377 lines
50 KiB
XML
377 lines
50 KiB
XML
<document id="8BC83444087F8710AC9BEB9C4B8595FD" ID-CLB-Dataset="25370" ID-DOI="10.11646/zootaxa.4674.4.4" ID-GBIF-Dataset="1e1be7bb-5fbe-4720-8846-ffbd532b6bb2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3465030" ID-ZooBank="BD3A0EE3-1F60-468B-9793-5E5EE55C6EBE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569830421185" checkinUser="plazi" docAuthor="Sokolov, Sergey G. & Gordeev, Ilya I." docDate="2019" docId="9B3EE44FFFFF944CEABB8CE06A7F6923" docLanguage="en" docName="zootaxa.4674.4.4.pdf" docOrigin="Zootaxa 4674 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asaccotrema vietnamiense Sokolov & Gordeev 2019, n. sp." docType="treatment" docUuid="23247B89-2433-4CAD-9247-7648C9AFF36D" docUuidSource="ZooBank" docVersion="11" lastPageNumber="455" masterDocId="67079C37FFFD9449EA2C8F256813683C" masterDocTitle="Asaccotrema vietnamiense n. gen.; n. sp. (Trematoda: Monorchioidea), a new aberrant representative of lissorchiid trematodes from the sidestripe rasbora, Rasbora paviana Tirant (Actinopterygii: Cyprinidae), Vietnam" masterLastPageNumber="462" masterPageNumber="451" pageNumber="453" updateTime="1698756485094" updateUser="ExternalLinkService">
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<mods:mods id="A9E9DD2BE564E04782D489C1B6D572F0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1DE2130A4807FBAF02729353315D9146">
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<mods:title id="2A92AFE9A100ED1A571035DD93650E3D">Asaccotrema vietnamiense n. gen.; n. sp. (Trematoda: Monorchioidea), a new aberrant representative of lissorchiid trematodes from the sidestripe rasbora, Rasbora paviana Tirant (Actinopterygii: Cyprinidae), Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="AF288E22E0F55F17BC21C985F8067642" type="personal">
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<mods:role id="A08217CF9C876B6437DA1B4FEC6D5F97">
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<mods:roleTerm id="207DA398FD3651296924A376EA4F09FC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2AC2676B117400DDFFDB279A68575741">Sokolov, Sergey G.</mods:namePart>
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</mods:name>
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<mods:name id="67EA89FA694CCFFF63C5CCB19A8D2F19" type="personal">
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<mods:role id="3920640B704308FF640E68E9740DA6A2">
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<mods:roleTerm id="778B52E7A06575FE862CF9F49A93126E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B426958E7DA07739B0798063C3A5800B">Gordeev, Ilya I.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FDD68E6143DFB05979F925196262F244">text</mods:typeOfResource>
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<mods:titleInfo id="FC2FA7E4C345CEF16F7FDB097A847133">
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<mods:title id="C99D1056B9597F96EE0A728CD45B3CAB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="70D4F64139BE0E96498824A83F24A5B1">
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<mods:date id="842404414DE8FE44B9E4B622D6AD894F">2019</mods:date>
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<mods:detail id="9119B73EC5F1FEB78AAFD3D6D0D55732" type="pubDate">
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<mods:number id="4F3ED7D9AE719FAFA1583DE0F619196A">2019-09-26</mods:number>
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</mods:detail>
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<mods:detail id="50FF6FF3E15AE36891543D64080F19FF" type="volume">
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<mods:number id="07AFA05B798F71ABC6FDB8176E2F39D3">4674</mods:number>
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</mods:detail>
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<mods:detail id="35196AF87C18F4D7EF1BA916805E3AA1" type="issue">
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<mods:number id="1632FE0DBD20FCB1C658829C6E1DF40A">4</mods:number>
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<mods:classification id="DD90DF9B9970DD5C60A0F558754B3E05">journal article</mods:classification>
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<mods:identifier id="8E0444AF555C041E0AE49F4FA825C32F" type="CLB-Dataset">25370</mods:identifier>
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<mods:identifier id="FCB748971AA2FA4E67FFCB738A22F4D7" type="DOI">10.11646/zootaxa.4674.4.4</mods:identifier>
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<mods:identifier id="74FB16B5FD72FAA7BA9EF10210E16E48" type="GBIF-Dataset">1e1be7bb-5fbe-4720-8846-ffbd532b6bb2</mods:identifier>
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<mods:identifier id="C8352DB125A5E52934E2F4010F3C87D5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9B152324C377F81E825A6AF00B7BD74C" type="Zenodo-Dep">3465030</mods:identifier>
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<mods:identifier id="54A41BCA6071B184EAEB442CF4C45359" type="ZooBank">BD3A0EE3-1F60-468B-9793-5E5EE55C6EBE</mods:identifier>
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</mods:mods>
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<treatment id="9B3EE44FFFFF944CEABB8CE06A7F6923" ID-DOI="http://doi.org/10.5281/zenodo.5619668" ID-GBIF-Taxon="159608994" ID-Zenodo-Dep="5619668" LSID="urn:lsid:zoobank.org:act:23247B89-2433-4CAD-9247-7648C9AFF36D" httpUri="http://treatment.plazi.org/id/9B3EE44FFFFF944CEABB8CE06A7F6923" lastPageId="5" lastPageNumber="455" pageId="2" pageNumber="453">
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<subSubSection id="5B8D06D2FFFF944BEABB8CE06BAD6C3F" pageId="2" pageNumber="453" type="nomenclature">
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<paragraph id="13285559FFFF944BEABB8CE06A306BDC" blockId="2.[151,958,965,1063]" box="[151,547,965,992]" pageId="2" pageNumber="453">
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<heading id="4860E235FFFF944BEABB8CE06A306BDC" bold="true" box="[151,547,965,992]" fontSize="11" level="1" pageId="2" pageNumber="453" reason="1">
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<emphasis id="21E3894BFFFF944BEABB8CE06A306BDC" bold="true" box="[151,547,965,992]" pageId="2" pageNumber="453">
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<taxonomicName id="D4972EDAFFFF944BEABB8CE069CA6BE3" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[151,473,965,991]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
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<emphasis id="21E3894BFFFF944BEABB8CE069CA6BE3" bold="true" box="[151,473,965,991]" italics="true" pageId="2" pageNumber="453">Asaccotrema vietnamiense</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3AD03430FFFF944BEBCC8CE36A306BDC" box="[480,547,966,992]" pageId="2" pageNumber="453" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="13285559FFFF944BEABB8CCC6BAD6C3F" blockId="2.[151,958,965,1063]" box="[151,958,1001,1027]" pageId="2" pageNumber="453">
