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<document id="B2F395CDB1A611A5CB8CCC27F1772314" ID-CLB-Dataset="6504" ID-DOI="10.11646/zootaxa.4966.4.1" ID-GBIF-Dataset="878bc664-23c5-4b8f-92dc-df86aa07242d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4738667" ID-ZooBank="2B853FE6-18EF-42D8-B43E-587AFA88163E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620208493244" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Zanetti, Adriano" docDate="2021" docId="B43EBA76FF80EE3149AE4A85FEC8F9FD" docLanguage="en" docName="zootaxa.4966.4.1.pdf" docOrigin="Zootaxa 4966 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lesteva osellai Zanetti &amp; Shavrin 2021, sp.n." docType="treatment" docVersion="6" lastPageNumber="420" masterDocId="4807C20EFF92EE2249394A12FF93FF88" masterDocTitle="Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula" masterLastPageNumber="427" masterPageNumber="401" pageNumber="419" updateTime="1698938342712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C8D572CF376ACA85269BC4D4EA5E0C08" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="083732D6F7D17DE154D2D65E00C4AFF1">
<mods:title id="87DD8FC29FC4F0991F94DBCCBCF713ED">Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula</mods:title>
</mods:titleInfo>
<mods:name id="63131B19F89083F039A0267E49B4C1CA" type="personal">
<mods:role id="D4EDC7E7D069D0B40B835085A213B7E5">
<mods:roleTerm id="EE4EB9EABA7B592297BFD9F9BCD482AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="937667FB69652459E61B3D6E8B756DB6">Shavrin, Alexey V.</mods:namePart>
<mods:affiliation id="7D171E0A283BDE7C28C0C9ED38D4C687">Daugavpils University, Institute of Life Sciences and Technology, Coleopterological Research Center, Vienibas 13, LV- 5401. Daugavpils, Latvia.</mods:affiliation>
</mods:name>
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<mods:role id="400E4612A40F1CE11D39BA23DBCC1EB2">
<mods:roleTerm id="3183A25949A828588383AD685392C3DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CFF9B81CF2D4A6E81F97CEE719A4717">Zanetti, Adriano</mods:namePart>
<mods:affiliation id="FB2A618849F3727576633E8C156ED949">Museo Civico di Storia Naturale, Verona, Lungadige Porta Vittoria, 9, I- 37129, Verona, Italy.</mods:affiliation>
</mods:name>
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<mods:title id="C92DB91F1A5A0F83B01C2A126CBE82B1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D98D64C0C27B6A8390D1E449D8A3391C">
<mods:date id="A5E8D585013FFFE12F6456ED5BBC3138">2021</mods:date>
<mods:detail id="DA200388FE07F250921318B6FA4E40B9" type="pubDate">
<mods:number id="CA310356045C74C3DF320AA320FF1AB3">2021-05-05</mods:number>
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<mods:number id="3F4EB5345C746BD356E7B8F01D872A72">4966</mods:number>
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<mods:number id="7FB2FEF2FEBA2CF24D476D08A578FA97">4</mods:number>
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<mods:classification id="FF5D6EC931D08C3835C838B51439D0D8">journal article</mods:classification>
<mods:identifier id="5B63BDC9388B02626259EC964D4E70B2" type="CLB-Dataset">6504</mods:identifier>
<mods:identifier id="A0CDCBC1D3E104D476967FD5772F0299" type="DOI">10.11646/zootaxa.4966.4.1</mods:identifier>
<mods:identifier id="5E0D4D135299D39F00E032DAE3EC0EEB" type="GBIF-Dataset">878bc664-23c5-4b8f-92dc-df86aa07242d</mods:identifier>
<mods:identifier id="C29B73827B2E14BEC41BA3A711597CD3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8D79458695061571E1C7B256D6BF6897" type="Zenodo-Dep">4738667</mods:identifier>
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<treatment id="B43EBA76FF80EE3149AE4A85FEC8F9FD" ID-DOI="http://doi.org/10.5281/zenodo.4738675" ID-GBIF-Taxon="182387733" ID-Zenodo-Dep="4738675" LSID="urn:lsid:plazi:treatment:B43EBA76FF80EE3149AE4A85FEC8F9FD" httpUri="http://treatment.plazi.org/id/B43EBA76FF80EE3149AE4A85FEC8F9FD" lastPageId="19" lastPageNumber="420" pageId="18" pageNumber="419">
<subSubSection id="748D58EBFF80EE3049AE4A85FD40FF39" box="[151,723,151,178]" pageId="18" pageNumber="419" type="nomenclature">
<paragraph id="3C280B60FF80EE3049AE4A85FD40FF39" blockId="18.[151,723,151,213]" box="[151,723,151,178]" pageId="18" pageNumber="419">
<heading id="6760BC0CFF80EE3049AE4A85FD40FF39" bold="true" box="[151,723,151,178]" fontSize="11" level="1" pageId="18" pageNumber="419" reason="1">
<emphasis id="0EE3D772FF80EE3049AE4A85FD40FF39" bold="true" box="[151,723,151,178]" pageId="18" pageNumber="419">
<taxonomicName id="FB9770E3FF80EE3049AE4A85FD18FF39" ID-CoL="8Y5X9" authority="Zanetti &amp; Shavrin" authorityName="Zanetti &amp; Shavrin" authorityYear="2021" box="[151,651,151,178]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="osellai" status="sp. nov.">
<emphasis id="0EE3D772FF80EE3049AE4A85FF61FF39" bold="true" box="[151,242,151,177]" italics="true" pageId="18" pageNumber="419">Lesteva</emphasis>
(s.str.)
