNumber="596">550 sh</td>
<td id="036F5DC4FF8C0006D17F09FDFCE9CAE1" box="[817,893,1164,1187]" gridcol="3" gridrow="6" pageId="5" pageNumber="596">0.1</td>
<td id="036F5DC4FF8C0006D64409FDFBADCAE1" box="[1034,1081,1164,1187]" gridcol="4" gridrow="6" pageId="5" pageNumber="596">spin</td>
<td id="036F5DC4FF8C0006D6A809FDFA18CAE1" box="[1254,1420,1164,1187]" gridcol="5" gridrow="6" pageId="5" pageNumber="596">0.4</td>
</tr>
<tr id="40BE34B8FF8C0006D2D109C7FA18CA8E" box="[159,1420,1206,1228]" gridrow="7" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D109C7FE8FCA8E" box="[159,283,1206,1228]" gridcol="0" gridrow="7" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D109C7FE8DCA8E" box="[159,281,1206,1228]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="prolongatus">
<emphasis id="CCFAEAE8FF8CFFFCD2D109C7FE8DCA8E" box="[159,281,1206,1228]" italics="true" pageId="5" pageNumber="596">prolongatus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D32209C7FE45CA8E" box="[364,465,1206,1228]" gridcol="1" gridrow="7" pageId="5" pageNumber="596">750</td>
<td id="036F5DC4FF8C0006D06A09C7FD40CA8E" box="[548,724,1206,1228]" gridcol="2" gridrow="7" pageId="5" pageNumber="596">740 l</td>
<td id="036F5DC4FF8C0006D17F09C7FCE9CA8E" box="[817,893,1206,1228]" gridcol="3" gridrow="7" pageId="5" pageNumber="596">0.5</td>
<td id="036F5DC4FF8C0006D64409C7FBADCA8E" box="[1034,1081,1206,1228]" gridcol="4" gridrow="7" pageId="5" pageNumber="596">spin</td>
<td id="036F5DC4FF8C0006D6A809C7FA18CA8E" box="[1254,1420,1206,1228]" gridcol="5" gridrow="7" pageId="5" pageNumber="596">0.7</td>
</tr>
</table>
</paragraph>
<caption id="AAF16672FF8CFFFCD2D90982FE11CB4B" ID-Table-UUID="AAF16672FF8CFFFCD2D90982FE11CB4B" box="[151,389,1267,1289]" httpUri="http://table.plazi.org/id/AAF16672FF8CFFFCD2D90982FE11CB4B" pageId="5" pageNumber="596" targetBox="[159,1424,1316,1751]" targetIsTable="true" targetPageId="5">
<paragraph id="FE3136FAFF8CFFFCD2D90982FE11CB4B" blockId="5.[151,389,1267,1289]" box="[151,389,1267,1289]" pageId="5" pageNumber="596">
<emphasis id="CCFAEAE8FF8CFFFCD2D90982FE90CB4A" bold="true" box="[151,260,1267,1288]" pageId="5" pageNumber="596">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="FE3136FAFF8CFFFCD2D10829FA83C895" pageId="5" pageNumber="596">
<table id="8C8EC45AFF8C0006D2D10855FA04C895" box="[159,1424,1316,1751]" gridcols="6" gridrows="9" pageId="5" pageNumber="596">
<tr id="40BE34B8FF8C0006D2D10855FA04CBCE" box="[159,1424,1316,1420]" gridrow="0" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10855FE8FCBCE" box="[159,283,1316,1420]" gridcol="0" gridrow="0" pageId="5" pageNumber="596">Species</th>
<th id="036F5DC4FF8C0006D3190855FE4ECBCE" box="[343,474,1316,1420]" gridcol="1" gridrow="0" pageId="5" pageNumber="596">dorsum position ds-1 to gp-1</th>
<th id="036F5DC4FF8C0006D3B90855FD12CBCE" box="[503,646,1316,1420]" gridcol="2" gridrow="0" pageId="5" pageNumber="596">OC, presence, shape, cornea</th>
<th id="036F5DC4FF8C0006D0E90855FC8DCBCE" box="[679,793,1316,1420]" gridcol="3" gridrow="0" pageId="5" pageNumber="596">position of canaliculus</th>
<th id="036F5DC4FF8C0006D1750855FC23CBCE" box="[827,951,1316,1420]" gridcol="4" gridrow="0" pageId="5" pageNumber="596">male, dorsal platelets</th>
<th id="036F5DC4FF8C0006D1A50855FA04CBCE" box="[1003,1424,1316,1420]" gridcol="5" gridrow="0" pageId="5" pageNumber="596">female male position of pgs-1 relative number of pgs to distance GAGO</th>
</tr>
<tr id="40BE34B8FF8C0006D2D108D1FA04CBF4" box="[159,1424,1440,1462]" gridrow="1" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D108D1FE8FCBF4" box="[159,283,1440,1462]" gridcol="0" gridrow="1" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D108D1FF72CBF4" box="[159,230,1440,1462]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="omului">