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<uri id="6706595BFFFF944BEABB8CCC6BAD6C3F" box="[151,958,1001,1027]" pageId="2" pageNumber="453">
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urn:lsid:zoobank.org:act:
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<uuid id="67316F8CFFFF944BEB8B8CCC6BAD6C3F" box="[423,958,1001,1027]" pageId="2" pageNumber="453">23247B89-2433-4CAD-9247-7648C9AFF36D</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="5B8D06D2FFFF944BEABB8B2868F86C1B" box="[151,235,1037,1063]" pageId="2" pageNumber="453" type="description">
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<paragraph id="13285559FFFF944BEABB8B2868F86C1B" blockId="2.[151,958,965,1063]" box="[151,235,1037,1063]" pageId="2" pageNumber="453">
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(
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<figureCitation id="8BAC49DCFFFF944BEA8C8B2868F36C1B" box="[160,224,1037,1063]" captionStart="FIGURE 1" captionStartId="3.[151,250,1874,1899]" captionTargetBox="[251,1343,528,1843]" captionTargetId="figure@3.[235,1352,517,1854]" captionTargetPageId="3" captionText="FIGURE 1 A–C. Asaccotrema vietnamiense n. sp. A. Holotype, ventral view. B. Terminal genitalia, sinistro-lateral view. C. Ovarian complex, dorso-lateral view. Abbreviations: cg, cluster of prostate gland-cells; ej, ejaculatory duct; ga, common genital atrium; lc, Laurer’s, canal; mod, middle portion of oviduct; mt, metraterm; o, ovary; ot, ootype with Mehlis’ gland; ov, ovicapt; pp, pars prostatica; pod, proximal portion of oviduct; s, sphincter; sr, seminal receptacle; sv, seminal vesicle; tm, thin-walled membrane; vd, common vitelline reservoir and common vitelline duct. Scale bars: A, 0.4 mm; B, C, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3467867" httpUri="https://zenodo.org/record/3467867/files/figure.png" pageId="2" pageNumber="453">Fig. 1</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5B8D06D2FFFF944BEABB8B736A426DB3" pageId="2" pageNumber="453" type="materials_examined">
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||
<paragraph id="13285559FFFF944BEABB8B736B666C4C" blockId="2.[151,1437,1109,2036]" box="[151,885,1109,1136]" pageId="2" pageNumber="453">
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<emphasis id="21E3894BFFFF944BEABB8B7369076C4C" bold="true" box="[151,276,1110,1136]" pageId="2" pageNumber="453">
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||
<typeStatus id="CC2CEBFBFFFF944BEABB8B7368C16C4C" box="[151,210,1110,1136]" pageId="2" pageNumber="453">Type</typeStatus>
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||
host:
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||
</emphasis>
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<emphasis id="21E3894BFFFF944BEB378B7069C86C53" box="[283,475,1109,1135]" italics="true" pageId="2" pageNumber="453">Rasbora paviana</emphasis>
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Tirant (Actinopterygii:
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<taxonomicName id="D4972EDAFFFF944BE8CB8B706B7A6C4C" box="[743,873,1109,1136]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="453" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
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).
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</paragraph>
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<paragraph id="13285559FFFF944BEAEB8B5F6D426CA8" blockId="2.[151,1437,1109,2036]" box="[199,1361,1145,1172]" pageId="2" pageNumber="453">
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<materialsCitation id="A3FF5F04FFFF944BEAEB8B5F6D5E6CA8" ID-GBIF-Occurrence="2423311035" box="[199,1357,1145,1172]" collectorName="Da Brout" country="Vietnam" latitude="11.442778" location="Cat Tien National Park" longLatPrecision="21" longitude="107.42916" pageId="2" pageNumber="453" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="21E3894BFFFF944BEAEB8B5F697A6CA8" bold="true" box="[199,361,1145,1172]" pageId="2" pageNumber="453">
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||
<typeStatus id="CC2CEBFBFFFF944BEAEB8B5F69116CA8" box="[199,258,1146,1172]" pageId="2" pageNumber="453">Type</typeStatus>
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locality:
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||
</emphasis>
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The brook
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<collectorName id="BE62308FFFFF944BEBC78B5F6A416CA8" box="[491,594,1146,1172]" pageId="2" pageNumber="453">Da Brout</collectorName>
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in the
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<location id="16480382FFFF944BE8B38B5C6BB06CAF" LSID="urn:lsid:plazi:treatment:9B3EE44FFFFF944CEABB8CE06A7F6923:16480382FFFF944BE8B38B5C6BB06CAF" box="[671,931,1145,1171]" country="Vietnam" latitude="11.442778" longLatPrecision="21" longitude="107.42916" name="Cat Tien National Park" pageId="2" pageNumber="453">Cat Tien National Park</location>
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,
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<collectingCountry id="6B8015C9FFFF944BE9828B5C6C1E6CAF" box="[942,1037,1145,1171]" name="Vietnam" pageId="2" pageNumber="453">Vietnam</collectingCountry>
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||
(
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||
<geoCoordinate id="76A3339EFFFF944BEE308B5C6CB66CA8" box="[1052,1189,1145,1172]" degrees="11" direction="north" minutes="26" orientation="latitude" pageId="2" pageNumber="453" precision="15" seconds="34" value="11.442778">11°26′34″N</geoCoordinate>
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||
;
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||
<geoCoordinate id="76A3339EFFFF944BEE9C8B5C6D566CA8" box="[1200,1349,1145,1172]" degrees="107" direction="east" minutes="25" orientation="longitude" pageId="2" pageNumber="453" precision="15" seconds="45" value="107.42916">107°25′45″E</geoCoordinate>
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||
)
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||
</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="13285559FFFF944BEAEB8BB869C66C8B" blockId="2.[151,1437,1109,2036]" box="[199,469,1181,1208]" pageId="2" pageNumber="453">
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<emphasis id="21E3894BFFFF944BEAEB8BB869986C84" bold="true" box="[199,395,1181,1208]" pageId="2" pageNumber="453">Site of infection:</emphasis>
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Liver.
