treatments-xml/data/A5/41/5E/A5415E5FF402FFF39857FA44FC95FE49.xml

<document id="4C4DAD75C0C82C6A13D921E899D1BF3C" ID-CLB-Dataset="199795" ID-DOI="10.5852/ejt.2022.835.1909" ID-GBIF-Dataset="31a2a3ea-9d40-4178-8629-da4b275f37dc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7033400" ID-ZooBank="8B8432B1-C714-4179-8687-66902F4CBF53" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661823783728" checkinUser="felipe" docAuthor="Groll, Elisa Von &amp; Lopes-Andrade, Cristiano" docDate="2022" docId="A5415E5FF402FFF39857FA44FC95FE49" docLanguage="en" docName="ejt-835_1-97.pdf" docOrigin="European Journal of Taxonomy 835 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.835.1909" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cyparium lescheni von Groll &amp; Lopes-Andrade 2022, sp. nov." docType="treatment" docUuid="8A653AB2-6DEF-4FA6-A885-438BDAF11D5F" docUuidSource="ZooBank" docVersion="11" lastPageNumber="36" masterDocId="59782627F41AFFD09A20FFFAFFCCFFB2" masterDocTitle="Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species" masterLastPageNumber="97" masterPageNumber="1" pageNumber="25" updateTime="1699374449524" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5FF4C8079248DD3488C999410FB9DDD7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C0A4C6D8EDDAB3CD337F8904EF29A594">
<mods:title id="CA24A3007476D1B2963D4E7BC450330A">Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species</mods:title>
</mods:titleInfo>
<mods:name id="41E248A47DB39B0D5D0F0BE8D4D233B8" type="personal">
<mods:role id="0B6A9D21B51F5670076A4E57E53F95E6">
<mods:roleTerm id="B3AB1DA160DFAB50B1B4E1899D19E9F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3DE4D6C5AF21027B13ABC227EE0D40DB">Groll, Elisa Von</mods:namePart>
<mods:nameIdentifier id="73272F25C83A46E7C2BF2BBD0757431F" type="LSID">DC564233-82A9-4939-8B7A-F02C994BBA29</mods:nameIdentifier>
<mods:affiliation id="53C49FDA8B7B2881C9C53542CA84D220">Programa de Pós-Graduação em Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil. &amp; Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil.</mods:affiliation>
<mods:nameIdentifier id="3010FF52CD1ECDF3B1AA4C0A6B01C5AE" type="email">elisavgroll@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5C547422A3C3D58128FE2283DE273E74" type="personal">
<mods:role id="69E57E9CD467A06C9280551105BE9556">
<mods:roleTerm id="4D9CC949B3E1277BE5111CDC0B22270B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E24499ABE52A02A871A2E586357D1A7A">Lopes-Andrade, Cristiano</mods:namePart>
<mods:nameIdentifier id="C3AACBCC3414E107560E7FF7A1C9C7AC" type="LSID">525ABCB9-3334-407E-9084-F0784968D290</mods:nameIdentifier>
<mods:affiliation id="95643C8D6C3A69559DF6F9C6A78F944A">Programa de Pós-Graduação em Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil. &amp; Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil.</mods:affiliation>
<mods:nameIdentifier id="668367B674335EEB5511D5F56191F20E" type="email">ciidae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="09B8C83A2570F1DF187880F5C5CBAA91">text</mods:typeOfResource>
<mods:relatedItem id="E7B23A325138549E06EE5F38F493CB5D" type="host">
<mods:titleInfo id="07AE377523B40C504A2EC2B9EAAC5439">
<mods:title id="C88248ED9E08F4C1D5B269478E9540FC">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="0A9E6FF330CE9BAA6184E283257DA783">
<mods:date id="59D39C55988B6A901A1061C5988E6975">2022</mods:date>
<mods:detail id="FBCF7CE4FDF311A4F8040D73978268F8" type="pubDate">
<mods:number id="A69D199D03FAC93E35AFD27691B6469E">2022-08-24</mods:number>
</mods:detail>
<mods:detail id="08ACAFC8DE224AEFE17B1FC8B3296E02" type="volume">
<mods:number id="3A1039A7C17AD64D1F61B3628F1A9DF7">835</mods:number>
</mods:detail>
<mods:detail id="A516E498033A30DE4580CEEBE63C2845" type="issue">
<mods:number id="F25879B1FFF853D7A234C00302ACDC38">1</mods:number>
</mods:detail>
<mods:extent id="0ADE664776DB658D6455A506781AEEA6" unit="page">
<mods:start id="510BA2E35C4CAFCCD072BC8FB2E23910">1</mods:start>
<mods:end id="C75D309548097F78A83E5B3AA4F4D909">97</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="FA3545A98A9B3D246B7ED84E0BD2901C">
<mods:url id="28398C32A68F9986E42A5F5D3F0743BE">http://dx.doi.org/10.5852/ejt.2022.835.1909</mods:url>
</mods:location>
<mods:classification id="530805E661FE5C0041959A0732D1A416">journal article</mods:classification>
<mods:identifier id="9844423FE52E3C89E0CC92585EE64FA8" type="CLB-Dataset">199795</mods:identifier>
<mods:identifier id="713694F5637335C8C27B0388859BC12B" type="DOI">10.5852/ejt.2022.835.1909</mods:identifier>
<mods:identifier id="3D89E4189B89E72D99158C0CEBBFF414" type="GBIF-Dataset">31a2a3ea-9d40-4178-8629-da4b275f37dc</mods:identifier>
<mods:identifier id="CBE97687150CF2384C01D9346DFEF398" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="6976E47960D7B967C70EF1D604A47746" type="Zenodo-Dep">7033400</mods:identifier>
<mods:identifier id="5F62ED4DCA608A5FEF743AB3B3034766" type="ZooBank">8B8432B1-C714-4179-8687-66902F4CBF53</mods:identifier>
</mods:mods>
<treatment id="A5415E5FF402FFF39857FA44FC95FE49" ID-DOI="http://doi.org/10.5281/zenodo.7392017" ID-GBIF-Taxon="205699045" ID-Zenodo-Dep="7392017" LSID="urn:lsid:zoobank.org:act:8A653AB2-6DEF-4FA6-A885-438BDAF11D5F" httpUri="http://treatment.plazi.org/id/A5415E5FF402FFF39857FA44FC95FE49" lastPageId="35" lastPageNumber="36" pageId="24" pageNumber="25">
<subSubSection id="65F2BCC2F402FFC89857FA44FC71FA6B" box="[631,957,1470,1497]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="2D57EF49F402FFC89857FA44FC71FA6B" blockId="24.[358,1229,1470,1567]" box="[631,957,1470,1497]" pageId="24" pageNumber="25">
<heading id="761F5825F402FFC89857FA44FC71FA6B" box="[631,957,1470,1497]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="EAE894CAF402FFC89857FA44FC96FA6B" ID-CoL="9GK3Q" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[631,858,1470,1497]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF402FFC89857FA44FC96FA6B" bold="true" box="[631,858,1470,1497]" italics="true" pageId="24" pageNumber="25">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F402FFC89942FA45FC71FA6B" box="[866,957,1471,1497]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F402FFC89B46FA1BFBE7F9AD" pageId="24" pageNumber="25" type="description">
<paragraph id="2D57EF49F402FFC89B46FA1BFB01FA4E" blockId="24.[358,1229,1470,1567]" box="[358,1229,1505,1532]" pageId="24" pageNumber="25">
<uri id="5979E34BF402FFC89B46FA1BFB01FA4E" box="[358,1229,1505,1532]" pageId="24" pageNumber="25">
urn:lsid:zoobank.org:act: 
<uuid id="594ED59CF402FFC898A6FA1BFB01FA4E" box="[646,1229,1505,1532]" pageId="24" pageNumber="25">8A653AB2-6DEF-4FA6-A885-438BDAF11D5F</uuid>
</uri>
</paragraph>
<paragraph id="2D57EF49F402FFC8982AF9FFFBE7F9AD" blockId="24.[358,1229,1470,1567]" box="[522,1067,1541,1567]" pageId="24" pageNumber="25">
<figureCitation id="B5D3F3CCF402FFC8982AF9FFFD99F9AD" box="[522,597,1541,1567]" captionStart="Fig" captionStartId="7.[189,232,1799,1825]" captionTargetBox="[193,1384,280,1696]" captionTargetId="figure-11@7.[189,1398,265,1704]" captionTargetPageId="7" captionText="Fig. 4. Cyparium Erichson, 1845. Neotropical species. Illustrations based on the original descriptions. *Illustrations made based on photographs, specimens, or illustrations made by other authors; **size not provided in original description. Model: Cyparium collare Pic, 1920. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033408" httpUri="https://zenodo.org/record/7033408/files/figure.png" pageId="24" pageNumber="25">Figs 4</figureCitation>
, 
<figureCitation id="B5D3F3CCF402FFC89842F9FFFD61F9AD" box="[610,685,1541,1567]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId-0="26.[189,232,1924,1950]" captionStartId-1="27.[189,232,1886,1912]" captionStartId-2="28.[189,232,1797,1823]" captionStartId-3="29.[189,232,1742,1768]" captionStartId-4="30.[189,232,1761,1787]" captionStartId-5="31.[189,232,1725,1751]" captionStartId-6="32.[189,232,1820,1846]" captionStartId-7="34.[189,232,1805,1831]" captionTargetBox-0="[205,1394,748,1886]" captionTargetBox-1="[189,1398,964,1848]" captionTargetBox-2="[189,1394,273,1750]" captionTargetBox-3="[189,1398,282,1702]" captionTargetBox-4="[189,1393,275,1713]" captionTargetBox-5="[189,1398,265,1657]" captionTargetBox-6="[240,1398,267,1761]" captionTargetBox-7="[208,1398,300,1761]" captionTargetId-0="figure-153@26.[189,1398,720,1888]" captionTargetId-1="figure-226@27.[189,1398,964,1850]" captionTargetId-2="figure-17@28.[189,1398,267,1761]" captionTargetId-3="figure-12@29.[189,1398,270,1705]" captionTargetId-4="figure-17@30.[189,1399,265,1713]" captionTargetId-5="figure-11@31.[189,1398,265,1663]" captionTargetId-6="figure-16@32.[189,1398,267,1761]" captionTargetId-7="figure-17@34.[189,1398,268,1762]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionTargetPageId-5="31" captionTargetPageId-6="32" captionTargetPageId-7="34" captionText-0="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." captionText-1="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." captionText-2="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." captionText-3="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." captionText-4="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." captionText-5="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." captionText-6="Fig. 20. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–C. Aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D–E. Internal sac. D. Frontal view. E. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm." captionText-7="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7033428" figureDoi-1="http://doi.org/10.5281/zenodo.7033432" figureDoi-2="http://doi.org/10.5281/zenodo.7033436" figureDoi-3="http://doi.org/10.5281/zenodo.7033438" figureDoi-4="http://doi.org/10.5281/zenodo.7033440" figureDoi-5="http://doi.org/10.5281/zenodo.7033442" figureDoi-6="http://doi.org/10.5281/zenodo.7033444" figureDoi-7="http://doi.org/10.5281/zenodo.7033447" httpUri-0="https://zenodo.org/record/7033428/files/figure.png" httpUri-1="https://zenodo.org/record/7033432/files/figure.png" httpUri-2="https://zenodo.org/record/7033436/files/figure.png" httpUri-3="https://zenodo.org/record/7033438/files/figure.png" httpUri-4="https://zenodo.org/record/7033440/files/figure.png" httpUri-5="https://zenodo.org/record/7033442/files/figure.png" httpUri-6="https://zenodo.org/record/7033444/files/figure.png" httpUri-7="https://zenodo.org/record/7033447/files/figure.png" pageId="24" pageNumber="25">14–21</figureCitation>
, 
<figureCitation id="B5D3F3CCF402FFC8989AF9FFFD17F9AD" box="[698,731,1541,1567]" captionStart="Fig" captionStartId="65.[189,232,1952,1978]" captionTargetBox="[189,1398,1082,1910]" captionTargetId="figure-297@65.[189,1398,1082,1910]" captionTargetPageId="65" captionText="Fig. 46. A. Distribution map of species of Cyparium Erichson, 1845, South America. B. Mata da Biologia. C. Mata do Paraíso." figureDoi="http://doi.org/10.5281/zenodo.7033501" httpUri="https://zenodo.org/record/7033501/files/figure.png" pageId="24" pageNumber="25">46</figureCitation>
; Supp. file 1B, Supp. file 2A
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F402FFC89A9DF9BCFD33F8AE" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="2D57EF49F402FFC89A9DF9BCFEFAF9D2" blockId="24.[189,310,1606,1632]" box="[189,310,1606,1632]" pageId="24" pageNumber="25">
