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<document id="279232BFBD8F9B05636FDA82399E32A5" ID-DOI="10.5281/zenodo.211829" ID-GBIF-Dataset="0a2ef110-4ceb-4b6c-b660-8a711540f219" ID-ISSN="1175-5326" ID-Zenodo-Dep="211829" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460488580763" checkinUser="plazi" docAuthor="Kamitani, Satoshi, Higuchi, Toshiyuki &amp; Okudera, Shigeru" docDate="2012" docId="584D3F2DFFE0FFFF92DFFF575745C8FA" docLanguage="en" docName="zt03343p015.pdf" docOrigin="Zootaxa 3343" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pagaronia spinosa Y.Hori 1994" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="A4744755FFECFFF39248FFC05357CB22" masterDocTitle="Taxonomic study of the protecta - group in the genus Pagaronia Ball (Hemiptera, Cicadellidae, Evacanthinae) from Japan" masterLastPageNumber="15" masterPageNumber="1" pageNumber="13" updateTime="1698309644170" updateUser="plazi">
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<mods:title id="67242F714EEA7ECC2673041341A2D396">Taxonomic study of the protecta - group in the genus Pagaronia Ball (Hemiptera, Cicadellidae, Evacanthinae) from Japan</mods:title>
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<mods:namePart id="9D75F6EAF7B3BE636F2EBA753C029B39">Kamitani, Satoshi</mods:namePart>
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<mods:namePart id="1755019211B1540435B5B8E9F7609D7B">Higuchi, Toshiyuki</mods:namePart>
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<mods:namePart id="0ED5A56FEDA455A644DC998B34B06110">Okudera, Shigeru</mods:namePart>
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<treatment id="584D3F2DFFE0FFFF92DFFF575745C8FA" ID-DOI="http://doi.org/10.5281/zenodo.6493223" ID-GBIF-Taxon="119519036" ID-Zenodo-Dep="6493223" LSID="urn:lsid:plazi:treatment:584D3F2DFFE0FFFF92DFFF575745C8FA" httpUri="http://treatment.plazi.org/id/584D3F2DFFE0FFFF92DFFF575745C8FA" lastPageNumber="13" pageId="12" pageNumber="13">
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<paragraph id="D05B8E3BFFE0FFFF92DFFF575149CB93" blockId="12.[151,542,151,210]" box="[151,542,151,177]" pageId="12" pageNumber="13">
<heading id="8B133957FFE0FFFF92DFFF575149CB93" bold="true" box="[151,542,151,177]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="17E4F5B8FFE0FFFF92DFFF575149CB93" ID-CoL="8JXDH" authority="Y.Hori, 1994" authorityName="Y.Hori" authorityYear="1994" box="[151,542,151,177]" class="Insecta" family="Cicadellidae" genus="Pagaronia" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E2905229FFE0FFFF92DFFF575149CB93" bold="true" box="[151,542,151,177]" pageId="12" pageNumber="13">
<emphasis id="E2905229FFE0FFFF92DFFF57522FCB93" bold="true" box="[151,376,151,177]" italics="true" pageId="12" pageNumber="13">Pagaronia spinosa</emphasis>
<bibRefCitation id="B475F3CAFFE0FFFF93C8FF575149CB93" author="Hori" box="[384,542,151,177]" pageId="12" pageNumber="13" refString="Hori, Y. (1994) Three new species of the genus Pagaronia from Mt. Ena-san and Mt. Chausu-yama, central Japan (Homoptera: Cicadellidae). Transactions of the Shikoku Entomological Society, 20, 155 - 160." type="journal article" year="1994">Y.Hori, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D05B8E3BFFE0FFFF92DFFF7A521DCBF0" blockId="12.[151,542,151,210]" box="[151,330,186,210]" pageId="12" pageNumber="13">