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</paragraph>
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<paragraph id="13285559FFFF944BEAEB8BE46C8B6CC3" blockId="2.[151,1437,1109,2036]" pageId="2" pageNumber="453">
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<emphasis id="21E3894BFFFF944BEAEB8BE469A06CE0" bold="true" box="[199,435,1217,1244]" pageId="2" pageNumber="453">Deposited material:</emphasis>
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||
<materialsCitation id="A3FF5F04FFFF944BEB968BE46C8B6CC3" ID-GBIF-Occurrence="2423311034" collectionCode="IPEE" pageId="2" pageNumber="453" specimenCode="RAS #1314, RAS #1314-1316" specimenCount="2" typeStatus="holotype">
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<typeStatus id="CC2CEBFBFFFF944BEB968BE46A316CE7" box="[442,546,1217,1243]" pageId="2" pageNumber="453" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="7586CD9CFFFF944BE81C8BE46A7E6CE7" box="[560,621,1217,1243]" country="Russia" httpUri="http://grbio.org/cool/q8g7-dfji" name="A.N. Severtzov Institute of Ecology and Evolution, Russian Academy of Sciences" pageId="2" pageNumber="453">IPEE</collectionCode>
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<specimenCode id="4331FD22FFFF944BE8588BE46AEA6CE7" box="[628,761,1217,1243]" collectionCode="IPEE" country="Russia" httpUri="http://grbio.org/cool/q8g7-dfji" name="A.N. Severtzov Institute of Ecology and Evolution, Russian Academy of Sciences" pageId="2" pageNumber="453">RAS #1314</specimenCode>
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) and
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<typeStatus id="CC2CEBFBFFFF944BE91B8BE76BB16CE0" box="[823,930,1218,1244]" pageId="2" pageNumber="453" type="paratype">paratypes</typeStatus>
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(3)
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<collectionCode id="7586CD9CFFFF944BE9FD8BE76C1F6CE0" box="[977,1036,1218,1244]" country="Russia" httpUri="http://grbio.org/cool/q8g7-dfji" name="A.N. Severtzov Institute of Ecology and Evolution, Russian Academy of Sciences" pageId="2" pageNumber="453">IPEE</collectionCode>
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<specimenCode id="4331FD22FFFF944BEE3F8BE46CCF6CE7" box="[1043,1244,1217,1243]" collectionCode="IPEE" country="Russia" httpUri="http://grbio.org/cool/q8g7-dfji" name="A.N. Severtzov Institute of Ecology and Evolution, Russian Academy of Sciences" pageId="2" pageNumber="453">
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RAS #
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<date id="67297399FFFF944BEE738BE46CCF6CE7" box="[1119,1244,1217,1243]" pageId="2" pageNumber="453" value="1314" valueMax="1316">1314–1316</date>
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</specimenCode>
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(whole-mounted excysted progenetic metacercariae), v
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<emphasis id="21E3894BFFFF944BE8128BC06B3D6D3C" box="[574,814,1253,1280]" italics="true" pageId="2" pageNumber="453">oucher specimens (3)</emphasis>
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with isolated terminal genitalia.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="13285559FFFF944BEAEB8A2F6AC46D7B" blockId="2.[151,1437,1109,2036]" pageId="2" pageNumber="453">
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||
<emphasis id="21E3894BFFFF944BEAEB8A2F6A446D18" bold="true" box="[199,599,1289,1316]" pageId="2" pageNumber="453">Zoobank Life Science Identifier</emphasis>
|
||
: (
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||
<collectionCode id="7586CD9CFFFF944BE8588A2C6AA66D1F" box="[628,693,1289,1315]" pageId="2" pageNumber="453">LSID</collectionCode>
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||
) for
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||
<taxonomicName id="D4972EDAFFFF944BE8D58A2F6C396D1F" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[761,1066,1290,1316]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFFF944BE8D58A2F6C396D1F" box="[761,1066,1290,1316]" italics="true" pageId="2" pageNumber="453">Asaccotrema vietnamiense</emphasis>
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</taxonomicName>
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||
<emphasis id="21E3894BFFFF944BEE148A2F6C6D6D18" bold="true" box="[1080,1150,1290,1316]" pageId="2" pageNumber="453">
|
||
<taxonomicNameLabel id="3AD03430FFFF944BEE148A2F6C6D6D18" box="[1080,1150,1290,1316]" pageId="2" pageNumber="453" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is
|
||
<uri id="6706595BFFFF944BEE808A2C6AC16D7B" pageId="2" pageNumber="453">
|
||
urn:lsid:zoobank.org: act:
|
||
<uuid id="67316F8CFFFF944BEA918A086AC16D7B" box="[189,722,1325,1351]" pageId="2" pageNumber="453">23247B89-2433-4CAD-9247-7648C9AFF36D</uuid>
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||
</uri>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="13285559FFFF944BEAEB8A7769CB6D57" blockId="2.[151,1437,1109,2036]" box="[199,472,1361,1388]" pageId="2" pageNumber="453">
|
||
<emphasis id="21E3894BFFFF944BEAEB8A7769406D50" bold="true" box="[199,339,1362,1388]" pageId="2" pageNumber="453">Prevalence:</emphasis>
|
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1 of 1 host.