<heading id="761F5825F402FFC89A9DF9BCFEFAF9D2" bold="true" box="[189,310,1606,1632]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="1F9C335BF402FFC89A9DF9BCFEFAF9D2" bold="true" box="[189,310,1606,1632]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DF98FFD33F8AE" blockId="24.[188,1399,1653,1820]" pageId="24" pageNumber="25">
TL: 
<quantity id="EA1042ACF402FFC89ACFF98FFE55F93D" box="[239,409,1653,1680]" metricMagnitude="-3" metricUnit="m" metricValue="2.16" metricValueMax="2.25" metricValueMin="2.07" pageId="24" pageNumber="25" unit="mm" value="2.16" valueMax="2.25" valueMin="2.07">2.07–2.25 mm</quantity>
in males and 
<quantity id="EA1042ACF402FFC89817F98FFD2CF93D" box="[567,736,1653,1680]" metricMagnitude="-3" metricUnit="m" metricValue="2.225" metricValueMax="2.35" metricValueMin="2.1" pageId="24" pageNumber="25" unit="mm" value="2.225" valueMax="2.35" valueMin="2.1">2.10–2.35 mm</quantity>
in females. Brown (
<figureCitation id="B5D3F3CCF402FFC899EBF98FFBFBF93D" box="[971,1079,1653,1679]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="24" pageNumber="25">Fig. 14A</figureCitation>
). Hypomeron, metaventrite and intercoxal plates with strigulate microsculpture. Metaventrite coarsely punctate above intercoxal plates (
<figureCitation id="B5D3F3CCF402FFC89B30F941FEB2F967" box="[272,382,1723,1749]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="24" pageNumber="25">Figs 14B</figureCitation>
, 
<figureCitation id="B5D3F3CCF402FFC89BA9F941FE73F967" box="[393,447,1723,1749]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="24" pageNumber="25">17D</figureCitation>
). Tergite VIII in males triangular (
<figureCitation id="B5D3F3CCF402FFC8996CF941FC74F967" box="[844,952,1723,1749]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="24" pageNumber="25">Fig. 19D</figureCitation>
). Openings of aedeagus in dorsal view forming an acute angle (
<figureCitation id="B5D3F3CCF402FFC89BC3F924FD9DF94B" box="[483,593,1758,1785]" captionStart="Fig" captionStartId="32.[189,232,1820,1846]" captionTargetBox="[240,1398,267,1761]" captionTargetId="figure-16@32.[189,1398,267,1761]" captionTargetPageId="32" captionText="Fig. 20. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–C. Aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D–E. Internal sac. D. Frontal view. E. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033444" httpUri="https://zenodo.org/record/7033444/files/figure.png" pageId="24" pageNumber="25">Fig. 20C</figureCitation>
); internal sac with drop-like sclerites (
<figureCitation id="B5D3F3CCF402FFC89E3FF924FB7DF94B" box="[1055,1201,1758,1785]" captionStart="Fig" captionStartId="32.[189,232,1820,1846]" captionTargetBox="[240,1398,267,1761]" captionTargetId="figure-16@32.[189,1398,267,1761]" captionTargetPageId="32" captionText="Fig. 20. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–C. Aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D–E. Internal sac. D. Frontal view. E. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033444" httpUri="https://zenodo.org/record/7033444/files/figure.png" pageId="24" pageNumber="25">Fig. 20D–E</figureCitation>
). Tergite VIII of females with a posterior invagination (
<figureCitation id="B5D3F3CCF402FFC898A4F8F8FD3DF8AE" box="[644,753,1794,1820]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="24" pageNumber="25">Fig. 21A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F402FFC89A9DF8B9FC14F81D" pageId="24" pageNumber="25" type="etymology">
<paragraph id="2D57EF49F402FFC89A9DF8B9FE88F8EF" blockId="24.[189,324,1859,1885]" box="[189,324,1859,1885]" pageId="24" pageNumber="25">
<heading id="761F5825F402FFC89A9DF8B9FE88F8EF" bold="true" box="[189,324,1859,1885]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="1F9C335BF402FFC89A9DF8B9FE88F8EF" bold="true" box="[189,324,1859,1885]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DF888FC14F81D" blockId="24.[189,1399,1906,1967]" pageId="24" pageNumber="25">
In homage to Dr Richard A.B. Leschen (
<collectingCountry id="55FFAFD9F402FFC898B6F888FCF9F83E" box="[662,821,1906,1932]" name="New Zealand" pageId="24" pageNumber="25">New Zealand</collectingCountry>
Arthropod Collection) for his great contributions to the systematics of 
<taxonomicName id="EAE894CAF402FFC89B94F86FFDF4F81D" authorityName="Linnaeus" authorityYear="1758" box="[436,568,1941,1967]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
, especially regarding 
<taxonomicName id="EAE894CAF402FFC89918F86FFC1FF81D" authorityName="Latreille" authorityYear="1806" box="[824,979,1941,1967]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subFamily" subFamily="Scaphidiinae">Scaphidiinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F403FFC99A9DFEF2FEB2FBC3" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="2D57EF49F403FFC99A9DFEF2FE66FE90" blockId="25.[189,426,264,290]" box="[189,426,264,290]" pageId="25" pageNumber="26">
<heading id="761F5825F403FFC99A9DFEF2FE66FE90" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="1F9C335BF403FFC99A9DFEF2FE66FE90" bold="true" box="[189,426,264,290]" pageId="25" pageNumber="26">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F403FFC99ACAFECDFE08FE09" blockId="25.[189,1399,311,443]" pageId="25" pageNumber="26">
<materialsCitation id="9D80E514F403FFC99ACAFECDFE08FE09" ID-GBIF-Occurrence="3983014328" collectingDate="2019-11-19" collectionCode="EPTEA, CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="1F9C335BF403FFC99ACAFECDFE90FEE3" bold="true" box="[234,348,311,337]" pageId="25" pageNumber="26">
<typeStatus id="F25351EBF403FFC99ACAFECDFE90FEE3" box="[234,348,311,337]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="55FFAFD9F403FFC99A9DFEA0FEEAFEC6" box="[189,294,346,372]" name="Brazil" pageId="25" pageNumber="26">BRAZIL</collectingCountry>
• 
<specimenCount id="3BEE24C0F403FFC99B1EFEA1FE9BFEC7" box="[318,343,347,373]" pageId="25" pageNumber="26" type="male">♂</specimenCount>
; 
<collectingRegion id="EF2C21ABF403FFC99B44FEA0FDC8FEC6" box="[356,516,346,372]" country="Brazil" name="Minas Gerais" pageId="25" pageNumber="26">Minas Gerais</collectingRegion>
, 
<collectingMunicipality id="CD337533F403FFC9982FFEA0FDAEFEC6" box="[527,610,346,372]" pageId="25" pageNumber="26">Viçosa</collectingMunicipality>
, 
<collectionCode id="4BF9778CF403FFC9984DFEA0FD0BFEC6" box="[621,711,346,372]" pageId="25" pageNumber="26">EPTEA</collectionCode>
<location id="2837B992F403FFC998EEFEA1FC5CFEC6" LSID="urn:lsid:plazi:treatment:A5415E5FF402FFF39857FA44FC95FE49:2837B992F403FFC998EEFEA1FC5CFEC6" box="[718,912,346,373]" country="Brazil" municipality="Vicosa" name="Mata do Paraiso" pageId="25" pageNumber="26" stateProvince="Minas Gerais">Mata do Paraíso</location>
; 
<date id="5956C989F403FFC999BBFEA0FBF1FEC6" box="[923,1085,346,373]" pageId="25" pageNumber="26" value="2019-11-19">
<collectingDate id="49123061F403FFC999BBFEA0FBF1FEC6" box="[923,1085,346,373]" pageId="25" pageNumber="26" value="2019-11-19">19 Nov. 2019</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99E69FEA0FB6DFEC6" box="[1097,1185,346,372]" pageId="25" pageNumber="26">LabCol</collectorName>
leg.; “Fungo 30 \\ em 
<taxonomicName id="EAE894CAF403FFC99AC9FE84FE02FE25" box="[233,462,381,408]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="sylvaticus">
<emphasis id="1F9C335BF403FFC99AC9FE84FE02FE25" box="[233,462,381,408]" italics="true" pageId="25" pageNumber="26">Agaricus sylvaticus</emphasis>
</taxonomicName>
\\ 
<taxonomicName id="EAE894CAF403FFC99BCEFE84FBEDFE25" authority="von Groll &amp; Lopes-Andrade" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[494,1057,381,408]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="1F9C335BF403FFC99BCEFE84FD07FE25" box="[494,715,381,408]" italics="true" pageId="25" pageNumber="26">Cyparium lescheni</emphasis>
von Groll &amp; Lopes-Andrade
</taxonomicName>
<typeStatus id="F25351EBF403FFC99E09FE84FB16FE2A" box="[1065,1242,382,408]" pageId="25" pageNumber="26" type="holotype">HOLOTYPUS</typeStatus>
” [red paper]; 
<collectionCode id="4BF9778CF403FFC99A9DFE5BFEC0FE09" box="[189,268,417,443]" pageId="25" pageNumber="26">CELC</collectionCode>
. (Supp. file 1 
<collectionCode id="4BF9778CF403FFC99B87FE5BFE77FE09" box="[423,443,417,443]" pageId="25" pageNumber="26">B</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="2D57EF49F403FFC99ACAFE18FE73FBBE" blockId="25.[189,1399,482,1036]" pageId="25" pageNumber="26">
<materialsCitation id="9D80E514F403FFC99ACAFE18FDB7FDF1" ID-GBIF-Occurrence="3983014386" collectingDate="2015-02-15" collectionCode="T, CELC" collectorName="S. Aloquio &amp; A. Orsetti &amp; M. Bento" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="1F9C335BF403FFC99ACAFE18FEABFE4E" bold="true" box="[234,359,482,508]" pageId="25" pageNumber="26">
<typeStatus id="F25351EBF403FFC99ACAFE18FEABFE4E" box="[234,359,482,508]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="55FFAFD9F403FFC99A9DFDFFFEEAFDAD" box="[189,294,517,543]" name="Brazil" pageId="25" pageNumber="26">BRAZIL</collectingCountry>
• 
<specimenCount id="3BEE24C0F403FFC99B1CFDFFFEA5FD92" box="[316,361,517,544]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99B54FDFFFE7AFD92" box="[372,438,517,544]" pageId="25" pageNumber="26" type="female">4 ♀♀</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC998DDFDFFFCABFDAD" box="[765,871,517,543]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, “ 
<collectionCode id="4BF9778CF403FFC9995DFDFFFC5DFDAD" box="[893,913,517,543]" name="Tavera, Department of Geology and Geophysics" pageId="25" pageNumber="26">T</collectionCode>
. dos Gigantes”; 
<date id="5956C989F403FFC99E6DFDFFFB24FD92" box="[1101,1256,517,544]" pageId="25" pageNumber="26" value="2015-02-15">
<collectingDate id="49123061F403FFC99E6DFDFFFB24FD92" box="[1101,1256,517,544]" pageId="25" pageNumber="26" value="2015-02-15">15 Feb. 2015</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99ED3FDFFFABEFDAD" box="[1267,1394,517,543]" pageId="25" pageNumber="26">S. Aloquio</collectorName>
, 
<collectorName id="801D8A9FF403FFC99A9DFDD3FEF8FDF0" box="[189,308,552,579]" pageId="25" pageNumber="26">A. Orsetti</collectorName>
and 
<collectorName id="801D8A9FF403FFC99B55FDD3FE2BFDF1" box="[373,487,553,579]" pageId="25" pageNumber="26">M. Bento</collectorName>
leg.; 
<collectionCode id="4BF9778CF403FFC99810FDD3FDB7FDF1" box="[560,635,553,579]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC998BCFDD3FC92FDD4" ID-GBIF-Occurrence="3983014318" collectingDate="2015-02-27" collectionCode="T, CELC" collectorName="I. S. C. Pecci-Maddalena" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC998BCFDD3FD00FDF1" box="[668,716,553,579]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC99E11FDD2FB57FDF0" box="[1073,1179,552,578]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, “ 
<collectionCode id="4BF9778CF403FFC99E96FDD3FB06FDF1" box="[1206,1226,553,579]" name="Tavera, Department of Geology and Geophysics" pageId="25" pageNumber="26">T</collectionCode>
. da Madeira”; 
<date id="5956C989F403FFC99A9DFDB6FE93FDD4" box="[189,351,587,614]" pageId="25" pageNumber="26" value="2015-02-27">
<collectingDate id="49123061F403FFC99A9DFDB6FE93FDD4" box="[189,351,587,614]" pageId="25" pageNumber="26" value="2015-02-27">27 Feb. 2015</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99B4DFDB6FD4BFDD4" box="[365,647,588,614]" pageId="25" pageNumber="26">I.S.C. Pecci-Maddalena</collectorName>