(
<figureCitation id="48DF92BEFFE0FFFF92D7FF7A53B8CBF0" box="[159,239,186,210]" captionStart="FIGURE 4" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[198,1422,706,1872]" captionTargetId="figure@2.[151,1436,661,1926]" captionTargetPageId="2" captionText="FIGURE 4. Distributional map of the Pagaronia species belong to the protecta - group." httpUri="https://zenodo.org/record/211831/files/figure.png" pageId="12" pageNumber="13">Figs. 4</figureCitation>
,
<figureCitation id="48DF92BEFFE0FFFF92B2FF7A5216CBF0" box="[250,321,186,210]" captionStart="FIGURES 55 57" captionStartId="12.[151,264,1882,1905]" captionTargetBox="[166,1431,1069,1828]" captionTargetId="figure@12.[151,1436,1054,1845]" captionTargetPageId="12" captionText="FIGURES 55 57. Pagaronia spinosa, ɗ genitalia (55 57). — 55, pygofer in lateral view; 56 57, left inner pygofer process. Scales; 0.3 mm." httpUri="https://zenodo.org/record/211839/files/figure.png" pageId="12" pageNumber="13">5557</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="98FEDDB0FFE0FFFF92DFFEC35752C98F" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="D05B8E3BFFE0FFFF92DFFEC35121CA3A" blockId="12.[151,630,258,280]" box="[151,630,258,280]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="F0F4A915FFE0FFFF92DFFEC35121CA3A" box="[151,630,258,280]" pageId="12" pageNumber="13">
<taxonomicName id="17E4F5B8FFE0FFFF92DFFEC352A8CA3A" ID-CoL="8JXDH" authority="Hori, 1994: 156" authorityName="Hori" authorityPageNumber="156" authorityYear="1994" box="[151,511,258,280]" class="Insecta" family="Cicadellidae" genus="Pagaronia" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E2905229FFE0FFFF92DFFEC35201CA3A" box="[151,342,259,280]" italics="true" pageId="12" pageNumber="13">Pagaronia spinosa</emphasis>
<treatmentCitation id="5145A82AFFE0FFFF9314FEC252A8CA3A" author="Hori" box="[348,511,258,280]" page="156" pageId="12" pageNumber="13" year="1994">
<bibRefCitation id="B475F3CAFFE0FFFF9314FEC25299CA3A" author="Hori" box="[348,462,258,280]" pageId="12" pageNumber="13" refString="Hori, Y. (1994) Three new species of the genus Pagaronia from Mt. Ena-san and Mt. Chausu-yama, central Japan (Homoptera: Cicadellidae). Transactions of the Shikoku Entomological Society, 20, 155 - 160." type="journal article" year="1994">Hori, 1994</bibRefCitation>
: 156
</treatmentCitation>
</taxonomicName>
, figs. 814.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D05B8E3BFFE0FFFF92DFFE855254C9A5" blockId="12.[151,1436,325,984]" pageId="12" pageNumber="13">
<emphasis id="E2905229FFE0FFFF92DFFE855235CA7F" bold="true" box="[151,354,325,350]" pageId="12" pageNumber="13">Taxonomic note.</emphasis>
This leafhopper had been known from four localities in Aichi Prefecture, and it was previously reported that the fuscous marking of the fore wing is sometimes obscure (
<bibRefCitation id="B475F3CAFFE0FFFF9620FEAA57B8CAA0" author="Hori" box="[1128,1263,362,386]" pageId="12" pageNumber="13" refString="Hori, Y. (1994) Three new species of the genus Pagaronia from Mt. Ena-san and Mt. Chausu-yama, central Japan (Homoptera: Cicadellidae). Transactions of the Shikoku Entomological Society, 20, 155 - 160." type="journal article" year="1994">Hori, 1994</bibRefCitation>