<emphasis id="0EE3D772FF80EE3048774A85FE0FFF3A" bold="true" box="[334,412,151,178]" italics="true" pageId="18" pageNumber="419">osellai</emphasis>
Zanetti &amp; Shavrin
</taxonomicName>
,
<taxonomicNameLabel id="15D06A09FF80EE304BA14A85FD40FF39" box="[664,723,151,177]" pageId="18" pageNumber="419" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="748D58EBFF80EE3049AE4AA9FE0DFF5D" box="[151,414,187,213]" pageId="18" pageNumber="419" type="description">
<paragraph id="3C280B60FF80EE3049AE4AA9FE0DFF5D" blockId="18.[151,723,151,213]" box="[151,414,187,213]" pageId="18" pageNumber="419">
(
<figureCitation id="A4AC17E5FF80EE3049A64AA9FF7DFF5D" box="[159,238,187,213]" captionStart="FIGURE 8" captionStartId="7.[151,250,1235,1259]" captionTargetBox="[167,1436,181,1210]" captionTargetId="figure-19@7.[151,1436,181,1210]" captionTargetPageId="7" captionText="FIGURE 8. Distribution of Lesteva in the Iberian Peninsula: L. aterrima (stars), L. lusitana (black circles), L. mariei (white circles), L. mateui (black squares), L. osellai (black triangles)." figureDoi="http://doi.org/10.5281/zenodo.4738677" httpUri="https://zenodo.org/record/4738677/files/figure.png" pageId="18" pageNumber="419">Figs. 8</figureCitation>
,
<figureCitation id="A4AC17E5FF80EE3049C24AA9FED7FF5D" box="[251,324,187,213]" captionStart="FIGURES 29–31" captionStartId="18.[151,264,1683,1707]" captionTargetBox="[160,1436,1002,1656]" captionTargetId="figure-367@18.[151,1436,993,1660]" captionTargetPageId="18" captionText="FIGURES 29–31. Habitus of Lesteva: 29–30–L. osellai (29: male, Spain: Picos de Europa, 30: female, Portugal: Estrela), 31–L. pubescens (Portugal: Estrela). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4738689" httpUri="https://zenodo.org/record/4738689/files/figure.png" pageId="18" pageNumber="419">29–30</figureCitation>
,
<figureCitation id="A4AC17E5FF80EE3048764AA9FE04FF5D" box="[335,407,187,213]" captionStart="FIGURES 32–37" captionStartId="21.[151,264,1954,1978]" captionTargetBox="[207,1377,200,1920]" captionTargetId="figure-19@21.[193,1394,192,1930]" captionTargetPageId="21" captionText="FIGURES 32–37. Aedeagus of Lesteva osellai, paratypes (32–33: Portugal: Estrela, 34–37: Spain: Picos de Europa): 32, 34, 36–parameral view, 33, 35, 37–lateral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4738691" httpUri="https://zenodo.org/record/4738691/files/figure.png" pageId="18" pageNumber="419">32–37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="748D58EBFF80EE3049AE4B11FBA5FC41" pageId="18" pageNumber="419" type="materials_examined">
<paragraph id="3C280B60FF80EE3049AE4B11FBC7FEC9" blockId="18.[151,1437,258,970]" pageId="18" pageNumber="419">
<emphasis id="0EE3D772FF80EE3049AE4B11FE2BFE95" bold="true" box="[151,440,259,285]" pageId="18" pageNumber="419">Type material examined</emphasis>
.
<typeStatus id="E32CB5C2FF80EE3048FF4B11FDA2FE95" box="[454,561,259,285]" pageId="18" pageNumber="419" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2A91C0E9FF80EE304B054B16FDC7FE96" box="[572,596,260,286]" pageId="18" pageNumber="419" type="male">♂</specimenCount>
: ‘
<collectingCountry id="44804BF0FF80EE304B5D4B11FD5BFE95" box="[612,712,259,285]" name="Portugal" pageId="18" pageNumber="419">Portugal</collectingCountry>
| Estrela m 1950 |10.vii.[19]79 | Osella’, ‘
<typeStatus id="E32CB5C2FF80EE304DA34B11FAA0FE95" box="[1178,1331,259,285]" pageId="18" pageNumber="419" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="FB9770E3FF80EE304C7E4B16FA0EFE95" authorityName="Latreille" authorityYear="1797" box="[1351,1437,260,285]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="0EE3D772FF80EE304C7E4B16FA0EFE95" box="[1351,1437,260,285]" italics="true" pageId="18" pageNumber="419">Lesteva</emphasis>
</taxonomicName>
(s.str.) |
<taxonomicName id="FB9770E3FF80EE3049C84B35FEAFFEC9" authority="Shavrin &amp; Zanetti, 2021" authorityName="Shavrin &amp; Zanetti" authorityYear="2021" box="[241,316,295,321]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="osellai" status="sp. nov.">
<emphasis id="0EE3D772FF80EE3049C84B35FEAFFEC9" box="[241,316,295,321]" italics="true" pageId="18" pageNumber="419">osellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15D06A09FF80EE30487B4B3AFEE4FECA" box="[322,375,296,322]" pageId="18" pageNumber="419" rank="species">sp.n.</taxonomicNameLabel>
| Zanetti A. &amp; Shavrin A.V. des. 2020’ &lt;red, printed&gt; (
<collectionCode id="5A8693A5FF80EE304AE74B35FBDBFEC9" box="[990,1096,295,321]" pageId="18" pageNumber="419">MCSNV</collectionCode>
).