<emphasis id="CCFAEAE8FF8CFFFCD2D108D1FF72CBF4" box="[159,230,1440,1462]" italics="true" pageId="5" pageNumber="596">omului</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D31908D1FE4ECBF4" box="[343,474,1440,1462]" gridcol="1" gridrow="1" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B908D1FD12CBF4" box="[503,646,1440,1462]" gridcol="2" gridrow="1" pageId="5" pageNumber="596">co</td>
<td id="036F5DC4FF8C0006D0E908D1FC8DCBF4" box="[679,793,1440,1462]" gridcol="3" gridrow="1" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D17508D1FC23CBF4" box="[827,951,1440,1462]" gridcol="4" gridrow="1" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A508D1FA04CBF4" box="[1003,1424,1440,1462]" gridcol="5" gridrow="1" pageId="5" pageNumber="596">0.71.2 5882</td>
</tr>
<tr id="40BE34B8FF8C0006D2D108B8FA04CB9D" box="[159,1424,1481,1503]" gridrow="2" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D108B8FE8FCB9D" box="[159,283,1481,1503]" gridcol="0" gridrow="2" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D108BBFF78CB9D" box="[159,236,1482,1503]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8CFFFCD2D108BBFF78CB9D" box="[159,236,1482,1503]" italics="true" pageId="5" pageNumber="596">actenos</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D31908B8FE4ECB9D" box="[343,474,1481,1503]" gridcol="1" gridrow="2" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B908B8FD12CB9D" box="[503,646,1481,1503]" gridcol="2" gridrow="2" pageId="5" pageNumber="596">co</td>
<td id="036F5DC4FF8C0006D0E908B8FC8DCB9D" box="[679,793,1481,1503]" gridcol="3" gridrow="2" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D17508B8FC23CB9D" box="[827,951,1481,1503]" gridcol="4" gridrow="2" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A508B8FA04CB9D" box="[1003,1424,1481,1503]" gridcol="5" gridrow="2" pageId="5" pageNumber="596">0.8 6883</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10883FA04C84A" box="[159,1424,1522,1544]" gridrow="3" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10883FE8FC84A" box="[159,283,1522,1544]" gridcol="0" gridrow="3" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10883FE8FC84A" box="[159,283,1522,1544]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="discophorus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10883FE8FC84A" box="[159,283,1522,1544]" italics="true" pageId="5" pageNumber="596">discophorus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3190883FE4EC84A" box="[343,474,1522,1544]" gridcol="1" gridrow="3" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B90883FD12C84A" box="[503,646,1522,1544]" gridcol="2" gridrow="3" pageId="5" pageNumber="596">ov+co</td>
<td id="036F5DC4FF8C0006D0E90883FC8DC84A" box="[679,793,1522,1544]" gridcol="3" gridrow="3" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D1750883FC23C84A" box="[827,951,1522,1544]" gridcol="4" gridrow="3" pageId="5" pageNumber="596">pres</td>
<td id="036F5DC4FF8C0006D1A50883FA04C84A" box="[1003,1424,1522,1544]" gridcol="5" gridrow="3" pageId="5" pageNumber="596">2.8 66</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10B6DFA04C870" box="[159,1424,1564,1586]" gridrow="4" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10B6DFE8FC870" box="[159,283,1564,1586]" gridcol="0" gridrow="4" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10B6CFF7CC870" box="[159,232,1565,1586]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10B6CFF7CC870" box="[159,232,1565,1586]" italics="true" pageId="5" pageNumber="596">griseus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3190B6DFE4EC870" box="[343,474,1564,1586]" gridcol="1" gridrow="4" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B90B6DFD12C870" box="[503,646,1564,1586]" gridcol="2" gridrow="4" pageId="5" pageNumber="596">co</td>