<emphasis id="1F9C335BF403FFC998B2FDB6FD01FDD4" box="[658,717,588,614]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
leg.; 
<collectionCode id="4BF9778CF403FFC99933FDB6FC92FDD4" box="[787,862,588,614]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC9995CFDB6FBBEFD3B" ID-GBIF-Occurrence="3983014339" collectingDate="2015-03-13" collectionCode="CELC" collectorName="S. Aloquio &amp; A. Orsetti &amp; C. Lopes-Andrade &amp; M. Bento" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC9995CFDB6FC61FDD4" box="[892,941,588,614]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC99F28FDB6FABEFDD4" box="[1288,1394,588,614]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 
<date id="5956C989F403FFC99A9DFD95FE97FD38" box="[189,347,623,650]" pageId="25" pageNumber="26" value="2015-03-13">
<collectingDate id="49123061F403FFC99A9DFD95FE97FD38" box="[189,347,623,650]" pageId="25" pageNumber="26" value="2015-03-13">13 Mar. 2015</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99B45FD95FE28FD3B" box="[357,484,623,649]" pageId="25" pageNumber="26">S. Aloquio</collectorName>
, 
<collectorName id="801D8A9FF403FFC99BCDFD95FDAFFD3B" box="[493,611,623,649]" pageId="25" pageNumber="26">A. Orsetti</collectorName>
, 
<collectorName id="801D8A9FF403FFC9984DFD95FC88FD3B" box="[621,836,623,649]" pageId="25" pageNumber="26">C. Lopes-Andrade</collectorName>
and 
<collectorName id="801D8A9FF403FFC9995AFD95FC24FD3B" box="[890,1000,623,649]" pageId="25" pageNumber="26">M. Bento</collectorName>
leg.; 
<collectionCode id="4BF9778CF403FFC99E07FD95FBBEFD3B" box="[1063,1138,623,649]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC99EA8FD95FE7AFD7D" ID-GBIF-Occurrence="3983014331" collectingDate="2016-11-09" collectionCode="CELC" collectorName="I. Pecci-Maddalena &amp; C. Lopes-Andrade" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC99EA8FD95FB7AFD3B" box="[1160,1206,623,649]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC99B64FD68FE62FD1E" box="[324,430,658,684]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 
<date id="5956C989F403FFC99B9CFD68FD9FFD1E" box="[444,595,658,685]" pageId="25" pageNumber="26" value="2016-11-09">
<collectingDate id="49123061F403FFC99B9CFD68FD9FFD1E" box="[444,595,658,685]" pageId="25" pageNumber="26" value="2016-11-09">9 Nov. 2016</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99842FD69FC84FD1E" box="[610,840,658,685]" pageId="25" pageNumber="26">I. Pecci-Maddalena</collectorName>
and 
<collectorName id="801D8A9FF403FFC999A4FD68FB92FD1E" box="[900,1118,658,684]" pageId="25" pageNumber="26">C. Lopes-Andrade</collectorName>
leg.; “\\ ex 
<taxonomicName id="EAE894CAF403FFC99ED0FD68FE82FD7D" class="Agaricomycetes" family="Psathyrellaceae" genus="Psathyrella" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="candolleana">
<emphasis id="1F9C335BF403FFC99ED0FD68FE82FD7D" italics="true" pageId="25" pageNumber="26">Psathyrella candolleana</emphasis>
</taxonomicName>
”; 
<collectionCode id="4BF9778CF403FFC99B4BFD4FFE7AFD7D" box="[363,438,693,719]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC99BF1FD4FFB12FD41" ID-GBIF-Occurrence="3983014308" collectingDate="2019-11-19" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="18" specimenCount-female="10" specimenCount-male="8" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC99BF1FD4FFDDAFD62" box="[465,534,693,720]" pageId="25" pageNumber="26" type="male">5 ♂♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99804FD4CFDABFD62" box="[548,615,694,720]" pageId="25" pageNumber="26" type="female">7 ♀♀</specimenCount>
(
<specimenCount id="3BEE24C0F403FFC99858FD4FFD72FD62" box="[632,702,693,720]" pageId="25" pageNumber="26" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC998ECFD4FFD35FD62" box="[716,761,693,720]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, entirely dissected, preserved in glycerin; 
<specimenCount id="3BEE24C0F403FFC99ECFFD4FFAD2FD62" box="[1263,1310,693,720]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99F0CFD4FFABCFD62" box="[1324,1392,693,720]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
, abdomen dissected, preserved in glycerin); same collection data as for 
<typeStatus id="F25351EBF403FFC99E3AFD22FB48FD40" box="[1050,1156,728,754]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 
<collectionCode id="4BF9778CF403FFC99EB3FD23FB12FD41" box="[1171,1246,729,755]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC99EDDFD23FD0BFCA4" ID-GBIF-Occurrence="3983014363" collectingDate="2019-11-19" collectionCode="CAMB" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC99EDDFD23FAFCFD41" box="[1277,1328,729,755]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
; 
<specimenCount id="3BEE24C0F403FFC99F1FFD23FAA3FD41" box="[1343,1391,729,755]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC99BDBFD06FDA9FCA4" box="[507,613,764,790]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 
<collectionCode id="4BF9778CF403FFC99851FD06FD0BFCA4" box="[625,711,764,790]" pageId="25" pageNumber="26">CAMB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC998C0FD06FAA9FCA4" ID-GBIF-Occurrence="3983014313" box="[736,1381,764,790]" collectingDate="2019-11-19" collectionCode="CERPE" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC998C0FD06FCC3FCA4" box="[736,783,764,790]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
; 
<specimenCount id="3BEE24C0F403FFC9993BFD06FC8BFCA4" box="[795,839,764,790]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC99EB4FD06FB32FCA4" box="[1172,1278,764,790]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 
<collectionCode id="4BF9778CF403FFC99F2AFD06FAA9FCA4" box="[1290,1381,764,790]" pageId="25" pageNumber="26">CERPE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC99A9DFCE5FC98FCEE" ID-GBIF-Occurrence="3983014389" collectingDate="2019-11-21" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="25" pageNumber="26" specimenCount="4" specimenCount-female="1" specimenCount-male="2" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="3BEE24C0F403FFC99A9DFCE5FECEFC8B" box="[189,258,799,825]" pageId="25" pageNumber="26" type="male">2 ♂♂</specimenCount>
; 
<specimenCount id="3BEE24C0F403FFC99B2FFCE5FEF0FC8B" box="[271,316,799,825]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99B6BFCE5FE6CFC8B" box="[331,416,799,825]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
; same collection data as for 
<typeStatus id="F25351EBF403FFC998CFFCE5FC95FC8B" box="[751,857,799,825]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 
<date id="5956C989F403FFC99946FCE5FBC5FC8B" box="[870,1033,799,825]" pageId="25" pageNumber="26" value="2019-11-21">
<collectingDate id="49123061F403FFC99946FCE5FBC5FC8B" box="[870,1033,799,825]" pageId="25" pageNumber="26" value="2019-11-21">21 Nov. 2019</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99E36FCE5FBA2FC8B" box="[1046,1134,799,825]" pageId="25" pageNumber="26">LabCol</collectorName>
leg.; “Fungo 06 \\ Em 
<taxonomicName id="EAE894CAF403FFC99A9DFCB8FE6BFCEE" box="[189,423,834,860]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="dulcidulus">
<emphasis id="1F9C335BF403FFC99A9DFCB8FE6BFCEE" box="[189,423,834,860]" italics="true" pageId="25" pageNumber="26">Agaricus dulcidulus</emphasis>
</taxonomicName>
e 
<taxonomicName id="EAE894CAF403FFC99B9FFCB9FD20FCEE" box="[447,748,834,860]" class="Agaricomycetes" family="Agaricaceae" genus="Leucocoprinus" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="brebissoni">
<emphasis id="1F9C335BF403FFC99B9FFCB9FD20FCEE" box="[447,748,834,860]" italics="true" pageId="25" pageNumber="26">Leucocoprinus brebissoni</emphasis>
</taxonomicName>
”; 
<collectionCode id="4BF9778CF403FFC99929FCB8FC98FCEE" box="[777,852,834,860]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC9994AFCB8FB06FCCD" ID-GBIF-Occurrence="3983014335" collectingDate="2019-11-20" collectionCode="CELC" collectorName="Vicosa &amp; Recanto das Cigarras &amp; LabCol" country="Brazil" location="Mata da Biol." pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="3BEE24C0F403FFC9994AFCB8FC67FCEF" box="[874,939,834,861]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
; 
<collectorName id="801D8A9FF403FFC99998FCB8FBC7FCEE" box="[952,1035,834,860]" pageId="25" pageNumber="26">Viçosa</collectorName>
, 
<collectorName id="801D8A9FF403FFC99E35FCB9FAC2FCEE" box="[1045,1294,834,861]" pageId="25" pageNumber="26">Recanto das Cigarras</collectorName>
, 
<location id="2837B992F403FFC99F39FCB9FF36FCCD" LSID="urn:lsid:plazi:treatment:A5415E5FF402FFF39857FA44FC95FE49:2837B992F403FFC99F39FCB9FF36FCCD" country="Brazil" name="Mata da Biol." pageId="25" pageNumber="26">Mata da Biol.</location>
; 
<date id="5956C989F403FFC99B27FC9FFE65FCCD" box="[263,425,869,896]" pageId="25" pageNumber="26" value="2019-11-20">
<collectingDate id="49123061F403FFC99B27FC9FFE65FCCD" box="[263,425,869,896]" pageId="25" pageNumber="26" value="2019-11-20">20 Nov. 2019</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99B95FC9FFDC1FCCD" box="[437,525,869,895]" pageId="25" pageNumber="26">LabCol</collectorName>
leg.; “Fungo 29 \\ Em 
<taxonomicName id="EAE894CAF403FFC9993CFC9FFBF5FCCD" box="[796,1081,869,895]" class="Agaricomycetes" family="Entolomataceae" genus="Entoloma" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="subGenus" subGenus="Inocephalus">
<emphasis id="1F9C335BF403FFC9993CFC9FFC42FCCD" box="[796,910,869,895]" italics="true" pageId="25" pageNumber="26">Entoloma</emphasis>
(
<emphasis id="1F9C335BF403FFC99980FC9FFBE3FCCD" box="[928,1071,869,895]" italics="true" pageId="25" pageNumber="26">Inocephalus</emphasis>
)
</taxonomicName>
sp.”; 
<collectionCode id="4BF9778CF403FFC99E5FFC9FFB06FCCD" box="[1151,1226,869,895]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC99EC3FC9FFB74FC74" ID-GBIF-Occurrence="3983014378" collectingDate="2019-11-20" collectionCode="CELC" collectorName="Vicosa &amp; Recanto das Cigarras &amp; LabCol" country="Brazil" location="Mata da Biol." pageId="25" pageNumber="26" specimenCount="15" specimenCount-female="4" specimenCount-male="11" typeStatus="paratype">
<specimenCount id="3BEE24C0F403FFC99EC3FC9FFAEBFC32" box="[1251,1319,869,896]" pageId="25" pageNumber="26" type="male">8 ♂♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99F14FC9FFABAFC32" box="[1332,1398,869,896]" pageId="25" pageNumber="26" type="female">3 ♀♀</specimenCount>
(
<specimenCount id="3BEE24C0F403FFC99AE9FC73FF3AFC11" box="[201,246,905,931]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, entirely dissected, preserved in glycerin; 
<specimenCount id="3BEE24C0F403FFC998CBFC73FCFDFC11" box="[747,817,905,931]" pageId="25" pageNumber="26" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC9991EFC73FCA0FC11" box="[830,876,905,931]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, abdomen dissected, preserved in glycerin); same collection data as for preceding; “Fungo 08; 
<collectionCode id="4BF9778CF403FFC99929FC56FC98FC74" box="[777,852,940,966]" pageId="25" pageNumber="26">CELC</collectionCode>
\\ Em 
<taxonomicName id="EAE894CAF403FFC99982FC56FB9FFC74" box="[930,1107,940,966]" class="Agaricomycetes" family="Psathyrellaceae" genus="Psathyrella" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF403FFC99982FC56FBE4FC74" box="[930,1064,940,966]" italics="true" pageId="25" pageNumber="26">Psathyrella</emphasis>
sp.