</paragraph>
<paragraph id="3C280B60FF80EE3049FE4B5EFBA5FC41" blockId="18.[151,1437,258,970]" pageId="18" pageNumber="419">
<typeStatus id="E32CB5C2FF80EE3049FE4B5EFEABFEEE" box="[199,312,332,358]" pageId="18" pageNumber="419" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2A91C0E9FF80EE30487B4B59FE1DFEEE" box="[322,398,331,358]" pageId="18" pageNumber="419" type="male">23 ♂♂</specimenCount>
(
<specimenCount id="2A91C0E9FF80EE3048A54B59FDC2FEED" box="[412,593,331,357]" pageId="18" pageNumber="419" type="generic">three specimens</specimenCount>
dissected): ‘Portugal | Estrela m 1950 | 10.vii.[19]79 Osella’ &lt;handwritten&gt; (
<specimenCount id="2A91C0E9FF80EE3049994B7DFF76FE02" box="[160,229,367,394]" pageId="18" pageNumber="419" type="male">2 ♂♂</specimenCount>
: cZan,
<specimenCount id="2A91C0E9FF80EE3048014B7DFE19FE02" box="[312,394,367,394]" pageId="18" pageNumber="419" type="male">21 ♂♂</specimenCount>
:
<collectionCode id="5A8693A5FF80EE3048AF4B7DFE6CFE01" box="[406,511,367,393]" pageId="18" pageNumber="419">MCSNV</collectionCode>
);
<specimenCount id="2A91C0E9FF80EE304B2A4B7DFDACFE02" box="[531,575,367,394]" pageId="18" pageNumber="419" type="male">1 ♂</specimenCount>
(dissected): ‘Portugal | Estrela 1950 | m 10-VII-79 | Osella’ &lt;handwritten&gt;, ‘
<taxonomicName id="FB9770E3FF80EE3049A24B81FF64FE25" authorityName="Latreille" authorityYear="1797" box="[155,247,403,429]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">Lesteva</taxonomicName>
[handwritten] | prope [handwritten] | lusitana [handwritten] | det. A. Zanetti 197’ &lt;printed&gt;, ‘
<taxonomicName id="FB9770E3FF80EE304C084B81FA1EFE25" authorityName="Latreille" authorityYear="1797" box="[1329,1421,403,429]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">Lesteva</taxonomicName>
| lusitanica Lohse | Det. Lohse’ &lt;printed&gt;, ‘Coll. | G.A. Lohse | MHNG-1994’ &lt;printed&gt; (
<collectionCode id="5A8693A5FF80EE304DB84BA5FB4EFE59" box="[1153,1245,439,465]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="18" pageNumber="419" type="Museum">MHNG</collectionCode>
);
<specimenCount id="2A91C0E9FF80EE304DC84BA5FA8FFE5A" box="[1265,1308,439,466]" pageId="18" pageNumber="419" type="male">1 ♂</specimenCount>
(dissected; right antennomeres 10-11 missing): ‘Portugal | Estrela
<quantity id="FB6FA685FF80EE304A2D4BC9FCF1FE7E" box="[788,866,475,502]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="18" pageNumber="419" unit="m" value="1950.0">1950m</quantity>
|
<date id="48292DA0FF80EE304A454BC9FB96FE7D" box="[892,1029,475,501]" pageId="18" pageNumber="419" value="1979-07-10">10.VII.1979</date>
| Osella’&lt;handwritten&gt; (cSh);
<specimenCount id="2A91C0E9FF80EE304C564BC9FA0FFE7E" box="[1391,1436,475,502]" pageId="18" pageNumber="419" type="male">1 ♂</specimenCount>
(aedeagus mounted),
<specimenCount id="2A91C0E9FF80EE3048BC4BEDFE43FD92" box="[389,464,511,538]" pageId="18" pageNumber="419" type="female">31 ♀♀</specimenCount>
(
<specimenCount id="2A91C0E9FF80EE3048E74BEDFD17FD91" box="[478,644,511,537]" pageId="18" pageNumber="419" type="generic">two specimens</specimenCount>
without head and pronotum): ‘Portugal | Estrela 1950 | m 10.VII.[19]79 | Osella’&lt;handwritten&gt; (
<specimenCount id="2A91C0E9FF80EE3048964831FE48FDB6" box="[431,475,547,574]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
: cZan,
<specimenCount id="2A91C0E9FF80EE304B084831FD11FDB6" box="[561,642,547,574]" pageId="18" pageNumber="419" type="female">30 ♀♀</specimenCount>
:
<collectionCode id="5A8693A5FF80EE304BA94831FD6AFDB5" box="[656,761,547,573]" pageId="18" pageNumber="419">MCSNV</collectionCode>
);
<specimenCount id="2A91C0E9FF80EE304A344831FCC3FDB6" box="[781,848,547,574]" pageId="18" pageNumber="419" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="2A91C0E9FF80EE304A674831FC18FDB6" box="[862,907,547,574]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
: ‘Espagne | Picos de | Europa’ &lt;handwritten&gt;, ‘Covadonga | Lago Enol |
<quantity id="FB6FA685FF80EE30488A4855FD9AFDEA" box="[435,521,583,610]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="419" unit="m" value="1600.0">1600 m</quantity>
. 3.VI.90’&lt;handwritten&gt; (
<specimenCount id="2A91C0E9FF80EE304A1F4855FCDDFDEA" box="[806,846,583,610]" pageId="18" pageNumber="419" type="male">1 ♂</specimenCount>
,
<specimenCount id="2A91C0E9FF80EE304A614855FC11FDEA" box="[856,898,583,610]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
: cZan;