<td id="036F5DC4FF8C0006D0E90B6DFC8DC870" box="[679,793,1564,1586]" gridcol="3" gridrow="4" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D1750B6DFC23C870" box="[827,951,1564,1586]" gridcol="4" gridrow="4" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A50B6DFA04C870" box="[1003,1424,1564,1586]" gridcol="5" gridrow="4" pageId="5" pageNumber="596">0.9 ca 90</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10B34FA04C819" box="[159,1424,1605,1627]" gridrow="5" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10B34FE8FC819" box="[159,283,1605,1627]" gridcol="0" gridrow="5" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10B34FF61C819" box="[159,245,1605,1627]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="leptopus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10B34FF61C819" box="[159,245,1605,1627]" italics="true" pageId="5" pageNumber="596">leptopus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3190B34FE4EC819" box="[343,474,1605,1627]" gridcol="1" gridrow="5" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B90B34FD12C819" box="[503,646,1605,1627]" gridcol="2" gridrow="5" pageId="5" pageNumber="596">co</td>
<td id="036F5DC4FF8C0006D0E90B34FC8DC819" box="[679,793,1605,1627]" gridcol="3" gridrow="5" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D1750B34FC23C819" box="[827,951,1605,1627]" gridcol="4" gridrow="5" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A50B34FA04C819" box="[1003,1424,1605,1627]" gridcol="5" gridrow="5" pageId="5" pageNumber="596">1.2 ca 70</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10B1FFA04C8C7" box="[159,1424,1646,1669]" gridrow="6" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10B1FFE8FC8C7" box="[159,283,1646,1669]" gridcol="0" gridrow="6" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10B1EFE83C8C7" box="[159,279,1647,1669]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="longiunguis">
<emphasis id="CCFAEAE8FF8CFFFCD2D10B1EFE83C8C7" box="[159,279,1647,1669]" italics="true" pageId="5" pageNumber="596">longiunguis</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3190B1FFE4EC8C7" box="[343,474,1646,1669]" gridcol="1" gridrow="6" pageId="5" pageNumber="596">level</td>
<td id="036F5DC4FF8C0006D3B90B1FFD12C8C7" box="[503,646,1646,1669]" gridcol="2" gridrow="6" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D0E90B1FFC8DC8C7" box="[679,793,1646,1669]" gridcol="3" gridrow="6" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D1750B1FFC23C8C7" box="[827,951,1646,1669]" gridcol="4" gridrow="6" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A50B1FFA04C8C7" box="[1003,1424,1646,1669]" gridcol="5" gridrow="6" pageId="5" pageNumber="596">adj GA ca 105</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10BE9FA04C8EC" box="[159,1424,1688,1710]" gridrow="7" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10BE9FE8FC8EC" box="[159,283,1688,1710]" gridcol="0" gridrow="7" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10BE8FE8CC8EC" box="[159,280,1689,1710]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="magniporus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10BE8FE8CC8EC" box="[159,280,1689,1710]" italics="true" pageId="5" pageNumber="596">magniporus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3190BE9FE4EC8EC" box="[343,474,1688,1710]" gridcol="1" gridrow="7" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B90BE9FD12C8EC" box="[503,646,1688,1710]" gridcol="2" gridrow="7" pageId="5" pageNumber="596">co</td>