</taxonomicName>
”; 
<collectionCode id="4BF9778CF403FFC99E4DFC56FB74FC74" box="[1133,1208,940,966]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="9D80E514F403FFC99EF0FC56FE77FBBE" ID-GBIF-Occurrence="3983014354" collectingDate="2021-10-15" collectionCode="CELC" collectorName="E. von Groll &amp; A. Orsetti" country="Brazil" county="da Biologia" location="Vicosa" municipality="Mata da Biologia" pageId="25" pageNumber="26" specimenCount="18" specimenCount-female="11" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="3BEE24C0F403FFC99EF0FC56FAD8FC74" box="[1232,1300,940,966]" pageId="25" pageNumber="26" type="male">6 ♂♂</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99F00FC56FABCFC74" box="[1312,1392,940,966]" pageId="25" pageNumber="26" type="female">11 ♀♀</specimenCount>
, 
<specimenCount id="3BEE24C0F403FFC99A9DFC35FEDEFC5B" box="[189,274,975,1001]" pageId="25" pageNumber="26" type="generic">1 spec.</specimenCount>
; 
<location id="2837B992F403FFC99B03FC35FEBAFC5B" LSID="urn:lsid:plazi:treatment:A5415E5FF402FFF39857FA44FC95FE49:2837B992F403FFC99B03FC35FEBAFC5B" box="[291,374,975,1001]" country="Brazil" county="da Biologia" municipality="Mata da Biologia" name="Vicosa" pageId="25" pageNumber="26">Viçosa</location>
, 
<collectingMunicipality id="CD337533F403FFC99BA4FC35FD95FC5B" box="[388,601,975,1001]" pageId="25" pageNumber="26">
Mata 
<collectingCounty id="C43697C5F403FFC99BEAFC35FD95FC5B" box="[458,601,975,1001]" pageId="25" pageNumber="26">da Biologia</collectingCounty>
</collectingMunicipality>
; 
<date id="5956C989F403FFC99847FC35FCC5FC5B" box="[615,777,975,1002]" pageId="25" pageNumber="26" value="2021-10-15">
<collectingDate id="49123061F403FFC99847FC35FCC5FC5B" box="[615,777,975,1002]" pageId="25" pageNumber="26" value="2021-10-15">15 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="801D8A9FF403FFC99938FC35FC63FC5B" box="[792,943,975,1001]" pageId="25" pageNumber="26">E. von Groll</collectorName>
and 
<collectorName id="801D8A9FF403FFC999CCFC35FBAEFC5B" box="[1004,1122,975,1001]" pageId="25" pageNumber="26">A. Orsetti</collectorName>
leg.; “Fungo 20 \\ Em 
<taxonomicName id="EAE894CAF403FFC99A9DFC08FE9EFBBE" box="[189,338,1010,1036]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF403FFC99A9DFC08FEEBFBBE" box="[189,295,1010,1036]" italics="true" pageId="25" pageNumber="26">Agaricus</emphasis>
sp.
</taxonomicName>
”; 
<collectionCode id="4BF9778CF403FFC99B4CFC08FE77FBBE" box="[364,443,1010,1036]" pageId="25" pageNumber="26">CELC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="2D57EF49F403FFC99A9DFBC9FEB2FBC3" blockId="25.[189,1398,1075,1137]" pageId="25" pageNumber="26">
All 
<typeStatus id="F25351EBF403FFC99ACFFBCEFE93FBFC" box="[239,351,1076,1102]" pageId="25" pageNumber="26" type="paratype">paratypes</typeStatus>
additionally labelled “ 
<taxonomicName id="EAE894CAF403FFC9985AFBCEFB08FBFF" authority="von Groll &amp; Lopes-Andrade" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[634,1220,1075,1102]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="1F9C335BF403FFC9985AFBCEFC90FBFF" box="[634,860,1075,1102]" italics="true" pageId="25" pageNumber="26">Cyparium lescheni</emphasis>
von Groll &amp; Lopes-Andrade
</taxonomicName>
<typeStatus id="F25351EBF403FFC99EF1FBCEFABAFBFC" box="[1233,1398,1076,1102]" pageId="25" pageNumber="26" type="paratype">PARATYPUS</typeStatus>
[yellow paper]”.
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F403FFF19A9DFB62FD7BF9A6" lastPageId="33" lastPageNumber="34" pageId="25" pageNumber="26" type="description">
<paragraph id="2D57EF49F403FFC99A9DFB62FE82FB00" blockId="25.[189,334,1176,1202]" box="[189,334,1176,1202]" pageId="25" pageNumber="26">
<heading id="761F5825F403FFC99A9DFB62FE82FB00" bold="true" box="[189,334,1176,1202]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="1F9C335BF403FFC99A9DFB62FE82FB00" bold="true" box="[189,334,1176,1202]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F403FFC99A9DFB3DFEFCFAB6" blockId="25.[189,1398,1223,1284]" pageId="25" pageNumber="26">
MEASUREMENTS (
<typeStatus id="F25351EBF403FFC99BA2FB3DFE21FB53" box="[386,493,1223,1249]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, in mm). TL 2.32, PL 0.76, PA 0.70, PB 1.40, EW 0.86, EL 1.62, IS 0.20, HW 0.62.
</paragraph>
<paragraph id="2D57EF49F403FFC99A9DFAD1FEF2F9AB" blockId="25.[189,1401,1323,1561]" pageId="25" pageNumber="26">
COLORATION. Iridescent. Brown; antennomeres I–VI, clypeus, mouthparts, tarsi yellow (
<figureCitation id="B5D3F3CCF403FFC99EDAFAD1FABEFAF7" box="[1274,1394,1323,1349]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Figs 14A</figureCitation>
, 
<figureCitation id="B5D3F3CCF403FFC99A9DFAB4FF38FADB" box="[189,244,1358,1385]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">15A</figureCitation>
). Variation: few 
<typeStatus id="F25351EBF403FFC99B98FAB5FDE4FADB" box="[440,552,1359,1385]" pageId="25" pageNumber="26" type="paratype">paratypes</typeStatus>
with pronotum and hypomeron reddish brown (
<figureCitation id="B5D3F3CCF403FFC99E41FAB4FB23FADB" box="[1121,1263,1358,1385]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Fig. 14D–F</figureCitation>
); scutellum reddish brown or black (
<figureCitation id="B5D3F3CCF403FFC99BFFFA88FD81FA3E" box="[479,589,1394,1420]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Fig. 14D</figureCitation>
); meso- and metathorax reddish brown, each sclerite lighter laterally (
<figureCitation id="B5D3F3CCF403FFC99AE6FA6FFEFCFA1D" box="[198,304,1429,1455]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Fig. 14E</figureCitation>
); elytra blackish (
<figureCitation id="B5D3F3CCF403FFC99820FA6FFDA1FA1D" box="[512,621,1429,1455]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Fig. 14D</figureCitation>
); epipleuron dark ochre; coxae and trochanters dark ochre; femora dark ochre, apex lighter; tibiae dark ochre, lighter anteriorly and posteriorly; tarsi yellow (
<figureCitation id="B5D3F3CCF403FFC99EDCFA42FAA5FA60" box="[1276,1385,1464,1490]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Fig. 14E</figureCitation>
); tergite VI and VII blackish; tergite VIII yellow; ventrite 1 dark ochre, 2–4 dark yellow, 5 and 6 yellow (
<figureCitation id="B5D3F3CCF403FFC99AE6FA04FEFDF9AB" box="[198,305,1534,1561]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="25" pageNumber="26">Fig. 14E</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F403FFC99A9DF9BAFBFDF82A" blockId="25.[188,1399,1600,1944]" pageId="25" pageNumber="26">
HEAD. Punctation dense, fine (
<figureCitation id="B5D3F3CCF403FFC9983CF9BAFD45F9E9" box="[540,649,1600,1627]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">Fig. 15A</figureCitation>
). Eyes long (
<figureCitation id="B5D3F3CCF403FFC99900F9BAFC41F9E9" box="[800,909,1600,1627]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">Fig. 15A</figureCitation>
). Labrum sub-rectangular, lateral margins smoothly rounded, not well delimited apically; central margin sub-straight; sclerotized portion inwardly curved; lateral setae considerably extending from the margins of the labrum; densely porose centrally (
<figureCitation id="B5D3F3CCF403FFC99AE7F953FEFEF976" box="[199,306,1705,1732]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">Fig. 15D</figureCitation>
). Mandibles strongly curved; subapical serrations on left mandible conspicuous (
<figureCitation id="B5D3F3CCF403FFC99EC1F953FAA6F976" box="[1249,1386,1705,1732]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">Fig. 15E–F</figureCitation>
). Maxillary palps, galea and lacinia moderate pubescent, elongated (
<figureCitation id="B5D3F3CCF403FFC999ECF937FBF5F95A" box="[972,1081,1741,1768]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">Fig. 15G</figureCitation>
). Mentum straight apically and rounded laterally; sides well delimited (
<figureCitation id="B5D3F3CCF403FFC9989AF90AFCEAF8B8" box="[698,806,1776,1802]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="25" pageNumber="26">Fig. 16A</figureCitation>
). Setae of labial palpomere II extending palpomere III; palpomere III longish with long apical setae (
<figureCitation id="B5D3F3CCF403FFC99933F8E9FC4FF89F" box="[787,899,1811,1837]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="25" pageNumber="26">Fig. 16A</figureCitation>
). Hypopharynx with narrow and rounded sclerotized-plate (
<figureCitation id="B5D3F3CCF403FFC99BB2F8CCFDE9F8E2" box="[402,549,1846,1872]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="25" pageNumber="26">Fig. 16A–B</figureCitation>
). Post gena microsculptured with transversal lines; gular pores sparse; gula long and sharp (
<figureCitation id="B5D3F3CCF403FFC99B96F8A3FDEEF8C1" box="[438,546,1881,1907]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="25" pageNumber="26">Fig. 16C</figureCitation>
). Antennal club very distinct; antennomere XI longish, hexagonal, more or less rounded apically; remarkably different between sexes (
<figureCitation id="B5D3F3CCF403FFC999B6F887FBE8F82A" box="[918,1060,1917,1944]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="25" pageNumber="26">Fig. 15B–C</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F400FFCA9A9DFEF3FDE1FE02" blockId="26.[188,1399,265,432]" pageId="26" pageNumber="27">
PROTHORAX. Pronotum smooth, shining; punctuation dense, fine; pubescence short, fine (
<figureCitation id="B5D3F3CCF400FFCA9EF9FEF3FAA6FE91" box="[1241,1386,265,291]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="26" pageNumber="27">Fig. 16D–E</figureCitation>
); transverse, sub-straight laterally, forming an obtuse angle at lateral areas of posterior margin (
<figureCitation id="B5D3F3CCF400FFCA9F20FED6FAA6FEF4" box="[1280,1386,300,326]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="26" pageNumber="27">Fig. 16E</figureCitation>
). Hypomeron with strigulate microsculpture. Notosternal suture straight, inward directed (
<figureCitation id="B5D3F3CCF400FFCA9EDBFEB5FAA5FEDB" box="[1275,1385,335,361]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="26" pageNumber="27">Fig. 16F</figureCitation>
). Profurca somewhat thick and elongated, slightly extending half length of foramen (
<figureCitation id="B5D3F3CCF400FFCA9EA0FE88FB20FE3E" box="[1152,1260,370,396]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="26" pageNumber="27">Fig. 16G</figureCitation>
). Prosternal process acuminated (
<figureCitation id="B5D3F3CCF400FFCA9B92FE6CFDECFE02" box="[434,544,406,432]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="26" pageNumber="27">Fig. 17A</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F400FFCB9A9DFE22FB11FE44" blockId="26.[187,1399,471,639]" lastBlockId="27.[189,1401,265,503]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, 
<bibRefCitation id="497992B8F400FFCA99DAFE2DFAC8FE40" author="Leschen R. A. B. &amp; Lobl I." box="[1018,1284,471,498]" pageId="26" pageNumber="27" pagination="1 - 63" refId="ref32588" refString="Leschen R. A. B. &amp; Lobl I. 2005. Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1 - 63. https: // doi. org / 10.1649 / 0010 - 065 X (2005) 059 [0001: PACOSS] 2.0. CO; 2" type="journal article" year="2005">Leschen &amp; Löbl 2005</bibRefCitation>
) strongly wavy (
<figureCitation id="B5D3F3CCF400FFCA9B31FE00FE4DFDA7" box="[273,385,506,533]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="26" pageNumber="27">Fig. 17B</figureCitation>
). Anterior phragma large and straight (
<figureCitation id="B5D3F3CCF400FFCA9943FE00FC1FFDA7" box="[867,979,506,533]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="26" pageNumber="27">Fig. 17C</figureCitation>
). Mesanepisternum with strigulate microsculpture. Procoxal rests triangular; slightly curved posteriorly (
<figureCitation id="B5D3F3CCF400FFCA99C3FDE4FB83FD8A" box="[995,1103,542,568]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="26" pageNumber="27">Fig. 17D</figureCitation>
). Mesoventral lines wavy and obtuse; median lines moderately wavy; area between median and mesoventral lines thin (
<figureCitation id="B5D3F3CCF400FFCA9EDBFDBBFAABFDE9" box="[1275,1383,577,603]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="26" pageNumber="27">Fig. 17D</figureCitation>
). Mesoventral process long, slightly curved at top and base, forming a not well marked ridge (
<figureCitation id="B5D3F3CCF400FFCA9ED9FD9EFAAFFDCC" box="[1273,1379,612,638]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="26" pageNumber="27">Fig. 17E</figureCitation>
). METATHORAX. Metanotum with alacrista somewhat enlarged anteriorly and turned to anterior portion; scutoscutellar suture very strongly curved; median membranous area remarkably wide, and short (
<figureCitation id="B5D3F3CCF401FFCB9AE6FEB5FEFFFEDB" box="[198,307,335,361]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="27" pageNumber="28">Fig. 17F</figureCitation>
). Metaventrite with strigulate microsculpture; punctuation sparse, fine; coarsely punctate above intercoxal plates (
<figureCitation id="B5D3F3CCF401FFCB9BC8FE88FD95FE3F" box="[488,601,370,397]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="27" pageNumber="28">Fig. 17D</figureCitation>
). Mesocoxal line forming a smooth angle between coxal cavities (
<figureCitation id="B5D3F3CCF401FFCB9AE6FE6CFEF9FE02" box="[198,309,406,432]" captionStart="Fig" captionStartId="29.[189,232,1742,1768]" captionTargetBox="[189,1398,282,1702]" captionTargetId="figure-12@29.[189,1398,270,1705]" captionTargetPageId="29" captionText="Fig. 17. Cyparium lescheni sp. nov., paratypes, ♂ (CELC). A. Prothorax, lateral view B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="27" pageNumber="28">Fig. 17D</figureCitation>
). Metanepisternum and metepimeron with imbricate microsculpture. Intercoxal plates with strigulate microsculpture. Metendosternite with curved furcal arms; ‘stalk ridge’ exceeding half length of stalk (
<figureCitation id="B5D3F3CCF401FFCB9B05FE26FE5FFE44" box="[293,403,476,502]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">Fig. 18A</figureCitation>
); ventral longitudinal flange long and narrow in lateral view (
<figureCitation id="B5D3F3CCF401FFCB9E44FE26FB03FE44" box="[1124,1231,476,502]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">Fig. 18B</figureCitation>
).