<specimenCount id="2A91C0E9FF80EE304AEB4855FC6EFDEA" box="[978,1021,583,610]" pageId="18" pageNumber="419" type="male">1 ♂</specimenCount>
: cSh);
<specimenCount id="2A91C0E9FF80EE304D7E4855FBFCFDEA" box="[1095,1135,583,610]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
(pronotum and mediobasal portion of left elytron damaged): ‘Espagne | Picos de Europa | (Oviedo)’ &lt;handwritten&gt;, ‘Covadonga | Lago Enol |
<quantity id="FB6FA685FF80EE3049AE489DFF7AFD22" box="[151,233,655,682]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="419" unit="m" value="1200.0">1200 m</quantity>
2.VI.[19]90’ &lt;handwritten&gt; (cZ);
<specimenCount id="2A91C0E9FF80EE304B5B489DFD18FD22" box="[610,651,655,682]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
: ‘E Cord.Cantabrica |Pto. de Cubilla |
<quantity id="FB6FA685FF80EE304D15489DFBE8FD21" box="[1068,1147,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="419" unit="m" value="1400.0">1400m</quantity>
Spritzmoos | 08.06.[19]91 [P.] Wunderle’ &lt;printed&gt;, ‘
<taxonomicName id="FB9770E3FF80EE3048FE48A1FDB0FD45" authorityName="Latreille" authorityYear="1797" box="[455,547,691,717]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">Lesteva</taxonomicName>
[handwritten] | det.P.Wunderle’ &lt;printed&gt;, ‘?coiffaiti’ &lt;handwritten&gt; (cW);
<specimenCount id="2A91C0E9FF80EE304CB748A1FF3FFD7A" pageId="18" pageNumber="419" type="male">1 ♂</specimenCount>
(aedeagus glued on the same card under the specimen): ‘Pto. de Pajares, | Spain. | G. C. C.’ &lt;printed&gt;, ‘coiffaiti [handwritten] | det.Dr.G.A.Lohse 197’ &lt;printed&gt;, ‘Coll. | G.A. Lohse | MHNG-1994’ &lt;printed&gt; (
<collectionCode id="5A8693A5FF80EE304DC548E9FACBFC9D" box="[1276,1368,763,789]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="18" pageNumber="419" type="Museum">MHNG</collectionCode>
);
<specimenCount id="2A91C0E9FF80EE304C5648E9FA0FFC9E" box="[1391,1436,763,790]" pageId="18" pageNumber="419" type="female">1 ♀</specimenCount>
(without left antennomeres 3-11 and right antenna, without apical part of the abdomen): ‘
<collectingCountry id="44804BF0FF80EE304D65490DFB21FCB1" box="[1116,1202,799,825]" name="Spain" pageId="18" pageNumber="419">SPAIN</collectingCountry>
.Leon, | Pto. de Tarna, |
<quantity id="FB6FA685FF80EE30499A4951FF75FCD5" box="[163,230,835,861]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="419" unit="km" value="17.0">17km</quantity>
.N.Riano, |
<quantity id="FB6FA685FF80EE30485B4951FE21FCD5" box="[354,434,835,861]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="18" pageNumber="419" unit="m" value="1490.0">1490m</quantity>
,
<date id="48292DA0FF80EE3048874951FDA1FCD6" box="[446,562,835,862]" pageId="18" pageNumber="419" value="1965-07-01">1.vii.1965</date>
’ &lt;printed&gt;, ‘Headwaters Rio Nalon.’ &lt;printed&gt;, ‘Among stones and | grit: fast stream.’ &lt;printed&gt;, ‘J.Balfour-Browne. | B.M. 1965-461.’ &lt;printed&gt;, ‘coiffaiti [handwritten] | det.Dr.G.A.Lohse 19’ &lt;printed&gt;, ‘Coll. | G.A. Lohse | MHNG-1994’ &lt;printed&gt; (
<collectionCode id="5A8693A5FF80EE304A254999FCEBFC2D" box="[796,888,907,933]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="18" pageNumber="419" type="Museum">MHNG</collectionCode>
). All
<typeStatus id="E32CB5C2FF80EE304A8A499EFB8DFC2E" box="[947,1054,908,934]" pageId="18" pageNumber="419" type="paratype">paratypes</typeStatus>
with additional red printed labels: ‘
<typeStatus id="E32CB5C2FF80EE3049A249BDFEB8FC41" box="[155,299,943,969]" pageId="18" pageNumber="419" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="FB9770E3FF80EE30480649A2FE06FC41" authorityName="Latreille" authorityYear="1797" box="[319,405,944,969]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="0EE3D772FF80EE30480649A2FE06FC41" box="[319,405,944,969]" italics="true" pageId="18" pageNumber="419">Lesteva</emphasis>
</taxonomicName>
(s.str.) |
<taxonomicName id="FB9770E3FF80EE3048CC49BDFDD3FC41" authority="Shavrin &amp; Zanetti, 2021" authorityName="Shavrin &amp; Zanetti" authorityYear="2021" box="[501,576,943,969]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="osellai" status="sp. nov.">
<emphasis id="0EE3D772FF80EE3048CC49BDFDD3FC41" box="[501,576,943,969]" italics="true" pageId="18" pageNumber="419">osellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15D06A09FF80EE304B7E49A2FDEFFC42" box="[583,636,944,970]" pageId="18" pageNumber="419" rank="species">sp.n.</taxonomicNameLabel>
| Zanetti A. &amp; Shavrin A.V. des. 2020’.