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||
</paragraph>
|
||
<paragraph id="13285559FFFF944BEAEB8A506A426DB3" blockId="2.[151,1437,1109,2036]" box="[199,593,1397,1424]" pageId="2" pageNumber="453">
|
||
<emphasis id="21E3894BFFFF944BEAEB8A5069296DAC" bold="true" box="[199,314,1397,1424]" pageId="2" pageNumber="453">Intensity:</emphasis>
|
||
8 worms/host specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B8D06D2FFFF944AEAEB8ABC699D69CB" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453" type="description">
|
||
<paragraph id="13285559FFFF944AEAEB8ABC699D69CB" blockId="2.[151,1437,1109,2036]" lastBlockId="3.[151,1437,153,503]" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453">
|
||
<emphasis id="21E3894BFFFF944BEAEB8ABC69486D88" bold="true" box="[199,347,1433,1460]" pageId="2" pageNumber="453">Description:</emphasis>
|
||
(Based on
|
||
<specimenCount id="05919ED0FFFF944BEBCC8ABF6A7E6D88" box="[480,621,1433,1460]" pageId="2" pageNumber="453" type="generic">7 specimens</specimenCount>
|
||
). Progenetic metacercariae. Body elongate-oval, length 775–875 [800], maximum width 275–350 [288] in middle third of body. Forebody 33.3–38.7 [34.4]% of body length, occasionally (in one of four specimens) 45.7 %. Tegument spinous. Oral sucker rounded, 125–138 × 138–150 [125 × 138]. Ventral sucker rounded, 125–150 × 138–163 [125 × 138]. Oral sucker to ventral sucker width ratio 1: 0.92–1.08 [1: 1]. Mouth opening subterminal. Prepharynx very short. Pharynx muscular, 55–61 × 49–61 [55 × 49], esophagus 122–159 [128]; intestinal bifurcation at level of anterior part of ventral sucker or just anterior to anterior edge of ventral sucker, occasionally (in one of four specimens) at some distance anterior to ventral sucker. Ceca extending to anterior end of testis. Testis single, slightly irregular, median, 73–153 × 55–67 [92 × 67], postovarian, in middle part of hindbody. Cirrus-sac absent. Seminal vesicle saccular proximally and tubular distally, surrounded by numerous prostate gland-cells; extending to ovary, occasionally (in one of four specimens) to midlevel of ventral sucker. Pars prostatica tubular, not clearly separated from seminal vesicle, penetrated by ducts of prostate gland-cells, and terminating with sphincter. Main mass of prostate gland-cells lies free in parenchyma. Terminal portion of prostate gland-cells cluster, which includes ducts of prostate gland-cells and distal gland-cells, covered with thin-walled open-ended membrane. Distal end of membrane connected to proximal edge of wall of ejaculatory duct. Ejaculatory duct long, covered with gland-cells; communicating with common genital atrium. Inner cytoplasmic layer of wall of ejaculatory duct forms numerous finger-like protrusions. Common genital atrium distinct. Genital pore dextrosublateral, at about of midlevel of esophagus. Ovary entire, transverse-oval, median, 31–49 × 61–73 [43 × 61], in middle part of hindbody, contiguous with testis, exceptionally (in one of four specimens) at level of posterior edge of ventral sucker and distinctly separated from testis. Muscular ovicapt distinct. Laurer’s canal opens in dorsal pore at level of ovary. Canalicular seminal receptacle conspicuous, sacсular. Oviduct connects to common vitelline duct before becoming oötype. Common vitelline reservoir ventral to ovary. Oötype enclosed in Mehlis’ gland opens into uterus. Uterine seminal receptacle not detected. Vitellarium follicular; subglobular follicles in two short lateral preovarian fields of 7–9 each, lying in anterior half of hindbody, exceptionally (in one of four specimens) overlapping ovary and ventral sucker to some extent. Uterine coils between ventral sucker and posterior end of testis. Metraterm well-developed, ventral to ejaculatory duct, rather dilated and filled with numerous eggs. Eggs numerous, ovate, operculate, 15 × 10–12. Excretory vesicle tubular, extending to anterior edge of testis; excretory pore subterminal at posterior extremity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="47E805D1FFFE944AEABB88776CE06FC6" ID-DOI="http://doi.org/10.5281/zenodo.3467867" ID-Zenodo-Dep="3467867" httpUri="https://zenodo.org/record/3467867/files/figure.png" pageId="3" pageNumber="454" startId="3.[151,250,1874,1899]" targetBox="[251,1343,528,1843]" targetPageId="3">
|
||
<paragraph id="13285559FFFE944AEABB88776CE06FC6" blockId="3.[151,1436,1874,2043]" pageId="3" pageNumber="454">
|
||
<emphasis id="21E3894BFFFE944AEABB8877695F6F57" bold="true" box="[151,332,1874,1899]" pageId="3" pageNumber="454">FIGURE 1 A–C.</emphasis>
|
||
<taxonomicName id="D4972EDAFFFE944AEB7888766A706F56" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[340,611,1875,1898]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="454" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFFE944AEB7888766A706F56" box="[340,611,1875,1898]" italics="true" pageId="3" pageNumber="454">Asaccotrema vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFFE944AE84788776AD66F57" bold="true" box="[619,709,1874,1899]" pageId="3" pageNumber="454">
|
||
<taxonomicNameLabel id="3AD03430FFFE944AE84788776AB56F56" box="[619,678,1874,1898]" pageId="3" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
|
||
A.
|
||
</emphasis>
|
||
Holotype, ventral view.
|
||
<emphasis id="21E3894BFFFE944AE9E988776BCF6F56" bold="true" box="[965,988,1874,1898]" pageId="3" pageNumber="454">B.</emphasis>
|
||
Terminal genitalia, sinistro-lateral view.
|
||
<emphasis id="21E3894BFFFE944AEFA888776D8F6F57" bold="true" box="[1412,1436,1874,1899]" pageId="3" pageNumber="454">C.</emphasis>
|
||
Ovarian complex, dorso-lateral view. Abbreviations: cg, cluster of prostate gland-cells; ej, ejaculatory duct; ga, common genital atrium; lc, Laurer’s, canal; mod, middle portion of oviduct; mt, metraterm; o, ovary; ot, ootype with Mehlis’ gland; ov, ovicapt; pp, pars prostatica; pod, proximal portion of oviduct; s, sphincter; sr, seminal receptacle; sv, seminal vesicle; tm, thin-walled membrane; vd, common vitelline reservoir and common vitelline duct. Scale bars: A, 0.4 mm; B, C, 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="5B8D06D2FFF9944DEAEB8FBC69C368EB" pageId="4" pageNumber="455" type="etymology">
|
||
<paragraph id="13285559FFF9944DEAEB8FBC69C368EB" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="455">
|
||
<emphasis id="21E3894BFFF9944DEAEB8FBC6941688F" bold="true" box="[199,338,153,179]" pageId="4" pageNumber="455">Etymology:</emphasis>
|
||
The specific epithet ‘
|
||
<taxonomicName id="D4972EDAFFF9944DE8658FBF6AC8688F" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[585,731,154,179]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense">
|
||
<emphasis id="21E3894BFFF9944DE8658FBF6AC8688F" box="[585,731,154,179]" italics="true" pageId="4" pageNumber="455">vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
’ is intended to mark the country (
|
||
<collectingCountry id="6B8015C9FFF9944DEE708FBC6CAC6888" box="[1116,1215,153,180]" name="Vietnam" pageId="4" pageNumber="455">Vietnam</collectingCountry>
|
||
) where this species was found for the first time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B8D06D2FFF9944CEAEB8FC46A7F6923" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455" type="discussion">