</paragraph>
<caption id="7997BFC1F400FFCA9A9DF87EFBC3F857" ID-DOI="http://doi.org/10.5281/zenodo.7033428" ID-Zenodo-Dep="7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1924,1950]" targetBox="[205,1394,748,1886]" targetPageId="26">
<paragraph id="2D57EF49F400FFCA9A9DF87EFBC3F857" blockId="26.[189,1398,1924,2021]" pageId="26" pageNumber="27">
<emphasis id="1F9C335BF400FFCA9A9DF87EFED7F82C" bold="true" box="[189,283,1924,1950]" pageId="26" pageNumber="27">Fig. 14.</emphasis>
<taxonomicName id="EAE894CAF400FFCA9B03F87FFDCCF82C" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[291,512,1924,1951]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF400FFCA9B03F87FFDCCF82C" box="[291,512,1924,1951]" italics="true" pageId="26" pageNumber="27">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F400FFCA9829F87FFDA8F82D" box="[521,612,1925,1951]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF400FFCA984CF87EFD69F82C" bold="true" box="[620,677,1924,1950]" pageId="26" pageNumber="27">A–C</emphasis>
. Holotype, ♂ (CELC). 
<emphasis id="1F9C335BF400FFCA999CF87EFC3EF82C" bold="true" box="[956,1010,1924,1950]" pageId="26" pageNumber="27">D–F</emphasis>
. Paratype, ♂ (CELC). 
<emphasis id="1F9C335BF400FFCA9F24F87EFAD5F82C" bold="true" box="[1284,1305,1924,1950]" pageId="26" pageNumber="27">A</emphasis>
. Dorsal view. 
<emphasis id="1F9C335BF400FFCA9B25F852FED5F870" bold="true" box="[261,281,1960,1986]" pageId="26" pageNumber="27">B</emphasis>
. Ventral view. 
<emphasis id="1F9C335BF400FFCA9BF3F85DFE2AF873" bold="true" box="[467,486,1959,1985]" pageId="26" pageNumber="27">C</emphasis>
. Lateral view. 
<emphasis id="1F9C335BF400FFCA98BCF852FD7DF870" bold="true" box="[668,689,1960,1986]" pageId="26" pageNumber="27">D</emphasis>
. Dorsal view. 
<emphasis id="1F9C335BF400FFCA9943F852FCBBF870" bold="true" box="[867,887,1960,1986]" pageId="26" pageNumber="27">E</emphasis>
. Ventral view. 
<emphasis id="1F9C335BF400FFCA9E11F852FB8DF870" bold="true" box="[1073,1089,1960,1986]" pageId="26" pageNumber="27">F</emphasis>
. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="2D57EF49F401FFCB9A9DFDE4FB0AFD77" blockId="27.[189,1398,542,709]" pageId="27" pageNumber="28">
WINGS. Elytra slightly wider than long, partially covering tergite VI (
<figureCitation id="B5D3F3CCF401FFCB9E03FDE4FB43FD8A" box="[1059,1167,542,568]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="27" pageNumber="28">Fig 14A</figureCitation>
); basal and lateral lines punctate (
<figureCitation id="B5D3F3CCF401FFCB9B53FDBBFE2EFDE9" box="[371,482,577,603]" captionStart="Fig" captionStartId="28.[189,232,1797,1823]" captionTargetBox="[189,1394,273,1750]" captionTargetId="figure-17@28.[189,1398,267,1761]" captionTargetPageId="28" captionText="Fig. 16. Cyparium lescheni sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D–E. Pronotum, dorsal view. D. Holotype, ♂ (CELC). E. Paratype, ♂ (CELC). F–G. Paratype, ♂ (CELC), prothorax. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="27" pageNumber="28">Fig. 16D</figureCitation>
); sutural lines dashed; adsutural area with a row of setae; six rows of coarse punctures (not including sutural line) (
<figureCitation id="B5D3F3CCF401FFCB98AAFD9EFCECFDCD" box="[650,800,612,639]" captionStart="Fig" captionStartId="26.[189,232,1924,1950]" captionTargetBox="[205,1394,748,1886]" captionTargetId="figure-153@26.[189,1398,720,1888]" captionTargetPageId="26" captionText="Fig. 14. Cyparium lescheni sp. nov. A–C. Holotype, ♂ (CELC). D–F. Paratype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7033428" httpUri="https://zenodo.org/record/7033428/files/figure.png" pageId="27" pageNumber="28">Figs 14A, D</figureCitation>
, 
<figureCitation id="B5D3F3CCF401FFCB990DFD9EFCAEFDCC" box="[813,866,612,638]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">18C</figureCitation>
); apical coarse punctation sparse; apical fine punctation dense (
<figureCitation id="B5D3F3CCF401FFCB9BB5FD7DFDC8FD13" box="[405,516,647,673]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">Fig. 18D</figureCitation>
); apical serrations almost inconspicuous (
<figureCitation id="B5D3F3CCF401FFCB99D4FD7DFBAEFD13" box="[1012,1122,647,673]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">Fig. 18D</figureCitation>
); pubescence short and fine. Epipleuron with a line of coarse punctures. Hind wings fully developed (
<figureCitation id="B5D3F3CCF401FFCB9E6FFD50FB75FD77" box="[1103,1209,682,709]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">Fig. 18E</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F401FFCB9A9DFD16FD5DFCDF" blockId="27.[189,1398,747,877]" pageId="27" pageNumber="28">
LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Femora strongly fusiform (
<figureCitation id="B5D3F3CCF401FFCB9AE5FCF4FE91FC9A" box="[197,349,782,808]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="27" pageNumber="28">Fig. 18F–H</figureCitation>
). Profemora with inconspicuous punctuation; mesofemora with sparse and coarse punctuation; metafemora with fine and shallow punctuation. Mesotibiae densely spinose, spines fine; metatibiae sparsely spinose, spines fine.
</paragraph>
<caption id="7997BFC1F401FFCB9A9DF8A4FD48F851" ID-DOI="http://doi.org/10.5281/zenodo.7033432" ID-Zenodo-Dep="7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1886,1912]" targetBox="[189,1398,964,1848]" targetPageId="27">
<paragraph id="2D57EF49F401FFCB9A9DF8A4FD48F851" blockId="27.[189,1399,1886,2019]" pageId="27" pageNumber="28">
<emphasis id="1F9C335BF401FFCB9A9DF8A4FED1F8CA" bold="true" box="[189,285,1886,1912]" pageId="27" pageNumber="28">Fig. 15.</emphasis>
<taxonomicName id="EAE894CAF401FFCB9B08F8A5FDC4F8CA" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[296,520,1886,1913]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF401FFCB9B08F8A5FDC4F8CA" box="[296,520,1886,1913]" italics="true" pageId="27" pageNumber="28">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F401FFCB9832F8A5FDBCF8CB" box="[530,624,1887,1913]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF401FFCB985AF8A4FD43F8CA" bold="true" box="[634,655,1886,1912]" pageId="27" pageNumber="28">A</emphasis>
. Holotype, ♂ (CELC), frontal view. 
<emphasis id="1F9C335BF401FFCB9E72F8A5FBAAF8CB" bold="true" box="[1106,1126,1887,1913]" pageId="27" pageNumber="28">B</emphasis>
. Paratype, ♂ (CELC), antennae. 
<emphasis id="1F9C335BF401FFCB9B14F87BFE85F829" bold="true" box="[308,329,1921,1947]" pageId="27" pageNumber="28">C</emphasis>
. Paratype, ♀ (CELC), antennae. 
<emphasis id="1F9C335BF401FFCB98EFF87BFCC5F829" bold="true" box="[719,777,1921,1947]" pageId="27" pageNumber="28">D–G</emphasis>
. Paratype, ♂ (CELC). 
<emphasis id="1F9C335BF401FFCB9E38F878FBE1F82E" bold="true" box="[1048,1069,1922,1948]" pageId="27" pageNumber="28">D</emphasis>
. Labrum. 
<emphasis id="1F9C335BF401FFCB9E87F878FB77F82E" bold="true" box="[1191,1211,1922,1948]" pageId="27" pageNumber="28">E</emphasis>
. Left mandible. 
<emphasis id="1F9C335BF401FFCB9A9DF85FFF03F80D" bold="true" box="[189,207,1957,1983]" pageId="27" pageNumber="28">F</emphasis>
. Right mandible. 
<emphasis id="1F9C335BF401FFCB9B8AF85FFE0DF80D" bold="true" box="[426,449,1957,1983]" pageId="27" pageNumber="28">G</emphasis>
. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm.
</paragraph>
</caption>
<caption id="7997BFC1F406FFCC9A9DF8FFFB07F838" ID-DOI="http://doi.org/10.5281/zenodo.7033436" ID-Zenodo-Dep="7033436" httpUri="https://zenodo.org/record/7033436/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1797,1823]" targetBox="[189,1394,273,1750]" targetPageId="28">
<paragraph id="2D57EF49F406FFCC9A9DF8FFFB07F838" blockId="28.[189,1399,1797,1930]" pageId="28" pageNumber="29">
<emphasis id="1F9C335BF406FFCC9A9DF8FFFEEFF8AD" bold="true" box="[189,291,1797,1823]" pageId="28" pageNumber="29">Fig. 16.</emphasis>
<taxonomicName id="EAE894CAF406FFCC9B13F8FCFDD4F8AD" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[307,536,1797,1824]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF406FFCC9B13F8FCFDD4F8AD" box="[307,536,1797,1824]" italics="true" pageId="28" pageNumber="29">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F406FFCC9808F8FCFD47F892" box="[552,651,1798,1824]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF406FFCC98BAF8FFFD1FF8AD" bold="true" box="[666,723,1797,1823]" pageId="28" pageNumber="29">A–C</emphasis>
. Paratype, ♂ (CELC). 
<emphasis id="1F9C335BF406FFCC9E24F8FFFBD5F8AD" bold="true" box="[1028,1049,1797,1823]" pageId="28" pageNumber="29">A</emphasis>
. Labium. 
<emphasis id="1F9C335BF406FFCC9E83F8FCFB7BF892" bold="true" box="[1187,1207,1798,1824]" pageId="28" pageNumber="29">B</emphasis>
. Hypopharynx. 
<emphasis id="1F9C335BF406FFCC9A9DF8D2FF1EF8F0" bold="true" box="[189,210,1832,1858]" pageId="28" pageNumber="29">C</emphasis>
. Head, ventral view. 
<emphasis id="1F9C335BF406FFCC9BEFF8D3FDCAF8F1" bold="true" box="[463,518,1833,1859]" pageId="28" pageNumber="29">D–E</emphasis>
. Pronotum, dorsal view. 
<emphasis id="1F9C335BF406FFCC990EF8D3FC8FF8F1" bold="true" box="[814,835,1833,1859]" pageId="28" pageNumber="29">D</emphasis>
. Holotype, ♂ (CELC). 
<emphasis id="1F9C335BF406FFCC9E7AF8D3FBA2F8F1" bold="true" box="[1114,1134,1833,1859]" pageId="28" pageNumber="29">E</emphasis>
. Paratype, ♂ (CELC). 
<emphasis id="1F9C335BF406FFCC9A9DF8B6FF38F8D4" bold="true" box="[189,244,1868,1894]" pageId="28" pageNumber="29">F–G</emphasis>
. Paratype, ♂ (CELC), prothorax. 