</paragraph>
</subSubSection>
<caption id="68E85BE8FF80EE3049AE4C81FD1FF947" ID-DOI="http://doi.org/10.5281/zenodo.4738689" ID-Zenodo-Dep="4738689" httpUri="https://zenodo.org/record/4738689/files/figure.png" pageId="18" pageNumber="419" startId="18.[151,264,1683,1707]" targetBox="[160,1436,1002,1656]" targetPageId="18">
<paragraph id="3C280B60FF80EE3049AE4C81FD1FF947" blockId="18.[151,1436,1683,1744]" pageId="18" pageNumber="419">
<emphasis id="0EE3D772FF80EE3049AE4C81FEC0F923" bold="true" box="[151,339,1683,1707]" pageId="18" pageNumber="419">FIGURES 29–31.</emphasis>
Habitus of
<taxonomicName id="FB9770E3FF80EE3048FE4C86FD87F923" authorityName="Latreille" authorityYear="1797" box="[455,532,1684,1707]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="0EE3D772FF80EE3048FE4C86FD87F923" box="[455,532,1684,1707]" italics="true" pageId="18" pageNumber="419">Lesteva</emphasis>
</taxonomicName>
: 29–30–
<taxonomicName id="FB9770E3FF80EE304B544C86FD59F923" authorityName="Shavrin &amp; Zanetti" authorityYear="2021" box="[621,714,1683,1707]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="osellai">
<emphasis id="0EE3D772FF80EE304B544C86FD59F923" box="[621,714,1683,1707]" italics="true" pageId="18" pageNumber="419">L. osellai</emphasis>
</taxonomicName>
(29: male, Spain: Picos de Europa, 30: female, Portugal: Estrela), 31–
<taxonomicName id="FB9770E3FF80EE304CB14C86FF6CF947" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="0EE3D772FF80EE304CB14C86FA05F923" box="[1416,1430,1684,1707]" italics="true" pageId="18" pageNumber="419">L</emphasis>
.
<emphasis id="0EE3D772FF80EE3049AE4CA5FF6CF947" box="[151,255,1719,1743]" italics="true" pageId="18" pageNumber="419">pubescens</emphasis>
</taxonomicName>
(Portugal: Estrela). Scale bar: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="748D58EBFF80EE3149FE4CEFFC05FA15" lastPageId="19" lastPageNumber="420" pageId="18" pageNumber="419" type="description">
<paragraph id="3C280B60FF80EE3049FE4CEFFAC5F8E8" blockId="18.[151,1437,1789,2032]" pageId="18" pageNumber="419">
<emphasis id="0EE3D772FF80EE3049FE4CEFFECAF89F" bold="true" box="[199,345,1789,1815]" pageId="18" pageNumber="419">Description.</emphasis>
Measurements (n=50): HW: 0.67–0.80; HL: 0.52–0.60; OL: 0.18–0.25; LT: 0.10–0.12; AL (mean): 2.38; PL: 0.72–0.85; PWMax: 0.82–0.97; PWMin: 0.60–0.70; EL: 1.50–1.80; ESL: 1.35–1.60; EW: 1.27–1.57; MTbL: 1.02; MTrL: 0.47; AW: 1.22–1.57; AedL: 0.77–1.02; FL: 3.15–3.35; TL: 4.10–5.05 (
<typeStatus id="E32CB5C2FF80EE304D984D54FA9AF8E8" box="[1185,1289,1862,1888]" pageId="18" pageNumber="419" type="holotype">holotype</typeStatus>
: 4.10).