<td id="036F5DC4FF8C0006D0E90BE9FC8DC8EC" box="[679,793,1688,1710]" gridcol="3" gridrow="7" pageId="5" pageNumber="596">?</td>
<td id="036F5DC4FF8C0006D1750BE9FC23C8EC" box="[827,951,1688,1710]" gridcol="4" gridrow="7" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A50BE9FA04C8EC" box="[1003,1424,1688,1710]" gridcol="5" gridrow="7" pageId="5" pageNumber="596">0.9 7074</td>
</tr>
<tr id="40BE34B8FF8C0006D2D10BB0FA04C895" box="[159,1424,1729,1751]" gridrow="8" pageId="5" pageNumber="596">
<th id="036F5DC4FF8C0006D2D10BB0FE8FC895" box="[159,283,1729,1751]" gridcol="0" gridrow="8" pageId="5" pageNumber="596">
<taxonomicName id="398E4D79FF8CFFFCD2D10BB0FE8DC895" box="[159,281,1729,1751]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="prolongatus">
<emphasis id="CCFAEAE8FF8CFFFCD2D10BB0FE8DC895" box="[159,281,1729,1751]" italics="true" pageId="5" pageNumber="596">prolongatus</emphasis>
</taxonomicName>
</th>
<td id="036F5DC4FF8C0006D3190BB0FE4EC895" box="[343,474,1729,1751]" gridcol="1" gridrow="8" pageId="5" pageNumber="596">ant</td>
<td id="036F5DC4FF8C0006D3B90BB0FD12C895" box="[503,646,1729,1751]" gridcol="2" gridrow="8" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D0E90BB0FC8DC895" box="[679,793,1729,1751]" gridcol="3" gridrow="8" pageId="5" pageNumber="596">integ</td>
<td id="036F5DC4FF8C0006D1750BB0FC23C895" box="[827,951,1729,1751]" gridcol="4" gridrow="8" pageId="5" pageNumber="596">abs</td>
<td id="036F5DC4FF8C0006D1A50BB0FA04C895" box="[1003,1424,1729,1751]" gridcol="5" gridrow="8" pageId="5" pageNumber="596">1.1 72</td>
</tr>
</table>
</paragraph>
<paragraph id="FE3136FAFF8CFFFFD2890A6FFF4BCFC5" blockId="5.[151,1436,1822,2027]" lastBlockId="6.[151,1436,151,607]" lastPageId="6" lastPageNumber="597" pageId="5" pageNumber="596">
Some taxonomic characters listed by
<bibRefCitation id="9A1F4B0BFF8CFFFCD03F0A6FFC8EC974" author="Bartsch" box="[625,794,1822,1846]" pageId="5" pageNumber="596" refString="Bartsch, I. (2011) Halacarus socius (Acari: Halacaridae), description of the male and diagnoses of species of the Halacarus actenos group. Zootaxa, 2800, 18 - 40." type="journal article" year="2011">Bartsch (2011)</bibRefCitation>
are scored for
<taxonomicName id="398E4D79FF8CFFFCD1860A6FFB00C974" box="[968,1172,1822,1846]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="omului" status="sp. nov.">
<emphasis id="CCFAEAE8FF8CFFFCD1860A6FFB00C974" box="[968,1172,1822,1846]" italics="true" pageId="5" pageNumber="596">Halacarus omului</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7C95793FF8CFFFCD6D30A6EFB6CC975" box="[1181,1272,1823,1847]" pageId="5" pageNumber="596" rank="species">
<emphasis id="CCFAEAE8FF8CFFFCD6D30A6EFB6CC975" bold="true" box="[1181,1272,1823,1847]" pageId="5" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
and the seven species mentioned above in
<tableCitation id="B30C0341FF8CFFFCD39F0A33FDB0C918" box="[465,548,1858,1882]" captionStart="TABLE 1" captionStartId="5.[151,239,614,638]" captionTargetBox="[159,1420,916,1228]" captionTargetPageId="5" captionText="TABLE 1. Character state distributions in species of the Halacarus actenos group (cf. Bartsch, 2011) that share the coexistence of ds- 5 with gp- 4 and ds- 6 with gp- 5. Abbreviations used here are as follows: abs, absent; ant, anterior; adj, adjacent to; co, cornea, ocular plate reduced; level, at same level; l, long; integ, striated integument; ling, linguiform; m, medium-sized; ov + co, ovate ocular plate with cornea; pres, present; post, posterior; rem, removed; sh, short; slen, slender; spin, spiniform; trian, triangular;?, no data available." httpUri="http://table.plazi.org/id/AAF16672FF8CFFFCD2D90F17FCBFCCBF" pageId="5" pageNumber="596" tableUuid="AAF16672FF8CFFFCD2D90F17FCBFCCBF">Table 1</tableCitation>
.