<emphasis id="1F9C335BF406FFCC98A7F8B6FD55F8D4" bold="true" box="[647,665,1868,1894]" pageId="28" pageNumber="29">F</emphasis>
. Ventral view. 
<emphasis id="1F9C335BF406FFCC996CF8B6FCAFF8D4" bold="true" box="[844,867,1868,1894]" pageId="28" pageNumber="29">G</emphasis>
. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.2 mm.
</paragraph>
</caption>
<caption id="7997BFC1F407FFCD9A9DF934FC14F8E1" ID-DOI="http://doi.org/10.5281/zenodo.7033438" ID-Zenodo-Dep="7033438" httpUri="https://zenodo.org/record/7033438/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1742,1768]" targetBox="[189,1398,282,1702]" targetPageId="29">
<paragraph id="2D57EF49F407FFCD9A9DF934FC14F8E1" blockId="29.[188,1399,1742,1875]" pageId="29" pageNumber="30">
<emphasis id="1F9C335BF407FFCD9A9DF934FED0F95B" bold="true" box="[189,284,1742,1769]" pageId="29" pageNumber="30">Fig. 17.</emphasis>
<taxonomicName id="EAE894CAF407FFCD9B05F935FDC8F95B" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[293,516,1743,1769]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF407FFCD9B05F935FDC8F95B" box="[293,516,1743,1769]" italics="true" pageId="29" pageNumber="30">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F407FFCD982DF935FDA0F95B" box="[525,620,1743,1769]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, paratypes, ♂ (CELC). 
<emphasis id="1F9C335BF407FFCD99A8F934FC51F95A" bold="true" box="[904,925,1742,1768]" pageId="29" pageNumber="30">A</emphasis>
. Prothorax, lateral view 
<emphasis id="1F9C335BF407FFCD9EE6F934FB31F95A" bold="true" box="[1222,1277,1742,1768]" pageId="29" pageNumber="30">B–C</emphasis>
. Scutellar plate. 
<emphasis id="1F9C335BF407FFCD9B24F908FED4F8BE" bold="true" box="[260,280,1778,1804]" pageId="29" pageNumber="30">B</emphasis>
. Dorsal view. 
<emphasis id="1F9C335BF407FFCD9B9DF908FE1EF8BE" bold="true" box="[445,466,1778,1804]" pageId="29" pageNumber="30">C</emphasis>
. Apically slanted view. 
<emphasis id="1F9C335BF407FFCD98C7F908FCD2F8BE" bold="true" box="[743,798,1778,1804]" pageId="29" pageNumber="30">D–E</emphasis>
. Meso- and metathorax. 
<emphasis id="1F9C335BF407FFCD9E1FF908FB98F8BE" bold="true" box="[1087,1108,1778,1804]" pageId="29" pageNumber="30">D</emphasis>
. Ventral view. 
<emphasis id="1F9C335BF407FFCD9F21F908FAD9F8BE" bold="true" box="[1281,1301,1778,1804]" pageId="29" pageNumber="30">E</emphasis>
. Lateral view. 
<emphasis id="1F9C335BF407FFCD9B23F8EFFED9F89D" bold="true" box="[259,277,1813,1839]" pageId="29" pageNumber="30">F</emphasis>
. Metanotum. Specimens collected at Mata do Paraíso, Viçosa (A–E) and Mata da Biologia (F), Viçosa (MG, Brazil). Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm.
</paragraph>
</caption>
<caption id="7997BFC1F404FFCE9A9DF91BFE83F8D4" ID-DOI="http://doi.org/10.5281/zenodo.7033440" ID-Zenodo-Dep="7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1761,1787]" targetBox="[189,1393,275,1713]" targetPageId="30">
<paragraph id="2D57EF49F404FFCE9A9DF91BFE83F8D4" blockId="30.[188,1399,1761,1894]" pageId="30" pageNumber="31">
<emphasis id="1F9C335BF404FFCE9A9DF91BFED3F949" bold="true" box="[189,287,1761,1787]" pageId="30" pageNumber="31">Fig. 18.</emphasis>
<taxonomicName id="EAE894CAF404FFCE9B0BF918FDC1F94E" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[299,525,1762,1788]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF404FFCE9B0BF918FDC1F94E" box="[299,525,1762,1788]" italics="true" pageId="30" pageNumber="31">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F404FFCE9839F918FDB7F94E" box="[537,635,1762,1788]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (CELC). 
<emphasis id="1F9C335BF404FFCE99B8F91BFC1CF949" bold="true" box="[920,976,1761,1787]" pageId="30" pageNumber="31">A–B</emphasis>
. Metendosternite. 
<emphasis id="1F9C335BF404FFCE9E95F91BFB06F949" bold="true" box="[1205,1226,1761,1787]" pageId="30" pageNumber="31">A</emphasis>
. Dorsal view. 
<emphasis id="1F9C335BF404FFCE9A9DF8FFFF1DF8AD" bold="true" box="[189,209,1797,1823]" pageId="30" pageNumber="31">B</emphasis>
. Lateral view. 
<emphasis id="1F9C335BF404FFCE9BAAF8FFFE0FF8AD" bold="true" box="[394,451,1797,1823]" pageId="30" pageNumber="31">C–D</emphasis>
. Elytron. 
<emphasis id="1F9C335BF404FFCE981FF8FFFD98F8AD" bold="true" box="[575,596,1797,1823]" pageId="30" pageNumber="31">C</emphasis>
. Entire. 
<emphasis id="1F9C335BF404FFCE98E1F8FFFD1AF8AD" bold="true" box="[705,726,1797,1823]" pageId="30" pageNumber="31">D</emphasis>
. Apex. 
<emphasis id="1F9C335BF404FFCE991AF8FFFC80F8AD" bold="true" box="[826,844,1797,1823]" pageId="30" pageNumber="31">E</emphasis>
. Hind wing. 
<emphasis id="1F9C335BF404FFCE99D0F8FFFBE4F8AD" bold="true" box="[1008,1064,1797,1823]" pageId="30" pageNumber="31">F–H</emphasis>
. Legs. 
<emphasis id="1F9C335BF404FFCE9EA5F8FFFB5BF8AD" bold="true" box="[1157,1175,1797,1823]" pageId="30" pageNumber="31">F</emphasis>
. Fore. 
<emphasis id="1F9C335BF404FFCE9ED1F8FFFAC4F8AD" bold="true" box="[1265,1288,1797,1823]" pageId="30" pageNumber="31">G</emphasis>
. Middle. 
<emphasis id="1F9C335BF404FFCE9A9DF8D2FF18F8F0" bold="true" box="[189,212,1832,1858]" pageId="30" pageNumber="31">H</emphasis>
. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm.
</paragraph>
</caption>
<caption id="7997BFC1F405FFCF9A9DF947FC75F8F0" ID-DOI="http://doi.org/10.5281/zenodo.7033442" ID-Zenodo-Dep="7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1725,1751]" targetBox="[189,1398,265,1657]" targetPageId="31">
<paragraph id="2D57EF49F405FFCF9A9DF947FC75F8F0" blockId="31.[185,1399,1725,1858]" pageId="31" pageNumber="32">
<emphasis id="1F9C335BF405FFCF9A9DF947FED4F96A" bold="true" box="[189,280,1725,1752]" pageId="31" pageNumber="32">Fig. 19.</emphasis>
<taxonomicName id="EAE894CAF405FFCF9B3EF944FE34F96A" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[286,504,1726,1752]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF405FFCF9B3EF944FE34F96A" box="[286,504,1726,1752]" italics="true" pageId="31" pageNumber="32">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F405FFCF9BDEF944FD9AF96A" box="[510,598,1726,1752]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF405FFCF9879F947FD5EF965" bold="true" box="[601,658,1725,1751]" pageId="31" pageNumber="32">A–C</emphasis>
. Holotype, ♂ (CELC). 
<emphasis id="1F9C335BF405FFCF9981F947FC10F965" bold="true" box="[929,988,1725,1751]" pageId="31" pageNumber="32">D–G</emphasis>
. Paratype ♂ (CELC). 
<emphasis id="1F9C335BF405FFCF9EFBF947FB3CF965" bold="true" box="[1243,1264,1725,1751]" pageId="31" pageNumber="32">A</emphasis>
. Abdomen, dorsal view. 
<emphasis id="1F9C335BF405FFCF9B78F91BFEA0F949" bold="true" box="[344,364,1761,1787]" pageId="31" pageNumber="32">B</emphasis>
. Abdomen, ventral view. 
<emphasis id="1F9C335BF405FFCF9886F91BFD77F949" bold="true" box="[678,699,1761,1787]" pageId="31" pageNumber="32">C</emphasis>
. Ventrite 1. 
<emphasis id="1F9C335BF405FFCF9974F91BFCA5F949" bold="true" box="[852,873,1761,1787]" pageId="31" pageNumber="32">D</emphasis>
. Tergite VIII. 
<emphasis id="1F9C335BF405FFCF9E3BF91BFBE3F949" bold="true" box="[1051,1071,1761,1787]" pageId="31" pageNumber="32">E</emphasis>
. Terminalia. 
<emphasis id="1F9C335BF405FFCF9EF0F91BFB2EF949" bold="true" box="[1232,1250,1761,1787]" pageId="31" pageNumber="32">F</emphasis>
. Tergite IX. 
<emphasis id="1F9C335BF405FFCF9A9DF8FEFF18F8AC" bold="true" box="[189,212,1796,1822]" pageId="31" pageNumber="32">G</emphasis>
. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm.
</paragraph>
</caption>
<caption id="7997BFC1F43AFFF09A9DF8E6FC87F8CF" ID-DOI="http://doi.org/10.5281/zenodo.7033444" ID-Zenodo-Dep="7033444" httpUri="https://zenodo.org/record/7033444/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1820,1846]" targetBox="[240,1398,267,1761]" targetPageId="32">
<paragraph id="2D57EF49F43AFFF09A9DF8E6FC87F8CF" blockId="32.[189,1399,1820,1917]" pageId="32" pageNumber="33">
<emphasis id="1F9C335BF43AFFF09A9DF8E6FED0F884" bold="true" box="[189,284,1820,1846]" pageId="32" pageNumber="33">Fig. 20.</emphasis>
<taxonomicName id="EAE894CAF43AFFF09B05F8E7FDCFF884" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[293,515,1820,1847]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF43AFFF09B05F8E7FDCFF884" box="[293,515,1820,1847]" italics="true" pageId="32" pageNumber="33">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F43AFFF0982CF8E7FDA7F885" box="[524,619,1821,1847]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (CELC). 
<emphasis id="1F9C335BF43AFFF0995BF8E6FC78F884" bold="true" box="[891,948,1820,1846]" pageId="32" pageNumber="33">A–C</emphasis>
. Aedeagi. 
<emphasis id="1F9C335BF43AFFF09E15F8E6FB86F884" bold="true" box="[1077,1098,1820,1846]" pageId="32" pageNumber="33">A</emphasis>
. Lateral view. 
<emphasis id="1F9C335BF43AFFF09EDFF8E6FADDF884" bold="true" box="[1279,1297,1820,1846]" pageId="32" pageNumber="33">B</emphasis>
. Frontal view. 
<emphasis id="1F9C335BF43AFFF09B24F8C5FED5F8EB" bold="true" box="[260,281,1855,1881]" pageId="32" pageNumber="33">C</emphasis>
. Dorsal view. 
<emphasis id="1F9C335BF43AFFF09BE7F8BAFE31F8E8" bold="true" box="[455,509,1856,1882]" pageId="32" pageNumber="33">D–E</emphasis>
. Internal sac. 
<emphasis id="1F9C335BF43AFFF09886F8BAFD76F8E8" bold="true" box="[678,698,1856,1882]" pageId="32" pageNumber="33">D</emphasis>
. Frontal view. 
<emphasis id="1F9C335BF43AFFF0994FF8BAFC4EF8E8" bold="true" box="[879,898,1856,1882]" pageId="32" pageNumber="33">E</emphasis>
. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="2D57EF49F43BFFF19A9DFEF3FD13FE39" blockId="33.[186,1398,265,395]" pageId="33" pageNumber="34">
ABDOMEN. Tergite VI and VII with imbricate microsculpture (
<figureCitation id="B5D3F3CCF43BFFF19953FEF3FC11FE91" box="[883,989,265,291]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19A</figureCitation>
).Tergite VII trapezoidal (
<figureCitation id="B5D3F3CCF43BFFF19EDFFEF3FAA4FE91" box="[1279,1384,265,291]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19A</figureCitation>
), triangular in some 
<typeStatus id="F25351EBF43BFFF19BB9FED6FDC5FEF4" box="[409,521,300,326]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
with tergite VIII not exposed; punctation sparse, fine; pubescence sparse, fine. Ventrites 1–5 sparsely and finely punctate; pubescence sparse, fine; with strigulate microsculpture (
<figureCitation id="B5D3F3CCF43BFFF19AE6FE8BFE98FE39" box="[198,340,369,395]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19B–C</figureCitation>
). Metacoxal lines finely punctate.