</paragraph>
<paragraph id="3C280B60FF80EE3149FE4D78FB74FE25" blockId="18.[151,1437,1789,2032]" lastBlockId="19.[151,1437,151,1653]" lastPageId="19" lastPageNumber="420" pageId="18" pageNumber="419">
Body brown, with paler elytra and apical abdominal tergites in some specimens; antennae and legs yellow-brown; mouthparts and tarsi yellow. Punctation of head dense, moderately coarse and deep, sparser in middle, and slightly finer around eyes (some
<typeStatus id="E32CB5C2FF80EE3048944DA0FD8BF844" box="[429,536,1970,1996]" pageId="18" pageNumber="419" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="44804BF0FF80EE304B674DA0FD0DF844" box="[606,670,1970,1996]" name="Spain" pageId="18" pageNumber="419">Spain</collectingCountry>
with oval impunctate portion between ocelli); punctation of neck as that of head, with impunctate longitudinal band in middle; punctation of pronotum usually coarser and deeper than that of head, sometimes deeper and coarser in medioapical third, finer and sparser in middle and mediobasal third (some
<typeStatus id="E32CB5C2FF81EE3149DA4AA9FEDDFF5D" box="[227,334,187,213]" pageId="19" pageNumber="420" type="paratype">paratypes</typeStatus>
from Estrela with indistinct impunctate longitudinal band); scutellum with several fine punctures; punctation of elytra similar to that on pronotum, but sparser, slightly finer and denser in parascutellar portion and finer and sparser along suture; abdominal tergites with fine, moderately dense, and sometimes indistinct punctation. Body glossy, mostly without microsculpture; basal portion of neck with coarse, dense, isodiametric sculpture, basal portion of scutellum with fine transverse meshes, abdominal tergites with distinct transverse microsculpture on tergites IV–VI and finer, isodiametric microreticulation on tergites VII–VIII. Forebody covered with dense and moderately long semierect setation; abdomen with fine and dense setae. Habitus as in
<figureCitation id="A4AC17E5FF81EE314D624B81FB70FE25" box="[1115,1251,403,429]" captionStart="FIGURES 29–31" captionStartId="18.[151,264,1683,1707]" captionTargetBox="[160,1436,1002,1656]" captionTargetId="figure-367@18.[151,1436,993,1660]" captionTargetPageId="18" captionText="FIGURES 29–31. Habitus of Lesteva: 29–30–L. osellai (29: male, Spain: Picos de Europa, 30: female, Portugal: Estrela), 31–L. pubescens (Portugal: Estrela). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4738689" httpUri="https://zenodo.org/record/4738689/files/figure.png" pageId="19" pageNumber="420">Figs. 29–30</figureCitation>
.
</paragraph>
<paragraph id="3C280B60FF81EE3149FE4BA5FD4EFD45" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">Head 1.2–1.3 times as broad as long, distinctly elevated in middle; middle portion separated from lateral portions by deep longitudinal depression, markedly deeper between middle length of eyes; eyes large and convex, about twice as long as temples (lateral view), gradually narrowing toward neck; ocelli large, distance between ocelli subequal to distance between ocellus and posterior margin of eye. Antennae moderately short, reaching basal portion of elytra when reclined, with slightly thickened and moderately short antennomeres 4–10; basal antennomere about twice as long as wide, 2 about half as wide as basal antennomere, 3 slightly longer than 2, 4 slightly broader and longer than 3, 5–7 slightly longer than 4, 8–10 slightly shorter and broader than 7, apical antennomere about 1.3 times as long as 10, in apical third sharply narrowed toward apex.</paragraph>
<paragraph id="3C280B60FF81EE3149FE48C5FB91FCB1" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
Pronotum convex, slightly broader than long and head, widest slightly before middle, from their widely rounded lateral margins gradually narrowed toward subparallel latero-basal margins, with obtuse hind angles; mediobasal third of some
<typeStatus id="E32CB5C2FF81EE31480D490DFE0CFCB1" box="[308,415,799,825]" pageId="19" pageNumber="420" type="paratype">paratypes</typeStatus>
with distinct, transverse, moderately deep impression.
</paragraph>
<paragraph id="3C280B60FF81EE3149FE4951FA01FC09" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
Elytra distinctly convex, slightly longer than broad, twice as long as pronotum, slightly widened toward hind angles; apical margins widely rounded; mediobasal surface of elytra in some
<typeStatus id="E32CB5C2FF81EE314AC14975FBF0FC09" box="[1016,1123,871,897]" pageId="19" pageNumber="420" type="paratype">paratypes</typeStatus>
with indistinct depression.
</paragraph>
<paragraph id="3C280B60FF81EE3149FE4999FD60FC41" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">Metatarsus more than twice as long as metatibia, with basal metatarsomere about as long as combined length of tarsomeres 2–3 and about as long as apical tarsomere.</paragraph>
<paragraph id="3C280B60FF81EE3149FE49C1FDE2FBBD" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">Abdomen slightly narrower or about as broad as elytra, with paired tomentose spots in the middle of abdominal tergites IV and V, more transverse in tergite IV, smaller and round in tergite V, with distinct narrow palisade fringe on apical margin of abdominal tergite VII.</paragraph>