|
||
<paragraph id="13285559FFF9944DEAEB8FC469D6695B" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="455">
|
||
<emphasis id="21E3894BFFF9944DEAEB8FC4692E68C7" bold="true" box="[199,317,225,251]" pageId="4" pageNumber="455">Remarks:</emphasis>
|
||
Many features, like living in a freshwater environment, the spinous tegument, the sublateral genital pore, the follicular vitellarium organized in two short symmetrical fields, rather short intestinal branches, the presence of one testis and its position allow us to place
|
||
<taxonomicName id="D4972EDAFFF9944DE8DF8E0C6BB5697F" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[755,934,297,323]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFF9944DE8DF8E0C6BB5697F" box="[755,934,297,323]" italics="true" pageId="4" pageNumber="455">A. vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF9944DE99C8E0C6BE2697F" bold="true" box="[944,1009,297,323]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DE99C8E0C6BE2697F" box="[944,1009,297,323]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the subfamily
|
||
<taxonomicName id="D4972EDAFFF9944DEE938E0C6D8A697F" authorityName="Szidat" authorityYear="1943" box="[1215,1433,297,323]" class="Trematoda" family="Monorchiidae" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="subFamily" subFamily="Asymphylodorinae">Asymphylodorinae</taxonomicName>
|
||
,
|
||
<taxonomicName id="D4972EDAFFF9944DEABB8E6869D3695B" authority="(Bray 2008)" authorityName="Bray" authorityYear="2008" baseAuthorityName="Bray" baseAuthorityYear="2008" box="[151,448,333,359]" class="Trematoda" family="Lissorchiidae" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="family">
|
||
Lissorchiidae (
|
||
<bibRefCitation id="770628A8FFF9944DEB118E6869AA695B" author="Bray, R. A." box="[317,441,333,359]" pageId="4" pageNumber="455" pagination="177 - 186" refId="ref5568" refString="Bray, R. A. (2008) Family Lissorchiidae Magath, 1917. In: Bray, R. A., Gibson, D. I. & Jones, A. (Eds.), Keys to the Trematoda. Vol. 3. CAB International and The Natural History Museum, Wallingford, pp. 177 - 186. https: // doi. org / 10.2478 / s 11686 - 009 - 0021 - 7" type="book chapter" year="2008">Bray 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="13285559FFF9944DEAEB8E546D7B6A27" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="455">
|
||
This subfamily is currently composed of four genera:
|
||
<taxonomicName id="D4972EDAFFF9944DE9068E546C7869B7" authority="Looss, 1899" authorityName="Looss" authorityYear="1899" box="[810,1131,369,395]" class="Trematoda" family="Monorchiidae" genus="Asymphylodora" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="21E3894BFFF9944DE9068E546BC969B7" box="[810,986,369,395]" italics="true" pageId="4" pageNumber="455">Asymphylodora</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DE9CE8E546C7869B7" author="Looss, A." box="[994,1131,369,395]" pageId="4" pageNumber="455" pagination="521 - 784" refId="ref7246" refString="Looss, A. (1899) Weitere Beitrage zur Kenntniss der Trematoden-Fauna Aegyptens, zugleich Versuch einer naturlichen Gliederung des Genus Distomum Retzius. Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere, 12, 521 - 784. [in German] https: // doi. org / 10.5962 / bhl. part. 2037" type="journal article" year="1899">Looss, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
s. lato,
|
||
<taxonomicName id="D4972EDAFFF9944DEEEA8E5469346993" authority="Gupta, 1955" authorityName="Gupta" authorityYear="1955" class="Trematoda" family="Lissorchiidae" genus="Brahamputrotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="21E3894BFFF9944DEEEA8E546D8F69B7" box="[1222,1436,369,395]" italics="true" pageId="4" pageNumber="455">Brahamputrotrema</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DEABB8EB069346993" author="Gupta, S. P." box="[151,295,404,431]" pageId="4" pageNumber="455" pagination="1 - 80" refId="ref6553" refString="Gupta, S. P. (1955) Trematode parasites of fresh water fishes. Indian Journal of Helminthology, 5, 1 - 80." type="journal article" year="1955">Gupta, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="21E3894BFFF9944DEB198EB069C06993" box="[309,467,405,431]" italics="true" pageId="4" pageNumber="455">Prosovitellina</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DEBF08EB06A756993" author="Wang, X." box="[476,614,404,431]" pageId="4" pageNumber="455" pagination="1 - 10" refId="ref8808" refString="Wang, X. - y. (1985) On trematodes of Monorchiidae Odhner, 1911, and the establishment of one new subfamily and two new genera. Jiangxi Science, 2, 1 - 10. [in Chinese]" type="journal article" year="1985">Wang, 1985</bibRefCitation>
|
||
and
|
||
<taxonomicName id="D4972EDAFFF9944DE88E8EB36C406993" authority="Bray, 2008" authorityName="Bray" authorityYear="2008" box="[674,1107,405,431]" class="Trematoda" family="Lissorchiidae" genus="Wangxiyunia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="21E3894BFFF9944DE88E8EB36B206993" box="[674,819,406,431]" italics="true" pageId="4" pageNumber="455">Wangxiyunia</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DE9118EB06BA86993" author="Bray, R. A." box="[829,955,405,431]" pageId="4" pageNumber="455" pagination="177 - 186" refId="ref5568" refString="Bray, R. A. (2008) Family Lissorchiidae Magath, 1917. In: Bray, R. A., Gibson, D. I. & Jones, A. (Eds.), Keys to the Trematoda. Vol. 3. CAB International and The Natural History Museum, Wallingford, pp. 177 - 186. https: // doi. org / 10.2478 / s 11686 - 009 - 0021 - 7" type="book chapter" year="2008">Bray, 2008</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="770628A8FFF9944DE9E08EB06C586993" author="Bray, R. A." box="[972,1099,405,431]" pageId="4" pageNumber="455" pagination="177 - 186" refId="ref5568" refString="Bray, R. A. (2008) Family Lissorchiidae Magath, 1917. In: Bray, R. A., Gibson, D. I. & Jones, A. (Eds.), Keys to the Trematoda. Vol. 3. CAB International and The Natural History Museum, Wallingford, pp. 177 - 186. https: // doi. org / 10.2478 / s 11686 - 009 - 0021 - 7" type="book chapter" year="2008">Bray 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Representatives of all these genera, like all the other lissorchiids (members of the subfamily
|
||
<taxonomicName id="D4972EDAFFF9944DE9548E9C6C0769E8" authorityName="Magath" authorityYear="1917" box="[888,1044,441,468]" class="Trematoda" family="Monorchiidae" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="subFamily" subFamily="Lissorchiinae">Lissorchiinae</taxonomicName>
|
||
), have a well-developed cirrus-sac and their genital pore is positioned approximately at the level of the ventral sucker. In contrast,
|
||
<taxonomicName id="D4972EDAFFF9944DEE9B8EF868F66A27" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFF9944DEE9B8EF868F66A27" italics="true" pageId="4" pageNumber="455">Asaccotrema vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF9944DEAC08D2469386A27" bold="true" box="[236,299,513,539]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DEAC08D2469386A27" box="[236,299,513,539]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
does not have a cirrus-sac, and the genital pore is positioned near the midlevel of the esophagus.