</paragraph>
<paragraph id="2D57EF49F43BFFF19ACAFE48FEF9FE7E" blockId="33.[187,1399,434,738]" box="[234,309,434,460]" pageId="33" pageNumber="34">
<emphasis id="1F9C335BF43BFFF19ACAFE48FEF9FE7E" bold="true" box="[234,309,434,460]" pageId="33" pageNumber="34">Males</emphasis>
</paragraph>
<paragraph id="2D57EF49F43BFFF19A9DFE2FFCACFD53" blockId="33.[187,1399,434,738]" pageId="33" pageNumber="34">
MEASUREMENTS (n = 1, 
<typeStatus id="F25351EBF43BFFF19BCDFE2FFD9BFE5D" box="[493,599,469,495]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
; in mm). Antennomeres (length(width)): 0.14(0.05), 0.10(0.04), 0.08(0.03), 0.07(0.04), 0.06(0.04), 0.06(0.05), 0.09(0.08), 0.08(0.08), 0.09(0.10), 0.09(0.11), 0.14(0.10); (n = 8, including the 
<typeStatus id="F25351EBF43BFFF19B8FFDE0FDD5FD86" box="[431,537,538,564]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
, unless otherwise specified; in mm): TL 2.07–2.32 (mean = 2.18, standard deviation ± 0.08), PL 0.74–0.80 (0.78 ± 0.02), PA 0.66–0.70 (0.69 ± 0.02), PB 1.30–1.40 (1.36 ± 0.04), SL (n = 7) 0.07–0.15 (0.11 ± 0.02), SW (n = 7) 0.09–0.14 (0.11 ± 0.02), EI 1.24–1.36 (1.30 ± 0.04), EL 1.44–1.62 (1.53 ± 0.08), EW 0.76–0.86 (0.81 ± 0.03), EH 0.50–0.62 (0.55 ± 0.04), HW 0.59–0.65 (0.62 ± 0.02), IS 0.15–0.20 (0.17 ± 0.02), WA 0.12–0.19 (0.14 ± 0.02), MC 0.55–0.64 (0.60 ± 0.03), MB 0.23–0.30 (0.27 ± 0.02), VL 0.40–0.49 (0.45 ± 0.03).
</paragraph>
<paragraph id="2D57EF49F43BFFF19A9DFCF2FC78FC40" blockId="33.[187,1399,776,1011]" pageId="33" pageNumber="34">
Antennae shorter than in females; club less distinct than in females (
<figureCitation id="B5D3F3CCF43BFFF1999DFCF2FBE9FC91" box="[957,1061,776,803]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="33" pageNumber="34">Fig. 15B</figureCitation>
). Pro- and mesotarsomeres I– III enlarged, with tenet setae (
<figureCitation id="B5D3F3CCF43BFFF19835FCD1FD6DFCF7" box="[533,673,811,837]" captionStart="Fig" captionStartId="30.[189,232,1761,1787]" captionTargetBox="[189,1393,275,1713]" captionTargetId="figure-17@30.[189,1399,265,1713]" captionTargetPageId="30" captionText="Fig. 18. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033440" httpUri="https://zenodo.org/record/7033440/files/figure.png" pageId="33" pageNumber="34">Fig. 18F–G</figureCitation>
). Tergite VIII heptagonal, acuminated posteriorly; punctuation inconspicuous; subglabrous (
<figureCitation id="B5D3F3CCF43BFFF19830FCB7FDB1FCDA" box="[528,637,845,872]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19D</figureCitation>
). Tergite IX with rectangular ventral struts (
<figureCitation id="B5D3F3CCF43BFFF19E5DFCB7FACBFCDA" box="[1149,1287,845,872]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19E–F</figureCitation>
). Sternite VIII rectangular, with a projection (
<figureCitation id="B5D3F3CCF43BFFF19850FC8AFD2DFC38" box="[624,737,880,906]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19G</figureCitation>
). Sternite IX straight, constricted centrally and thicker apically (
<figureCitation id="B5D3F3CCF43BFFF19B08FC69FE43FC1F" box="[296,399,915,941]" captionStart="Fig" captionStartId="31.[189,232,1725,1751]" captionTargetBox="[189,1398,265,1657]" captionTargetId="figure-11@31.[189,1398,265,1663]" captionTargetPageId="31" captionText="Fig. 19. Cyparium lescheni sp. nov.A–C. Holotype, ♂ (CELC). D–G. Paratype ♂ (CELC). A. Abdomen, dorsal view. B. Abdomen, ventral view. C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (A–C) and Mata da Biologia (D–G), Viçosa (MG, Brazil). Scale bars: A–B = 0.2 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033442" httpUri="https://zenodo.org/record/7033442/files/figure.png" pageId="33" pageNumber="34">Fig. 19F</figureCitation>
). Aedeagus sclerotized, apex of median lobe short; openings in dorsal view forming an acute angle (
<figureCitation id="B5D3F3CCF43BFFF19B70FC4FFE2EFC62" box="[336,482,949,976]" captionStart="Fig" captionStartId="32.[189,232,1820,1846]" captionTargetBox="[240,1398,267,1761]" captionTargetId="figure-16@32.[189,1398,267,1761]" captionTargetPageId="32" captionText="Fig. 20. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–C. Aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D–E. Internal sac. D. Frontal view. E. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033444" httpUri="https://zenodo.org/record/7033444/files/figure.png" pageId="33" pageNumber="34">Fig. 20A–C</figureCitation>
); internal sac with weak irregular sclerites, with drop-like sclerites (
<figureCitation id="B5D3F3CCF43BFFF19EDEFC4FFF1EFC40" captionStart="Fig" captionStartId="32.[189,232,1820,1846]" captionTargetBox="[240,1398,267,1761]" captionTargetId="figure-16@32.[189,1398,267,1761]" captionTargetPageId="32" captionText="Fig. 20. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–C. Aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D–E. Internal sac. D. Frontal view. E. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033444" httpUri="https://zenodo.org/record/7033444/files/figure.png" pageId="33" pageNumber="34">Fig. 20D– E</figureCitation>
); parameres short, enlarged apically in lateral view (
<figureCitation id="B5D3F3CCF43BFFF19919FC22FC6BFC40" box="[825,935,984,1010]" captionStart="Fig" captionStartId="32.[189,232,1820,1846]" captionTargetBox="[240,1398,267,1761]" captionTargetId="figure-16@32.[189,1398,267,1761]" captionTargetPageId="32" captionText="Fig. 20. Cyparium lescheni sp. nov., paratype, ♂ (CELC). A–C. Aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D–E. Internal sac. D. Frontal view. E. Dorsal view. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033444" httpUri="https://zenodo.org/record/7033444/files/figure.png" pageId="33" pageNumber="34">Fig. 20A</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F43BFFF19ACAFBE3FE9DFB81" blockId="33.[186,1399,1049,1318]" box="[234,337,1049,1075]" pageId="33" pageNumber="34">
<emphasis id="1F9C335BF43BFFF19ACAFBE3FE9DFB81" bold="true" box="[234,337,1049,1075]" pageId="33" pageNumber="34">Females</emphasis>
</paragraph>
<paragraph id="2D57EF49F43BFFF19A9DFBC6FA9FFA94" blockId="33.[186,1399,1049,1318]" pageId="33" pageNumber="34">
MEASUREMENTS (n= 1, 
<typeStatus id="F25351EBF43BFFF19B9AFBC6FDEEFBE4" box="[442,546,1084,1110]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
; in mm).Antennomeres (length(width)):0.14(0.05), 0.11(0.05), 0.11(0.04), 0.06(0.04), 0.06(0.05), 0.05(0.06), 0.10(0.11), 0.09(0.13), 0.07(0.13), 0.08(0.14), 0.12(0.12); (n = 7, 
<typeStatus id="F25351EBF43BFFF19A9DFB78FEFCFB2E" box="[189,304,1154,1180]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
; in mm): TL 2.10–2.35 (mean = 2.26, standard deviation ± 0.09), PL 0.76–0.86 (0.81 ± 0.04), PA 0.66–0.74 (0.71 ± 0.02), PB 1.34–1.50 (1.43 ± 0.06), SL 0.09–0.13 (0.11 ± 0.01), SW 0.12–0.14 (0.13 ± 0.01), EI 1.30–1.40 (1.35 ± 0.04), EL 1.48–1.66 (1.58 ± 0.06), EW 0.76–0.84 (0.81 ± 0.03), EH 0.52–0.65 (0.60 ± 0.04), HW 0.61–0.66 (0.63 ± 0.02), IS 0.18–0.20 (0.18 ± 0.01), WA 0.13–0.15 (0.14 ± 0.00), MC 0.57–0.65 (0.62 ± 0.03), MB 0.25–0.32 (0.30 ± 0.02), VL 0.43–0.52 (0.50 ± 0.03).
</paragraph>
<paragraph id="2D57EF49F43BFFF19A9DFAB7FD7BF9A6" blockId="33.[188,1400,1356,1556]" pageId="33" pageNumber="34">
Antennae larger than in males; antennal club more distinct (
<figureCitation id="B5D3F3CCF43BFFF199AAFAB6FC3BFAD5" box="[906,1015,1356,1383]" captionStart="Fig" captionStartId="27.[189,232,1886,1912]" captionTargetBox="[189,1398,964,1848]" captionTargetId="figure-226@27.[189,1398,964,1850]" captionTargetPageId="27" captionText="Fig. 15. Cyparium lescheni sp. nov. A. Holotype, ♂ (CELC), frontal view. B. Paratype, ♂ (CELC), antennae. C. Paratype, ♀ (CELC), antennae. D–G. Paratype, ♂ (CELC). D. Labrum. E. Left mandible. F. Right mandible. G. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 0.2 mm; D–G = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033432" httpUri="https://zenodo.org/record/7033432/files/figure.png" pageId="33" pageNumber="34">Fig. 15C</figureCitation>
). Tergite VIII hexagonal, with a posterior invagination; punctuation inconspicuous; subglabrous (
<figureCitation id="B5D3F3CCF43BFFF199E9FA95FBF5FA3B" box="[969,1081,1391,1417]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="33" pageNumber="34">Fig. 21A</figureCitation>
). Sternite VIII rectangular with a projection (
<figureCitation id="B5D3F3CCF43BFFF19BBCFA68FDC6FA1E" box="[412,522,1426,1452]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="33" pageNumber="34">Fig. 21B</figureCitation>
). Vagina membranous; bursa copulatrix with sclerites on wall (
<figureCitation id="B5D3F3CCF43BFFF19F25FA68FABDFA1E" box="[1285,1393,1426,1452]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="33" pageNumber="34">Fig. 21C</figureCitation>
, Supp. file 2A). Vaginal plate with an apical sclerite, more or less developed (
<figureCitation id="B5D3F3CCF43BFFF19E13FA4EFB0DFA7D" box="[1075,1217,1460,1487]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="33" pageNumber="34">Fig. 21C–D</figureCitation>
, Supp. file 2A). Two filiform spermatheca (Supp. file 2A). Distal gonocoxites straight, thin (
<figureCitation id="B5D3F3CCF43BFFF19E09FA2DFB79FA43" box="[1065,1205,1495,1521]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="33" pageNumber="34">Fig. 21C–E</figureCitation>
); gonostyli long, somewhat thick, and parallel (
<figureCitation id="B5D3F3CCF43BFFF1983DFA00FD66F9A6" box="[541,682,1530,1556]" captionStart="Fig" captionStartId="34.[189,232,1805,1831]" captionTargetBox="[208,1398,300,1761]" captionTargetId="figure-17@34.[189,1398,268,1762]" captionTargetPageId="34" captionText="Fig. 21. Cyparium lescheni sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="33" pageNumber="34">Fig. 21C–E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F43BFFF19A9DF9C1FE1DF959" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="2D57EF49F43BFFF19A9DF9C1FE8CF9E7" blockId="33.[189,320,1595,1621]" box="[189,320,1595,1621]" pageId="33" pageNumber="34">
<heading id="761F5825F43BFFF19A9DF9C1FE8CF9E7" bold="true" box="[189,320,1595,1621]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="1F9C335BF43BFFF19A9DF9C1FE8CF9E7" bold="true" box="[189,320,1595,1621]" pageId="33" pageNumber="34">Host fungi</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F43BFFF19A9DF993FE1DF959" blockId="33.[189,1399,1641,1771]" pageId="33" pageNumber="34">
Adults were collected from 
<taxonomicName id="EAE894CAF43BFFF19804F993FC2EF931" authority="(Fr.) Maire" authorityName="Maire" baseAuthorityName="Fr." box="[548,994,1641,1667]" class="Agaricomycetes" family="Psathyrellaceae" genus="Psathyrella" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="candolleana">
<emphasis id="1F9C335BF43BFFF19804F993FC86F931" box="[548,842,1641,1667]" italics="true" pageId="33" pageNumber="34">Psathyrella candolleana</emphasis>
(Fr.) Maire
</taxonomicName>
(
<taxonomicName id="EAE894CAF43BFFF199D9F993FBB7F931" box="[1017,1147,1641,1667]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
, 
<taxonomicName id="EAE894CAF43BFFF19EAFF993FA8BF931" box="[1167,1351,1641,1667]" class="Agaricomycetes" family="Psathyrellaceae" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="family">Psathyrellaceae</taxonomicName>
) (1 record, 1 individual), 
<taxonomicName id="EAE894CAF43BFFF19BE4F971FDBFF914" box="[452,627,1675,1702]" class="Agaricomycetes" family="Psathyrellaceae" genus="Psathyrella" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF43BFFF19BE4F971FD86F917" box="[452,586,1675,1701]" italics="true" pageId="33" pageNumber="34">Psathyrella</emphasis>
sp.