<paragraph id="3C280B60FF81EE3149FE4E2DFAEDFB4D" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
Male (
<figureCitation id="A4AC17E5FF81EE3148284E2DFEF4FBD1" box="[273,359,1087,1113]" captionStart="FIGURES 29–31" captionStartId="18.[151,264,1683,1707]" captionTargetBox="[160,1436,1002,1656]" captionTargetId="figure-367@18.[151,1436,993,1660]" captionTargetPageId="18" captionText="FIGURES 29–31. Habitus of Lesteva: 29–30–L. osellai (29: male, Spain: Picos de Europa, 30: female, Portugal: Estrela), 31–L. pubescens (Portugal: Estrela). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4738689" httpUri="https://zenodo.org/record/4738689/files/figure.png" pageId="19" pageNumber="420">Fig. 29</figureCitation>
). Apical margins of abdominal tergite VIII slightly and sternite VIII deeply concave. Aedeagus with wide basal portion, gradually narrowed toward moderately narrow, long median lobe, in apical third sharply narrowed toward subacute apex; parameres narrow, slightly reaching apex of median lobe, with four short apical setae; internal sac moderately wide and long (
<figureCitation id="A4AC17E5FF81EE314BA14EB9FCD8FB4E" box="[664,843,1195,1222]" captionStart="FIGURES 32–37" captionStartId="21.[151,264,1954,1978]" captionTargetBox="[207,1377,200,1920]" captionTargetId="figure-19@21.[193,1394,192,1930]" captionTargetPageId="21" captionText="FIGURES 32–37. Aedeagus of Lesteva osellai, paratypes (32–33: Portugal: Estrela, 34–37: Spain: Picos de Europa): 32, 34, 36–parameral view, 33, 35, 37–lateral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4738691" httpUri="https://zenodo.org/record/4738691/files/figure.png" pageId="19" pageNumber="420">Figs. 32, 34, 36</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="A4AC17E5FF81EE314DF04EB9FAE9FB4D" box="[1225,1402,1194,1221]" captionStart="FIGURES 32–37" captionStartId="21.[151,264,1954,1978]" captionTargetBox="[207,1377,200,1920]" captionTargetId="figure-19@21.[193,1394,192,1930]" captionTargetPageId="21" captionText="FIGURES 32–37. Aedeagus of Lesteva osellai, paratypes (32–33: Portugal: Estrela, 34–37: Spain: Picos de Europa): 32, 34, 36–parameral view, 33, 35, 37–lateral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4738691" httpUri="https://zenodo.org/record/4738691/files/figure.png" pageId="19" pageNumber="420">Figs. 33, 35, 37</figureCitation>
.
</paragraph>
<paragraph id="3C280B60FF81EE3149FE4EDDFB16FB61" blockId="19.[151,1437,151,1653]" box="[199,1157,1231,1257]" pageId="19" pageNumber="420">
Female (
<figureCitation id="A4AC17E5FF81EE3148104EDDFEEEFB61" box="[297,381,1231,1257]" captionStart="FIGURES 29–31" captionStartId="18.[151,264,1683,1707]" captionTargetBox="[160,1436,1002,1656]" captionTargetId="figure-367@18.[151,1436,993,1660]" captionTargetPageId="18" captionText="FIGURES 29–31. Habitus of Lesteva: 29–30–L. osellai (29: male, Spain: Picos de Europa, 30: female, Portugal: Estrela), 31–L. pubescens (Portugal: Estrela). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4738689" httpUri="https://zenodo.org/record/4738689/files/figure.png" pageId="19" pageNumber="420">Fig. 30</figureCitation>
). Apical margins of abdominal tergite VIII and sternite VIII rounded.
</paragraph>
<paragraph id="3C280B60FF81EE3149FE4EE0FC05FA15" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
<emphasis id="0EE3D772FF81EE3149FE4EE0FE23FA85" bold="true" box="[199,432,1266,1293]" pageId="19" pageNumber="420">Comparative notes.</emphasis>
Based on the general body shape, coloration, robust antennomeres and moderately sparse and large punctation of the pronotum,
<taxonomicName id="FB9770E3FF81EE314B724F05FD27FAB9" authority="Shavrin &amp; Zanetti, 2021" authorityName="Shavrin &amp; Zanetti" authorityYear="2021" box="[587,692,1303,1329]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="osellai" status="sp. nov.">
<emphasis id="0EE3D772FF81EE314B724F05FDC8FAB8" box="[587,603,1303,1328]" italics="true" pageId="19" pageNumber="420">L</emphasis>
.
<emphasis id="0EE3D772FF81EE314B504F05FD27FAB9" box="[617,692,1303,1329]" italics="true" pageId="19" pageNumber="420">osellai</emphasis>
</taxonomicName>
<emphasis id="0EE3D772FF81EE314B854F05FD60FAB9" bold="true" box="[700,755,1303,1329]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="15D06A09FF81EE314B854F05FD60FAB9" box="[700,755,1303,1329]" pageId="19" pageNumber="420" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="FB9770E3FF81EE314AB34F05FB90FAB9" authorityName="Lohse" authorityYear="1955" box="[906,1027,1303,1329]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="lusitana">
<emphasis id="0EE3D772FF81EE314AB34F05FC09FAB8" box="[906,922,1303,1328]" italics="true" pageId="19" pageNumber="420">L</emphasis>
.
<emphasis id="0EE3D772FF81EE314A904F05FB90FAB9" box="[937,1027,1303,1329]" italics="true" pageId="19" pageNumber="420">lusitana</emphasis>
</taxonomicName>
,
<taxonomicName id="FB9770E3FF81EE314D2B4F05FB59FAB9" authorityName=": Shavrin" authorityYear="2014" box="[1042,1226,1303,1329]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="longoelytrata">
<emphasis id="0EE3D772FF81EE314D2B4F05FBB1FAB8" box="[1042,1058,1303,1328]" italics="true" pageId="19" pageNumber="420">L</emphasis>
.
