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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.529.6139" ID-GBIF-Dataset="db8513fd-e2ce-4f63-a857-3530566adade" ID-PMC="PMC4668899" ID-Pensoft-Pub="1313-2970-529-1" ID-PubMed="26692792" ID-ZBK="AD4323B4913C447A88A7CE05EC8862A3" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-529-1" ModsDocOrigin="ZooKeys 529" ModsDocTitle="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)" checkinTime="1451243868489" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Budha, Prem B., Naggs, Fred, Backeljau, Thierry &amp; Asami, Takahiro" docDate="2015" docId="CD0F63365E733B4C171EA0667E36C8D1" docLanguage="en" docName="ZooKeys 529: 1-70" docOrigin="ZooKeys 529" docSource="http://dx.doi.org/10.3897/zookeys.529.6139" docTitle="Plectopylis hanleyi Godwin-Austen 1879" docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="E206FFD8FFF5FFFCFFF16833B258FB6B" masterDocTitle="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)" masterLastPageNumber="70" masterPageNumber="1" pageNumber="45" updateTime="1668160861707" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Budha, Prem B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naggs, Fred</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Backeljau, Thierry</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asami, Takahiro</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>529</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>70</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.529.6139</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.529.6139</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-529-1</mods:identifier>
<mods:identifier type="ZBK">AD4323B4913C447A88A7CE05EC8862A3</mods:identifier>
<mods:identifier type="ZooBank">AD4323B4913C447A88A7CE05EC8862A3</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152062799" LSID="urn:lsid:plazi:treatment:CD0F63365E733B4C171EA0667E36C8D1" httpUri="http://treatment.plazi.org/id/CD0F63365E733B4C171EA0667E36C8D1" lastPageId="45" lastPageNumber="45" pageId="44" pageNumber="45">
<subSubSection pageId="44" pageNumber="45" type="multiple">
<paragraph pageId="44" pageNumber="45">Taxon classification Animalia Pulmonata Plectopylidae</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="nomenclature">
<paragraph pageId="44" pageNumber="45">
<taxonomicName authority="Godwin-Austen, 1879 b" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi Godwin-Austen, 1879b</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="reference_group">
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
1879b
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
Godwin-Austen: The Annals and Magazine of Natural History, 5 (4): 164. [
<normalizedToken originalValue="“Sikkim?”">&quot;Sikkim?&quot;</normalizedToken>
].
</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
1897c
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
, - Gude: Science Gossip, 4: 11.
</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
1899a
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
, - Gude: Science Gossip, 5: 240.
</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
1899c
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis (Endothyra) hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi" subGenus="Endothyra">Plectopylis (Endothyra) hanleyi</taxonomicName>
, - Gude: Science Gossip, 6: 148.
</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
1899d
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis (Endothyra) hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi" subGenus="Endothyra">Plectopylis (Endothyra) hanleyi</taxonomicName>
, - Gude: Science Gossip, 6: 175, 176.
</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
1914b
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis (Endothyra) hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi" subGenus="Endothyra">Plectopylis (Endothyra) hanleyi</taxonomicName>
, - Gude: The Fauna of British India
<normalizedToken originalValue="…">...</normalizedToken>
: 73, 77.
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="original description">
<paragraph pageId="44" pageNumber="45">Original description.</paragraph>
<paragraph pageId="44" pageNumber="45">
&quot;Shell sinistral, depressedly conoid, openly umbilicated, probably hirsute when young. Sculpture coarse, irregular, transverse ridges. Colour uniform ochraceous. Spire conoidal; apex blunt, smooth. Suture well marked. Whorls six, close-wound, convex. Aperture semicircular, diagonal; peristome somewhat thickened, white, with a thin callus on the parietal margin, not to the extent of a ridge. Size - major diam. 5.5, minor diam. 5.0, alt. 3.0 millims. Parietal vertical lamina simple; palatal
<normalizedToken originalValue="plicæ">plicae</normalizedToken>
in two rows, four long in front, four short behind, and one basal long. The shell is very distinct; it has somewhat the form of
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis plectostoma" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="plectostoma">Plectopylis plectostoma</taxonomicName>
, but is not so angular on the periphery, while the internal plication is quite different, besides being so very much smaller in size.&quot;
</paragraph>
</subSubSection>
<subSubSection lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="remarks">
<paragraph pageId="44" pageNumber="45">Remarks.</paragraph>
<paragraph lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
In the original description
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="The Annals and Magazine of Natural History" pageId="47" pageNumber="48" pagination="163 - 164" title="Description of two new Species of Plectopylis, a Subgenus of Helicidae." url="10.1080/00222937908679807" volume="5" year="1879 b">Godwin-Austen (1879b)</bibRefCitation>
wrote that the holotype is &quot;in the collection of Mr. Sylvanus Hanley&quot;. In
<normalizedToken originalValue="Godwin-Austens">Godwin-Austen's</normalizedToken>
copy of Gude (1914, page 77), Godwin-Austen has written &quot;In my collection&quot;. The holotype, however, was not found in the collection of the NHM. Only one NHM specimen was found labelled
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
, and this is annotated with a question mark (&quot;Sikkim, Rarhichu, H. H. Godwin-Austen colln.&quot;). However, this specimen is very similar to the type specimen of
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis blanda" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="blanda">Plectopylis blanda</taxonomicName>
, and is not identical with the single shell in
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="The Annals and Magazine of Natural History" pageId="47" pageNumber="48" pagination="163 - 164" title="Description of two new Species of Plectopylis, a Subgenus of Helicidae." url="10.1080/00222937908679807" volume="5" year="1879 b">
<normalizedToken originalValue="Godwin-Austens">Godwin-Austen's</normalizedToken>
(1879b)
</bibRefCitation>
description, because it has only 4.75 whorls (the holotype of
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
has six). Moreover,
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="The Annals and Magazine of Natural History" pageId="47" pageNumber="48" pagination="163 - 164" title="Description of two new Species of Plectopylis, a Subgenus of Helicidae." url="10.1080/00222937908679807" volume="5" year="1879 b">Godwin-Austen (1879b)</bibRefCitation>
described the palatal lamellation, whereas the above mentioned specimen is intact, therefore the inner lamellae and plicae could not be observed. Some parts of
<normalizedToken originalValue="Hanleys">Hanley's</normalizedToken>
collection are housed in the
<pageBreakToken pageId="45" pageNumber="46" start="start">Leeds</pageBreakToken>
Museum and in the Manchester Museum. The former were contacted and confirmed that the holotype was not deposited there. The catalogue of the type specimens of the Manchester Museum (McGhee 2008) did not list
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
. Since the holotype of
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
seems to be lost, and the description is not sufficient to diagnose the species (although it matches with
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella blanda" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="blanda">Endothyrella blanda</taxonomicName>
),
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis hanleyi" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="hanleyi">Plectopylis hanleyi</taxonomicName>
is considered to be a nomen dubium.
</paragraph>
</subSubSection>
</treatment>
</document>