<taxonomicName id="398E4D79FF8CFFFCD07E0A32FC96C919" box="[560,770,1859,1883]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8CFFFCD07E0A32FC96C919" box="[560,770,1859,1883]" italics="true" pageId="5" pageNumber="596">Halacarus actenos</emphasis>
</taxonomicName>
differs from the present new species in having two setae on trochanter IV instead of one, by the absence of outlying setae on GA in the male, and by the gp-1 placed on the lateral edge of AD (
<bibRefCitation id="9A1F4B0BFF8CFFFCD3C60AFBFDBBC9E0" author="Bartsch" box="[392,559,1930,1954]" pageId="5" pageNumber="596" refString="Bartsch, I. (1980) Halacaridae (Acari) aus der Bucht von Morlaix (Bretagne). Acarologia, 21, 34 - 45." type="journal article" year="1980">Bartsch, 1980</bibRefCitation>
).
<taxonomicName id="398E4D79FF8CFFFCD0080AFAFCB3C9E1" box="[582,807,1931,1955]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="leptopus">
<emphasis id="CCFAEAE8FF8CFFFCD0080AFAFCB3C9E1" box="[582,807,1931,1955]" italics="true" pageId="5" pageNumber="596">Halacarus leptopus</emphasis>
</taxonomicName>
is distinguishable form
<taxonomicName id="398E4D79FF8CFFFCD6040AFDFB2BC9E1" box="[1098,1215,1931,1955]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="omului">
<emphasis id="CCFAEAE8FF8CFFFCD6040AFDFB2BC9E1" box="[1098,1215,1931,1955]" italics="true" pageId="5" pageNumber="596">H. omului</emphasis>
</taxonomicName>
by the absence of ventromedial bipectinate setae on genu II as well as a single such seta on tibiae II (
<bibRefCitation id="9A1F4B0BFF8CFFFCD6240ADFFA84C984" author="Bartsch" box="[1130,1296,1966,1990]" pageId="5" pageNumber="596" refString="Bartsch, I. (2002) Halacarids from the Great Meteor Seamount (Northeastern Atlantic). Description of new species of the genera Agauopsis, Atelopsalis and Halacarus and redescription of H. spiniger Bartsch (Arachnida: Acari: Halacaridae). Mitteilungen aus dem Hamburgischen Zoologische Museum und Institut, 99, 29 - 45." type="journal article" year="2002">Bartsch, 2002</bibRefCitation>
).
<taxonomicName id="398E4D79FF8CFFFCD7680ADFFEB4C9A9" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="discophorus">
<emphasis id="CCFAEAE8FF8CFFFCD7680ADFFEB4C9A9" italics="true" pageId="5" pageNumber="596">Halacarus discophorus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="398E4D79FF8CFFFCD3F60AA5FDBEC9A9" box="[440,554,2003,2027]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="omului">
<emphasis id="CCFAEAE8FF8CFFFCD3F60AA5FDBEC9A9" box="[440,554,2003,2027]" italics="true" pageId="5" pageNumber="596">H. omului</emphasis>
</taxonomicName>
by having a pair of sclerites among gp-4 and gp-
<quantity id="39769B1FFF8CFFFCD62E0AA3FB1FC9A8" box="[1120,1163,2002,2026]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="596" unit="in" value="5.0">5 in</quantity>
the male, the triangular outline of posterior margin of AD, oval OC with the cornea, and four pairs of outlying pgs on GA in the male instead of a single pair of such setae (
<bibRefCitation id="9A1F4B0BFF8FFFFFD00E0DCDFD76CE96" author="Bartsch" box="[576,738,188,212]" pageId="6" pageNumber="597" refString="Bartsch, I. (1993) Halacarus (Halacaridae, Acari) from south-western Australia. In: Wells, F. E., Walker, D. I., Kirkman, H. &amp; Lethbridge, R. (Eds.), The marine fauna and flora of Rottnest Island, Western Australia. Western Australian Museum, Perth, pp. 19 - 43." type="book chapter" year="1993">Bartsch, 1993</bibRefCitation>
).