<emphasis id="0EE3D772FF81EE314D094F05FB59FAB9" box="[1072,1226,1303,1329]" italics="true" pageId="19" pageNumber="420">longoelytrata</emphasis>
</taxonomicName>
<emphasis id="0EE3D772FF81EE314DF34F05FB5DFAB9" box="[1226,1230,1303,1329]" italics="true" pageId="19" pageNumber="420">,</emphasis>
and
<taxonomicName id="FB9770E3FF81EE314C3E4F05FA06FAB9" authorityName=": Zanetti" authorityYear="1987" box="[1287,1429,1303,1329]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="0EE3D772FF81EE314C3E4F05FA84FAB8" box="[1287,1303,1303,1328]" italics="true" pageId="19" pageNumber="420">L</emphasis>
.
<emphasis id="0EE3D772FF81EE314C1C4F05FA06FAB9" box="[1317,1429,1303,1329]" italics="true" pageId="19" pageNumber="420">monticola</emphasis>
</taxonomicName>
. Regarding body size, the shape of the apical margins of the elytra, the length of the apical metatarsomere (not longer than basal metatarsomere), this species is similar to
<taxonomicName id="FB9770E3FF81EE314BE34F4DFCC2FAF1" authorityName="Lohse" authorityYear="1955" box="[730,849,1375,1401]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="lusitana">
<emphasis id="0EE3D772FF81EE314BE34F4DFD79FAF0" box="[730,746,1375,1400]" italics="true" pageId="19" pageNumber="420">L</emphasis>
.
<emphasis id="0EE3D772FF81EE314BCE4F4DFCC2FAF1" box="[759,849,1375,1401]" italics="true" pageId="19" pageNumber="420">lusitana</emphasis>
</taxonomicName>
and
<taxonomicName id="FB9770E3FF81EE314ABF4F4DFB87FAF1" authorityName=": Zanetti" authorityYear="1987" box="[902,1044,1375,1401]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="0EE3D772FF81EE314ABF4F4DFC05FAF0" box="[902,918,1375,1400]" italics="true" pageId="19" pageNumber="420">L</emphasis>
.
<emphasis id="0EE3D772FF81EE314A9D4F4DFB87FAF1" box="[932,1044,1375,1401]" italics="true" pageId="19" pageNumber="420">monticola</emphasis>
</taxonomicName>
. It differs from them by the shorter antennomeres 8–10, a narrower median lobe, and shorter parameres.
</paragraph>
</subSubSection>
<subSubSection id="748D58EBFF81EE3149FE4FB4FEA3F9A5" pageId="19" pageNumber="420" type="distribution">
<paragraph id="3C280B60FF81EE3149FE4FB4FE28FA6D" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
<emphasis id="0EE3D772FF81EE3149FE4FB4FEF2FA48" bold="true" box="[199,353,1446,1472]" pageId="19" pageNumber="420">Distribution.</emphasis>
The species is known from several localities in Serra da Estrela,
<collectingCountry id="44804BF0FF81EE314D6C4FB5FB24FA49" box="[1109,1207,1447,1473]" name="Portugal" pageId="19" pageNumber="420">Portugal</collectingCountry>
, and the Cantabrian mountains,
<collectingCountry id="44804BF0FF81EE3148234FD9FEC9FA6D" box="[282,346,1483,1509]" name="Spain" pageId="19" pageNumber="420">Spain</collectingCountry>
(
<figureCitation id="A4AC17E5FF81EE3148504FD9FE3EFA6D" box="[361,429,1483,1509]" captionStart="FIGURE 8" captionStartId="7.[151,250,1235,1259]" captionTargetBox="[167,1436,181,1210]" captionTargetId="figure-19@7.[151,1436,181,1210]" captionTargetPageId="7" captionText="FIGURE 8. Distribution of Lesteva in the Iberian Peninsula: L. aterrima (stars), L. lusitana (black circles), L. mariei (white circles), L. mateui (black squares), L. osellai (black triangles)." figureDoi="http://doi.org/10.5281/zenodo.4738677" httpUri="https://zenodo.org/record/4738677/files/figure.png" pageId="19" pageNumber="420">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="3C280B60FF81EE3149FE4FFDFEA3F9A5" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
<emphasis id="0EE3D772FF81EE3149FE4FFDFE15F980" bold="true" box="[199,390,1518,1545]" pageId="19" pageNumber="420">Natural history.</emphasis>
Specimens were collected at elevations from
<quantity id="FB6FA685FF81EE314AAB4FFDFBD5F981" box="[914,1094,1518,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.95" metricValueMin="1.2" pageId="19" pageNumber="420" unit="m" value="1575.0" valueMax="1950.0" valueMin="1200.0">1200 to 1950 m</quantity>
a.s.l. Detailed ecological data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="748D58EBFF81EE3149FE4C25FEC8F9FD" pageId="19" pageNumber="420" type="etymology">
<paragraph id="3C280B60FF81EE3149FE4C25FEC8F9FD" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="420">
<emphasis id="0EE3D772FF81EE3149FE4C25FEDDF9D9" bold="true" box="[199,334,1591,1617]" pageId="19" pageNumber="420">Etymology.</emphasis>
Patronymic, the species is named to honour our colleague, G. Osella (Verona,
<collectingCountry id="44804BF0FF81EE314DF64C25FA95F9DA" box="[1231,1286,1591,1618]" name="Italy" pageId="19" pageNumber="420">Italy</collectingCountry>
), collector of the
<typeStatus id="E32CB5C2FF81EE3149F94C49FF63F9FD" box="[192,240,1627,1653]" pageId="19" pageNumber="420">type</typeStatus>
material.
</paragraph>
</subSubSection>
</treatment>
</document>
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