|
||
</paragraph>
|
||
<paragraph id="13285559FFF9944DEAEB8D016AB46ABB" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="455">
|
||
<emphasis id="21E3894BFFF9944DEAEB8D0169DF6A03" bold="true" box="[199,460,548,575]" pageId="4" pageNumber="455">Phylogenetic position,</emphasis>
|
||
PCR amplification of the
|
||
<emphasis id="21E3894BFFF9944DE8DF8D006B766A03" box="[755,869,549,575]" italics="true" pageId="4" pageNumber="455">28S rRNA</emphasis>
|
||
gene produced a 1300-bp fragment for the
|
||
<taxonomicName id="D4972EDAFFF9944DEF678D0069126A5F" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFF9944DEF678D0069126A5F" italics="true" pageId="4" pageNumber="455">A. vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF9944DEB248D6C69556A5F" bold="true" box="[264,326,585,611]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DEB248D6C69556A5F" box="[264,326,585,611]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimen. After assembly and alignment procedures were carried out, the resulting
|
||
<emphasis id="21E3894BFFF9944DEEDC8D6C6D726A5F" box="[1264,1377,585,611]" italics="true" pageId="4" pageNumber="455">28S rRNA</emphasis>
|
||
gene sequence from these species was 1135 bp long.
|
||
</paragraph>
|
||
<paragraph id="13285559FFF9944DEAEB8DB46ADC6B2B" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="455">
|
||
In both the ML and BI analyses,
|
||
<taxonomicName id="D4972EDAFFF9944DE8108DB46AFD6A97" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[572,750,657,683]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFF9944DE8108DB46AFD6A97" box="[572,750,657,683]" italics="true" pageId="4" pageNumber="455">A. vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF9944DE8DA8DB46B266A97" bold="true" box="[758,821,657,683]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DE8DA8DB46B266A97" box="[758,821,657,683]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears as a member of the well-supported
|
||
<taxonomicName id="D4972EDAFFF9944DEF078DB468D56AF3" baseAuthorityName="Bray" baseAuthorityYear="2008" class="Trematoda" family="Lissorchiidae" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="family">Lissorchiidae</taxonomicName>
|
||
clade, which also includes
|
||
<taxonomicName id="D4972EDAFFF9944DEBD08D906BF86AF3" authority="Barnhart & Powell, 1979" authorityName="Barnhart & Powell" authorityYear="1979" box="[508,1003,693,719]" class="Trematoda" family="Lissorchiidae" genus="Lissorchis" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="species" species="kritskyi">
|
||
<emphasis id="21E3894BFFF9944DEBD08D906ADB6AF3" box="[508,712,693,719]" italics="true" pageId="4" pageNumber="455">Lissorchis kritskyi</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DE8FC8D906BF86AF3" author="Barnhart, M. C. & Powell, E. C." box="[720,1003,693,719]" pageId="4" pageNumber="455" pagination="47 - 51" refId="ref5293" refString="Barnhart, M. C. & Powell, E. C. (1979) Lissorchis kritskyi sp. n. (Digenea: Lissorchiidae) from the River Carpsucker, Carpiodes carpio (Rafinesque). Proceedings of the Helminthological Society of Washington, 46, 47 - 51." type="journal article" year="1979">Barnhart & Powell, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D4972EDAFFF9944DEE0E8D906A5F6ACF" authority="Besprozvanykh, Ermolenko & Atopkin, 2012" authorityName="Besprozvanykh, Ermolenko & Atopkin" authorityYear="2012" class="Trematoda" family="Monorchiidae" genus="Asymphylodora" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="species" species="perccotti">
|
||
<emphasis id="21E3894BFFF9944DEE0E8D906D2E6AF3" box="[1058,1341,693,719]" italics="true" pageId="4" pageNumber="455">Asymphylodora perccotti</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DEF688D906A5F6ACF" author="Besprozvanykh, V. V & Ermolenko, A. V. & Atopkin, D. M." pageId="4" pageNumber="455" pagination="235 - 241" refId="ref5394" refString="Besprozvanykh, V. V, Ermolenko, A. V. & Atopkin, D. M. (2012) The life cycle of Asymphylodora perccotti sp n. (Trematoda: Lissorchiidae) in the Russian Southern Far East. Parasitology International, 61, 235 - 241. https: // doi. org / 10.1016 / j. parint. 2011.10.001" type="journal article" year="2012">Besprozvanykh, Ermolenko & Atopkin, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="8BAC49DCFFF9944DE8728DFC6AB66ACF" box="[606,677,729,755]" captionStart="FIGURE 2" captionStartId="4.[151,250,1565,1590]" captionTargetBox="[157,1430,807,1539]" captionTargetId="figure@4.[151,1436,805,1545]" captionTargetPageId="4" captionText="FIGURE 2. Phylogram showing the phylogenetic relationships of Asaccotrema vietnamiense n. sp. based on the analysis of the 28S rRNA gene partial sequences dataset. Node labels display ML/BI support." figureDoi="http://doi.org/10.5281/zenodo.3467869" httpUri="https://zenodo.org/record/3467869/files/figure.png" pageId="4" pageNumber="455">Fig. 2</figureCitation>
|
||
). However, the phylogenetic relationships between representatives within the
|
||
<taxonomicName id="D4972EDAFFF9944DEB238DD869B56B2B" baseAuthorityName="Bray" baseAuthorityYear="2008" box="[271,422,765,791]" class="Trematoda" family="Lissorchiidae" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="family">Lissorchiidae</taxonomicName>
|
||
clade are poorly resolved.