<taxonomicName id="398E4D79FF8FFFFFD0B80DCDFC6CCE96" box="[758,1016,188,212]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="longiunguis">
<emphasis id="CCFAEAE8FF8FFFFFD0B80DCDFC6CCE96" box="[758,1016,188,212]" italics="true" pageId="6" pageNumber="597">Halacarus longiunguis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="398E4D79FF8FFFFFD6C00DCCFB6BCE96" box="[1166,1279,188,212]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="omului">
<emphasis id="CCFAEAE8FF8FFFFFD6C00DCCFB6BCE96" box="[1166,1279,188,212]" italics="true" pageId="6" pageNumber="597">H. omului</emphasis>
</taxonomicName>
in the lack of corneae, the presence of the accessory process on the lateral claws of tarsi III-IV, and only three ventral setae on tarsi IV (
<bibRefCitation id="9A1F4B0BFF8FFFFFD2B40C75FE08CF5E" author="Bartsch" box="[250,412,260,284]" pageId="6" pageNumber="597" refString="Bartsch, I. (2007) Halacarus longiunguis Police, 1909 (Acari: Halacaridae), a new record a century later, re-description and notes on Mediterranean Halacarus species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14, 393 - 403." type="journal article" year="2007">Bartsch, 2007</bibRefCitation>
).
<taxonomicName id="398E4D79FF8FFFFFD3E00C75FD25CF5E" box="[430,689,260,284]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="magniporus">
<emphasis id="CCFAEAE8FF8FFFFFD3E00C75FD25CF5E" box="[430,689,260,284]" italics="true" pageId="6" pageNumber="597">Halacarus magniporus</emphasis>
</taxonomicName>
is discriminated from
<taxonomicName id="398E4D79FF8FFFFFD1E30C74FB89CF5E" box="[941,1053,260,284]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="omului">
<emphasis id="CCFAEAE8FF8FFFFFD1E30C74FB89CF5E" box="[941,1053,260,284]" italics="true" pageId="6" pageNumber="597">H. omului</emphasis>
</taxonomicName>
by the protruded posterior margin of AD and the strong boat-shaped spine on P3 (
<bibRefCitation id="9A1F4B0BFF8FFFFFD0E40C56FCD5CF7D" author="Krantz" box="[682,833,295,319]" pageId="6" pageNumber="597" refString="Krantz, G. W. (1973) Four new predatory species of Halacaridae (Acari: Prostigmata) from Oregon, with remarks on their distribution in the intertidal mussel habitats (Pelecypoda: Mytilidae). Annals of the Entomological Society of America, 66, 975 - 985." type="journal article" year="1973">Krantz, 1973</bibRefCitation>
).
<taxonomicName id="398E4D79FF8FFFFFD11B0C59FBCDCF02" box="[853,1113,296,320]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="prolongatus">
<emphasis id="CCFAEAE8FF8FFFFFD11B0C59FBCDCF02" box="[853,1113,296,320]" italics="true" pageId="6" pageNumber="597">Halacarus prolongatus</emphasis>
</taxonomicName>
differs from the present new species in the convex posterior margin of AD and the absence of OC and corneae in the ocular area (
<bibRefCitation id="9A1F4B0BFF8FFFFFD7700C3DFF47CFC5" author="Bartsch" pageId="6" pageNumber="597" refString="Bartsch, I. (1996) Halacarid mites (Acari) from the Mid-Atlantic Ridge. New records. Cahiers de Biologie Marine, 37, 159 - 167." type="journal article" year="1996">Bartsch, 1996</bibRefCitation>
).
</paragraph>
<paragraph id="FE3136FAFF8FFFFFD2890CE5FAFCCC1D" blockId="6.[151,1436,151,607]" pageId="6" pageNumber="597">
<bibRefCitation id="9A1F4B0BFF8FFFFFD2890CE5FEFBCFEE" author="Newell" box="[199,367,404,428]" pageId="6" pageNumber="597" refString="Newell, I. M. (1947) A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection, 10 (3), 1 - 232." type="journal article" year="1947">Newell (1947)</bibRefCitation>
assigned
<taxonomicName id="398E4D79FF8FFFFFD3A70CE4FDF2CFEE" box="[489,614,405,428]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8FFFFFD3A70CE4FDF2CFEE" box="[489,614,405,428]" italics="true" pageId="6" pageNumber="597">H. actenos</emphasis>
</taxonomicName>
to Florida,
<collectingCountry id="8699766AFF8FFFFFD0B50CE5FCA2CFEE" box="[763,822,404,428]" name="United States of America" pageId="6" pageNumber="597">USA</collectingCountry>
, based on the study of a male, a female and three deutonymphs. Except for the deutonymph, all of the specimens had lost their distal leg segments before Newells examination.