). Through our research, eleven localities in three prefectures are newly recorded, and some variations in the shape of male genitalia are recognizable. This species is unique in the structure of the male genitalia. The inner pygofer process is very long and the short ventral branch is forked near the base of the process. The shaft of the aedeagus is slender and has a very long spine-like ventral projection (
<figureCitation id="48DF92BEFFE0FFFF90A4FE3F50D7C935" box="[748,896,511,535]" captionStart="FIGURES 55 57" captionStartId="12.[151,264,1882,1905]" captionTargetBox="[166,1431,1069,1828]" captionTargetId="figure@12.[151,1436,1054,1845]" captionTargetPageId="12" captionText="FIGURES 55 57. Pagaronia spinosa, ɗ genitalia (55 57). — 55, pygofer in lateral view; 56 57, left inner pygofer process. Scales; 0.3 mm." httpUri="https://zenodo.org/record/211839/files/figure.png" pageId="12" pageNumber="13">Figs. 55, 56</figureCitation>
). Two males collected from northeasternmost localities, Bungui Pass and Ôkawara, are slightly different from the others in the pygofer lobe and inner pygofer process; the pygofer lobe is shorter and wider; the inner process has no ventral branch and is directed dorsally (
<figureCitation id="48DF92BEFFE0FFFF92D7FDAF53A2C9A5" box="[159,245,623,647]" captionStart="FIGURES 55 57" captionStartId="12.[151,264,1882,1905]" captionTargetBox="[166,1431,1069,1828]" captionTargetId="figure@12.[151,1436,1054,1845]" captionTargetPageId="12" captionText="FIGURES 55 57. Pagaronia spinosa, ɗ genitalia (55 57). — 55, pygofer in lateral view; 56 57, left inner pygofer process. Scales; 0.3 mm." httpUri="https://zenodo.org/record/211839/files/figure.png" pageId="12" pageNumber="13">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="D05B8E3BFFE0FFFF9281FD555752C98F" blockId="12.[151,1436,325,984]" box="[201,1029,660,686]" pageId="12" pageNumber="13">
<emphasis id="E2905229FFE0FFFF9281FD5552EDC98C" bold="true" box="[201,442,661,686]" pageId="12" pageNumber="13">Body length (mean).</emphasis>
ď,
<quantity id="171C23DEFFE0FFFF93AAFD55513EC98F" box="[482,617,661,685]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="12" pageNumber="13" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.59.5 mm</quantity>
(
<quantity id="171C23DEFFE0FFFF9030FD555184C98F" box="[632,723,661,685]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="12" pageNumber="13" unit="mm" value="8.8">8.8 mm</quantity>
); Ψ, 9.0
<quantity id="171C23DEFFE0FFFF9171FD5550D9C98F" box="[825,910,661,685]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="12" pageNumber="13" unit="mm" value="9.6">9.6 mm</quantity>
(
<quantity id="171C23DEFFE0FFFF91D5FD5550AFC98F" box="[925,1016,661,685]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="12" pageNumber="13" unit="mm" value="9.4">9.4 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="98FEDDB0FFE0FFFF9281FD7A51A3C890" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="D05B8E3BFFE0FFFF9281FD7A51A3C890" blockId="12.[151,1436,325,984]" pageId="12" pageNumber="13">
<emphasis id="E2905229FFE0FFFF9281FD7A52E9C9F1" bold="true" box="[201,446,698,723]" pageId="12" pageNumber="13">Additional material.</emphasis>
[Nagano Pref.] 1ď 2Ψ (SUU), Bungui Pass, Haseichi-nose, Ina,