|
||
</paragraph>
|
||
<caption id="47E805D1FFF9944DEABB89386BBC6E65" ID-DOI="http://doi.org/10.5281/zenodo.3467869" ID-Zenodo-Dep="3467869" httpUri="https://zenodo.org/record/3467869/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,250,1565,1590]" targetBox="[157,1430,807,1539]" targetPageId="4">
|
||
<paragraph id="13285559FFF9944DEABB89386BBC6E65" blockId="4.[151,1436,1565,1626]" pageId="4" pageNumber="455">
|
||
<emphasis id="21E3894BFFF9944DEABB893869016E0A" bold="true" box="[151,274,1565,1590]" pageId="4" pageNumber="455">FIGURE 2.</emphasis>
|
||
Phylogram showing the phylogenetic relationships of
|
||
<taxonomicName id="D4972EDAFFF9944DE91B893B6C576E09" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[823,1092,1566,1589]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFF9944DE91B893B6C576E09" box="[823,1092,1566,1589]" italics="true" pageId="4" pageNumber="455">Asaccotrema vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF9944DEE6689386C906E09" bold="true" box="[1098,1155,1565,1589]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DEE6689386C906E09" box="[1098,1155,1565,1589]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
based on the analysis of the
|
||
<emphasis id="21E3894BFFF9944DEABB896468ED6E65" box="[151,254,1601,1625]" italics="true" pageId="4" pageNumber="455">28S rRNA</emphasis>
|
||
gene partial sequences dataset. Node labels display ML/BI support.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="13285559FFF9944CEAEB89A26A7F6923" blockId="4.[151,1437,1671,2022]" lastBlockId="5.[151,1437,153,287]" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455">
|
||
<emphasis id="21E3894BFFF9944DEAEB89A269566E9E" bold="true" box="[199,325,1671,1698]" pageId="4" pageNumber="455">Discussion</emphasis>
|
||
. Among the lissorchiids progenesis at the metacercarial stage is not a unique feature of the species of concern, since this phenomenon is a characteristic of a number of representatives of the genus
|
||
<taxonomicName id="D4972EDAFFF9944DEEC0898969336ED6" authority="Looss, 1899" authorityName="Sokolov & Gordeev" authorityYear="2019" class="Trematoda" family="Monorchiidae" genus="Asymphylodora" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="genus" sensu="lato">
|
||
<emphasis id="21E3894BFFF9944DEEC089896D8F6EFA" box="[1260,1436,1708,1734]" italics="true" pageId="4" pageNumber="455">Asymphylodora</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DEABB89F569336ED6" author="Looss, A." box="[151,288,1744,1770]" pageId="4" pageNumber="455" pagination="521 - 784" refId="ref7246" refString="Looss, A. (1899) Weitere Beitrage zur Kenntniss der Trematoden-Fauna Aegyptens, zugleich Versuch einer naturlichen Gliederung des Genus Distomum Retzius. Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere, 12, 521 - 784. [in German] https: // doi. org / 10.5962 / bhl. part. 2037" type="journal article" year="1899">Looss, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF9944DEB0B89F5698C6ED6" box="[295,415,1744,1770]" italics="true" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DEB0B89F569886ED6" box="[295,411,1744,1770]" pageId="4" pageNumber="455" sensu="lato">sensu lato</taxonomicNameLabel>
|
||
.
|
||
</emphasis>
|
||
The terminal genitalia and other reproductive organs of progenetic metacercariae of
|
||
<taxonomicName id="D4972EDAFFF9944DEF7A89F569196F32" authorityName="Sokolov & Gordeev" authorityYear="2019" class="Trematoda" family="Monorchiidae" genus="Asymphylodora" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="21E3894BFFF9944DEF7A89F569196F32" italics="true" pageId="4" pageNumber="455">Asymphylodora</emphasis>
|
||
</taxonomicName>
|
||
spp. are morphologically identical to those in the adult stage (Serkova & Bychowsky 1940;
|
||
<bibRefCitation id="770628A8FFF9944DEF3F89D168C26F0E" author="Biguet, J. & Deblock, S. & Capron, A." pageId="4" pageNumber="455" pagination="525 - 542" refId="ref5460" refString="Biguet, J., Deblock, S. & Capron, A. (1956) Description d'une metacercaire progenetique du genre Asymphylodora Loos, 1899, decouverte chez Bythinia leachi Sheppard dans le nord de la France. Annales de Parasitologie Humaine et Comparee, 31, 525 - 542. [in French] https: // doi. org / 10.1051 / parasite / 1956315525" type="journal article" year="1956">
|
||
Biguet
|
||
<emphasis id="21E3894BFFF9944DEF4889D168C26F0E" italics="true" pageId="4" pageNumber="455">et al. 1956</emphasis>
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DEAF3883D69956F0E" author="Stunkard, H. W." box="[223,390,1815,1842]" pageId="4" pageNumber="455" pagination="562 - 581" refId="ref8361" refString="Stunkard, H. W. (1959) The morphology and life-history of the digenetic trematode, Asymphylodora amnicolae n. sp.; the possible significance of progenesis for the phylogeny of the Digenea. Biological Bulletin, 117, 562 - 581. https: // doi. org / 10.2307 / 1538867" type="journal article" year="1959">Stunkard 1959</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DEBBE883D6AAE6F0D" author="Goodman, J. D. & Panesar, T. S." box="[402,701,1815,1842]" pageId="4" pageNumber="455" pagination="204 - 209" refId="ref6488" refString="Goodman, J. D. & Panesar, T. S. (1976) Parasymphylodora manteri n. sp., an adult trematode from a fresh-water pulmonate snail, Biomphalaria choanomphala choanomphala, from Lake Victoria, Uganda. Transactions of the American Microscopical Society, 95, 204 - 209. https: // doi. org / 10.2307 / 3225065" type="journal article" year="1976">Goodman & Panesar 1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DE8E5883D6B646F0E" author="Kulakova, A. P." box="[713,887,1816,1842]" pageId="4" pageNumber="455" pagination="78 - 81" refId="ref6922" refString="Kulakova, A. P. (1982) New data on Asymphylodora progenetica - Parasymphylodora progenetica (Trematoda, Monorchidae). Parasitologiya, 16, 78 - 81. [in Russian]" type="journal article" year="1982">Kulakova 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DE9AF883D6C1D6F0E" author="Kudlai, O. S." box="[899,1038,1816,1842]" pageId="4" pageNumber="455" pagination="559 - 562" refId="ref6873" refString="Kudlai, O. S. (2010) The first record of Asymphylodora progenetica (Trematoda, Monorchiidae) in Ukraine. Vestnik zoologii, 44, 559 - 562. https: // doi. org / 10.2478 / v 10058 - 010 - 0036 - 4" type="journal article" year="2010">Kudlai 2010</bibRefCitation>
|
||
<emphasis id="21E3894BFFF9944DEE22883D6C0C6F0E" box="[1038,1055,1816,1842]" italics="true" pageId="4" pageNumber="455">).</emphasis>
|
||
The terminal genitalia in the nonprogenetic metacercariae of
|
||
<taxonomicName id="D4972EDAFFF9944DEBFE88196A916F6A" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[466,642,1852,1878]" class="Trematoda" family="Monorchiidae" genus="Asymphylodora" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="21E3894BFFF9944DEBFE88196A916F6A" box="[466,642,1852,1878]" italics="true" pageId="4" pageNumber="455">Asymphylodora</emphasis>