<taxonomicName id="398E4D79FF8FFFFFD3760CADFE2ACFB6" box="[312,446,476,500]" class="Arachnida" family="Halacaridae" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="family">Halacaridae</taxonomicName>
species are known to be shared by Caribbean Sea and Brazilian coastline (e.g.
<taxonomicName id="398E4D79FF8FFFFFD2D90F71FDE0CC55" authority="Newell, 1947" authorityName="Newell" authorityYear="1947" box="[151,628,511,536]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="CCFAEAE8FF8FFFFFD2D90F71FE54CC5A" box="[151,448,512,536]" italics="true" pageId="6" pageNumber="597">Copidognathus floridensis</emphasis>
(
<bibRefCitation id="9A1F4B0BFF8FFFFFD3810C8EFDFFCC55" author="Newell" box="[463,619,511,535]" pageId="6" pageNumber="597" refString="Newell, I. M. (1947) A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection, 10 (3), 1 - 232." type="journal article" year="1947">Newell, 1947</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="398E4D79FF8FFFFFD0CE0F71FC39CC55" authority="Bartsch, 1984" authorityName="Bartsch" authorityYear="1984" box="[640,941,511,536]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="CCFAEAE8FF8FFFFFD0CE0F71FC9DCC5A" box="[640,777,512,536]" italics="true" pageId="6" pageNumber="597">C. modestus</emphasis>
<bibRefCitation id="9A1F4B0BFF8FFFFFD15F0C8EFC39CC55" author="Bartsch" box="[785,941,511,535]" pageId="6" pageNumber="597" refString="Bartsch, I. (1984) New species of the genus Copidognathus (Halacaridae) from the Caribbean region. Studies on the fauna of Curacao and other Caribbean islands, 67, 1 - 14." type="journal article" year="1984">Bartsch, 1984</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="398E4D79FF8FFFFFD1AA0F71FA12CC55" authority="Bartsch &amp; Iliffe, 1985" authorityName="Bartsch &amp; Iliffe" authorityYear="1985" box="[996,1414,511,536]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="longispinus">
<emphasis id="CCFAEAE8FF8FFFFFD1AA0F71FB12CC5A" box="[996,1158,512,536]" italics="true" pageId="6" pageNumber="597">C. longispinus</emphasis>
<bibRefCitation id="9A1F4B0BFF8FFFFFD6C00C8EFA12CC55" author="Bartsch" box="[1166,1414,511,535]" pageId="6" pageNumber="597" refString="Bartsch, I. &amp; Iliffe, T. M. (1985) The halacarid fauna (Halacaridae, Acari) of Bermudas caves. Stygologia, 1, 300 - 321." type="journal article" year="1985">Bartsch &amp; Iliffe, 1985</bibRefCitation>
</taxonomicName>
<bibRefCitation id="9A1F4B0BFF8FFFFFD2D90F54FE01CC7E" author="Pepato" box="[151,405,548,572]" pageId="6" pageNumber="597" refString="Pepato, A. R. &amp; Tiago, C. G. (2005) New species and new occurrences of Copidognathus (Acari, Halacaridae) from the northern littoral zone of Sao Paulo State (Brazil). Zootaxa, 1083, 1 - 35." type="journal article" year="2005">Pepato &amp; Tiago, 2005</bibRefCitation>
). Nevertheless, similar to other
<taxonomicName id="398E4D79FF8FFFFFD0B00F54FCECCC7E" box="[766,888,549,572]" class="Arachnida" family="Halacaridae" genus="Halacarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="actenos">
<emphasis id="CCFAEAE8FF8FFFFFD0B00F54FCECCC7E" box="[766,888,549,572]" italics="true" pageId="6" pageNumber="597">H. actenos</emphasis>
</taxonomicName>
specimens, Newells material has the trochanter chaetotaxy of 1, 1, 2, 2 (
<bibRefCitation id="9A1F4B0BFF8FFFFFD3E80F36FDD2CC1D" author="Bartsch" box="[422,582,583,607]" pageId="6" pageNumber="597" refString="Bartsch, I. (2011) Halacarus socius (Acari: Halacaridae), description of the male and diagnoses of species of the Halacarus actenos group. Zootaxa, 2800, 18 - 40." type="journal article" year="2011">Bartsch, 2011</bibRefCitation>
), whereas the species described here has a 1, 1, 2, 1 trochanter formula.
</paragraph>
</subSubSection>
</treatment>
</document>