</taxonomicName>
(1, 11), 
<taxonomicName id="EAE894CAF43BFFF198FFF976FCBEF914" box="[735,882,1676,1702]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF43BFFF198FFF976FC85F914" box="[735,841,1676,1702]" italics="true" pageId="33" pageNumber="34">Agaricus</emphasis>
sp.
</taxonomicName>
(1, 18; von Groll 
<emphasis id="1F9C335BF43BFFF19E72F976FB41F917" box="[1106,1165,1675,1701]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
2021: figs 1–4), 
<taxonomicName id="EAE894CAF43BFFF19F7DF976FE6BF97A" authority="Schulzer" authorityName="Schulzer" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="dulcidulus">
<emphasis id="1F9C335BF43BFFF19F7DF976FEFBF97A" italics="true" pageId="33" pageNumber="34">A. dulcidulus</emphasis>
Schulzer
</taxonomicName>
(1, 4), 
<taxonomicName id="EAE894CAF43BFFF19820F955FD32F97A" authority="Schaeff." authorityName="Schaeff." box="[512,766,1710,1737]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="sylvaticus">
<emphasis id="1F9C335BF43BFFF19820F955FD59F97A" box="[512,661,1710,1737]" italics="true" pageId="33" pageNumber="34">A. sylvaticus</emphasis>
Schaeff.
</taxonomicName>
(1, 20) and 
<taxonomicName id="EAE894CAF43BFFF199B8F954FB7AF97A" box="[920,1206,1710,1736]" class="Agaricomycetes" family="Entolomataceae" genus="Entoloma" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="subGenus" subGenus="Inocephalus">
<emphasis id="1F9C335BF43BFFF199B8F954FBC6F97A" box="[920,1034,1710,1736]" italics="true" pageId="33" pageNumber="34">Entoloma</emphasis>
(
<emphasis id="1F9C335BF43BFFF19E3DF954FB60F97A" box="[1053,1196,1710,1736]" italics="true" pageId="33" pageNumber="34">Inocephalus</emphasis>
)
</taxonomicName>
sp. (
<taxonomicName id="EAE894CAF43BFFF19ED0F954FABEF97A" box="[1264,1394,1710,1736]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
, 
<taxonomicName id="EAE894CAF43BFFF19A9EF92BFEB7F959" authorityName="Kotl. &amp; Pouzar" authorityYear="1972" box="[190,379,1745,1771]" class="Agaricomycetes" family="Entolomataceae" kingdom="Fungi" order="Agaricales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="family">Entolomataceae</taxonomicName>
) (1, 2).
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F43BFFF39A9DF8E8FBE8FEDA" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="discussion">
<paragraph id="2D57EF49F43BFFF19A9DF8E8FEE3F89E" blockId="33.[189,303,1810,1836]" box="[189,303,1810,1836]" pageId="33" pageNumber="34">
<heading id="761F5825F43BFFF19A9DF8E8FEE3F89E" bold="true" box="[189,303,1810,1836]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="1F9C335BF43BFFF19A9DF8E8FEE3F89E" bold="true" box="[189,303,1810,1836]" pageId="33" pageNumber="34">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F43BFFF39A9DF8BAFBE8FEDA" blockId="33.[189,1399,1856,2021]" lastBlockId="35.[189,1399,265,361]" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
Measurements similar to those of 
<taxonomicName id="EAE894CAF43BFFF1987EF8BAFD2EF8E8" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[606,738,1856,1882]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF43BFFF1987EF8BAFD2EF8E8" box="[606,738,1856,1882]" italics="true" pageId="33" pageNumber="34">C. newtoni</emphasis>
</taxonomicName>
, except for the comparatively longer antennae. It was found co-occurring with 
<taxonomicName id="EAE894CAF43BFFF19BCBF899FDA0F8CE" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[491,620,1891,1917]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF43BFFF19BCBF899FDA0F8CE" box="[491,620,1891,1917]" italics="true" pageId="33" pageNumber="34">C. newtoni</emphasis>
</taxonomicName>
in the same host fungi. However, they differ at first view by the coloration: 
<taxonomicName id="EAE894CAF43BFFF19B67F87FFE00F82D" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[327,460,1925,1951]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF43BFFF19B67F87FFE00F82D" box="[327,460,1925,1951]" italics="true" pageId="33" pageNumber="34">C. lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F43BFFF19BF7F87CFDF9F812" box="[471,565,1926,1952]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
is brown with few individuals of the same coloration as 
<taxonomicName id="EAE894CAF43BFFF19ED4F87FFABBF82D" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[1268,1399,1925,1951]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF43BFFF19ED4F87FFABBF82D" box="[1268,1399,1925,1951]" italics="true" pageId="33" pageNumber="34">C. newtoni</emphasis>
</taxonomicName>
(pronotum reddish brown and elytra black); a few individuals of 
<taxonomicName id="EAE894CAF43BFFF19998F852FBF4F870" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[952,1080,1960,1986]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF43BFFF19998F852FBF4F870" box="[952,1080,1960,1986]" italics="true" pageId="33" pageNumber="34">C. newtoni</emphasis>
</taxonomicName>
are brown, but darker than 
<taxonomicName id="EAE894CAF43BFFF19A9DF831FEF2F856" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[189,318,1994,2021]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="1F9C335BF43BFFF19A9DF831FEF2F856" box="[189,318,1994,2021]" italics="true" pageId="33" pageNumber="34">C. lescheni</emphasis>
</taxonomicName>
. They also differ by the microsculpture of the metaventrite: present in 
<taxonomicName id="EAE894CAF43BFFF19E52F831FB3FF856" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[1138,1267,1994,2021]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="1F9C335BF43BFFF19E52F831FB3FF856" box="[1138,1267,1994,2021]" italics="true" pageId="33" pageNumber="34">C. lescheni</emphasis>
</taxonomicName>
and absent in 
<taxonomicName id="EAE894CAF439FFF39AFFFEF3FEADFE91" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[223,353,265,291]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF439FFF39AFFFEF3FEADFE91" box="[223,353,265,291]" italics="true" pageId="35" pageNumber="36">C. newtoni</emphasis>
</taxonomicName>
. Tergite VIII in males is triangular in 
<taxonomicName id="EAE894CAF439FFF3991AFEF3FC0CFE91" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[826,960,265,291]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="1F9C335BF439FFF3991AFEF3FC0CFE91" box="[826,960,265,291]" italics="true" pageId="35" pageNumber="36">C. lescheni</emphasis>
</taxonomicName>
and straight apically in 
<taxonomicName id="EAE894CAF439FFF39ECCFEF3FAA3FE91" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[1260,1391,265,291]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF439FFF39ECCFEF3FAA3FE91" box="[1260,1391,265,291]" italics="true" pageId="35" pageNumber="36">C. newtoni</emphasis>
</taxonomicName>
. Parameres in 
<taxonomicName id="EAE894CAF439FFF39B40FED6FE2DFEF7" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[352,481,300,326]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF439FFF39B40FED6FE2DFEF7" box="[352,481,300,326]" italics="true" pageId="35" pageNumber="36">C. newtoni</emphasis>
</taxonomicName>
are longer and the sclerite of the internal sac is also different. Tergite VIII in females has an invagination in 
<taxonomicName id="EAE894CAF439FFF39809FEB5FD67FEDA" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[553,683,334,361]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="1F9C335BF439FFF39809FEB5FD67FEDA" box="[553,683,334,361]" italics="true" pageId="35" pageNumber="36">C. lescheni</emphasis>
</taxonomicName>
, while in 
<taxonomicName id="EAE894CAF439FFF39901FEB5FC6CFEDA" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[801,928,335,361]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="1F9C335BF439FFF39901FEB5FC6CFEDA" box="[801,928,335,361]" italics="true" pageId="35" pageNumber="36">C. newtoni</emphasis>
</taxonomicName>
it is entire.
</paragraph>
</subSubSection>
<caption id="7997BFC1F438FFF29A9DF8F7FCAEF8DC" ID-DOI="http://doi.org/10.5281/zenodo.7033447" ID-Zenodo-Dep="7033447" httpUri="https://zenodo.org/record/7033447/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1805,1831]" targetBox="[208,1398,300,1761]" targetPageId="34">
<paragraph id="2D57EF49F438FFF29A9DF8F7FCAEF8DC" blockId="34.[186,1399,1805,1902]" pageId="34" pageNumber="35">
<emphasis id="1F9C335BF438FFF29A9DF8F7FED4F895" bold="true" box="[189,280,1805,1831]" pageId="34" pageNumber="35">Fig. 21.</emphasis>
<taxonomicName id="EAE894CAF438FFF29B3DF8F4FE34F895" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[285,504,1805,1832]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1F9C335BF438FFF29B3DF8F4FE34F895" box="[285,504,1805,1832]" italics="true" pageId="34" pageNumber="35">Cyparium lescheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F438FFF29BDDF8F4FD94F89A" box="[509,600,1806,1832]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (CELC). 
<emphasis id="1F9C335BF438FFF29979F8F7FCA2F895" bold="true" box="[857,878,1805,1831]" pageId="34" pageNumber="35">A</emphasis>
. Tergite VIII. 
<emphasis id="1F9C335BF438FFF29E2EF8F7FBEEF895" bold="true" box="[1038,1058,1805,1831]" pageId="34" pageNumber="35">B</emphasis>
. Sternite VIII. 
<emphasis id="1F9C335BF438FFF29EEDF8F7FB2EF895" bold="true" box="[1229,1250,1805,1831]" pageId="34" pageNumber="35">C</emphasis>
. Terminalia. 
<emphasis id="1F9C335BF438FFF29A9AF8CAFF03F8F8" bold="true" box="[186,207,1840,1866]" pageId="34" pageNumber="35">D</emphasis>
. Sclerite of vaginal plate. 
<emphasis id="1F9C335BF438FFF29BDEF8CAFDDEF8F8" bold="true" box="[510,530,1840,1866]" pageId="34" pageNumber="35">E</emphasis>
. Ovipositor. Specimen collected at Mata da Biologia, Viçosa (MG, Brazil). Scale bars: A–B, E = 0.1 mm; C = 0.2 mm; D = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="65F2BCC2F439FFF39A9DFE75FC95FE49" pageId="35" pageNumber="36" type="distribution">
<paragraph id="2D57EF49F439FFF39A9DFE75FE9AFE1B" blockId="35.[189,342,399,425]" box="[189,342,399,425]" pageId="35" pageNumber="36">
<heading id="761F5825F439FFF39A9DFE75FE9AFE1B" bold="true" box="[189,342,399,425]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="1F9C335BF439FFF39A9DFE75FE9AFE1B" bold="true" box="[189,342,399,425]" pageId="35" pageNumber="36">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F439FFF39A9DFE45FC95FE49" blockId="35.[187,1398,446,508]" pageId="35" pageNumber="36">
Known only from Mata da Biologia and Mata do Paraíso, campus of the Universidade Federal de Viçosa, Viçosa, state of 
<collectingRegion id="EF2C21ABF439FFF39B57FE1BFDDBFE49" box="[375,535,481,507]" country="Brazil" name="Minas Gerais" pageId="35" pageNumber="36">Minas Gerais</collectingRegion>
, Southeast 
<collectingCountry id="55FFAFD9F439FFF398BCFE1BFD28FE49" box="[668,740,481,507]" name="Brazil" pageId="35" pageNumber="36">Brazil</collectingCountry>
(
<figureCitation id="B5D3F3CCF439FFF398D4FE1BFC80FE49" box="[756,844,481,507]" captionStart="Fig" captionStartId="65.[189,232,1952,1978]" captionTargetBox="[189,1398,1082,1910]" captionTargetId="figure-297@65.[189,1398,1082,1910]" captionTargetPageId="65" captionText="Fig. 46. A. Distribution map of species of Cyparium Erichson, 1845, South America. B. Mata da Biologia. C. Mata do Paraíso." figureDoi="http://doi.org/10.5281/zenodo.7033501" httpUri="https://zenodo.org/record/7033501/files/figure.png" pageId="35" pageNumber="36">Fig. 46</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>