<date id="A45AA8FBFFE0FFFF96EEFD7A5661C9F0" box="[1190,1334,698,722]" pageId="12" pageNumber="13" value="2006-07-04">4. VII. 2006</date>
, S. Okudera; 1ď (
<collectionCode id="B6F516FEFFE0FFFF934DFD1F5202C9D5" box="[261,341,735,759]" pageId="12" pageNumber="13">ELKU</collectionCode>
), Ôkawara, Ôshika,
<date id="A45AA8FBFFE0FFFF9071FD1F519EC9D5" box="[569,713,735,759]" pageId="12" pageNumber="13" value="2010-06-11">11. VI. 2010</date>
, S. Okudera; 1ď (SUU), Chisato, Achi,
<date id="A45AA8FBFFE0FFFF96C1FD1F565CC9D5" box="[1161,1291,735,759]" pageId="12" pageNumber="13" value="2005-06-08">8. VI. 2005</date>
, S. Okudera; 4ď 5Ψ (SUU), Namiai, Achi,
<date id="A45AA8FBFFE0FFFF93A5FCC5512BC83F" box="[493,636,773,797]" pageId="12" pageNumber="13" value="2006-07-06">6. VII. 2006</date>
, S. Okudera; 1ď (SUU), Wagô, Anan,
<date id="A45AA8FBFFE0FFFF9674FCC55795C83F" box="[1084,1218,773,797]" pageId="12" pageNumber="13" value="2005-06-08">8. VI. 2005</date>
, S. Okudera. [Gifu Pref.] 1ď 5Ψ (
<collectionCode id="B6F516FEFFE0FFFF9370FCEA52DFC860" box="[312,392,810,834]" pageId="12" pageNumber="13">ELKU</collectionCode>
), Yamaokachô-tashiro, Ena,
<date id="A45AA8FBFFE0FFFF9086FCEA5002C860" box="[718,853,810,834]" pageId="12" pageNumber="13" value="2009-05-28">28. V. 2009</date>
, S. Okudera. [Aichi Pref.] 2ď 3Ψ (SUU), Furikusa, Tôei,
<date id="A45AA8FBFFE0FFFF9292FC8F5236C845" box="[218,353,847,871]" pageId="12" pageNumber="13" value="2005-06-08">8. VI. 2005</date>
, S. Okudera; 4ď 1Ψ (
<collectionCode id="B6F516FEFFE0FFFF9015FC8F51FAC845" box="[605,685,847,871]" pageId="12" pageNumber="13">ELKU</collectionCode>
), Higashi-sonome, Tôei,
<date id="A45AA8FBFFE0FFFF9186FC8F5701C845" box="[974,1110,847,871]" pageId="12" pageNumber="13" value="2009-05-27">27. V. 2009</date>
, S. Okudera; 3ď 4Ψ (SUU), Damine, Shitara,
<date id="A45AA8FBFFE0FFFF932AFCB552ADC8AF" box="[354,506,885,909]" pageId="12" pageNumber="13" value="2007-06-26">26. VI. 2007</date>
, S. Okudera; 2ď (
<collectionCode id="B6F516FEFFE0FFFF9098FCB55077C8AF" box="[720,800,885,909]" pageId="12" pageNumber="13">ELKU</collectionCode>
), Ônumachô, Toyota,
<date id="A45AA8FBFFE0FFFF966AFCB557FAC8AF" box="[1058,1197,885,909]" pageId="12" pageNumber="13" value="2009-05-28">28. V. 2009</date>
, S. Okudera; 4ď 1Ψ (
<collectionCode id="B6F516FEFFE0FFFF92D7FC5A53B8C890" box="[159,239,922,946]" pageId="12" pageNumber="13">ELKU</collectionCode>
), Ôdairachô, Toyota,
<date id="A45AA8FBFFE0FFFF93A8FC5A5134C890" box="[480,611,922,946]" pageId="12" pageNumber="13" value="2009-05-28">28. V. 2009</date>
, S. Okudera.
</paragraph>
</subSubSection>
<subSubSection id="98FEDDB0FFE0FFFF928FFC7F5745C8FA" box="[199,1042,959,984]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="D05B8E3BFFE0FFFF928FFC7F5745C8FA" blockId="12.[151,1436,325,984]" box="[199,1042,959,984]" pageId="12" pageNumber="13">
<emphasis id="E2905229FFE0FFFF928FFC7F5237C8FA" bold="true" box="[199,352,959,984]" pageId="12" pageNumber="13">Distribution.</emphasis>
<collectingCountry id="A8F3CEABFFE0FFFF932FFC0052F0C8F5" box="[359,423,960,983]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
(central Honshu: Nagano, Gifu and Aichi Prefectures).
</paragraph>
</subSubSection>
</treatment>
</document>