|
||
</taxonomicName>
|
||
spp. and other lissorchiids are non-functioning, but structurally comparable with the those of the adult worms
|
||
<emphasis id="21E3894BFFF9944DE87688456A706F46" box="[602,611,1888,1914]" italics="true" pageId="4" pageNumber="455">(</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DE84888456B726F46" author="Macy, R. W. & English R. G." box="[612,865,1887,1914]" pageId="4" pageNumber="455" pagination="509 - 512" refId="ref7374" refString="Macy, R. W. & English R. G. (1975) On the life cycle of Palaeorchis problematicus Macy and Berntzen (n. comb.) (Trematoda: Monorchiidae) from Oregon. American Midland Naturalist, 94, 509 - 512. https: // doi. org / 10.2307 / 2424449" type="journal article" year="1975">Macy & English 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DE94288456CC06F46" author="Filimonova, L. V. & Shalyapina, V. I." box="[878,1235,1888,1914]" pageId="4" pageNumber="455" pagination="157 - 166" refId="ref6446" refString="Filimonova, L. V. & Shalyapina, V. I. (1979) Metacercariae in the snails Bithynia inflata in Severnaya Kulunda lakes. Trudy Gelminthologicheskoi laboratorii AN SSSR, 29, 157 - 166 [in Russian]" type="journal article" year="1979">Filimonova & Shalyapina 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DEECC884568C16FA2" author="Besprozvannykh, V. V." pageId="4" pageNumber="455" pagination="346 - 351" refId="ref5345" refString="Besprozvannykh, V. V. (2005) Life-cycles of trematods Parasymphilodora japonica (Yamaguti, 1938) and P. markewitschi (Kulakowskaja, 1947) (Monorcidae) in conditions of Primorsky Region. Parasitologiya, 39, 346 - 351. [in Russian]" type="journal article" year="2005">Besprozvannykh 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="770628A8FFF9944DEAF188A1696E6FA1" author="Shimazu, T." box="[221,381,1923,1950]" pageId="4" pageNumber="455" pagination="1 - 22" refId="ref8241" refString="Shimazu, T. (2016) Digeneans parasitic in freshwater fishes (Osteichthyes) of Japan VI. Lissorchiidae. Bulletin of the National Museum of Nature and Science, Series A, 42, 1 - 22." type="journal article" year="2016">Shimazu 2016</bibRefCitation>
|
||
<emphasis id="21E3894BFFF9944DEB5188A169956FA2" box="[381,390,1924,1950]" italics="true" pageId="4" pageNumber="455">)</emphasis>
|
||
. Metacercariae of
|
||
<taxonomicName id="D4972EDAFFF9944DE87A88A16BEE6FA2" authority="Schell, 1973" authorityName="Schell" authorityYear="1973" box="[598,1021,1924,1950]" class="Trematoda" family="Lissorchiidae" genus="Neopaleorchis" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="455" phylum="Platyhelminthes" rank="species" species="catostomi">
|
||
<emphasis id="21E3894BFFF9944DE87A88A16B786FA1" box="[598,875,1924,1950]" italics="true" pageId="4" pageNumber="455">Neopaleorchis catostomi</emphasis>
|
||
<bibRefCitation id="770628A8FFF9944DE95D88A16BEE6FA2" author="Schell, S. C." box="[881,1021,1924,1950]" pageId="4" pageNumber="455" pagination="463 - 468" refId="ref7944" refString="Schell, S. C. (1973) The life history of Neopaleorchis catostomi gen. et sp. n. (Trematoda: Monorchiidae), an intestinal parasite of the Coarsescale sucker, Catostomus macrocheilus Girard. Journal of Parasitology, 59, 463 - 468. https: // doi. org / 10.2307 / 3278773" type="journal article" year="1973">Schell, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
are an exception, but only because in this species the cirrus-sac and metraterm are present as poorly differentiated primordia (
|
||
<bibRefCitation id="770628A8FFF9944DEE4A888D6CE36FFE" author="Schell, S. C." box="[1126,1264,1960,1986]" pageId="4" pageNumber="455" pagination="463 - 468" refId="ref7944" refString="Schell, S. C. (1973) The life history of Neopaleorchis catostomi gen. et sp. n. (Trematoda: Monorchiidae), an intestinal parasite of the Coarsescale sucker, Catostomus macrocheilus Girard. Journal of Parasitology, 59, 463 - 468. https: // doi. org / 10.2307 / 3278773" type="journal article" year="1973">Schell 1973</bibRefCitation>
|
||
). In view of the above-mentioned facts, we have no doubt that the
|
||
<emphasis id="21E3894BFFF9944DE8E488E96B2C6FD9" box="[712,831,1996,2021]" italics="true" pageId="4" pageNumber="455">progenetic</emphasis>
|
||
metacercariae
|
||
<emphasis id="21E3894BFFF9944DE9CB88E96CA76FD9" box="[999,1204,1996,2022]" italics="true" pageId="4" pageNumber="455">
|
||
of
|
||
<taxonomicName id="D4972EDAFFF9944DEE2F88E96CA76FD9" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[1027,1204,1996,2022]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="455" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">A. vietnamiense</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="21E3894BFFF9944DEE9788E96CEA6FDA" bold="true" box="[1211,1273,1996,2022]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="3AD03430FFF9944DEE9788E96CEA6FDA" box="[1211,1273,1996,2022]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
demonstrate a definitive morphology for the terminal genitalia, typical of the adult hermaphroditic generation of this species. Such a morphology with respect to the terminal genitalia, unique for lissorchiids, allows us to describe these specimens not only as a new species, but as a new genus.
|
||
<emphasis id="21E3894BFFF8944CE8868FC46AC768C7" box="[682,724,225,251]" italics="true" pageId="5" pageNumber="456">The</emphasis>
|
||
description
|
||
<taxonomicName id="D4972EDAFFF8944CE94C8FC46C0168C7" authority="Sokolov & Gordeev, 2019" authorityName="Sokolov & Gordeev" authorityYear="2019" box="[864,1042,225,251]" class="Actinopterygii" family="Cyprinidae" genus="Asaccotrema" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="456" phylum="Chordata" rank="species" species="vietnamiense" status="sp. nov.">
|
||
<emphasis id="21E3894BFFF8944CE94C8FC46C0168C7" box="[864,1042,225,251]" italics="true" pageId="5" pageNumber="456">A. vietnamiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21E3894BFFF8944CEE358FC46C4A68C7" bold="true" box="[1049,1113,225,251]" pageId="5" pageNumber="456">
|
||
<taxonomicNameLabel id="3AD03430FFF8944CEE358FC46C4A68C7" box="[1049,1113,225,251]" pageId="5" pageNumber="456" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
means an amendment to the diagnosis of the
|
||
<taxonomicName id="D4972EDAFFF8944CEB7C8E2069F46923" baseAuthorityName="Bray" baseAuthorityYear="2008" box="[336,487,261,287]" class="Trematoda" family="Lissorchiidae" kingdom="Animalia" order="Plagiorchiida" pageId="5" pageNumber="456" phylum="Platyhelminthes" rank="family">Lissorchiidae</taxonomicName>
|
||
is required.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |